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<document id="04CAE116CAD14900F1B02DB097F43E2A" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFFB735D32FAF9E4E66FFDED" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Microsphaerotherium anjozorobe Wesener 2009, new species" docType="treatment" docUuid="F357BE2F-8270-4906-B1E9-6FDA7C8D30EC" docUuidSource="ZooBank" docVersion="4" lastPageNumber="14" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="9" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="14D8AB952BFE0EB4D0A1161703A3BB85" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D4B1A472C2CCF359B8178CDB7EB9130D">
<mods:title id="BC2F9DF5D0797836846BDDCEB7778831">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
</mods:titleInfo>
<mods:name id="37E03055090D98FEC53AE6FB8F5A4057" type="personal">
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</mods:role>
<mods:namePart id="B949F944610EEA9C6EF71F663F5AED70">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="949E0D015A04E04E50D9F73D349E6206">text</mods:typeOfResource>
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<mods:titleInfo id="3C8BA3FC009C80806D990753F45C8DE5">
<mods:title id="3C9C844B36C592FE765282780537CC62">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4B78177DAF7ED153B7803783BC5B60DA">
<mods:date id="6182A46869047D816E480AE72491473E">2009</mods:date>
<mods:detail id="C479B9C501C3093D72697E6253C36011" type="pubDate">
<mods:number id="C6D6F19A07AE7C99869401F5E1B2E524">2009-05-11</mods:number>
</mods:detail>
<mods:detail id="AF7ED35EBA5BC0CFD8F64AC2AD0F85B6" type="volume">
<mods:number id="5A415B96EC9AA8C18305A08F8B54CAAD">2097</mods:number>
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<mods:detail id="E8FE05F1EB44710AD3168D2C0CB507DD" type="issue">
<mods:number id="4C70152B706619A24D54DB0D73010B82">1</mods:number>
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<mods:url id="5091D72477C8EC781B99EDF8F3202A8F">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
</mods:location>
<mods:classification id="46B8AF662E9CE25F411424DF967DD193">journal article</mods:classification>
<mods:identifier id="F05081D7580FE642174C063BDC4F08CE" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="E95770263BB840747696E8CA9DD03C63" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="529FEBFC7DF0652521FF566DE1F46BE5" type="Zenodo-Dep">5316536</mods:identifier>
<mods:identifier id="5D5B7A98A03E0EF404C06A12A60AF8EC" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
</mods:mods>
<treatment id="03FFBE1DFFFB735D32FAF9E4E66FFDED" LSID="urn:lsid:zoobank.org:act:F357BE2F-8270-4906-B1E9-6FDA7C8D30EC" httpUri="http://treatment.plazi.org/id/03FFBE1DFFFB735D32FAF9E4E66FFDED" lastPageId="13" lastPageNumber="14" pageId="8" pageNumber="9">
<subSubSection id="C34C5C80FFFB735832FAF9E4E1CBF9A1" box="[151,716,1630,1656]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BE90F0BFFFB735832FAF9E4E1CBF9A1" blockId="8.[151,716,1630,1656]" box="[151,716,1630,1656]" pageId="8" pageNumber="9">
<heading id="D0A1B867FFFB735832FAF9E4E1CBF9A1" bold="true" box="[151,716,1630,1656]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="B922D319FFFB735832FAF9E4E1CBF9A1" bold="true" box="[151,716,1630,1656]" pageId="8" pageNumber="9">
<taxonomicName id="4C567488FFFB735832FAF9E4E12AF9A1" ID-CoL="436VS" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,557,1630,1656]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="anjozorobe" status="sp. nov.">
<emphasis id="B922D319FFFB735832FAF9E4E12AF9A1" bold="true" box="[151,557,1630,1656]" italics="true" pageId="8" pageNumber="9">Microsphaerotherium anjozorobe</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2116E62FFFB73583051F9E4E1CBF9A1" box="[572,716,1630,1656]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFFB735832FAF91DE05CF881" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BE90F0BFFFB735832FAF91DE0E8F919" blockId="8.[151,1436,1703,2035]" box="[151,1007,1703,1729]" pageId="8" pageNumber="9">
<emphasis id="B922D319FFFB735832FAF91DE287F918" bold="true" box="[151,384,1703,1729]" pageId="8" pageNumber="9">Derivatio nominis:</emphasis>
