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<document id="9C2176FC6A38A3BC5F7F26E2F74C95FC" ID-DOI="10.11646/zootaxa.2472.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="6507307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629679035506" checkinUser="felipe" docAuthor="Andersen, Trond, Saether, Ole A. &amp; Mendes, Humberto F." docDate="2010" docId="03C0582D991EFFA6BB8BFF6000676B0E" docLanguage="en" docName="zt02472p077.pdf" docOrigin="Zootaxa 2472 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2472.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Allocladius neobilobulatus Andersen &amp; Saether &amp; Mendes 2010, comb. n." docType="treatment" docVersion="4" lastPageNumber="17" masterDocId="FFF920559913FFB6BB1CFFF7045F683C" masterDocTitle="Neotropical Allocladius Kieffer, 1913 and Pseudosmittia Edwards, 1932 (Diptera: Chironomidae) 2472" masterLastPageNumber="77" masterPageNumber="1" pageNumber="14" updateTime="1699094922473" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="986AED18D9E016D13C0724BE462CC095" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2365986C5C987FCE3DD16BA724109C0C">Neotropical Allocladius Kieffer, 1913 and Pseudosmittia Edwards, 1932 (Diptera: Chironomidae) 2472</mods:title>
</mods:titleInfo>
<mods:name id="9057FF0A4C2DAADB3CE8AD55DE059C95" type="personal">
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<mods:namePart id="AFB1300EA219E6AA05817214FA8EFB32">Andersen, Trond</mods:namePart>
</mods:name>
<mods:name id="C9027178AF8C64D67F4881564E66559F" type="personal">
<mods:role id="E508862E0EF01D785598B10062756A91">
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<mods:namePart id="86974F88C898ACAF13DAFE23B3AC842B">Saether, Ole A.</mods:namePart>
</mods:name>
<mods:name id="81165834A88D8D126A0606652F0D9832" type="personal">
<mods:role id="49176F2D930DFA923A1E99B2BD5A9B6C">
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</mods:role>
<mods:namePart id="7D79DCA60878F24AD8F8531338B37AC2">Mendes, Humberto F.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1F64DD583680BD743D6F5D797FCF03DA">text</mods:typeOfResource>
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<mods:title id="A2A0D9F77B90D0606063AE4DDCB7BF07">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E59D4E6B72EED8748C599A5E2075B9CA">
<mods:date id="993DAB88022AA6787D19DEBD5209548A">2010</mods:date>
<mods:detail id="DE35C86D17DBA614C4E68E3CC0682B8C" type="pubDate">
<mods:number id="2A07B4D1200C6B19007633817310FB3B">2010-05-14</mods:number>
</mods:detail>
<mods:detail id="EC5D68A255AAE42CAED6521A7C72970B" type="volume">
<mods:number id="C9BC9550CD73448DA96069C0CF65E30D">2472</mods:number>
</mods:detail>
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<mods:number id="8725BC515040D5B906CD91CA2AD9B2C3">1</mods:number>
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<mods:url id="53BE044CF091D1CD23D7D19717FECB0D">https://biotaxa.org/Zootaxa/article/view/zootaxa.2472.1.1</mods:url>
</mods:location>
<mods:classification id="7A8C12D3B76C4D1527248920465656DE">journal article</mods:classification>
<mods:identifier id="354F77034C41DB0D6AC236A63262D584" type="DOI">10.11646/zootaxa.2472.1.1</mods:identifier>
<mods:identifier id="A383DFB38C0D212146663CE80DBC5FD5" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="2A8EDAE6FADC53BF9E8728F4785DA0DB" type="Zenodo-Dep">6507307</mods:identifier>
</mods:mods>
<treatment id="03C0582D991EFFA6BB8BFF6000676B0E" ID-DOI="http://doi.org/10.5281/zenodo.6509008" ID-Zenodo-Dep="6509008" LSID="urn:lsid:plazi:treatment:03C0582D991EFFA6BB8BFF6000676B0E" httpUri="http://treatment.plazi.org/id/03C0582D991EFFA6BB8BFF6000676B0E" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
<subSubSection id="C373BAB0991EFFBBBB8BFF6006F2688D" box="[151,685,151,177]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BD6E93B991EFFBBBB8BFF6006F2688D" blockId="13.[151,685,151,212]" box="[151,685,151,177]" pageId="13" pageNumber="14">
<heading id="D09E5E57991EFFBBBB8BFF6006F2688D" bold="true" box="[151,685,151,177]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B91D3529991EFFBBBB8BFF6006F2688D" bold="true" box="[151,685,151,177]" pageId="13" pageNumber="14">
<taxonomicName id="4C6992B8991EFFBBBB8BFF600664688D" authority="(Paggi)" authorityName="Andersen &amp; Saether &amp; Mendes" authorityYear="2010" baseAuthorityName="Paggi" box="[151,571,151,177]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="neobilobulatus" status="comb. nov.">
<emphasis id="B91D3529991EFFBBBB8BFF600585688D" bold="true" box="[151,474,151,177]" italics="true" pageId="13" pageNumber="14">Allocladius neobilobulatus</emphasis>
(Paggi)
</taxonomicName>
<taxonomicNameLabel id="A22E8852991EFFBBB95EFF6006F2688D" box="[578,685,151,177]" pageId="13" pageNumber="14" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373BAB0991EFFBBBB8BFF4D057268E8" box="[151,301,186,212]" pageId="13" pageNumber="14" type="description">
<paragraph id="8BD6E93B991EFFBBBB8BFF4D057268E8" blockId="13.[151,685,151,212]" box="[151,301,186,212]" pageId="13" pageNumber="14">
(
<figureCitation id="1352F5BE991EFFBBBB83FF4D057A68E8" box="[159,293,186,212]" captionStart-0="FIGURES 27–33" captionStart-1="FIGURES 34–37" captionStartId-0="14.[151,269,1755,1779]" captionStartId-1="15.[151,269,1962,1986]" captionTargetBox-0="[243,1338,159,1696]" captionTargetBox-1="[274,1276,417,1927]" captionTargetId-0="figure-117@14.[226,1360,135,1723]" captionTargetId-1="figure-127@15.[226,1360,375,1963]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 27–33. Allocladius neobilobulatus (Paggi, 1993) comb. n., male. 27—tentorium, stipes, and cibarial pump; 28—palp; 29—thorax; 30—wing; 31—hypopygium, dorsal aspect; 32—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 33—pars ventralis." captionText-1="FIGURES 34–37. Allocladius neobilobulatus (Paggi, 1993) comb. n., female pupa. 34—frontal apotome; 35—thorax; 36—tergites; 37—sternites with detail of the papilla on the genital sac." figureDoi-0="http://doi.org/10.5281/zenodo.6507317" figureDoi-1="http://doi.org/10.5281/zenodo.6507319" httpUri-0="https://zenodo.org/record/6507317/files/figure.png" httpUri-1="https://zenodo.org/record/6507319/files/figure.png" pageId="13" pageNumber="14">Figs 27–37</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C373BAB0991EFFBBBB8BFEF206D36920" box="[151,652,261,284]" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8BD6E93B991EFFBBBB8BFEF206D36920" blockId="13.[151,652,261,284]" box="[151,652,261,284]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB79CE15991EFFBBBB8BFEF206D36920" box="[151,652,261,284]" pageId="13" pageNumber="14">
