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<document id="325F8EA80C3AAA38352F14CE41E75FBB" ID-DOI="10.5281/zenodo.198427" ID-GBIF-Dataset="1c32eb21-6978-41da-b589-c98453d4f572" ID-ISSN="1175-5326" ID-Zenodo-Dep="198427" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460297110161" checkinUser="plazi" docAuthor="Short, Andrew E. Z. &amp; García, Mauricio" docDate="2010" docId="03CEA91F8C36B90FA2C06572FBF8F060" docLanguage="en" docName="zt02635p031.pdf" docOrigin="Zootaxa 2635" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Oocyclus petra Short &amp; Garcia, sp. n." docType="treatment" docVersion="12" lastPageNumber="20" masterDocId="FFF7D1678C27B91CA2576660FFD8F557" masterDocTitle="A review of the Oocyclus Sharp of Venezuela with description of twelve new species (Coleoptera: Hydrophilidae: Laccobiini)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="18" updateTime="1698252212253" updateUser="plazi">
<mods:mods id="FB6546BA091E954C8C947C3FE718507D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DEA373E7FCE0A807718D634C59725BEA">
<mods:title id="ECBD2F82594F1DB6DB4ABF17583DBDA0">A review of the Oocyclus Sharp of Venezuela with description of twelve new species (Coleoptera: Hydrophilidae: Laccobiini)</mods:title>
</mods:titleInfo>
<mods:name id="FD61C3AB23385348EBE11C9686FC6326" type="personal">
<mods:role id="A3E9906585FAFF72E963BED4330BCBEC">
<mods:roleTerm id="0BEB4D952DBC3A389F64FDED8A955ADF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A876ED16DA48AE909EC90D5D26467191">Short, Andrew E. Z.</mods:namePart>
</mods:name>
<mods:name id="411E9A263134ABFB9D8D3E44A34B8E67" type="personal">
<mods:role id="8848A672AD7CFFE9B2C41296E5D4931C">
<mods:roleTerm id="0459487AD7E8227C4C6A98B9D3BAF7B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D94971F790AE804803CA9B6BC015680">García, Mauricio</mods:namePart>
</mods:name>
<mods:typeOfResource id="7F03BB70AB0CFFBD9F108AE9C906CE79">text</mods:typeOfResource>
<mods:relatedItem id="1CA57D1FB0DE0F88C74D82C958659320" type="host">
<mods:titleInfo id="369C9C35E357334E4C2AA1DC9559B163">
<mods:title id="459943768E1B021885F1C1FC7E439CB3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F2B03BE1B9D2A6A90F530E9F43272E09">
<mods:date id="CC80A314AFF5A678ADDB3A037A07DEFD">2010</mods:date>
<mods:detail id="8FAF24D45DF2B1FB8A8C651541B2B065" type="volume">
<mods:number id="04B8A31FF42C921FE20184E2C1BD56C5">2635</mods:number>
</mods:detail>
<mods:extent id="2BE1B0ABC2022986AF54278D54CD96A8" unit="page">
<mods:start id="89E7C00E48DD811A7FFA59A09598A349">1</mods:start>
<mods:end id="29FA51C958B97BD1FE382601AC098A3D">31</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="51CAA2B1E0F4B40F61A7D90258B6079E">journal article</mods:classification>
<mods:identifier id="DC8045BA1032F9786190638AF4DF4F7B" type="DOI">10.5281/zenodo.198427</mods:identifier>
<mods:identifier id="69C10BB1289945065CC6841B635F0379" type="GBIF-Dataset">1c32eb21-6978-41da-b589-c98453d4f572</mods:identifier>
<mods:identifier id="83CB6B76BF556CB3CCEF1D64FC5FE774" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="26C9F3CBF029830BB71682819D0D6594" type="Zenodo-Dep">198427</mods:identifier>
</mods:mods>
<treatment id="03CEA91F8C36B90FA2C06572FBF8F060" ID-DOI="http://doi.org/10.5281/zenodo.6205306" ID-GBIF-Taxon="119399266" ID-Zenodo-Dep="6205306" LSID="urn:lsid:plazi:treatment:03CEA91F8C36B90FA2C06572FBF8F060" httpUri="http://treatment.plazi.org/id/03CEA91F8C36B90FA2C06572FBF8F060" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="C37D4B828C36B90DA2C06572FE53F618" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BD818098C36B90DA2C06572FD86F67B" blockId="17.[151,606,786,847]" box="[151,606,786,812]" pageId="17" pageNumber="18">
<heading id="D090AF658C36B90DA2C06572FD86F67B" bold="true" box="[151,606,786,812]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="B913C41B8C36B90DA2C06572FD86F67B" bold="true" box="[151,606,786,812]" pageId="17" pageNumber="18">
<taxonomicName id="4C67638A8C36B90DA2C06572FDCDF67B" ID-CoL="49RWH" authority="Short &amp; Garcia" authorityName="Short &amp; Garcia" box="[151,533,786,812]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="petra" status="sp. nov.">
<emphasis id="B913C41B8C36B90DA2C06572FE93F67B" bold="true" box="[151,331,786,812]" italics="true" pageId="17" pageNumber="18">Oocyclus petra</emphasis>
Short &amp; García
</taxonomicName>
<taxonomicNameLabel id="A22079608C36B90DA04B6572FD86F67B" box="[540,606,786,812]" pageId="17" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD818098C36B90DA2C06555FE53F618" blockId="17.[151,606,786,847]" box="[151,395,821,847]" pageId="17" pageNumber="18">
(
<figureCitation id="135C048C8C36B90DA2C86555FF28F618" box="[159,240,821,847]" captionStart="FIGURE 4" captionStartId="9.[151,255,1458,1482]" captionTargetBox="[165,1429,214,1420]" captionTargetId="figure@9.[151,1436,197,1437]" captionTargetPageId="9" captionText="FIGURE 4. Dorsal and lateral habitus of Oocyclus species found in the Venezuelan Guiana Shield region. A) O. petra sp. n., B) O. coromoto sp. n.; C) O. floccus sp. n. Scale bar = 2.0 mm." httpUri="https://zenodo.org/record/198431/files/figure.png" pageId="17" pageNumber="18">Figs. 4</figureCitation>
A, 6A, 7, 10)
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C36B90EA2C0651AFA45F7B5" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BD818098C36B90EA2C0651AFA86F7C0" blockId="17.[151,1438,890,2036]" lastBlockId="18.[151,1437,152,2007]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B913C41B8C36B90DA2C0651AFC47F6C3" bold="true" box="[151,927,890,916]" pageId="17" pageNumber="18">
<typeStatus id="54DCA6AB8C36B90DA2C0651AFF00F6C3" box="[151,216,890,916]" pageId="17" pageNumber="18">Type</typeStatus>
material.