<emphasis id="B922D319FFFB735833EAF910E102F918" box="[391,517,1706,1729]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C567488FFFB735833EAF910E106F918" authorityName="Wesener" authorityYear="2009" box="[391,513,1706,1729]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="anjozorobe">anjozorobe</taxonomicName>
,
</emphasis>
noun in apposition, refers to the
<typeStatus id="54EDB1A9FFFB73583108F913E094F919" box="[869,915,1705,1728]" pageId="8" pageNumber="9">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BE90F0BFFFB735832ABF970E0A9F8EB" blockId="8.[151,1436,1703,2035]" pageId="8" pageNumber="9">
<emphasis id="B922D319FFFB735832ABF970E298F93D" bold="true" box="[198,415,1738,1764]" pageId="8" pageNumber="9">Studied material:</emphasis>
<materialsCitation id="3B3E0556FFFB735833CBF970E0A9F8EB" collectingDate="2000-12-05" collectingDateMax="2000-12-13" collectingDateMin="2000-12-05" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="1300" latitude="-18.473333" location="Andranomay" longLatPrecision="21" longitude="47.96" pageId="8" pageNumber="9" specimenCode="BLF 2370" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
<emphasis id="B922D319FFFB735833CBF970E11FF93D" bold="true" box="[422,536,1738,1764]" pageId="8" pageNumber="9">
<typeStatus id="54EDB1A9FFFB735833CBF970E11FF93D" box="[422,536,1738,1764]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
: 1 M (
<quantity id="4CAEA2EEFFFB7358300AF970E1BEF93D" box="[615,697,1738,1764]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="mm" value="15.0">15 mm</quantity>
long),
<emphasis id="B922D319FFFB73583161F970E0C0F93D" bold="true" box="[780,967,1738,1764]" pageId="8" pageNumber="9">
<collectionCode id="ED4797CEFFFB73583161F970E040F93D" box="[780,839,1738,1764]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="9" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFFB73583120F970E0C0F93D" box="[845,967,1738,1764]" collectionCode="BLF" pageId="8" pageNumber="9">BLF 2370</specimenCode>
</emphasis>
,
<collectingCountry id="F3414F9BFFFB735831BBF970E76FF93D" box="[982,1128,1738,1764]" name="Madagascar" pageId="8" pageNumber="9">Madagascar</collectingCountry>
:
<collectingRegion id="4992C1E9FFFB7358361FF970E690F93D" box="[1138,1431,1738,1764]" country="Madagascar" name="Antananarivo" pageId="8" pageNumber="9">Province d'Antananarivo</collectingRegion>
,
<quantity id="4CAEA2EEFFFB735832FAF94BE3DDF8D2" box="[151,218,1777,1803]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="km" value="3.0">3 km</quantity>
41° NE
<location id="8E8959D0FFFB73583337F94BE2F3F8D2" LSID="urn:lsid:plazi:treatment:03FFBE1DFFFB735D32FAF9E4E66FFDED:8E8959D0FFFB73583337F94BE2F3F8D2" box="[346,500,1777,1803]" country="Madagascar" latitude="-18.473333" longLatPrecision="21" longitude="47.96" name="Andranomay" pageId="8" pageNumber="9" stateProvince="Antananarivo">Andranomay</location>
,
<locationDeviation id="9A5B1629FFFB73583064F94BE080F8D2" box="[521,903,1777,1803]" location="Anjozorobe" pageId="8" pageNumber="9">
<quantity id="4CAEA2EEFFFB73583064F94BE177F8D2" box="[521,624,1777,1803]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="8" pageNumber="9" unit="km" value="11.5">11.5 km</quantity>
147°SSE Anjozorobe
</locationDeviation>
, montane rainforest,
<quantity id="4CAEA2EEFFFB735836FAF94BE7FCF8D2" box="[1175,1275,1777,1803]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="9" unit="m" value="1300.0">
<elevation id="007BE838FFFB735836FAF94BE7FCF8D2" box="[1175,1275,1777,1803]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="9" unit="m" value="1300.0">1300 m</elevation>
</quantity>
,
<geoCoordinate id="EE6269CCFFFB7358377DF94BE690F8D2" box="[1296,1431,1777,1803]" degrees="18" direction="south" minutes="28" orientation="latitude" pageId="8" pageNumber="9" precision="15" seconds="24" value="-18.473333">18°28'24&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFFFB735832FAF8A2E227F8EB" box="[151,288,1816,1842]" degrees="47" direction="east" minutes="57" orientation="longitude" pageId="8" pageNumber="9" precision="15" seconds="36" value="47.96">47°57'36&quot;E</geoCoordinate>
, coll.