<treatmentCitation id="0AC8CF2A991EFFBBBB8BFEF206D76920" author="Paggi, A. C." box="[151,648,261,284]" page="172" pageId="13" pageNumber="14" year="1993">
<taxonomicName id="4C6992B8991EFFBBBB8BFEF206D76920" ID-CoL="4PBMB" authority="Paggi, 1993: 172" authorityName="Paggi" authorityPageNumber="172" authorityYear="1993" box="[151,648,261,284]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="neobilobulata">
<emphasis id="B91D3529991EFFBBBB8BFEF205976920" box="[151,456,261,284]" italics="true" pageId="13" pageNumber="14">Pseudosmittia neobilobulata</emphasis>
<bibRefCitation id="EFF894CA991EFFBBBAD3FEF206D76920" author="Paggi, A. C." box="[463,648,261,284]" pageId="13" pageNumber="14" pagination="171 - 174" refId="ref44092" refString="Paggi, A. C. (1993). Redescription of Pseudosmittia bilobulata (Edw.) n. comb. (= Spaniotoma (Smittia) bilobulata Edwards 1931) and description of P. neobilobulata sp. n. (Diptera: Chironomidae). Annales de Limnologie, 29 (2), 171 - 174." type="journal article" year="1993">Paggi, 1993: 172</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C373BAB0991EFFA6BB8BFEBD04BE6A4B" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BD6E93B991EFFBBBB8BFEBD008769C6" blockId="13.[151,1436,330,1296]" pageId="13" pageNumber="14">
<emphasis id="B91D3529991EFFBBBB8BFEBD05D36958" bold="true" box="[151,396,330,356]" pageId="13" pageNumber="14">Material examined.</emphasis>
<materialsCitation id="3B01E366991EFFBBBA88FEBD04B469B6" collectingDate="1989-06-15" collectionCode="ILLP" collectorName="A. Paggi" country="Argentina" location="Laguna de Lobos" pageId="13" pageNumber="14" specimenCount="2" specimenCount-male="2" stateProvince="Ciudad Autonoma de Buenos Aires" typeStatus="paratype">
<emphasis id="B91D3529991EFFBBBA88FEBD06B76958" bold="true" box="[404,744,330,356]" pageId="13" pageNumber="14">
<collectingCountry id="F37EA9AB991EFFBBBA88FEBD0677695F" box="[404,552,330,356]" name="Argentina" pageId="13" pageNumber="14">ARGENTINA</collectingCountry>
:
<collectingRegion id="49AD27D9991EFFBBB927FEBD06BD6958" box="[571,738,330,356]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="13" pageNumber="14">Buenos Aires</collectingRegion>
,
</emphasis>
<location id="8EB6BFE0991EFFBBB9ECFEBD079E6958" LSID="urn:lsid:plazi:treatment:03C0582D991EFFA6BB8BFF6000676B0E:8EB6BFE0991EFFBBB9ECFEBD079E6958" box="[752,961,330,356]" country="Argentina" name="Laguna de Lobos" pageId="13" pageNumber="14" stateProvince="Ciudad Autonoma de Buenos Aires">Laguna de Lobos</location>
,
<specimenCount id="9D6F22B2991EFFBBB8D2FEBD00416958" box="[974,1054,330,356]" pageId="13" pageNumber="14" type="male">2 male</specimenCount>
<typeStatus id="54D25799991EFFBBBF3BFEBD00C56958" box="[1063,1178,330,356]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
,
<date id="FFD7CFFB991EFFBBBFBBFEBD01796958" box="[1191,1318,330,356]" pageId="13" pageNumber="14" value="1989-06-15">
<collectingDate id="EF933613991EFFBBBFBBFEBD01796958" box="[1191,1318,330,356]" pageId="13" pageNumber="14" value="1989-06-15">15.vi.1989</collectingDate>
</date>
,
<collectorName id="269C8CED991EFFBBBE28FEBD01C36958" box="[1332,1436,330,356]" pageId="13" pageNumber="14">A. Paggi</collectorName>
(
<collectionCode id="ED7871FE991EFFBBBB83FE8704BC69B6" box="[159,227,368,394]" pageId="13" pageNumber="14">ILLP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B01E366991EFFBBBBE6FE9806CC6993" collectingDate="1998-11-18" collectionCode="ZMBN" collectorName="T. Andersen" country="Chile" elevation="212" latitude="-35.15285" location="Rio Claro South of Molina" longLatPrecision="1" longitude="-71.28423" municipality="Region" pageId="13" pageNumber="14" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B91D3529991EFFBBBBE6FE9805B569B5" bold="true" box="[250,490,367,393]" pageId="13" pageNumber="14">
<collectingCountry id="F37EA9AB991EFFBBBBE6FE98051569B4" box="[250,330,367,393]" name="Chile" pageId="13" pageNumber="14">CHILE</collectingCountry>
:
<collectingMunicipality id="6BB27341991EFFBBBA43FE9805E869B5" box="[351,439,367,393]" pageId="13" pageNumber="14">Región</collectingMunicipality>
VI,
</emphasis>
<location id="8EB6BFE0991EFFBBBAE8FE87071F69B6" LSID="urn:lsid:plazi:treatment:03C0582D991EFFA6BB8BFF6000676B0E:8EB6BFE0991EFFBBBAE8FE87071F69B6" box="[500,832,368,394]" country="Chile" latitude="-35.15285" longLatPrecision="1" longitude="-71.28423" municipality="Region" name="Rio Claro South of Molina" pageId="13" pageNumber="14">Río Claro South of Molina</location>
,
<geoCoordinate id="EE5D8FFC991EFFBBB853FE87078169B6" box="[847,990,368,394]" degrees="35" direction="south" minutes="09.171" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="-35.15285">35º09.171'S</geoCoordinate>
,
<geoCoordinate id="EE5D8FFC991EFFBBB8F2FE8700DA69B6" box="[1006,1157,368,394]" degrees="71" direction="west" minutes="17.054" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-71.28423">71º17.054'W</geoCoordinate>
,
<elevation id="00440E08991EFFBBBF89FE87017969B6" box="[1173,1318,368,394]" metricMagnitude="2" metricUnit="m" metricValue="2.12" pageId="13" pageNumber="14" unit="m" value="212.0">
<quantity id="4C9144DE991EFFBBBF89FE8700BC69B6" box="[1173,1251,368,394]" metricMagnitude="2" metricUnit="m" metricValue="2.12" pageId="13" pageNumber="14" unit="m" value="212.0">212 m</quantity>
a.s.l.