<typeStatus id="54DCA6AB8C36B90DA335651AFE38F6C3" box="[354,480,890,916]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F37058998C36B90DA3B9651AFD6AF6C3" box="[494,690,890,916]" name="Venezuela" pageId="17" pageNumber="18">VENEZUELA</collectingCountry>
: Amazonas State:
</emphasis>
male, “
<collectingCountry id="F37058998C36B90DA1A8651AFB66F6C3" box="[1023,1214,890,916]" name="Venezuela" pageId="17" pageNumber="18">VENEZUELA</collectingCountry>
: Amazonas State/
<geoCoordinate id="EE537ECE8C36B90DA2C065C0FEFBF6ED" box="[151,291,928,954]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="5.386783">5°23.207’N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C36B90DA37865C0FE16F6ED" box="[303,462,928,954]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="-67.615364">67°36.922’W</geoCoordinate>
,
<quantity id="4C9FB5EC8C36B90DA38E65C0FDF9F6ED" box="[473,545,928,954]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="17" pageNumber="18" unit="m" value="125.0">125 m</quantity>
/ Tobogan de la Selva;
<date id="FFD93EC98C36B90DA17C65C0FC41F6ED" box="[811,921,928,954]" pageId="17" pageNumber="18" value="2009-01-14">14.i.2009</date>
/ leg. Short, García, Miller, &amp; Joly/ wet rock covered w/ detritus/ VZ09-0114-01F; upstream slide”, “[barcode]/
<collectionCode id="ED7680CC8C36B90DA1A165A5FB93F688" box="[1014,1099,965,991]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="17" pageNumber="18">SEMC</collectionCode>
0853777/ KUNHM-ENT”, “
<typeStatus id="54DCA6AB8C36B90DA2F6658AFE9FF153" box="[161,327,1002,1028]" pageId="17" pageNumber="18" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C67638A8C36B90DA30E658AFE2FF154" box="[345,503,1002,1027]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B913C41B8C36B90DA30E658AFE2FF154" box="[345,503,1002,1027]" italics="true" pageId="17" pageNumber="18">OOCYCLUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C67638A8C36B90DA05E658AFD94F154" box="[521,588,1002,1027]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="petra" status="sp. nov.">
<emphasis id="B913C41B8C36B90DA05E658AFD94F154" box="[521,588,1002,1027]" italics="true" pageId="17" pageNumber="18">petra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22079608C36B90DA002658AFD57F153" box="[597,655,1002,1028]" pageId="17" pageNumber="18" rank="species">sp.n.</taxonomicNameLabel>
/ des.
<bibRefCitation id="EFF665F88C36B90DA088658AFC34F153" author="Short" box="[735,1004,1002,1028]" pageId="17" pageNumber="18" refString="Hall, W. E. &amp; Short, A. E. Z. (2010) A new species of Hydroscapha LeConte from northwestern Venezuela (Coleoptera: Myxophaga: Hydroscaphidae). Zootaxa, 2358, 49 - 56." type="journal article" year="2010">Short &amp; García 2010</bibRefCitation>
” (
<collectionCode id="ED7680CC8C36B90DA65F658AFB87F153" LSID="urn:lsid:biocol.org:col:33834" box="[1032,1119,1002,1028]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="17" pageNumber="18">MIZA</collectionCode>
).
<typeStatus id="54DCA6AB8C36B90DA622658AFB23F153" box="[1141,1275,1002,1028]" pageId="17" pageNumber="18" type="paratype">
<emphasis id="B913C41B8C36B90DA622658AFB23F153" bold="true" box="[1141,1275,1002,1028]" pageId="17" pageNumber="18">Paratypes</emphasis>
</typeStatus>
(1279 exs.):
<emphasis id="B913C41B8C36B90DA2C0626FFDE8F17E" bold="true" box="[151,560,1039,1065]" pageId="17" pageNumber="18">
<collectingCountry id="F37058998C36B90DA2C0626FFE8AF17E" box="[151,338,1039,1065]" name="Venezuela" pageId="17" pageNumber="18">VENEZUELA</collectingCountry>
: Amazonas State:
</emphasis>
<emphasis id="B913C41B8C36B90DA06F6270FCE9F17E" box="[568,817,1040,1065]" italics="true" pageId="17" pageNumber="18">Tobogan de la Selva:</emphasis>
<date id="FFD93EC98C36B90DA16E6270FC6AF17D" box="[825,946,1040,1066]" pageId="17" pageNumber="18" value="1986-02-19">19.ii.1986</date>
, leg. P.J. Spangler, collection #1 (84 exs.,
<collectionCode id="ED7680CC8C36B90DA2C06255FF2BF118" LSID="urn:lsid:biocol.org:col:34871" box="[151,243,1077,1103]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<date id="FFD93EC98C36B90DA3506255FE5CF118" box="[263,388,1077,1103]" pageId="17" pageNumber="18" value="1986-02-22">22.ii.1986</date>
, leg. P.J. Spangler, collection #6 (361 exs.,
<collectionCode id="ED7680CC8C36B90DA1C46255FC37F118" LSID="urn:lsid:biocol.org:col:34871" box="[915,1007,1077,1103]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<date id="FFD93EC98C36B90DA6536255FB58F118" box="[1028,1152,1077,1103]" pageId="17" pageNumber="18" value="1986-02-22">22.ii.1986</date>
, leg. P.J. Spangler, side arm of stream, collection #7 (8 exs.,
<collectionCode id="ED7680CC8C36B90DA003623AFD77F123" LSID="urn:lsid:biocol.org:col:34871" box="[596,687,1114,1140]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<date id="FFD93EC98C36B90DA093623AFC98F123" box="[708,832,1114,1140]" pageId="17" pageNumber="18" value="1986-02-22">22.ii.1986</date>
, leg. P.J. Spangler, edge of lower falls, “bare rock streambed in sunlight; beneath wet leaves at margin” collection #9 (21 exs.,
<collectionCode id="ED7680CC8C36B90DA61762E0FB44F1CD" LSID="urn:lsid:biocol.org:col:34871" box="[1088,1180,1152,1178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<date id="FFD93EC98C36B90DA6E562E0FAE8F1CD" box="[1202,1328,1152,1178]" pageId="17" pageNumber="18" value="1986-02-23">23.ii.1986</date>
, leg. P.J. Spangler, sandy margins, collection #10 (11 exs.,
<collectionCode id="ED7680CC8C36B90DA08C62C5FCEDF1E8" LSID="urn:lsid:biocol.org:col:34871" box="[731,821,1189,1215]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<date id="FFD93EC98C36B90DA11E62C5FC1BF1E8" box="[841,963,1189,1215]" pageId="17" pageNumber="18" value="1986-02-24">24.ii.1986</date>