<collectorName id="26A36ADDFFFB7358330AF8A2E2B5F8EB" box="[359,434,1816,1842]" pageId="8" pageNumber="9">Fisher</collectorName>
,
<collectorName id="26A36ADDFFFB735833D0F8A2E12DF8EB" box="[445,554,1816,1842]" pageId="8" pageNumber="9">Griswold</collectorName>
<emphasis id="B922D319FFFB7358305FF8A2E16AF8E8" box="[562,621,1816,1841]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
<date id="FFE829CBFFFB73583019F8A2E013F8EB" box="[628,788,1816,1842]" pageId="8" pageNumber="9" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">
<collectingDate id="EFACD023FFFB73583019F8A2E013F8EB" box="[628,788,1816,1842]" pageId="8" pageNumber="9" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">5-13.xii.2000</collectingDate>
</date>
,
<collectingMethod id="5217771CFFFB7358314DF8A2E0ADF8EB" box="[800,938,1816,1842]" pageId="8" pageNumber="9">pitfall traps</collectingMethod>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFFFB735832ABF884E05CF881" blockId="8.[151,1436,1703,2035]" box="[198,859,1854,1880]" pageId="8" pageNumber="9">
<materialsCitation id="3B3E0556FFFB735832ABF884E050F881" box="[198,855,1854,1880]" collectingDate="2000-12-05" collectingDateMax="2000-12-13" collectingDateMin="2000-12-05" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="1300" latitude="-18.473333" location="Andranomay" longLatPrecision="21" longitude="47.96" pageId="8" pageNumber="9" specimenCode="BLF 2371" specimenCount="1" stateProvince="Antananarivo" typeStatus="paratype">
<emphasis id="B922D319FFFB735832ABF884E23FF881" bold="true" box="[198,312,1854,1880]" pageId="8" pageNumber="9">
<typeStatus id="54EDB1A9FFFB735832ABF884E23FF881" box="[198,312,1854,1880]" pageId="8" pageNumber="9" type="paratype">Paratype</typeStatus>
</emphasis>
: 1 M,
<emphasis id="B922D319FFFB735833EBF884E145F881" bold="true" box="[390,578,1854,1880]" pageId="8" pageNumber="9">
<collectionCode id="ED4797CEFFFB735833EBF884E2C7F881" box="[390,448,1854,1880]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="9" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFFB735833A5F884E145F881" box="[456,578,1854,1880]" collectionCode="BLF" pageId="8" pageNumber="9">BLF 2371</specimenCode>
</emphasis>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFFB735832ABF8DFE79EF8A6" box="[198,1177,1893,1919]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BE90F0BFFFB735832ABF8DFE79EF8A6" blockId="8.[151,1436,1703,2035]" box="[198,1177,1893,1919]" pageId="8" pageNumber="9">
<emphasis id="B922D319FFFB735832ABF8DFE26EF8A6" bold="true" box="[198,361,1893,1919]" pageId="8" pageNumber="9">Distribution:</emphasis>
This species is only known from the Andranomay highlands (Fig. 5).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFFB735C32ABF831E079FCCF" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BE90F0BFFFB735932ABF831E77DFEAA" blockId="8.[151,1436,1703,2035]" lastBlockId="9.[151,1436,152,371]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="B922D319FFFB735832ABF831E24FF87C" bold="true" box="[198,328,1931,1957]" pageId="8" pageNumber="9">Diagnosis:</emphasis>
Small, minute, up to
<quantity id="4CAEA2EEFFFB7358303FF836E1A1F87F" box="[594,678,1932,1958]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="mm" value="15.0">15 mm</quantity>
long. Color shiny brownish (Fig. 1A). Legs, antenna and head castaneous-brown. Texture of tergites polished and shiny. First antennomere with groove, antennomeres 1–3 with sclerotized teeth (Fig. 3A). Disc with 7–9 apical cones (Fig. 3A). Movable finger of posterior telopods slender, conical and elongated, weakly curved, posterior side with
<emphasis id="B922D319FFFA735931A5FF22E704FF68" box="[968,1027,152,177]" italics="true" pageId="9" pageNumber="10">circa</emphasis>
four large crenulated, sclerotized teeth and three thin spines (Fig. 2E). Fixed finger basally with single spine, at mid-length strongly curved towards movable finger. Male harp with single, weakly developed stridulation rib (Fig. 2A). Third podomere of anterior telopods shorter than first podomere (Fig. 2A). Single, very long locking carina on the anal shield, suture marking borders of two fused carinae still visible (Fig. 1B). Anal shield of slightly rectangular shape, completely covered with dense field of short (sensory?) hair and few very long hairs.
</paragraph>
<paragraph id="8BE90F0BFFFA735932FAF923E2F7F884" blockId="9.[151,1436,1689,1885]" pageId="9" pageNumber="10">
<emphasis id="B922D319FFFA735932FAF923E229F968" bold="true" box="[151,302,1689,1713]" pageId="9" pageNumber="10">FIGURE 001</emphasis>
.