</elevation>
,
<specimenCount id="9D6F22B2991EFFBBBE2BFE8701C869B6" box="[1335,1431,368,394]" pageId="13" pageNumber="14" type="male">2 males</specimenCount>
,
<date id="FFD7CFFB991EFFBBBB8BFE6205486993" box="[151,279,405,431]" pageId="13" pageNumber="14" value="1998-11-18">
<collectingDate id="EF933613991EFFBBBB8BFE6205486993" box="[151,279,405,431]" pageId="13" pageNumber="14" value="1998-11-18">18.xi.1998</collectingDate>
</date>
,
<collectingMethod id="5228912C991EFFBBBA3EFE6205D66993" box="[290,393,405,431]" pageId="13" pageNumber="14">hand net</collectingMethod>
,
<collectorName id="269C8CED991EFFBBBA8FFE62067E6993" box="[403,545,405,431]" pageId="13" pageNumber="14">T. Andersen</collectorName>
(
<collectionCode id="ED7871FE991EFFBBB92CFE6206D46993" box="[560,651,405,431]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="13" pageNumber="14">ZMBN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B01E366991EFFBBB982FE6206C769E8" collectingDate="1996-02-14" collectionCode="ZMBN" collectorName="T. Andersen" country="Chile" location="Laguna Cea" municipality="Region" pageId="13" pageNumber="14" specimenCount="4" specimenCount-male="4" typeStatus="paratype">
<emphasis id="B91D3529991EFFBBB982FE62077B6993" bold="true" box="[670,804,405,431]" pageId="13" pageNumber="14">
<collectingMunicipality id="6BB27341991EFFBBB982FE6206AA6993" box="[670,757,405,431]" pageId="13" pageNumber="14">Región</collectingMunicipality>
XI,
</emphasis>
<location id="8EB6BFE0991EFFBBB837FE6207E56993" LSID="urn:lsid:plazi:treatment:03C0582D991EFFA6BB8BFF6000676B0E:8EB6BFE0991EFFBBB837FE6207E56993" box="[811,954,405,431]" country="Chile" municipality="Region" name="Laguna Cea" pageId="13" pageNumber="14">Laguna Cea</location>
,
<locationDeviation id="9A64F019991EFFBBB8D9FE62016F6993" box="[965,1328,405,431]" location="Southwest of Coihaique" pageId="13" pageNumber="14">
<quantity id="4C9144DE991EFFBBB8D9FE6200516993" box="[965,1038,405,431]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="km" value="20.0">20 km</quantity>
Southwest of Coihaique
</locationDeviation>
,
<specimenCount id="9D6F22B2991EFFBBBE27FE6201C86993" box="[1339,1431,405,431]" pageId="13" pageNumber="14" type="male">4 males</specimenCount>
,
<date id="FFD7CFFB991EFFBBBB8BFE4D054F69E8" box="[151,272,442,468]" pageId="13" pageNumber="14" value="1996-02-14">
<collectingDate id="EF933613991EFFBBBB8BFE4D054F69E8" box="[151,272,442,468]" pageId="13" pageNumber="14" value="1996-02-14">14.ii.1996</collectingDate>
</date>
,
<collectingMethod id="5228912C991EFFBBBA02FE4D05D869E8" box="[286,391,442,468]" pageId="13" pageNumber="14">hand net</collectingMethod>
,
<collectorName id="269C8CED991EFFBBBA88FE4D067B69E8" box="[404,548,442,468]" pageId="13" pageNumber="14">T. Andersen</collectorName>
(
<collectionCode id="ED7871FE991EFFBBB929FE4D06CE69E8" box="[565,657,442,468]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="13" pageNumber="14">ZMBN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B01E366991EFFBBB9B9FE4D008B69C6" collectingDate="1976-07-14" collectionCode="ZSM" collectorName="E. J. Fittkau" country="Bolivia" elevation="3" location="Between Oruru" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Cochabamba" typeStatus="paratype">
<emphasis id="B91D3529991EFFBBB9B9FE4D074669E8" bold="true" box="[677,793,442,468]" pageId="13" pageNumber="14">
<collectingCountry id="F37EA9AB991EFFBBB9B9FE4D075069EF" box="[677,783,442,468]" name="Bolivia" pageId="13" pageNumber="14">BOLIVIA</collectingCountry>
:
</emphasis>
<location id="8EB6BFE0991EFFBBB83EFE4D078869E8" LSID="urn:lsid:plazi:treatment:03C0582D991EFFA6BB8BFF6000676B0E:8EB6BFE0991EFFBBB83EFE4D078869E8" box="[802,983,442,468]" country="Bolivia" name="Between Oruru" pageId="13" pageNumber="14" stateProvince="Cochabamba">Between Oruru</location>
and
<collectingRegion id="49AD27D9991EFFBBBF0FFE4D00F069E8" box="[1043,1199,442,468]" country="Bolivia" name="Cochabamba" pageId="13" pageNumber="14">Cochabamba</collectingRegion>
, side arm of a dam,
<elevation id="00440E08991EFFBBBB8BFE17056969C6" box="[151,310,480,506]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="14" unit="m" value="3.0">
<quantity id="4C9144DE991EFFBBBB8BFE1704A869C6" box="[151,247,480,506]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="14" unit="m" value="3.0">3.000 m</quantity>
a.s.l.
</elevation>
,
<specimenCount id="9D6F22B2991EFFBBBA5FFE1705CA69C6" box="[323,405,480,506]" pageId="13" pageNumber="14" type="male">1 male</specimenCount>
, 1 mature male pupa, 2 pupal exuviae,
<date id="FFD7CFFB991EFFBBB843FE1707B869C6" box="[863,999,480,506]" pageId="13" pageNumber="14" value="1976-07-14">
<collectingDate id="EF933613991EFFBBB843FE1707B869C6" box="[863,999,480,506]" pageId="13" pageNumber="14" value="1976-07-14">14.vii.1976</collectingDate>
</date>
,
<collectorName id="269C8CED991EFFBBB8E8FE17002569C6" box="[1012,1146,480,506]" pageId="13" pageNumber="14">E.J. Fittkau</collectorName>
(
<collectionCode id="ED7871FE991EFFBBBF96FE17009369C6" box="[1162,1228,480,506]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="13" pageNumber="14" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD6E93B991EFFBBBBDAFDF207756AB3" blockId="13.[151,1436,330,1296]" pageId="13" pageNumber="14">
<emphasis id="B91D3529991EFFBBBBDAFDF205806A23" bold="true" box="[198,479,517,543]" pageId="13" pageNumber="14">Diagnostic characters.</emphasis>
The two broad, large and lobe-like pars ventralis will separate the male imago from all other
<taxonomicName id="4C6992B8991EFFBBBA58FDDD06726A7F" box="[324,557,554,580]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91D3529991EFFBBBA58FDDD05966A7F" box="[324,457,554,579]" italics="true" pageId="13" pageNumber="14">Allocladius</emphasis>
species.
</taxonomicName>
The pupa is very similar to that of the Holarctic
<taxonomicName id="4C6992B8991EFFBBBF65FDDD01366A78" authority="(Kieffer)" baseAuthorityName="Kieffer" baseAuthorityYear="1921" box="[1145,1385,554,580]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="nanseni">
<emphasis id="B91D3529991EFFBBBF65FDDD00A96A7F" box="[1145,1270,554,579]" italics="true" pageId="13" pageNumber="14">A. nanseni</emphasis>
(Kieffer)
</taxonomicName>
, but can be distinguished by having a papilla on the genital sac which is nearly twice as long as wide and more numerous spinules on the posterior sternal conjunctives.
</paragraph>
<paragraph id="8BD6E93B991EFFBBBBDAFD6D010B6AE6" blockId="13.[151,1436,330,1296]" pageId="13" pageNumber="14">
<emphasis id="B91D3529991EFFBBBBDAFD6D055A6A88" bold="true" box="[198,261,666,692]" pageId="13" pageNumber="14">Male</emphasis>
(n = 8–10, except when otherwise stated). Total length 1.85–2.74,
<quantity id="4C9144DE991EFFBBBF0FFD6D00226A88" box="[1043,1149,666,692]" metricMagnitude="-3" metricUnit="m" metricValue="2.11" pageId="13" pageNumber="14" unit="mm" value="2.11">2.11 mm</quantity>
. Wing length 1.08–1.67,
<quantity id="4C9144DE991EFFBBBB8BFD37055D6AE6" box="[151,258,704,730]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="14" unit="mm" value="1.25">1.25 mm</quantity>
. Total length / wing length 1.45–1.82, 1.66. Wing length / length of profemur 2.82–3.16, 2.96.
</paragraph>
<paragraph id="8BD6E93B991EFFBBBBDAFD1206496AC3" blockId="13.[151,1436,330,1296]" box="[198,534,741,767]" pageId="13" pageNumber="14">
<emphasis id="B91D3529991EFFBBBBDAFD1205116AC2" box="[198,334,741,766]" italics="true" pageId="13" pageNumber="14">Coloration.</emphasis>
Brownish black.