, leg. P.J. Spangler, collection #13 (1 ex.,
<collectionCode id="ED7680CC8C36B90DA2C062AAFF2CF1B3" LSID="urn:lsid:biocol.org:col:34871" box="[151,244,1226,1252]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<date id="FFD93EC98C36B90DA35D62AAFE52F1B3" box="[266,394,1226,1252]" pageId="17" pageNumber="18" value="1986-02-24">24.ii.1986</date>
, leg. P.J. Spangler, in potholes, collection #13: (4 exs.,
<collectionCode id="ED7680CC8C36B90DA66B62AAFB41F1B3" LSID="urn:lsid:biocol.org:col:34871" box="[1084,1177,1226,1252]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<date id="FFD93EC98C36B90DA6F862AAFAF7F1B3" box="[1199,1327,1226,1252]" pageId="17" pageNumber="18" value="1986-02-25">25.ii.1986</date>
, leg. P.J. Spangler, collection #15 (6 exs.,
<collectionCode id="ED7680CC8C36B90DA0436290FDB6F05D" LSID="urn:lsid:biocol.org:col:34871" box="[532,622,1264,1290]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<date id="FFD93EC98C36B90DA0D56290FD23F05D" box="[642,763,1264,1290]" pageId="17" pageNumber="18" value="1986-02-25">25.ii.1986</date>
, leg. P.J. Spangler, leaves from stream, collection #16 (84 exs.,
<collectionCode id="ED7680CC8C36B90DA2836375FEF6F078" LSID="urn:lsid:biocol.org:col:34871" box="[212,302,1301,1327]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<date id="FFD93EC98C36B90DA3156375FE6BF078" box="[322,435,1301,1327]" pageId="17" pageNumber="18" value="1989-01-25">25.i.1989</date>
, upper seep, leg. P.J. Spangler, R.A. Faitoute, &amp; C.B. Barr (1 ex.,
<collectionCode id="ED7680CC8C36B90DA6EF6375FACAF078" LSID="urn:lsid:biocol.org:col:34871" box="[1208,1298,1301,1327]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<date id="FFD93EC98C36B90DA7716375FA4FF078" box="[1318,1431,1301,1327]" pageId="17" pageNumber="18" value="1989-01-26">26.i.1989</date>
, P.J. Spangler, R.A. Faitoute, &amp; C.B. Barr, seep at upper shelter (41 exs.,
<collectionCode id="ED7680CC8C36B90DA1B5635AFBE4F003" LSID="urn:lsid:biocol.org:col:34871" box="[994,1084,1338,1364]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
); Tobogan:
<date id="FFD93EC98C36B90DA692635AFA9EF003" box="[1221,1350,1338,1364]" pageId="17" pageNumber="18" value="1992-08-04">4.viii.1992</date>
, (1 ex.,
<collectionCode id="ED7680CC8C36B90DA2C06300FF3FF02D" LSID="urn:lsid:biocol.org:col:33834" box="[151,231,1376,1402]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="17" pageNumber="18">MIZA</collectionCode>
); Tobogan:
<date id="FFD93EC98C36B90DA3276300FE0BF02D" box="[368,467,1376,1402]" pageId="17" pageNumber="18" value="2006-01-05">5.i.2006</date>
, leg. A. Short &amp; P. Torres, sandy margin of stream, AS-06-011a (6 exs.); Tobogan:
<date id="FFD93EC98C36B90DA2C063E5FF23F0C8" box="[151,251,1413,1439]" pageId="17" pageNumber="18" value="2006-01-05">5.i.2006</date>
, leg. A. Short, rock pools/wet leaves, AS-06-011c (27 exs.); Tobogan:
<date id="FFD93EC98C36B90DA60463E5FB60F0C8" box="[1107,1208,1413,1439]" pageId="17" pageNumber="18" value="2006-01-05">5.i.2006</date>
, leg. A. Short &amp; P. Torres, wet rock with leaves and detritus, AS-06-011e (197 exs.);
<date id="FFD93EC98C36B90DA1C963CAFBD9F093" box="[926,1025,1450,1476]" pageId="17" pageNumber="18" value="2006-01-07">7.i.2006</date>
, leg. A. Short, AS-06-017 (2 exs.);
<date id="FFD93EC98C36B90DA2C063B0FECFF0BD" box="[151,279,1488,1514]" pageId="17" pageNumber="18" value="2007-09-12">12.ix.2007</date>
, leg. A. Short, M. García, L. Joly, washing main slide, AS-07-007b (8 exs.); Tobogan:
<date id="FFD93EC98C36B90DA77263B0FA4FF0BD" box="[1317,1431,1488,1514]" pageId="17" pageNumber="18" value="2004-01-24">24.i.2004</date>
, leg. M. García (13 exs.);
<date id="FFD93EC98C36B90DA3976395FDE9F358" box="[448,561,1525,1551]" pageId="17" pageNumber="18" value="2009-01-14">14.i.2009</date>
, leg. Short, Miller, &amp; Joly, margin of mail slide, VZ09-0114-01A (8 exs.);
<date id="FFD93EC98C36B90DA2C0607AFED3F363" box="[151,267,1562,1588]" pageId="17" pageNumber="18" value="2009-01-14">14.i.2009</date>
, leg. A. Short, clumps of wet leaves on rock, VZ09-0114-01D (5 exs); same data as
<typeStatus id="54DCA6AB8C36B90DA753607AFAB4F363" box="[1284,1388,1562,1588]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
(83 exs.);
<emphasis id="B913C41B8C36B90DA2B36020FD53F30E" box="[228,651,1600,1625]" italics="true" pageId="17" pageNumber="18">other localities in Amazonas State</emphasis>
:
<quantity id="4C9FB5EC8C36B90DA0CB6020FD33F30D" box="[668,747,1600,1626]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="18" unit="km" value="35.0">35 km</quantity>
SE Puerto Ayacucho, Gavilan area,
<date id="FFD93EC98C36B90DA6F86020FAF7F30D" box="[1199,1327,1600,1626]" pageId="17" pageNumber="18" value="1986-02-20">20.ii.1986</date>
, leg. P.J. Spangler &amp; W. Sanchez, in masses of foam, collection #3 (20 exs.,
<collectionCode id="ED7680CC8C36B90DA1F86005FBD1F328" LSID="urn:lsid:biocol.org:col:34871" box="[943,1033,1637,1663]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
);
<quantity id="4C9FB5EC8C36B90DA6496005FBB1F328" box="[1054,1129,1637,1663]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="17" pageNumber="18" unit="km" value="37.0">37 km</quantity>
SE Puerto Ayacucho, Rio Gavilan,
<date id="FFD93EC98C36B90DA35060EAFE5DF3F3" box="[263,389,1674,1700]" pageId="17" pageNumber="18" value="1986-02-20">20.ii.1986</date>
, leg. P.J. Spangler &amp; W. Sanchez, collection #2 (3 exs.,
<collectionCode id="ED7680CC8C36B90DA66F60EAFB4DF3F3" LSID="urn:lsid:biocol.org:col:34871" box="[1080,1173,1674,1700]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
); ca.