<emphasis id="B922D319FFFA73593356F923E249F968" bold="true" box="[315,334,1689,1713]" pageId="9" pageNumber="10">A</emphasis>
:
<taxonomicName id="4C567488FFFA73593331F923E1C5F969" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[348,706,1689,1712]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="anjozorobe" status="sp. nov.">
<emphasis id="B922D319FFFA73593331F923E1C5F969" box="[348,706,1689,1712]" italics="true" pageId="9" pageNumber="10">Microsphaerotherium anjozorobe</emphasis>
</taxonomicName>
<emphasis id="B922D319FFFA735930A4F923E002F968" bold="true" box="[713,773,1689,1713]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A2116E62FFFA735930A4F923E002F968" box="[713,773,1689,1713]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFFA7359317EF923E075F969" box="[787,882,1689,1712]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
, photograph;
<emphasis id="B922D319FFFA73593669F923E711F968" bold="true" box="[1028,1046,1689,1713]" pageId="9" pageNumber="10">B</emphasis>
:
<taxonomicName id="4C567488FFFA73593649F923E7B9F969" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1060,1214,1689,1712]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="anjozorobe" status="sp. nov.">
<emphasis id="B922D319FFFA73593649F923E7B9F969" box="[1060,1214,1689,1712]" italics="true" pageId="9" pageNumber="10">M. anjozorobe</emphasis>
</taxonomicName>
<emphasis id="B922D319FFFA735936A8F923E606F968" bold="true" box="[1221,1281,1689,1713]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A2116E62FFFA735936A8F923E606F968" box="[1221,1281,1689,1713]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFFA73593762F923E669F969" box="[1295,1390,1689,1712]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
, left locking carina of anal shield;
<emphasis id="B922D319FFFA7359338EF901E2F1F90A" bold="true" box="[483,502,1723,1747]" pageId="9" pageNumber="10">C</emphasis>
:
<taxonomicName id="4C567488FFFA7359306AF906E1E9F90A" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[519,750,1724,1747]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="album" status="sp. nov.">
<emphasis id="B922D319FFFA7359306AF906E1E9F90A" box="[519,750,1724,1747]" italics="true" pageId="9" pageNumber="10">Zoosphaerium album</emphasis>
</taxonomicName>
<emphasis id="B922D319FFFA73593095F901E030F90A" bold="true" box="[760,823,1723,1747]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A2116E62FFFA73593095F901E030F90A" box="[760,823,1723,1747]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, photograph of living specimens from Zombitse, photo taken by Peter Oxford, all rights reserved. Abbreviations:
<emphasis id="B922D319FFFA73593178F964E045F92F" bold="true" box="[789,834,1758,1782]" pageId="9" pageNumber="10">12T</emphasis>
= 12th tergite;
<emphasis id="B922D319FFFA7359319DF964E749F92F" bold="true" box="[1008,1102,1758,1782]" pageId="9" pageNumber="10">
13T+
<collectingCountry id="F3414F9BFFFA73593646F964E749F92F" box="[1067,1102,1758,1782]" name="American Samoa" pageId="9" pageNumber="10">AS</collectingCountry>
</emphasis>
= 13th tergite fused with anal shield;
<emphasis id="B922D319FFFA73593288F8BBE200F8C0" bold="true" box="[229,263,1793,1817]" pageId="9" pageNumber="10">
<collectingCountry id="F3414F9BFFFA73593288F8BBE200F8C0" box="[229,263,1793,1817]" name="American Samoa" pageId="9" pageNumber="10">AS</collectingCountry>
</emphasis>
= Anal shield;
<emphasis id="B922D319FFFA735933C3F8BBE2BDF8C0" bold="true" box="[430,442,1793,1817]" pageId="9" pageNumber="10">c</emphasis>
= collum (modified first tergite);
<emphasis id="B922D319FFFA7359314BF8BBE032F8C0" bold="true" box="[806,821,1793,1817]" pageId="9" pageNumber="10">h</emphasis>
= head;
<emphasis id="B922D319FFFA735931F9F8BBE0BEF8C0" bold="true" box="[916,953,1793,1817]" pageId="9" pageNumber="10">LC</emphasis>
= locking carina;
<emphasis id="B922D319FFFA73593610F8BBE792F8C0" bold="true" box="[1149,1173,1793,1817]" pageId="9" pageNumber="10">pt</emphasis>
= paratergite;
<emphasis id="B922D319FFFA7359375EF8BBE63AF8C0" bold="true" box="[1331,1341,1793,1817]" pageId="9" pageNumber="10">s</emphasis>
= suture between 13th tergite fused with anal shield;
<emphasis id="B922D319FFFA735930EBF899E1B6F8E2" bold="true" box="[646,689,1827,1851]" pageId="9" pageNumber="10">th-s</emphasis>
= thoracic shield;
<emphasis id="B922D319FFFA735931E8F899E0D3F8E2" bold="true" box="[901,980,1827,1851]" pageId="9" pageNumber="10">th-s gr</emphasis>
= lateral depression (groove) of thoracic shield. Photographs not to scale.