</paragraph>
<paragraph id="8BD6E93B991EFFBBBBDAFCFD01D16B86" blockId="13.[151,1436,330,1296]" pageId="13" pageNumber="14">
<emphasis id="B91D3529991EFFBBBBDAFCFD05516B1F" box="[198,270,778,803]" italics="true" pageId="13" pageNumber="14">Head.</emphasis>
AR 0.82–1.23, 0.99. Ultimate flagellomere 281–443, 340 µm long. Temporal setae 4–9, 6; consisting of 0–3, 2 inner verticals; 0–3, 2 outer verticals and 0–4, 2 postorbitals. Clypeus with 4–9, 6 setae. Tentorium, stipes, and cibarial pump as in
<figureCitation id="1352F5BE991EFFBBB950FCA2069E6B53" box="[588,705,853,879]" captionStart="FIGURES 27–33" captionStartId="14.[151,269,1755,1779]" captionTargetBox="[243,1338,159,1696]" captionTargetId="figure-117@14.[226,1360,135,1723]" captionTargetPageId="14" captionText="FIGURES 27–33. Allocladius neobilobulatus (Paggi, 1993) comb. n., male. 27—tentorium, stipes, and cibarial pump; 28—palp; 29—thorax; 30—wing; 31—hypopygium, dorsal aspect; 32—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 33—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507317" httpUri="https://zenodo.org/record/6507317/files/figure.png" pageId="13" pageNumber="14">Figure 27</figureCitation>
. Tentorium 82–127, 106 µm long; 20–30, 24 µm wide. Stipes 93–127, 105 µm long; 27–39, 32 (6) µm wide. Palpomere (
<figureCitation id="1352F5BE991EFFBBB87DFC8D07E86BA8" box="[865,951,890,916]" captionStart="FIGURES 27–33" captionStartId="14.[151,269,1755,1779]" captionTargetBox="[243,1338,159,1696]" captionTargetId="figure-117@14.[226,1360,135,1723]" captionTargetPageId="14" captionText="FIGURES 27–33. Allocladius neobilobulatus (Paggi, 1993) comb. n., male. 27—tentorium, stipes, and cibarial pump; 28—palp; 29—thorax; 30—wing; 31—hypopygium, dorsal aspect; 32—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 33—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507317" httpUri="https://zenodo.org/record/6507317/files/figure.png" pageId="13" pageNumber="14">Fig. 28</figureCitation>
) lengths (in µm): 16–25, 20; 27–43, 35; 45–75, 56; 52–82, 59; 70–109, 83. Third palpomere with 1 lanceolate sensilla clavata; 9–11, 11 (7) µm long.
</paragraph>
<paragraph id="8BD6E93B991EFFBBBBDAFC3205756C16" blockId="13.[151,1436,330,1296]" pageId="13" pageNumber="14">
<emphasis id="B91D3529991EFFBBBBDAFC3205446BE2" box="[198,283,965,990]" italics="true" pageId="13" pageNumber="14">Thorax</emphasis>
(
<figureCitation id="1352F5BE991EFFBBBA37FC3205DD6BE3" box="[299,386,965,991]" captionStart="FIGURES 27–33" captionStartId="14.[151,269,1755,1779]" captionTargetBox="[243,1338,159,1696]" captionTargetId="figure-117@14.[226,1360,135,1723]" captionTargetPageId="14" captionText="FIGURES 27–33. Allocladius neobilobulatus (Paggi, 1993) comb. n., male. 27—tentorium, stipes, and cibarial pump; 28—palp; 29—thorax; 30—wing; 31—hypopygium, dorsal aspect; 32—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 33—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507317" httpUri="https://zenodo.org/record/6507317/files/figure.png" pageId="13" pageNumber="14">Fig. 29</figureCitation>
). Median antepronotal lobes well developed; antepronotum with 0–1, 1 lateral seta. Dorsocentrals 7–8, 7; acrostichals 4–8, 6, simple or scalpellate; prealars 2–3, 3; supraalar 0–1, 0. Scutellum with 5–8, 7 setae.
</paragraph>
<paragraph id="8BD6E93B991EFFBBBBDAFBC2010D6C4B" blockId="13.[151,1436,330,1296]" pageId="13" pageNumber="14">
<emphasis id="B91D3529991EFFBBBBDAFBC2055D6C72" box="[198,258,1077,1102]" italics="true" pageId="13" pageNumber="14">Wing</emphasis>
(
<figureCitation id="1352F5BE991EFFBBBA0EFBC205356C73" box="[274,362,1077,1103]" captionStart="FIGURES 27–33" captionStartId="14.[151,269,1755,1779]" captionTargetBox="[243,1338,159,1696]" captionTargetId="figure-117@14.[226,1360,135,1723]" captionTargetPageId="14" captionText="FIGURES 27–33. Allocladius neobilobulatus (Paggi, 1993) comb. n., male. 27—tentorium, stipes, and cibarial pump; 28—palp; 29—thorax; 30—wing; 31—hypopygium, dorsal aspect; 32—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 33—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507317" httpUri="https://zenodo.org/record/6507317/files/figure.png" pageId="13" pageNumber="14">Fig. 30</figureCitation>
). VR 1.27–1.72, 1.35. Anal lobe projecting. Costal extension 0–50, 13 µm long. R
<subScript id="17EDEB7E991EFFBBBE30FBB201196C6F" attach="left" box="[1324,1350,1093,1107]" fontSize="6" pageId="13" pageNumber="14">4+5</subScript>
ending above apex of M
<subScript id="17EDEB7E991EFFBBBA40FB9A05296C47" attach="left" box="[348,374,1133,1147]" fontSize="6" pageId="13" pageNumber="14">3+4</subScript>
; Cu
<subScript id="17EDEB7E991EFFBBBABBFB9A05F06C47" attach="left" box="[423,431,1133,1147]" fontSize="6" pageId="13" pageNumber="14">1</subScript>
, slightly sinuate. Brachiolum with 1 seta; R with 2–3, 2 setae; other veins bare.
</paragraph>
<paragraph id="8BD6E93B991EFFBBBBDAFB71074A6D2C" blockId="13.[151,1436,330,1296]" pageId="13" pageNumber="14">
<emphasis id="B91D3529991EFFBBBBDAFB7104A26CA3" box="[198,253,1158,1183]" italics="true" pageId="13" pageNumber="14">Legs</emphasis>
. Spur of fore tibia 34–47, 39 µm long; spurs of mid tibia 18–34, 21 µm and 11–19, 14 µm long; of hind tibia 31–43, 36 µm and 9–18, 14 µm long. Width at apex of fore tibia 22–29, 25 µm; of mid tibia 20–34, 25 µm; of hind tibia 27–45, 35 µm. Comb of 11–14, 13 setae; longest 20–40, 28 µm long; shortest 15–19, 16 µm long. Length and proportions of legs as in
<tableCitation id="C6EBDC80991EFFBBB9ABFB01074F6D2C" box="[695,784,1270,1296]" captionStart="TABLE 5" captionStartId="13.[151,241,1355,1379]" captionText="TABLE 5. Length (in µm) and proportions of legs of Allocladius neobilobulatus (Paggi), male (n = 9)." pageId="13" pageNumber="14">Table 5</tableCitation>
.
</paragraph>
<caption id="DF16B9B3991EFFBBBB8BFABC00BE6D5E" box="[151,1249,1355,1379]" pageId="13" pageNumber="14" startId="13.[151,241,1355,1379]">
<paragraph id="8BD6E93B991EFFBBBB8BFABC00BE6D5E" blockId="13.[151,1249,1355,1379]" box="[151,1249,1355,1379]" pageId="13" pageNumber="14">
<emphasis id="B91D3529991EFFBBBB8BFABC05546D5F" bold="true" box="[151,267,1355,1379]" pageId="13" pageNumber="14">TABLE 5.</emphasis>
Length (in µm) and proportions of legs of
<taxonomicName id="4C6992B8991EFFBBB9CBFABC00126D5E" authority="(Paggi)" authorityName="Andersen &amp; Saether &amp; Mendes" authorityYear="2010" baseAuthorityName="Andersen &amp; Saether &amp; Mendes" baseAuthorityYear="2010" box="[727,1101,1355,1378]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="neobilobulatus">
<emphasis id="B91D3529991EFFBBB9CBFABC07AA6D5E" box="[727,1013,1355,1378]" italics="true" pageId="13" pageNumber="14">Allocladius neobilobulatus</emphasis>
(Paggi)
</taxonomicName>
, male (n = 9).