<quantity id="4C9FB5EC8C36B90DA68160EAFAFBF3F3" box="[1238,1315,1674,1700]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="18" unit="km" value="15.0">15 km</quantity>
S. Puerto Ayacucho, nr. Campamento Canturama,
<geoCoordinate id="EE537ECE8C36B90DA02A60D0FCD0F39D" box="[637,776,1712,1738]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="5.510383">5°30.623'N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C36B90DA14260D0FC6BF39D" box="[789,947,1712,1738]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="-67.601814">67°36.109'W</geoCoordinate>
,
<quantity id="4C9FB5EC8C36B90DA19760D0FBD7F39D" box="[960,1039,1712,1738]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="100.0">100 m</quantity>
, isolated seepage on side of rock outcrop, leg. A. Short &amp; M. García, AS-07-009a (34 exs.); same locality but
<date id="FFD93EC98C36B90DA67A60B5FB75F3B8" box="[1069,1197,1749,1775]" pageId="17" pageNumber="18" value="2007-09-14">14.ix.2007</date>
, leg. A. Short &amp; M. García, AS-07-011X (4 exs.); between Puerto Ayacucho &amp; Samariapo, small stream in road culvert,
<date id="FFD93EC98C36B90DA763609AFA4FF243" box="[1332,1431,1786,1812]" pageId="17" pageNumber="18" value="2009-01-06">6.i.2009</date>
, AS-06-013 (7 exs.); between Puerto Ayacucho &amp; Gavilan, 5°32.803',
<geoCoordinate id="EE537ECE8C36B90DA1AE6140FB43F26D" box="[1017,1179,1824,1850]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="-67.44873">67°26.924'W</geoCoordinate>
,
<quantity id="4C9FB5EC8C36B90DA6FF6140FB32F26D" box="[1192,1258,1824,1850]" metricMagnitude="1" metricUnit="m" metricValue="7.7" pageId="17" pageNumber="18" unit="m" value="77.0">77 m</quantity>
, small stream,
<date id="FFD93EC98C36B90DA2C06125FF21F208" box="[151,249,1861,1887]" pageId="17" pageNumber="18" value="2009-01-06">6.i.2009</date>
, leg. A. Short, AS-06-014 (13 exs.); between Puerto Ayacucho &amp; Gavilan,
<geoCoordinate id="EE537ECE8C36B90DA63E6125FB29F208" box="[1129,1265,1861,1887]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="5.573467">5°34.408'N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C36B90DA6AA6125FA4FF208" box="[1277,1431,1861,1887]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="-67.504715">67°30.283'W</geoCoordinate>
,
<quantity id="4C9FB5EC8C36B90DA2C0610AFF0FF2D3" box="[151,215,1898,1924]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="17" pageNumber="18" unit="m" value="66.0">66 m</quantity>
,
<date id="FFD93EC98C36B90DA2B3610AFE91F2D3" box="[228,329,1898,1924]" pageId="17" pageNumber="18" value="2006-01-06">6.i.2006</date>
, small stream, leg. A. Short, AS-06-015 (24 exs.); Communidad Caño Gato on Rio Sipapo,
<date id="FFD93EC98C36B90DA2C061F0FF22F2FD" box="[151,250,1936,1962]" pageId="17" pageNumber="18" value="2006-01-07">7.i.2006</date>
, leg. A. Short &amp; P. Torres, AS-06-016 (1 ex.); S. Puerto Ayacucho, Campamento Nacamtur, isolated rock seepage on outcrop,
<date id="FFD93EC98C36B90DA38861D5FDB1F298" box="[479,617,1973,1999]" pageId="17" pageNumber="18" value="2007-09-14">14.ix.2007</date>
, leg. A. Short, AS-07-013X (68 exs.); Nr. Iboruwa,
<geoCoordinate id="EE537ECE8C36B90DA75261D5FA4FF298" box="[1285,1431,1973,1999]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="5.8069">5°48.414'N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C36B90DA2C061BAFEEAF2A3" box="[151,306,2010,2036]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="-67.43855">67°26.313'W</geoCoordinate>
,
<quantity id="4C9FB5EC8C36B90DA36B61BAFEAFF2A3" box="[316,375,2010,2036]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="18" unit="m" value="80.0">80 m</quantity>
,“Tobogancito”, rock pool with detritus,
<date id="FFD93EC98C36B90DA10261BAFC1EF2A3" box="[853,966,2010,2036]" pageId="17" pageNumber="18" value="2009-01-13">13.i.2009</date>
, leg. A. Short, VZ09-0113-02B (2 exs.); same data but VZ09-0113-
<date id="FFD93EC98C35B90EA3B766F8FDCEF5E5" box="[480,534,152,178]" pageId="18" pageNumber="19" value="2002-10">02X</date>
(5 exs.); River near confluence of Orinoco &amp; Sipapo rivers,
<geoCoordinate id="EE537ECE8C35B90EA75E66F8FA40F5E5" box="[1289,1432,152,178]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="5.0617833">5°03.707'N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C35B90EA2C066DDFEE2F580" box="[151,314,189,215]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="-67.779465">67°46.768'W</geoCoordinate>
, shallow backwaters on rock,
<date id="FFD93EC98C35B90EA0E266DDFCF5F580" box="[693,813,189,215]" pageId="18" pageNumber="19" value="2009-01-15">15.i.2009</date>
, 92 m, leg. A. Short, VZ09-0115-01C (5 exs.); S. Communidad Porvenir,
<geoCoordinate id="EE537ECE8C35B90EA3E96682FD95F5AB" box="[446,589,226,252]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="5.3419">5°20.514'N</geoCoordinate>
;
<geoCoordinate id="EE537ECE8C35B90EA00D6682FD24F5AB" box="[602,764,226,252]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="-67.75525">67°45.315'W</geoCoordinate>
,