</paragraph>
<paragraph id="8BE90F0BFFFA735932ABF835E69BF809" blockId="9.[151,1436,1935,2000]" pageId="9" pageNumber="10">
<emphasis id="B922D319FFFA735932ABF835E28BF870" bold="true" box="[198,396,1935,1961]" pageId="9" pageNumber="10">Similar species:</emphasis>
The shape and position of the locking carinae and the special-shaped setae on the anal shield marks
<taxonomicName id="4C567488FFFA73593356F80DE2EFF809" authorityName="Wesener" authorityYear="2009" box="[315,488,1975,2000]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="anjozorobe">
<emphasis id="B922D319FFFA73593356F80DE2EFF809" box="[315,488,1975,2000]" italics="true" pageId="9" pageNumber="10">M. anjozorobe</emphasis>
</taxonomicName>
as a member of
<taxonomicName id="4C567488FFFA735930D3F80DE66AF809" authority="Wesener &amp; van den Spiegel, 2007" authorityName="Wesener &amp; van den Spiegel" authorityYear="2007" box="[702,1389,1974,2000]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B922D319FFFA735930D3F80DE0BAF809" box="[702,957,1975,2000]" italics="true" pageId="9" pageNumber="10">Microsphaerotherium</emphasis>
Wesener &amp; van den Spiegel, 2007
</taxonomicName>
.
<taxonomicName id="4C567488FFFA73593711F80DE69BF809" box="[1404,1436,1975,2000]" pageId="9" pageNumber="10">
<emphasis id="B922D319FFFA73593711F80DE69BF809" box="[1404,1436,1975,2000]" italics="true" pageId="9" pageNumber="10">M.</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BE90F0BFFF9735A32FAFF22E277FF01" blockId="10.[151,1435,152,216]" pageId="10" pageNumber="11">
<emphasis id="B922D319FFF9735A32FAFF22E21DFF68" box="[151,282,152,177]" italics="true" pageId="10" pageNumber="11">anjozorobe</emphasis>
differs from
<taxonomicName id="4C567488FFF9735A33D9FF22E166FF68" authorityName="Wesener &amp; VandenSpiegel" authorityYear="2007" box="[436,609,152,177]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ivohibiense">
<emphasis id="B922D319FFF9735A33D9FF22E166FF68" box="[436,609,152,177]" italics="true" pageId="10" pageNumber="11">M. ivohibiense</emphasis>
</taxonomicName>
in the total absence of pits or hair on the tergites and in the absence of a sclerotized crest.
</paragraph>
<paragraph id="8BE90F0BFFF9735A32FAF8A6E25DF867" blockId="10.[151,1436,1820,1982]" pageId="10" pageNumber="11">
<emphasis id="B922D319FFF9735A32FAF8A6E228F8ED" bold="true" box="[151,303,1820,1844]" pageId="10" pageNumber="11">FIGURE 002</emphasis>
.
<taxonomicName id="4C567488FFF9735A3352F8A6E1A0F8EA" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[319,679,1820,1843]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="anjozorobe" status="sp. nov.">
<emphasis id="B922D319FFF9735A3352F8A6E1A0F8EA" box="[319,679,1820,1843]" italics="true" pageId="10" pageNumber="11">Microsphaerotherium anjozorobe</emphasis>
</taxonomicName>
<emphasis id="B922D319FFF9735A30C2F8A6E1E9F8ED" bold="true" box="[687,750,1820,1844]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A2116E62FFF9735A30C2F8A6E1E9F8ED" box="[687,750,1820,1844]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFF9735A3090F8A6E05AF8EA" box="[765,861,1820,1843]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
, roman numbers refer to number of podomere,
<emphasis id="B922D319FFF9735A3705F8A6E69BF8ED" bold="true" box="[1384,1436,1820,1844]" pageId="10" pageNumber="11">A–C</emphasis>
anterior telopods,
<emphasis id="B922D319FFF9735A3331F884E268F88F" bold="true" box="[348,367,1854,1878]" pageId="10" pageNumber="11">A</emphasis>
: anterior view;
<emphasis id="B922D319FFF9735A304DF884E135F88F" bold="true" box="[544,562,1854,1878]" pageId="10" pageNumber="11">B</emphasis>
: last two podomeres, posterior view;
<emphasis id="B922D319FFF9735A31BCF884E0E3F88F" bold="true" box="[977,996,1854,1878]" pageId="10" pageNumber="11">C</emphasis>
: last two podomeres, lateral view;
<emphasis id="B922D319FFF9735A370AF884E69BF88F" bold="true" box="[1383,1436,1854,1878]" pageId="10" pageNumber="11">D, E</emphasis>
posterior telopods,
<emphasis id="B922D319FFF9735A3301F8DBE278F8A0" bold="true" box="[364,383,1889,1913]" pageId="10" pageNumber="11">D</emphasis>