</paragraph>
</caption>
<paragraph id="8BD6E93B991EFFBBBA78FA72014E6F86" pageId="13" pageNumber="14">
<table id="F9691B9B991E0049BBB9FA7501006F87" box="[165,1375,1410,1979]" gridcols="5" gridrows="12" pageId="13" pageNumber="14">
<tr id="3559EB79991E0049BBB9FA7501006DA1" box="[165,1375,1410,1437]" gridrow="0" pageId="13" pageNumber="14" rowspan-0="1">
<th id="76888205991E0049BA78FA7506546DA1" box="[356,523,1410,1437]" gridcol="1" gridrow="0" pageId="13" pageNumber="14">fe</th>
<th id="76888205991E0049B99DFA7507206DA1" box="[641,895,1410,1437]" gridcol="2" gridrow="0" pageId="13" pageNumber="14">ti</th>
<th id="76888205991E0049B881FA75006F6DA1" box="[925,1072,1410,1437]" gridcol="3" gridrow="0" pageId="13" pageNumber="14">ta1</th>
<th id="76888205991E0049BF8FFA7501006DA1" box="[1171,1375,1410,1437]" gridcol="4" gridrow="0" pageId="13" pageNumber="14">ta2</th>
</tr>
<tr id="3559EB79991E0049BBB9FA4401006DF3" box="[165,1375,1459,1487]" gridrow="1" pageId="13" pageNumber="14">
<th id="76888205991E0049BBB9FA4404E56DF3" box="[165,186,1459,1487]" gridcol="0" gridrow="1" pageId="13" pageNumber="14">p1</th>
<td id="76888205991E0049BA78FA4406546DF3" box="[356,523,1459,1487]" gridcol="1" gridrow="1" pageId="13" pageNumber="14">367–522, 419</td>
<td id="76888205991E0049B99DFA4407206DF3" box="[641,895,1459,1487]" gridcol="2" gridrow="1" pageId="13" pageNumber="14">450–698, 536</td>
<td id="76888205991E0049B881FA44006F6DF3" box="[925,1072,1459,1487]" gridcol="3" gridrow="1" pageId="13" pageNumber="14">194–241, 216</td>
<td id="76888205991E0049BF8FFA4401006DF3" box="[1171,1375,1459,1487]" gridcol="4" gridrow="1" pageId="13" pageNumber="14">122–191, 144</td>
</tr>
<tr id="3559EB79991E0049BBB9FA1301006E3C" box="[165,1375,1508,1536]" gridrow="2" pageId="13" pageNumber="14">
<th id="76888205991E0049BBB9FA1304E56E3C" box="[165,186,1508,1536]" gridcol="0" gridrow="2" pageId="13" pageNumber="14">p2</th>
<td id="76888205991E0049BA78FA1306546E3C" box="[356,523,1508,1536]" gridcol="1" gridrow="2" pageId="13" pageNumber="14">414–605, 482</td>
<td id="76888205991E0049B99DFA1307206E3C" box="[641,895,1508,1536]" gridcol="2" gridrow="2" pageId="13" pageNumber="14">425–655, 511</td>
<td id="76888205991E0049B881FA13006F6E3C" box="[925,1072,1508,1536]" gridcol="3" gridrow="2" pageId="13" pageNumber="14">191–284, 227</td>
<td id="76888205991E0049BF8FFA1301006E3C" box="[1171,1375,1508,1536]" gridcol="4" gridrow="2" pageId="13" pageNumber="14">94–151, 113</td>
</tr>
<tr id="3559EB79991E0049BBB9F9E101006E0E" box="[165,1375,1558,1586]" gridrow="3" pageId="13" pageNumber="14">
<th id="76888205991E0049BBB9F9E104E56E0E" box="[165,186,1558,1586]" gridcol="0" gridrow="3" pageId="13" pageNumber="14">p3</th>
<td id="76888205991E0049BA78F9E106546E0E" box="[356,523,1558,1586]" gridcol="1" gridrow="3" pageId="13" pageNumber="14">432–623, 502</td>
<td id="76888205991E0049B99DF9E107206E0E" box="[641,895,1558,1586]" gridcol="2" gridrow="3" pageId="13" pageNumber="14">439–970, 546</td>
<td id="76888205991E0049B881F9E1006F6E0E" box="[925,1072,1558,1586]" gridcol="3" gridrow="3" pageId="13" pageNumber="14">227–540, 303</td>
<td id="76888205991E0049BF8FF9E101006E0E" box="[1171,1375,1558,1586]" gridcol="4" gridrow="3" pageId="13" pageNumber="14">112–284, 155</td>
</tr>
<tr id="3559EB79991E0049BBB9F9BF01006E5F" box="[165,1375,1608,1635]" gridrow="4" pageId="13" pageNumber="14" rowspan-0="1">
<td id="76888205991E0049BA78F9BF06546E5F" box="[356,523,1608,1635]" gridcol="1" gridrow="4" pageId="13" pageNumber="14">ta3</td>
<td id="76888205991E0049B99DF9BF07206E5F" box="[641,895,1608,1635]" gridcol="2" gridrow="4" pageId="13" pageNumber="14">ta4</td>
<td id="76888205991E0049B881F9BF006F6E5F" box="[925,1072,1608,1635]" gridcol="3" gridrow="4" pageId="13" pageNumber="14">ta5</td>
<td id="76888205991E0049BF8FF9BF01006E5F" box="[1171,1375,1608,1635]" gridcol="4" gridrow="4" pageId="13" pageNumber="14">LR</td>
</tr>
<tr id="3559EB79991E0049BBB9F98E01006EA9" box="[165,1375,1657,1685]" gridrow="5" pageId="13" pageNumber="14">
<th id="76888205991E0049BBB9F98E04E56EA9" box="[165,186,1657,1685]" gridcol="0" gridrow="5" pageId="13" pageNumber="14">p1</th>
<td id="76888205991E0049BA78F98E06546EA9" box="[356,523,1657,1685]" gridcol="1" gridrow="5" pageId="13" pageNumber="14">79–122, 95</td>
<td id="76888205991E0049B99DF98E07206EA9" box="[641,895,1657,1685]" gridcol="2" gridrow="5" pageId="13" pageNumber="14">43–65, 58</td>
<td id="76888205991E0049B881F98E006F6EA9" box="[925,1072,1657,1685]" gridcol="3" gridrow="5" pageId="13" pageNumber="14">36–68, 47</td>
<td id="76888205991E0049BF8FF98E01006EA9" box="[1171,1375,1657,1685]" gridcol="4" gridrow="5" pageId="13" pageNumber="14">0.37–0.48, 0.42</td>
</tr>
<tr id="3559EB79991E0049BBB9F95C01006EFB" box="[165,1375,1707,1735]" gridrow="6" pageId="13" pageNumber="14">
<th id="76888205991E0049BBB9F95C04E56EFB" box="[165,186,1707,1735]" gridcol="0" gridrow="6" pageId="13" pageNumber="14">p2</th>
<td id="76888205991E0049BA78F95C06546EFB" box="[356,523,1707,1735]" gridcol="1" gridrow="6" pageId="13" pageNumber="14">68–112, 84</td>
<td id="76888205991E0049B99DF95C07206EFB" box="[641,895,1707,1735]" gridcol="2" gridrow="6" pageId="13" pageNumber="14">36–76, 51</td>
<td id="76888205991E0049B881F95C006F6EFB" box="[925,1072,1707,1735]" gridcol="3" gridrow="6" pageId="13" pageNumber="14">36–61, 42</td>