<quantity id="4C9FB5EC8C35B90EA15E6682FC93F5AB" box="[777,843,226,252]" metricMagnitude="1" metricUnit="m" metricValue="8.7" pageId="18" pageNumber="19" unit="m" value="87.0">87 m</quantity>
,
<date id="FFD93EC98C35B90EA10F6682FC17F5AB" box="[856,975,226,252]" pageId="18" pageNumber="19" value="2009-01-15">15.i.2009</date>
, leg. Short &amp; García, pool in culvert, VZ09-0115-03A (1 ex.); W. Communidad La Danta,
<geoCoordinate id="EE537ECE8C35B90EA1506768FC57F475" box="[775,911,264,290]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="5.5752">5°34.512'N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C35B90EA1CC6768FBEDF475" box="[923,1077,264,290]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="-67.54163">67°32.498'W</geoCoordinate>
,
<quantity id="4C9FB5EC8C35B90EA6166768FB58F475" box="[1089,1152,264,290]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="19" unit="m" value="80.0">80 m</quantity>
, kick netting,
<date id="FFD93EC98C35B90EA7726768FA4FF475" box="[1317,1431,264,290]" pageId="18" pageNumber="19" value="2009-01-15">15.i.2009</date>
, leg. Short &amp; Camacho, VZ09-0115-04A (2 exs.); Stream along Rio Sipapo,
<geoCoordinate id="EE537ECE8C35B90EA64B674DFB7DF410" box="[1052,1189,301,327]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="4.9308167">4°55.849'N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C35B90EA6E6674DFA93F410" box="[1201,1355,301,327]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="-67.74408">67°44.645'W</geoCoordinate>
,
<quantity id="4C9FB5EC8C35B90EA70F674DFA40F410" box="[1368,1432,301,327]" metricMagnitude="1" metricUnit="m" metricValue="8.7" pageId="18" pageNumber="19" unit="m" value="87.0">87 m</quantity>
,
<date id="FFD93EC98C35B90EA2C06732FED0F43B" box="[151,264,338,364]" pageId="18" pageNumber="19" value="2009-01-16">16.i.2009</date>
, leg. Short, García, Camacho, Miller, &amp; Joly, VZ09-0116-
<date id="FFD93EC98C35B90EA1F96732FC39F43B" box="[942,993,338,364]" pageId="18" pageNumber="19" value="2002-10">02X</date>
(1 ex.);
<emphasis id="B913C41B8C35B90EA6146732FB36F43B" bold="true" box="[1091,1262,338,364]" pageId="18" pageNumber="19">Bolívar State:</emphasis>
Los Pijiguaos,
<geoCoordinate id="EE537ECE8C35B90EA2C06718FEFCF4C5" box="[151,292,376,402]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="6.5936165">6°35.617'N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C35B90EA3666718FE09F4C5" box="[305,465,376,402]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="-66.82063">66°49.238'W</geoCoordinate>
,
<quantity id="4C9FB5EC8C35B90EA3886718FDF8F4C5" box="[479,544,376,402]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="19" unit="m" value="80.0">80 m</quantity>
, outcrop/morichal, marginal algae on rock,
<date id="FFD93EC98C35B90EA66E6718FB76F4C5" box="[1081,1198,376,402]" pageId="18" pageNumber="19" value="2009-01-12">12.i.2009</date>
, leg. Short, García, Camacho, Miller, &amp; Joly, VZ09-0112-01B (16 exs.); same locality but leg. A. Short &amp; M. García,
<date id="FFD93EC98C35B90EA74067FDFA4FF4E0" box="[1303,1431,413,439]" pageId="18" pageNumber="19" value="2007-09-16">16.ix.2007</date>
, AS-07-015 (44 exs.); Rio Caripito,
<geoCoordinate id="EE537ECE8C35B90EA01967A2FD0FF48B" box="[590,727,450,476]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="6.58694">6.58694°N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C35B90EA0B367A2FCA7F48B" box="[740,895,450,476]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="-67.02912">67.02912°W</geoCoordinate>
, detrital pool,
<date id="FFD93EC98C35B90EA66467A2FB72F48B" box="[1075,1194,450,476]" pageId="18" pageNumber="19" value="2009-01-12">12.i.2009</date>
, leg. A. Short &amp; K. Miller, VZ09-0112-02B (1 ex.); ca.
<quantity id="4C9FB5EC8C35B90EA0686788FD52F755" box="[575,650,488,514]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="km" value="25.0">25 km</quantity>
E. El Burro,
<geoCoordinate id="EE537ECE8C35B90EA17E6788FC18F755" box="[809,960,488,514]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="6.21765">06°13.059'N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C35B90EA19A6788FBBFF755" box="[973,1127,488,514]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="-67.24112">67°14.467'W</geoCoordinate>
,
<quantity id="4C9FB5EC8C35B90EA6236788FB6BF755" box="[1140,1203,488,514]" metricMagnitude="1" metricUnit="m" metricValue="6.2" pageId="18" pageNumber="19" unit="m" value="62.0">62 m</quantity>
,
<date id="FFD93EC98C35B90EA6E86788FAE7F755" box="[1215,1343,488,514]" pageId="18" pageNumber="19" value="2007-09-15">15.ix.2007</date>
, leg. A. Short, AS-07-013 (2 exs.); same locality but
<date id="FFD93EC98C35B90EA0F5646DFCCBF770" box="[674,787,525,551]" pageId="18" pageNumber="19" value="2009-01-13">13.i.2009</date>
, leg. Short, García, Camacho, Miller, Joly, VZ09-0113- 01C (11 exs.); same date and locality but VZ09-0113-
<date id="FFD93EC98C35B90EA1136452FCA0F71B" box="[836,888,562,588]" pageId="18" pageNumber="19" value="2001-10">01X</date>
(2 exs.);
<date id="FFD93EC98C35B90EA1A46452FBA7F71B" box="[1011,1151,562,588]" pageId="18" pageNumber="19" value="2008-08-05">5.viii.2008</date>
, ca.