: anterior view;
<emphasis id="B922D319FFF9735A3058F8DBE140F8A0" bold="true" box="[565,583,1889,1913]" pageId="10" pageNumber="11">E</emphasis>
: posterior view. Abbreviations:
<emphasis id="B922D319FFF9735A31C2F8DBE0D1F8A0" bold="true" box="[943,982,1889,1913]" pageId="10" pageNumber="11">cav</emphasis>
= cavity;
<emphasis id="B922D319FFF9735A3620F8DBE771F8A0" bold="true" box="[1101,1142,1889,1913]" pageId="10" pageNumber="11">cr-t</emphasis>
= crenulated teeth;
<emphasis id="B922D319FFF9735A3737F8DBE685F8A0" bold="true" box="[1370,1410,1889,1913]" pageId="10" pageNumber="11">m-f</emphasis>
= movable finger;
<emphasis id="B922D319FFF9735A3321F83EE26EF845" bold="true" box="[332,361,1924,1948]" pageId="10" pageNumber="11">ml</emphasis>
= membranous lobe;
<emphasis id="B922D319FFF9735A3030F83EE1A5F845" bold="true" box="[605,674,1924,1948]" pageId="10" pageNumber="11">imm-f</emphasis>
= fixed finger;
<emphasis id="B922D319FFF9735A3138F83EE08FF845" bold="true" box="[853,904,1924,1948]" pageId="10" pageNumber="11">proc</emphasis>
= 2nd podomere process;
<emphasis id="B922D319FFF9735A36DCF83EE7D4F845" bold="true" box="[1201,1235,1924,1948]" pageId="10" pageNumber="11">SR</emphasis>
= stridulation rib. Scale bar =
<quantity id="4CAEA2EEFFF9735A3378F81DE250F867" box="[277,343,1959,1982]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFFF8735B32FAF82AE705F834" blockId="11.[151,1436,1936,2029]" pageId="11" pageNumber="12">
<emphasis id="B922D319FFF8735B32FAF82AE229F871" bold="true" box="[151,302,1936,1960]" pageId="11" pageNumber="12">FIGURE 003</emphasis>
.
<taxonomicName id="4C567488FFF8735B3351F82BE1A5F871" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[316,674,1937,1960]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="anjozorobe" status="sp. nov.">
<emphasis id="B922D319FFF8735B3351F82BE1A5F871" box="[316,674,1937,1960]" italics="true" pageId="11" pageNumber="12">Microsphaerotherium anjozorobe</emphasis>
</taxonomicName>
<emphasis id="B922D319FFF8735B30C4F82AE1E2F871" bold="true" box="[681,741,1936,1960]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="A2116E62FFF8735B30C4F82AE1E2F871" box="[681,741,1936,1960]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFF8735B309EF82BE054F871" box="[755,851,1937,1960]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
, SEM,
<emphasis id="B922D319FFF8735B31CEF82AE0B1F871" bold="true" box="[931,950,1936,1960]" pageId="11" pageNumber="12">A</emphasis>
: antenna, lateral view;
<emphasis id="B922D319FFF8735B36DFF82AE7C3F871" bold="true" box="[1202,1220,1936,1960]" pageId="11" pageNumber="12">B</emphasis>
: antennomeres 1–2, lateral view;
<emphasis id="B922D319FFF8735B3345F809E23CF812" bold="true" box="[296,315,1971,1995]" pageId="11" pageNumber="12">C</emphasis>
: right mandible, mesal view. Abbreviations:
<emphasis id="B922D319FFF8735B3143F809E052F812" bold="true" box="[814,853,1971,1995]" pageId="11" pageNumber="12">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FFF8735B367EF809E734F812" bold="true" box="[1043,1075,1971,1995]" pageId="11" pageNumber="12">Co</emphasis>
= condylus;
<emphasis id="B922D319FFF8735B36AAF809E7E0F812" bold="true" box="[1223,1255,1971,1995]" pageId="11" pageNumber="12">Gr</emphasis>
= groove;
<emphasis id="B922D319FFF8735B3708F809E684F812" bold="true" box="[1381,1411,1971,1995]" pageId="11" pageNumber="12">eT</emphasis>
= external tooth;
<emphasis id="B922D319FFF8735B3355F86FE25AF834" bold="true" box="[312,349,2005,2029]" pageId="11" pageNumber="12">mp</emphasis>
= molar plate;
<emphasis id="B922D319FFF8735B306DF86FE127F834" bold="true" box="[512,544,2005,2029]" pageId="11" pageNumber="12">pL</emphasis>
= pectinate lamella;
<emphasis id="B922D319FFF8735B316CF86FE037F834" bold="true" box="[769,816,2005,2029]" pageId="11" pageNumber="12">scl-t</emphasis>
= sclerotized teeth.
</paragraph>
<paragraph id="8BE90F0BFFFF735C32FAFD66E079FCCF" blockId="12.[151,1436,732,790]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32FAFD66E231FD2D" bold="true" box="[151,310,732,756]" pageId="12" pageNumber="13">FIGURE 004</emphasis>
.