<td id="76888205991E0049BF8FF95C01006EFB" box="[1171,1375,1707,1735]" gridcol="4" gridrow="6" pageId="13" pageNumber="14">0.38–0.50, 0.44</td>
</tr>
<tr id="3559EB79991E0049BBB9F92B01006EC4" box="[165,1375,1756,1784]" gridrow="7" pageId="13" pageNumber="14">
<th id="76888205991E0049BBB9F92B04E56EC4" box="[165,186,1756,1784]" gridcol="0" gridrow="7" pageId="13" pageNumber="14">p3</th>
<td id="76888205991E0049BA78F92B06546EC4" box="[356,523,1756,1784]" gridcol="1" gridrow="7" pageId="13" pageNumber="14">97–162, 123</td>
<td id="76888205991E0049B99DF92B07206EC4" box="[641,895,1756,1784]" gridcol="2" gridrow="7" pageId="13" pageNumber="14">47–126, 65</td>
<td id="76888205991E0049B881F92B006F6EC4" box="[925,1072,1756,1784]" gridcol="3" gridrow="7" pageId="13" pageNumber="14">36–72, 46</td>
<td id="76888205991E0049BF8FF92B01006EC4" box="[1171,1375,1756,1784]" gridcol="4" gridrow="7" pageId="13" pageNumber="14">0.47–0.55, 0.51</td>
</tr>
<tr id="3559EB79991E0049BBB9F8F901006F19" box="[165,1375,1806,1829]" gridrow="8" pageId="13" pageNumber="14" rowspan-0="1" rowspan-3="1">
<td id="76888205991E0049BA78F8F906546F19" box="[356,523,1806,1829]" gridcol="1" gridrow="8" pageId="13" pageNumber="14">BV</td>
<td id="76888205991E0049B99DF8F907206F19" box="[641,895,1806,1829]" gridcol="2" gridrow="8" pageId="13" pageNumber="14">SV</td>
<td id="76888205991E0049BF8FF8F901006F19" box="[1171,1375,1806,1829]" gridcol="4" gridrow="8" pageId="13" pageNumber="14">BR</td>
</tr>
<tr id="3559EB79991E0049BBB9F8CB01006F6B" box="[165,1375,1852,1879]" gridrow="9" pageId="13" pageNumber="14" rowspan-3="1">
<th id="76888205991E0049BBB9F8CB04E56F6B" box="[165,186,1852,1879]" gridcol="0" gridrow="9" pageId="13" pageNumber="14">p1</th>
<td id="76888205991E0049BA78F8CB06546F6B" box="[356,523,1852,1879]" gridcol="1" gridrow="9" pageId="13" pageNumber="14">3.11–3.77, 3.45</td>
<td id="76888205991E0049B99DF8CB07206F6B" box="[641,895,1852,1879]" gridcol="2" gridrow="9" pageId="13" pageNumber="14">4.04–4.88, 4.33</td>
<td id="76888205991E0049BF8FF8CB01006F6B" box="[1171,1375,1852,1879]" gridcol="4" gridrow="9" pageId="13" pageNumber="14">2.3–4.4, 3.0</td>
</tr>
<tr id="3559EB79991E0049BBB9F89A01006FB5" box="[165,1375,1901,1929]" gridrow="10" pageId="13" pageNumber="14" rowspan-3="1">
<th id="76888205991E0049BBB9F89A04E56FB5" box="[165,186,1901,1929]" gridcol="0" gridrow="10" pageId="13" pageNumber="14">p2</th>
<td id="76888205991E0049BA78F89A06546FB5" box="[356,523,1901,1929]" gridcol="1" gridrow="10" pageId="13" pageNumber="14">3.68–4.66, 4.24</td>
<td id="76888205991E0049B99DF89A07206FB5" box="[641,895,1901,1929]" gridcol="2" gridrow="10" pageId="13" pageNumber="14">4.18–5.17, 4.51</td>
<td id="76888205991E0049BF8FF89A01006FB5" box="[1171,1375,1901,1929]" gridcol="4" gridrow="10" pageId="13" pageNumber="14">2.8–4.4, 3.6</td>
</tr>
<tr id="3559EB79991E0049BBB9F86801006F87" box="[165,1375,1951,1979]" gridrow="11" pageId="13" pageNumber="14" rowspan-3="1">
<th id="76888205991E0049BBB9F86804E56F87" box="[165,186,1951,1979]" gridcol="0" gridrow="11" pageId="13" pageNumber="14">p3</th>
<td id="76888205991E0049BA78F86806546F87" box="[356,523,1951,1979]" gridcol="1" gridrow="11" pageId="13" pageNumber="14">3.28–3.88, 3.61</td>
<td id="76888205991E0049B99DF86807206F87" box="[641,895,1951,1979]" gridcol="2" gridrow="11" pageId="13" pageNumber="14">3.51–4.12, 3.78</td>
<td id="76888205991E0049BF8FF86801006F87" box="[1171,1375,1951,1979]" gridcol="4" gridrow="11" pageId="13" pageNumber="14">3.8–5.8, 4.4</td>
</tr>
</table>
</paragraph>
<caption id="DF16B9B3991DFFB8BB8BF92C07B06F05" ID-DOI="http://doi.org/10.5281/zenodo.6507317" ID-Zenodo-Dep="6507317" httpUri="https://zenodo.org/record/6507317/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,269,1755,1779]" targetBox="[243,1338,159,1696]" targetPageId="14">
<paragraph id="8BD6E93B991DFFB8BB8BF92C07B06F05" blockId="14.[151,1436,1755,1850]" pageId="14" pageNumber="15">
<emphasis id="B91D3529991DFFB8BB8BF92C05006ECF" bold="true" box="[151,351,1755,1779]" pageId="14" pageNumber="15">FIGURES 27–33.</emphasis>
<taxonomicName id="4C6992B8991DFFB8BA7AF92C077F6ECE" authority="(Paggi, 1993)" authorityName="Andersen &amp; Saether &amp; Mendes" authorityYear="2010" baseAuthorityName="Andersen &amp; Saether &amp; Mendes" baseAuthorityYear="2010" box="[358,800,1755,1778]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="neobilobulatus" status="comb. nov.">
<emphasis id="B91D3529991DFFB8BA7AF92C06DA6ECE" box="[358,645,1755,1778]" italics="true" pageId="14" pageNumber="15">Allocladius neobilobulatus</emphasis>
(
<bibRefCitation id="EFF894CA991DFFB8B989F92C07476ECE" author="Paggi, A. C." box="[661,792,1755,1778]" pageId="14" pageNumber="15" pagination="171 - 174" refId="ref44092" refString="Paggi, A. C. (1993). Redescription of Pseudosmittia bilobulata (Edw.) n. comb. (= Spaniotoma (Smittia) bilobulata Edwards 1931) and description of P. neobilobulata sp. n. (Diptera: Chironomidae). Annales de Limnologie, 29 (2), 171 - 174." type="journal article" year="1993">Paggi, 1993</bibRefCitation>
)
</taxonomicName>
<emphasis id="B91D3529991DFFB8B834F92C07CE6ECF" bold="true" box="[808,913,1755,1779]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A22E8852991DFFB8B834F92C07D46ECF" box="[808,907,1755,1779]" pageId="14" pageNumber="15" rank="species">comb. n.</taxonomicNameLabel>
,
</emphasis>
male.