<quantity id="4C9FB5EC8C35B90EA6ED6452FAD1F71B" box="[1210,1289,562,588]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="18" pageNumber="19" unit="km" value="14.0">14 km</quantity>
E Rio Aro,
<geoCoordinate id="EE537ECE8C35B90EA2C06438FEF4F725" box="[151,300,600,626]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="7.689889">7°41'23.6&quot;N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C35B90EA36F6438FE09F725" box="[312,465,600,626]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="-64.03223">64°1'56.0&quot;W</geoCoordinate>
,
<quantity id="4C9FB5EC8C35B90EA3896438FDF4F725" box="[478,556,600,626]" metricMagnitude="2" metricUnit="m" metricValue="1.34" pageId="18" pageNumber="19" unit="m" value="134.0">134 m</quantity>
, rock outcropping, leg. A. Short, AS-08-073 (3 exs.);
<quantity id="4C9FB5EC8C35B90EA6E56438FB37F725" box="[1202,1263,600,626]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="km" value="2.0">2 km</quantity>
E. Cuchivero,
<geoCoordinate id="EE537ECE8C35B90EA2C0641DFEF3F7C0" box="[151,299,637,663]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="7.496472">7°29'47.3&quot;N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C35B90EA360641DFE05F7C0" box="[311,477,637,663]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="-65.86244">65°51'44.8&quot;W</geoCoordinate>
,
<quantity id="4C9FB5EC8C35B90EA3BE641DFDF0F7C0" box="[489,552,637,663]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="19" unit="m" value="45.0">45 m</quantity>
,
<date id="FFD93EC98C35B90EA064641DFD6DF7C0" box="[563,693,637,663]" pageId="18" pageNumber="19" value="2008-08-06">6.viii.2008</date>
, leg. A. Short, M. García, &amp; L. Joly, AS-08-075 (38 exs.).
</paragraph>
<paragraph id="8BD818098C35B90EA29164C2FA45F7B5" blockId="18.[151,1437,152,2007]" pageId="18" pageNumber="19">
<emphasis id="B913C41B8C35B90EA29164C2FC0BF7EB" bold="true" box="[198,979,674,700]" pageId="18" pageNumber="19">
Additional material examined:
<collectingCountry id="F37058998C35B90EA00E64C2FCCCF7EB" box="[601,788,674,700]" name="Venezuela" pageId="18" pageNumber="19">VENEZUELA</collectingCountry>
: Bolívar State:
</emphasis>
Auyantepui, Camp A,
<date id="FFD93EC98C35B90EA6A564C2FA4FF7EB" box="[1266,1431,674,700]" pageId="18" pageNumber="19" value="1988-02-10" valueMax="1988-02-13" valueMin="1988-02-10">10–13.ii.1988</date>
,
<quantity id="4C9FB5EC8C35B90EA2C064A8FF2AF7B5" box="[151,242,712,738]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="18" pageNumber="19" unit="m" value="2100.0">2100 m</quantity>
, leg. L. Joly &amp; A. Chacon (1 ex.,
<collectionCode id="ED7680CC8C35B90EA02D64A8FD11F7B5" LSID="urn:lsid:biocol.org:col:33834" box="[634,713,712,738]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="18" pageNumber="19">MIZA</collectionCode>
), Ayantepui,
<quantity id="4C9FB5EC8C35B90EA13564A8FC65F7B5" box="[866,957,712,738]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="18" pageNumber="19" unit="m" value="1650.0">1650 m</quantity>
,
<date id="FFD93EC98C35B90EA19F64A8FB81F7B5" box="[968,1113,712,738]" pageId="18" pageNumber="19" value="1992-08-26">26.viii.1992</date>
, C.G. Trezzi (1 ex.,
<collectionCode id="ED7680CC8C35B90EA71764A8FA48F7B5" LSID="urn:lsid:biocol.org:col:33834" box="[1344,1424,712,738]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="18" pageNumber="19">MIZA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C35B90EA291648DFA44F6B0" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8BD818098C35B90EA291648DFB48F620" blockId="18.[151,1437,152,2007]" pageId="18" pageNumber="19">
<emphasis id="B913C41B8C35B90EA291648DFE9EF650" bold="true" box="[198,326,749,775]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
Larger species (
<figureCitation id="135C048C8C35B90EA04C648DFDB0F650" box="[539,616,749,775]" captionStart="FIGURE 4" captionStartId="9.[151,255,1458,1482]" captionTargetBox="[165,1429,214,1420]" captionTargetId="figure@9.[151,1436,197,1437]" captionTargetPageId="9" captionText="FIGURE 4. Dorsal and lateral habitus of Oocyclus species found in the Venezuelan Guiana Shield region. A) O. petra sp. n., B) O. coromoto sp. n.; C) O. floccus sp. n. Scale bar = 2.0 mm." httpUri="https://zenodo.org/record/198431/files/figure.png" pageId="18" pageNumber="19">Fig. 4</figureCitation>
A). Posterolateral corners of pronotum strongly spinose. Procoxae spinose. Elytra without rows of larger systematic or serial punctures. Elytral suture not raised in posterior third. Elytral margins not explanate, and without a marginal fringe of long setae. Ground elytral punctation bearing short setae. Elytra with mottled white patches along posterior margin of elytra.
</paragraph>
<paragraph id="8BD818098C35B90EA29165E2FA44F6B0" blockId="18.[151,1437,152,2007]" pageId="18" pageNumber="19">
This species may be distinguished from all other Venezuelan species by the short, fine setae of the ground punctation and the white marginal maculae on the elytra. The latter character is quite variable, and some dark specimens may appear to lack this feature. It is not strongly similar to any other Venezuelan
<taxonomicName id="4C67638A8C35B90EA69B65ADFAE1F6B1" box="[1228,1337,973,998]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B913C41B8C35B90EA69B65ADFAE1F6B1" box="[1228,1337,973,998]" italics="true" pageId="18" pageNumber="19">Oocyclus</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C35B90FA2926592FF06F67B" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="8BD818098C35B90EA2926592FDDEF165" blockId="18.[151,1437,152,2007]" pageId="18" pageNumber="19">
<emphasis id="B913C41B8C35B90EA2926592FEBAF15B" bold="true" box="[197,354,1010,1036]" pageId="18" pageNumber="19">Description.</emphasis>
<emphasis id="B913C41B8C35B90EA33C6592FDC3F15C" box="[363,539,1010,1035]" italics="true" pageId="18" pageNumber="19">Size and form.</emphasis>
Body length =
<quantity id="4C9FB5EC8C35B90EA08C6592FCB5F15B" box="[731,877,1010,1036]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.3" metricValueMin="3.7" pageId="18" pageNumber="19" unit="mm" value="4.5" valueMax="5.3" valueMin="3.7">3.7–5.3 mm</quantity>
(most specimens 4.0–5.0 mm). TL/GW= 1.7. Body oval, moderately convex.