<taxonomicName id="4C567488FFFF735C3321FD66E1BDFD2A" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[332,698,732,755]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anjozorobe" status="sp. nov.">
<emphasis id="B922D319FFFF735C3321FD66E1BDFD2A" box="[332,698,732,755]" italics="true" pageId="12" pageNumber="13">Microsphaerotherium anjozorobe</emphasis>
</taxonomicName>
<emphasis id="B922D319FFFF735C30A7FD66E008FD2D" bold="true" box="[714,783,732,756]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A2116E62FFFF735C30A7FD66E008FD2D" box="[714,783,732,756]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFFF735C3148FD66E082FD2A" box="[805,901,732,755]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
, SEM, underside of midbody left paratergite. Abbreviations:
<emphasis id="B922D319FFFF735C3351FD44E271FCCF" bold="true" box="[316,374,766,790]" pageId="12" pageNumber="13">Lock</emphasis>
= locking carina of tergite;
<emphasis id="B922D319FFFF735C30F2FD44E1DBFCCF" bold="true" box="[671,732,766,790]" pageId="12" pageNumber="13">Endo</emphasis>
= endotergum.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFFF735D32ABFCF2E66FFDED" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="8BE90F0BFFFF735C32ABFCF2E7F1FCBB" blockId="12.[151,1436,840,2026]" box="[198,1270,840,866]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32ABFCF2E267FCBB" bold="true" box="[198,352,840,866]" pageId="12" pageNumber="13">Description:</emphasis>
<emphasis id="B922D319FFFF735C3305FCF3E2F2FCBB" box="[360,501,841,866]" italics="true" pageId="12" pageNumber="13">Body length</emphasis>
: Length
<quantity id="4CAEA2EEFFFF735C3032FCF2E1B3FCBB" box="[607,692,840,866]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="mm" value="15.0">15 mm</quantity>
, width of thoracic shield
<quantity id="4CAEA2EEFFFF735C31B1FCF2E73FFCBB" box="[988,1080,840,866]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="12" pageNumber="13" unit="mm" value="5.9">5.9 mm</quantity>
, height
<quantity id="4CAEA2EEFFFF735C36F8FCF2E7F5FCBB" box="[1173,1266,840,866]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="12" pageNumber="13" unit="mm" value="3.9">3.9 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAFCD5E11CFC69" blockId="12.[151,1436,840,2026]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAFCD5E256FC51" box="[199,337,879,904]" italics="true" pageId="12" pageNumber="13">Coloration:</emphasis>
Legs, antenna, head, collum and tergites shiny castaneous brown (Fig. 1A). Natural coloration influenced by alcohol.
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAFC06E19EFC0F" blockId="12.[151,1436,840,2026]" box="[199,665,956,982]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAFC06E217FC0C" box="[199,272,956,981]" italics="true" pageId="12" pageNumber="13">Head:</emphasis>
Eyes with less than 28–32 ocelli.
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAFC59E1A7FB93" blockId="12.[151,1436,840,2026]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAFC59E238FC25" box="[199,319,995,1020]" italics="true" pageId="12" pageNumber="13">Antennae:</emphasis>
Length of antennomeres: 1&gt;2=3&gt;4=5&lt;&lt;6, sixth antennomere cylindrical, on tip bearing disc with 7–9 apical cones (Fig. 3A). First antennomere of greatest width and with groove (Fig. 3B). Sclerotized teeth at base of antennomeres 1–3 (Fig. 3A).
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAFBEDE080FB41" blockId="12.[151,1436,840,2026]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAFBEDE247FBA9" box="[199,320,1111,1136]" italics="true" pageId="12" pageNumber="13">Mandible:</emphasis>
Five rows of pectinate lamellae, number of teeth declining apically to proximally (Fig. 3C). Molar plate with deep groove, condylus well-rounded (Fig. 3C).
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAFB1FE103FB3C" blockId="12.[151,1436,840,2026]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAFB1FE296FB67" box="[199,401,1189,1214]" italics="true" pageId="12" pageNumber="13">Gnathochilarium</emphasis>
: Lateral of palpi four sensory cones, located together. Depression on posterior side of palpi with some sensory cones.
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAFB48E1AEFAD5" blockId="12.[151,1436,840,2026]" box="[199,681,1266,1292]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAFB48E220FAD2" box="[199,295,1266,1291]" italics="true" pageId="12" pageNumber="13">Collum:</emphasis>
Median part of collum glabrous.
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAFAA3E07EFAEB" blockId="12.[151,1436,840,2026]" box="[199,889,1304,1330]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAFAA3E279FAEB" box="[199,382,1305,1330]" italics="true" pageId="12" pageNumber="13">Thoracic shield</emphasis>
: Surface similar to other tergites (Fig. 1A).
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAFA85E7AFFA80" blockId="12.[151,1436,840,2026]" box="[199,1192,1343,1369]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAFA85E224FA81" box="[199,291,1343,1368]" italics="true" pageId="12" pageNumber="13">Tergites</emphasis>
: Texture glabrous and shiny. Paratergites projecting posteriorly (Figs 1A; 4).