<emphasis id="B91D3529991DFFB8B8C7F92C07A96ECF" bold="true" box="[987,1014,1755,1779]" pageId="14" pageNumber="15">27</emphasis>
—tentorium, stipes, and cibarial pump;
<emphasis id="B91D3529991DFFB8BB8BF90804ED6F2B" bold="true" box="[151,178,1791,1815]" pageId="14" pageNumber="15">28</emphasis>
—palp;
<emphasis id="B91D3529991DFFB8BA10F90805796F2B" bold="true" box="[268,294,1791,1815]" pageId="14" pageNumber="15">29</emphasis>
—thorax;
<emphasis id="B91D3529991DFFB8BA84F90805EC6F2B" bold="true" box="[408,435,1791,1815]" pageId="14" pageNumber="15">30</emphasis>
—wing;
<emphasis id="B91D3529991DFFB8B909F90806706F2B" bold="true" box="[533,559,1791,1815]" pageId="14" pageNumber="15">31</emphasis>
—hypopygium, dorsal aspect;
<emphasis id="B91D3529991DFFB8B898F90807C06F2B" bold="true" box="[900,927,1791,1815]" pageId="14" pageNumber="15">32</emphasis>
—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right;
<emphasis id="B91D3529991DFFB8B83FF8D507616F06" bold="true" box="[803,830,1826,1850]" pageId="14" pageNumber="15">33</emphasis>
—pars ventralis.
</paragraph>
</caption>
<paragraph id="8BD6E93B991DFFB9BBDAF8880596691E" blockId="14.[151,1436,1919,2020]" lastBlockId="15.[151,1436,152,364]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B91D3529991DFFB8BBDAF88805036FA4" box="[198,348,1919,1944]" italics="true" pageId="14" pageNumber="15">Hypopygium</emphasis>
(
<figureCitation id="1352F5BE991DFFB8BA71F88805A76FA5" box="[365,504,1919,1945]" captionStart="FIGURES 27–33" captionStartId="14.[151,269,1755,1779]" captionTargetBox="[243,1338,159,1696]" captionTargetId="figure-117@14.[226,1360,135,1723]" captionTargetPageId="14" captionText="FIGURES 27–33. Allocladius neobilobulatus (Paggi, 1993) comb. n., male. 27—tentorium, stipes, and cibarial pump; 28—palp; 29—thorax; 30—wing; 31—hypopygium, dorsal aspect; 32—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 33—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507317" httpUri="https://zenodo.org/record/6507317/files/figure.png" pageId="14" pageNumber="15">Figs 31–32</figureCitation>
). Anal point 11–43, 24 µm long, tapering to pointed apex; tergite IX with 10– 24, 15 setae including setae on anal point. Laterosternite IX with 2–5, 4 setae. Phallapodeme 59–79, 70 (7) µm long. Transverse sternapodeme 45–75, 57 µm long, with weak oral projections. Virga 9–18, 13 µm long; plate-like. Pars ventralis (
<figureCitation id="1352F5BE991CFFB9BAD5FF6F067F688E" box="[457,544,152,178]" captionStart="FIGURES 27–33" captionStartId="14.[151,269,1755,1779]" captionTargetBox="[243,1338,159,1696]" captionTargetId="figure-117@14.[226,1360,135,1723]" captionTargetPageId="14" captionText="FIGURES 27–33. Allocladius neobilobulatus (Paggi, 1993) comb. n., male. 27—tentorium, stipes, and cibarial pump; 28—palp; 29—thorax; 30—wing; 31—hypopygium, dorsal aspect; 32—hypopygium with anal point and tergite IX removed, dorsal aspect to the left, ventral aspect to the right; 33—pars ventralis." figureDoi="http://doi.org/10.5281/zenodo.6507317" httpUri="https://zenodo.org/record/6507317/files/figure.png" pageId="15" pageNumber="16">Fig. 33</figureCitation>
) double, 75–120, 93 µm long; 34–71, 42 µm wide; reaching to 0.50–0.76 gonocoxite length. Gonocoxite 140–179, 160 µm long. Inferior volsella reaching to 0.65–0.69 gonocoxite length; no apparent posterior lobe. Gonostylus 61–79, 68 µm long; megaseta 6–9, 8 µm long. HR 2.07–2.51, 2.36. HV 2.91–3.88, 3.22.
</paragraph>
<paragraph id="8BD6E93B991CFFB9BBDAFEDA00C46950" blockId="15.[151,1436,152,364]" pageId="15" pageNumber="16">
<emphasis id="B91D3529991CFFB9BBDAFEDA0558697B" bold="true" box="[198,263,301,327]" pageId="15" pageNumber="16">Pupa</emphasis>
(n = 2). Total length
<quantity id="4C9144DE991CFFB9B918FEDA06ED697B" box="[516,690,301,327]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.42" metricValueMin="3.38" pageId="15" pageNumber="16" unit="mm" value="3.4" valueMax="3.42" valueMin="3.38">3.38–3.42 mm</quantity>
. Exuviae tinged with brownish yellow along margins of cephalothorax and anal segments, particularly on margin of wing sheath and male genital sac.
</paragraph>
<caption id="DF16B9B3991CFFB9BB8BF85D07D66FDE" ID-DOI="http://doi.org/10.5281/zenodo.6507319" ID-Zenodo-Dep="6507319" httpUri="https://zenodo.org/record/6507319/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,269,1962,1986]" targetBox="[274,1276,417,1927]" targetPageId="15">
<paragraph id="8BD6E93B991CFFB9BB8BF85D07D66FDE" blockId="15.[151,1436,1962,2018]" pageId="15" pageNumber="16">
<emphasis id="B91D3529991CFFB9BB8BF85D05006FFE" bold="true" box="[151,351,1962,1986]" pageId="15" pageNumber="16">FIGURES 34–37.</emphasis>
<taxonomicName id="4C6992B8991CFFB9BA7AF85C077E6FFE" authority="(Paggi, 1993)" authorityName="Andersen &amp; Saether &amp; Mendes" authorityYear="2010" baseAuthorityName="Andersen &amp; Saether &amp; Mendes" baseAuthorityYear="2010" box="[358,801,1963,1986]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="neobilobulatus" status="comb. nov.">
<emphasis id="B91D3529991CFFB9BA7AF85C06D96FFE" box="[358,646,1963,1986]" italics="true" pageId="15" pageNumber="16">Allocladius neobilobulatus</emphasis>
(
<bibRefCitation id="EFF894CA991CFFB9B98AF85C07476FFE" author="Paggi, A. C." box="[662,792,1963,1986]" pageId="15" pageNumber="16" pagination="171 - 174" refId="ref44092" refString="Paggi, A. C. (1993). Redescription of Pseudosmittia bilobulata (Edw.) n. comb. (= Spaniotoma (Smittia) bilobulata Edwards 1931) and description of P. neobilobulata sp. n. (Diptera: Chironomidae). Annales de Limnologie, 29 (2), 171 - 174." type="journal article" year="1993">Paggi, 1993</bibRefCitation>
)
</taxonomicName>
<emphasis id="B91D3529991CFFB9B834F85D07CE6FFE" bold="true" box="[808,913,1962,1986]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="A22E8852991CFFB9B834F85D07D56FFE" box="[808,906,1962,1986]" pageId="15" pageNumber="16" rank="species">comb. n.</taxonomicNameLabel>
,
</emphasis>
female pupa.
<emphasis id="B91D3529991CFFB9BF30F85D00196FFE" bold="true" box="[1068,1094,1962,1986]" pageId="15" pageNumber="16">34</emphasis>
—frontal apotome;
<emphasis id="B91D3529991CFFB9BE07F85D016A6FFE" bold="true" box="[1307,1333,1962,1986]" pageId="15" pageNumber="16">35</emphasis>
—thorax;
<emphasis id="B91D3529991CFFB9BB8BF83D04ED6FDE" bold="true" box="[151,178,1994,2018]" pageId="15" pageNumber="16">36</emphasis>
—tergites;
<emphasis id="B91D3529991CFFB9BA34F83D051C6FDE" bold="true" box="[296,323,1994,2018]" pageId="15" pageNumber="16">37</emphasis>
—sternites with detail of the papilla on the genital sac.