</paragraph>
<paragraph id="8BD818098C35B90EA292625DFBABF060" blockId="18.[151,1437,152,2007]" pageId="18" pageNumber="19">
<emphasis id="B913C41B8C35B90EA292625DFECAF101" box="[197,274,1085,1110]" italics="true" pageId="18" pageNumber="19">Color.</emphasis>
Dorsum of head, pronotum, and elytral disc black with general faint iridescent green sheen. Anterolateral margins of pronotum broadly margined pale yellow on anterior two-thirds. Elytra without distinct iridescent maculae, however the iridescent sheen may be irregularly distributed, giving the appearance of subtle patches; posterior two-fifths of elytra along lateral margin with depigmented (translucent) region, giving the elytra yellow margined appearance. Maxillary and labial palps yellow. Mentum and stipes dark brown, not distinctly paler than ventral face of head. Legs, epipleura, lateral margins of prosternum yellowish brown. Abdominal ventrites dark brown with pale patches laterally on ventrites 1–4.
</paragraph>
<paragraph id="8BD818098C35B90EA2926323FEC9F335" blockId="18.[151,1437,152,2007]" pageId="18" pageNumber="19">
<emphasis id="B913C41B8C35B90EA2926323FED4F00B" box="[197,268,1347,1372]" italics="true" pageId="18" pageNumber="19">Head.</emphasis>
Ground punctation on labrum, clypeus and frons distinctly impressed, distance between punctures 1.5–3.0× the width of one puncture. Systematic punctures on labrum consisting of several indistinct punctures medially. Frons with irregular row of systematic punctures mesad of each eye, bearing fine erect setae. Clypeus with a few very indistinct systematic punctures along anterolateral margins, slightly larger than surrounding punctation. Some ground punctures on head bearing minute setae. Maxillary palps short, about as long as width of labrum; segment 2 slightly bulbous, apical segment slightly longer than penultimate. Labial palps three-fourths as long as width of mentum. Mentum quadrate, anterior margin convex, bearing fine punctures.
</paragraph>
<paragraph id="8BD818098C35B90FA292600DFC1BF580" blockId="18.[151,1437,152,2007]" lastBlockId="19.[151,1437,152,1335]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B913C41B8C35B90EA292600DFEFAF3D1" box="[197,290,1645,1670]" italics="true" pageId="18" pageNumber="19">Thorax.</emphasis>
Ground punctation on pronotum and elytra composed of distinct evenly distributed punctures of generally uniform size with very short, slightly thickened setae. Pronotal systematic punctures with short fine setae, similar to slightly larger in size to ground punctures, blending with larger ground punctures. Lateral margins of pronotum with short indistinct setae; margins of elytra with row of very short coarse spine-like spicules. Posterolateral corners of pronotum weakly to sharply spinose. Sutural punctation on elytra absent or unmodified from ground punctation. Elytra without distinct rows of larger systematic punctures, as they blend uniformly with ground punctation. Prosternum with strongly raised median carina elevated anteromedially, the elevation with two thick spine-like setae. Elevated process of mesoventrite narrow and elongate, more than twice as long as wide; with 5–6 thickened, coarse spine-like setae. Metaventrite with long, narrow glabrous area posteromedially, more than twice as long as wide, length of glabrous area more than two-thirds the length of the metaventrite. Procoxae with dense, fine short pubescence, and throughout with long, coarse spine-like setae. Protibiae each with 6 spine-like setae on dorsal face.
</paragraph>
<paragraph id="8BD818098C34B90FA2926682FA80F475" blockId="19.[151,1437,152,1335]" pageId="19" pageNumber="20">
<emphasis id="B913C41B8C34B90FA2926682FEE3F5AC" box="[197,315,226,251]" italics="true" pageId="19" pageNumber="20">Abdomen.</emphasis>
Ventrites with scattered setae of varying length; each ventrite with irregular rows of long erect setae which are more than twice as long as the longest setae on metaventrite. Aedeagus as in
<figureCitation id="135C048C8C34B90FA68E6768FAE3F475" box="[1241,1339,264,290]" captionStart="FIGURE 6" captionStartId="16.[151,255,1053,1077]" captionTargetBox="[155,1436,199,1024]" captionTargetId="figure@16.[151,1436,194,1032]" captionTargetPageId="16" captionText="FIGURE 6. Aedeagi of Oocyclus species. A) O. petra sp. n., B) O. galbus sp. n.; C) O. pico sp. n. D) O. bolivari sp. n. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/198433/files/figure.png" pageId="19" pageNumber="20">Figure 6</figureCitation>
A.
</paragraph>
<paragraph id="8BD818098C34B90FA292674DFDFBF48B" blockId="19.[151,1437,152,1335]" pageId="19" pageNumber="20">
<emphasis id="B913C41B8C34B90FA292674DFE9CF410" bold="true" box="[197,324,301,327]" pageId="19" pageNumber="20">Variation.</emphasis>
This species has the largest size variation among all species examined here. Even a single population may have individuals at opposite ends of this variation; it is not correlated with geography or time of year. The white margin and spots at the apex of the elytra is nearly always present but varies it its contrast with the surrounding dark coloration; in some specimens the spots are very darkened and thus may appear absent unless carefully examined.
</paragraph>
<paragraph id="8BD818098C34B90FA2926788FF06F67B" blockId="19.[151,1437,152,1335]" pageId="19" pageNumber="20">
We examined two female specimens from Ayantepui, many hundreds of kilometers to the east of the rest of the known populations, that we tentatively assign to this species but which we did not include as
<typeStatus id="54DCA6AB8C34B90FA773646DFA40F770" box="[1316,1432,525,551]" pageId="19" pageNumber="20" type="paratype">paratypes</typeStatus>
. These specimens agree in many aspects with the ‘typical’
<taxonomicName id="4C67638A8C34B90FA1326452FC15F71C" box="[869,973,562,587]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="petra">
<emphasis id="B913C41B8C34B90FA1326452FC15F71C" box="[869,973,562,587]" italics="true" pageId="19" pageNumber="20">
<collectingCountry id="F37058998C34B90FA1326452FCA6F71C" box="[869,894,562,587]" name="Iceland" pageId="19" pageNumber="20">O</collectingCountry>
. petra
</emphasis>
</taxonomicName>
specimens, but are the largest known examples, and have a much broader pale pronotal anterolateral margin. While these characters of size and color are often very variable within species (including
<taxonomicName id="4C67638A8C34B90FA169641DFC7DF7C1" box="[830,933,637,662]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="petra">
<emphasis id="B913C41B8C34B90FA169641DFC7DF7C1" box="[830,933,637,662]" italics="true" pageId="19" pageNumber="20">
<collectingCountry id="F37058998C34B90FA169641DFC8FF7C1" box="[830,855,637,662]" name="Iceland" pageId="19" pageNumber="20">O</collectingCountry>
. petra
</emphasis>
</taxonomicName>
), the fact that these two specimens were collected from different sites and times on the tepui, and yet still share the same outlying features, suggest that perhaps they may represent a distinct species. This possibility is further enhanced by geographic separation from other collecting events. More specimens, including males, are needed before a final determination can be made.