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAFADCE292FA14" blockId="12.[151,1436,840,2026]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAFADCE267FAA6" box="[199,352,1382,1407]" italics="true" pageId="12" pageNumber="13">Endotergum:</emphasis>
Inner area with small conical spines and long hair. Single row of large, circular cuticular impressions (Fig. 23A). Single row of marginal bristles, standing closely to one another, not reaching tergite margin (Figs 4, 23A).
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAFA60E184F9B1" blockId="12.[151,1436,840,2026]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAFA60E251FA2A" box="[199,342,1498,1523]" italics="true" pageId="12" pageNumber="13">Anal shield:</emphasis>
Well-rounded, neither bell-shaped nor tapered, in ventral view of rectangular shape. At least in the male completely covered by dense field of short and stout setae, each placed inside pit. Between those setae stand few, isolated, very long hairs which reach above anal shield margin. Ventral side of anal shield with single, long locking carina (Fig. 1B).
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAF9CFE681F905" blockId="12.[151,1436,840,2026]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAF9CFE20FF957" box="[199,264,1653,1678]" italics="true" pageId="12" pageNumber="13">Legs:</emphasis>
Tarsi of first pair of legs with three or four, second with four or five and third with seven ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 4–21 with curved claws, 9 or 10 ventral spines and single apical spine. Femur 2.5 and tarsi 4.8 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAF953E056F8DB" blockId="12.[151,1436,840,2026]" box="[199,849,1768,1794]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAF953E28BF8DB" box="[199,396,1769,1794]" italics="true" pageId="12" pageNumber="13">Stigmatic plates:</emphasis>
First lobe long, with well-rounded tip.
</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAF8B5E296F8F0" blockId="12.[151,1436,840,2026]" box="[199,401,1807,1833]" pageId="12" pageNumber="13">Female unknown</paragraph>
<paragraph id="8BE90F0BFFFF735C32AAF88CE127F8AF" blockId="12.[151,1436,840,2026]" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAF88CE2E2F896" box="[199,485,1846,1871]" italics="true" pageId="12" pageNumber="13">Male sexual characters</emphasis>
: Male gonopore covered by large, sclerotized, undivided and rounded plate. Apical part of plate membranous.
</paragraph>
<paragraph id="8BE90F0BFFFF735D32AAF839E00FFEFF" blockId="12.[151,1436,840,2026]" lastBlockId="13.[151,1436,152,564]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B922D319FFFF735C32AAF839E290F845" box="[199,407,1923,1948]" italics="true" pageId="12" pageNumber="13">Anterior telopods</emphasis>
(Figs 2A–C): Syncoxite glabrous, covered anteriorly with a few minute spines. Harp with single, long, but weakly developed stridulation rib. Second podomere process lobe-like and little curved with rounded edges, reaching up to half of third podomere's height; process with sclerotized spots on surface facing third podomere. Third podomere short, covered with numerous isolated hairs on both sides. Mesal margin towards cavity with
<emphasis id="B922D319FFFE735D338FFF05E11AFF01" box="[482,541,191,216]" italics="true" pageId="13" pageNumber="14">circa</emphasis>
four thin, sclerotized spines and in basal half covered with numerous small sclerotized spots. Lateral margin towards cavity with
<emphasis id="B922D319FFFE735D3161FF5FE040FF27" box="[780,839,229,254]" italics="true" pageId="13" pageNumber="14">circa</emphasis>
four crenulated, sclerotized teeth. Third podomere apically at mesal margin with larger, sclerotized spot.
</paragraph>
<paragraph id="8BE90F0BFFFE735D32AAFE88E66FFDED" blockId="13.[151,1436,152,564]" pageId="13" pageNumber="14">
<emphasis id="B922D319FFFE735D32AAFE88E2A2FE92" box="[199,421,306,331]" italics="true" pageId="13" pageNumber="14">Posterior telopods</emphasis>
(Figs 2D, E): Movable finger covered on both sides with few, long hairs. Finger weakly curved, more slender and cylindrical, length
<emphasis id="B922D319FFFE735D317DFEE3E04CFEAB" box="[784,843,345,370]" italics="true" pageId="13" pageNumber="14">circa</emphasis>
3.3 times width. Inner margin of movable finger only at apical half hollowed-out, with three small, sclerotized spines and weakly developed membranous lobe; posterior aspect with single row of
<emphasis id="B922D319FFFE735D302AFE1DE185FE19" box="[583,642,423,448]" italics="true" pageId="13" pageNumber="14">circa</emphasis>
four large, crenulated sclerotized teeth. Fixed finger basally with short, sclerotized spine. Finger long, ending slightly before tip of movable finger. Finger at mid-point strongly curved towards movable finger. Margin juxtaposed to movable finger covered apically with few minute sclerotized spots. First and second podomere glabrous. Inner horns of syncoxite with pointed, slender tip.
</paragraph>
</subSubSection>
</treatment>
</document>
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