</paragraph>
</caption>
<paragraph id="8BD6E93B9903FFA6BBDAFF6F06C468C3" blockId="16.[151,1436,152,818]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAFF6F0529688D" box="[198,374,152,177]" italics="true" pageId="16" pageNumber="17">Cephalothorax</emphasis>
(
<figureCitation id="1352F5BE9903FFA6BA99FF6F0651688E" box="[389,526,152,178]" captionStart="FIGURES 34–37" captionStartId="15.[151,269,1962,1986]" captionTargetBox="[274,1276,417,1927]" captionTargetId="figure-127@15.[226,1360,375,1963]" captionTargetPageId="15" captionText="FIGURES 34–37. Allocladius neobilobulatus (Paggi, 1993) comb. n., female pupa. 34—frontal apotome; 35—thorax; 36—tergites; 37—sternites with detail of the papilla on the genital sac." figureDoi="http://doi.org/10.5281/zenodo.6507319" httpUri="https://zenodo.org/record/6507319/files/figure.png" pageId="16" pageNumber="17">Figs 34–35</figureCitation>
). Frontal apotome with some rugulosity, without warts, with frontal setae. All cephalothoracic setae about 45–85 µm long. Distance between Dc
<subScript id="17EDEB7E9903FFA6B8B0FF3A07EB68E7" attach="left" box="[940,948,205,219]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
and Dc
<subScript id="17EDEB7E9903FFA6BF0FFF3A004468E7" attach="left" box="[1043,1051,205,219]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
9–10 µm, between Dc
<subScript id="17EDEB7E9903FFA6BE31FF3A016A68E7" attach="left" box="[1325,1333,205,219]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
and Dc
<subScript id="17EDEB7E9903FFA6BE88FF3A01C368E7" attach="right" box="[1428,1436,205,219]" fontSize="6" pageId="16" pageNumber="17">3</subScript>
14–25 µm, between Dc
<subScript id="17EDEB7E9903FFA6BAB5FF0205EE693F" attach="left" box="[425,433,245,259]" fontSize="6" pageId="16" pageNumber="17">3</subScript>
and Dc
<subScript id="17EDEB7E9903FFA6B910FF02064B693F" attach="left" box="[524,532,245,259]" fontSize="6" pageId="16" pageNumber="17">4</subScript>
50–55 µm.
</paragraph>
<paragraph id="8BD6E93B9903FFA6BBDAFEF904BE6A4B" blockId="16.[151,1436,152,818]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAFEF9056A691B" box="[198,309,270,295]" italics="true" pageId="16" pageNumber="17">Abdomen</emphasis>
(
<figureCitation id="1352F5BE9903FFA6BA59FEFA0591691B" box="[325,462,269,295]" captionStart="FIGURES 34–37" captionStartId="15.[151,269,1962,1986]" captionTargetBox="[274,1276,417,1927]" captionTargetId="figure-127@15.[226,1360,375,1963]" captionTargetPageId="15" captionText="FIGURES 34–37. Allocladius neobilobulatus (Paggi, 1993) comb. n., female pupa. 34—frontal apotome; 35—thorax; 36—tergites; 37—sternites with detail of the papilla on the genital sac." figureDoi="http://doi.org/10.5281/zenodo.6507319" httpUri="https://zenodo.org/record/6507319/files/figure.png" pageId="16" pageNumber="17">Figs 36–37</figureCitation>
). Tergite I bare; T II–VIII with strong anterior and posterior shagreen, with weaker shagreen covering most of remaining tergites; T IX with strong median shagreen covering most of segment. Sternites I–II bare; S III–V with weak median shagreen, S VI–VII with spinules in anterior 1/3; S VIII with spinules in anterior 1/2; S IX with lateral shagreen. Tergal conjunctive III/IV with 38–84 spinules, IV/V with 137–160, V/VI with 110–150, VI/VII with 90–115 spinules. Sternal conjunctive III/IV with 4–10 spinules, IV/V with 78–100, V/VI with 130–165, VI/VII with 155–200, VII/VIII with 110–119, VIII/IX with 43 spinules in male pupa, none in female pupa. Anal segment with single 27–37 µm long basal seta, three 27–37 µm long apical setae and single 18 µm long median seta. Genital sac of male overreaching anal segment by 96 µm; of female with apical papilla nearly twice as long as wide (papilla also indicated to varying degree in male).
</paragraph>
</subSubSection>
<subSubSection id="C373BAB09903FFA6BBDAFD75075B6AFE" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BD6E93B9903FFA6BBDAFD75075B6AFE" blockId="16.[151,1436,152,818]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAFD7505606AA0" bold="true" box="[198,319,642,668]" pageId="16" pageNumber="17">Remarks.</emphasis>
The double pars ventralis is unique among orthoclads. However, the pupa is hardly separable from that of the Holarctic
<taxonomicName id="4C6992B89903FFA6BAD6FD5F075F6AFE" authority="(Kieffer, 1926)" baseAuthorityName="Kieffer" baseAuthorityYear="1921" box="[458,768,680,706]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="nanseni">
<emphasis id="B91D35299903FFA6BAD6FD5F05836AFD" box="[458,476,680,705]" italics="true" pageId="16" pageNumber="17">A</emphasis>
.
<emphasis id="B91D35299903FFA6BAF6FD5F061A6AFD" box="[490,581,680,705]" italics="true" pageId="16" pageNumber="17">nanseni</emphasis>
(Kieffer, 1926)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C373BAB09903FFA6BBDAFD3A00676B0E" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BD6E93B9903FFA6BBDAFD3A00676B0E" blockId="16.[151,1436,152,818]" pageId="16" pageNumber="17">
<emphasis id="B91D35299903FFA6BBDAFD3A065C6ADB" bold="true" box="[198,515,717,743]" pageId="16" pageNumber="17">Distribution and biology.</emphasis>
The species is known from two localities in
<collectingCountry id="F37EA9AB9903FFA6BF3BFD3A00C46ADB" box="[1063,1179,717,743]" name="Argentina" pageId="16" pageNumber="17">Argentina</collectingCountry>
(
<bibRefCitation id="EFF894CA9903FFA6BFB1FD3A01656ADB" author="Paggi, A. C." box="[1197,1338,717,743]" pageId="16" pageNumber="17" pagination="171 - 174" refId="ref44092" refString="Paggi, A. C. (1993). Redescription of Pseudosmittia bilobulata (Edw.) n. comb. (= Spaniotoma (Smittia) bilobulata Edwards 1931) and description of P. neobilobulata sp. n. (Diptera: Chironomidae). Annales de Limnologie, 29 (2), 171 - 174." type="journal article" year="1993">Paggi 1993</bibRefCitation>
), a high mountain dam in
<collectingCountry id="F37EA9AB9903FFA6BA76FD04059F6B31" box="[362,448,755,781]" name="Bolivia" pageId="16" pageNumber="17">Bolivia</collectingCountry>
and two localities in southern and central
<collectingCountry id="F37EA9AB9903FFA6B8A0FD0407A46B31" box="[956,1019,755,781]" name="Chile" pageId="16" pageNumber="17">Chile</collectingCountry>
where it has been netted along the banks of lakes and rivers. The altitude range is from about 200 to
<quantity id="4C9144DE9903FFA6B886FCEF07A56B0E" box="[922,1018,792,818]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="17" unit="m" value="3.0">3.000 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>
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