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C34B90FA2926557FB0DF605" box="[197,1237,823,850]" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8BD818098C34B90FA2926557FB0DF605" blockId="19.[151,1437,152,1335]" box="[197,1237,823,850]" pageId="19" pageNumber="20">
<emphasis id="B913C41B8C34B90FA2926557FE8AF606" bold="true" box="[197,338,823,849]" pageId="19" pageNumber="20">Etymology.</emphasis>
<taxonomicName id="4C67638A8C34B90FA30E6558FE42F606" box="[345,410,824,849]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="26" phylum="Arthropoda" rank="species" species="petra">
<emphasis id="B913C41B8C34B90FA30E6558FE42F606" box="[345,410,824,849]" italics="true" pageId="19" pageNumber="20">Petra</emphasis>
</taxonomicName>
, a latinized Greek noun for a large rock. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C34B90FA292653DFDF5F6CB" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BD818098C34B90FA292653DFDF5F6CB" blockId="19.[151,1437,152,1335]" pageId="19" pageNumber="20">
<emphasis id="B913C41B8C34B90FA292653DFEB2F620" bold="true" box="[197,362,861,887]" pageId="19" pageNumber="20">Distribution.</emphasis>
Found in southern
<collectingCountry id="F37058998C34B90FA000653DFD0AF620" box="[599,722,861,887]" name="Venezuela" pageId="19" pageNumber="20">Venezuela</collectingCountry>
(States of Amazonas and Bolívar), along the northwestern edge of the
<collectingCountry id="F37058998C34B90FA37765E2FEACF6CB" box="[288,372,898,924]" name="Guyana" pageId="19" pageNumber="20">Guiana</collectingCountry>
Shield (
<figureCitation id="135C048C8C34B90FA38065E2FDC7F6CB" box="[471,543,898,924]" captionStart="FIGURE 7" captionStartId="22.[151,255,1419,1443]" captionTargetBox="[151,1431,198,1376]" captionTargetId="figure@22.[151,1436,197,1398]" captionTargetPageId="22" captionText="FIGURE 7. Distribution of Oocyclus species in Venezuela." httpUri="https://zenodo.org/record/198434/files/figure.png" pageId="19" pageNumber="20">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C34B90FA29265C7FBF8F060" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8BD818098C34B90FA29265C7FD9EF190" blockId="19.[151,1437,152,1335]" pageId="19" pageNumber="20">
<emphasis id="B913C41B8C34B90FA29265C7FEE6F696" bold="true" box="[197,318,935,961]" pageId="19" pageNumber="20">Remarks.</emphasis>
Within its range,
<taxonomicName id="4C67638A8C34B90FA05A65C8FDB6F696" box="[525,622,936,961]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="petra">
<emphasis id="B913C41B8C34B90FA05A65C8FDB6F696" box="[525,622,936,961]" italics="true" pageId="19" pageNumber="20">
<collectingCountry id="F37058998C34B90FA05A65C8FDFDF696" box="[525,549,936,961]" name="Iceland" pageId="19" pageNumber="20">O</collectingCountry>
. petra
</emphasis>
</taxonomicName>
is frequently encountered in nearly all seepage habitats, sometimes in exceedingly large numbers (e.g.
<figureCitation id="135C048C8C34B90FA04965ADFDAEF6B0" box="[542,630,973,999]" captionStart="FIGURE 10" captionStartId="25.[151,255,1843,1867]" captionTargetBox="[225,1361,943,1821]" captionTargetId="figure@25.[225,1361,943,1821]" captionTargetPageId="25" captionText="FIGURE 10. Habitat of Oocyclus species. A) Amazonas State, 15 km S. Puerto Ayacucho, collecting event AS- 07 - 009 a; locality and habitat of O. petra. B) Amazonas State, Tobogan de la Selva; habitat for O. petra. C) Amazonas State, Tobogan de la Selva, habitat for O. petra and O. coromoto." httpUri="https://zenodo.org/record/198437/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
). This is evidenced by the fact that well over 1000 specimens have been examined, including several series of hundreds of individuals. On more than one occasion in the dry season, so many individuals were observed on riparian rock seepages covered with fallen leaves that we simply could not have collected them all even if we had desired. Found in both wet and dry seasons, it is found in a variety of hygropetric habitats, including wet rock adjacent to streams as well as isolated, temporary seeps (
<figureCitation id="135C048C8C34B90FA33E62E8FE66F1F5" box="[361,446,1160,1186]" captionStart="FIGURE 10" captionStartId="25.[151,255,1843,1867]" captionTargetBox="[225,1361,943,1821]" captionTargetId="figure@25.[225,1361,943,1821]" captionTargetPageId="25" captionText="FIGURE 10. Habitat of Oocyclus species. A) Amazonas State, 15 km S. Puerto Ayacucho, collecting event AS- 07 - 009 a; locality and habitat of O. petra. B) Amazonas State, Tobogan de la Selva; habitat for O. petra. C) Amazonas State, Tobogan de la Selva, habitat for O. petra and O. coromoto." httpUri="https://zenodo.org/record/198437/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
A) that are hundreds of meters from the nearest permanent water source suggesting they may disperse with relative ease.
</paragraph>
<paragraph id="8BD818098C34B90FA29162B2FBF8F060" blockId="19.[151,1437,152,1335]" pageId="19" pageNumber="20">Of particular interest is that this species is one of the few (and perhaps only known) that occurs in streams that are not associated with hygropetric habitats. It is not uncommon to find the species (albeit, not the aforementioned large numbers) in sandy/detrital margins of regional streams.</paragraph>
</subSubSection>
</treatment>
</document>
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