448 lines
65 KiB
XML
448 lines
65 KiB
XML
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<document id="B36441649E4A2AEB36DD295C8D143C23" ID-CLB-Dataset="50742" ID-DOI="10.5281/zenodo.172563" ID-GBIF-Dataset="d325feef-2116-4c36-9571-ea93d68c89a4" ID-ISSN="11755326" ID-Zenodo-Dep="172563" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459758407235" checkinUser="plazi" docAuthor="Cuezzo, Maria Gabriela" docDate="2006" docId="03A4A163FFD6E373BA4040959B124F42" docLanguage="en" docName="zt01221p023.pdf" docOrigin="Zootaxa 1221" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Isomeria awa Cuezzo, 2006, n. sp." docType="treatment" docVersion="10" lastPageNumber="21" masterDocId="FF9DD91BFFC6E366BB48435199524E1C" masterDocTitle="On a new species of Isomeria Beck and redescription of species of Labyrinthus Beck from South America (Gastropoda: Stylommatophora: Camaenidae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="17" updateTime="1698218777145" updateUser="ExternalLinkService">
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<mods:mods id="9A943327DDB137DA1939CE4E1B276B87" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3C8AA68E5C686F8FEC968A2D763874BF">
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<mods:title id="60BFBD231123CE4C25496B4B5BB4DD8A">On a new species of Isomeria Beck and redescription of species of Labyrinthus Beck from South America (Gastropoda: Stylommatophora: Camaenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A81BAF7A238750E2D71BFCF034A4B74B" type="personal">
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<mods:role id="FC139C0F290B0EEAB282972C79CF66C4">
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<mods:roleTerm id="2BF3A86F02CD42E4E564FB041137EE0C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F47067C906E3A694832A0D1A102DE35B">Cuezzo, Maria Gabriela</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="56D40A350B3134411511D551AC884D5B">text</mods:typeOfResource>
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<mods:relatedItem id="B66C303AFADD0FBF58FF86152ED0340B" type="host">
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<mods:titleInfo id="ABFD5E7ADACE145C025032406660D371">
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<mods:title id="E072F57B84DE4716855104CB77C503A3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="890C41F9ECB236C2936A41BDDCF197DB">
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<mods:date id="50B2B23F745925FAF32F140416042CAE">2006</mods:date>
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<mods:detail id="933F7D65B21DB4FECCD3303BD8B21B8F" type="volume">
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<mods:number id="D0E411800C32A3E858ADA07C83961038">1221</mods:number>
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</mods:detail>
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<mods:extent id="28FD6E15BA322FD21ADBA1BB39FD0AA6" unit="page">
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<mods:start id="7CE5286A0F837FCC7C32575653E86EBC">1</mods:start>
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<mods:end id="94BD37786D2090D59DEE3DD6BD5C20BC">23</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="411FCD9ADC631224BAECB20377499430">journal article</mods:classification>
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<mods:identifier id="FE39764D1FBFAD73466195CEAB8420ED" type="CLB-Dataset">50742</mods:identifier>
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<mods:identifier id="0DCD668BF998491F3D4318B8ACAEE83F" type="DOI">10.5281/zenodo.172563</mods:identifier>
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<mods:identifier id="56FF6E9A849BEC53CCE4EDA5CA0E1281" type="GBIF-Dataset">d325feef-2116-4c36-9571-ea93d68c89a4</mods:identifier>
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<mods:identifier id="9C0E445BBC430F8C8675C460169B339C" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="BF46ED70BF7B4E21D475B12B21DC3333" type="Zenodo-Dep">172563</mods:identifier>
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</mods:mods>
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<treatment id="03A4A163FFD6E373BA4040959B124F42" ID-DOI="http://doi.org/10.5281/zenodo.6261415" ID-GBIF-Taxon="119344231" ID-Zenodo-Dep="6261415" LSID="urn:lsid:plazi:treatment:03A4A163FFD6E373BA4040959B124F42" httpUri="http://treatment.plazi.org/id/03A4A163FFD6E373BA4040959B124F42" lastPageId="21" lastPageNumber="21" pageId="16" pageNumber="17">
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<subSubSection id="C31743FEFFD6E376BA40409598E24A1D" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="8BB21075FFD6E376BA40409598A74DC2" blockId="16.[264,501,964,1025]" box="[264,501,964,990]" pageId="16" pageNumber="17">
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<heading id="D0FAA719FFD6E376BA40409598A74DC2" bold="true" box="[264,501,964,990]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<emphasis id="B979CC67FFD6E376BA40409598A74DC2" bold="true" box="[264,501,964,990]" pageId="16" pageNumber="17">
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<taxonomicName id="4C0D6BF6FFD6E376BA40409598FE4DC2" box="[264,428,964,990]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="awa" status="sp. nov.">
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<emphasis id="B979CC67FFD6E376BA40409598FE4DC2" bold="true" box="[264,428,964,990]" italics="true" pageId="16" pageNumber="17">Isomeria awa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24A711CFFD6E376BAFB409598A74DC2" box="[435,501,964,990]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB21075FFD6E376BA4040B698E24A1D" blockId="16.[264,501,964,1025]" box="[264,432,999,1025]" pageId="16" pageNumber="17">
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<figureCitation id="13360CF0FFD6E376BA4040B698E24A1D" box="[264,432,999,1025]" captionStart-0="FIGURES 36 – 38" captionStart-1="FIGURES 39 – 42" captionStart-2="FIGURES 43 – 48" captionStartId-0="17.[264,383,603,627]" captionStartId-1="18.[264,382,1580,1604]" captionStartId-2="19.[264,383,1729,1753]" captionTargetBox-0="[271,1315,283,575]" captionTargetBox-1="[279,1297,402,1535]" captionTargetBox-2="[271,1315,514,1704]" captionTargetId-0="figure@17.[271,1317,281,575]" captionTargetId-1="figure@18.[255,1333,390,1570]" captionTargetId-2="figure@19.[271,1317,514,1704]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionText-0="FIGURES 36 – 38: Isomeria awa n. sp. 36: Holotype, dorsal view, scale bar = 1 cm; 37: lateral view, scale bar = 1 cm; 38: ventral view, scale bar = 1 cm." captionText-1="FIGURES 39 – 42: Isomeria awa n. sp. 39: Genital system, ventral view, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; bcd: duct of bursa copulatrix; e: epiphallus; f: flagellum; fo: free oviduct; gp: glandular pouch; hd: hermaphroditic duct; pr: penis retractor muscle; ps: penis sheath; s: spermoviduct; v: vagina; vd: vas deferens; 40: Detail of distal genital system in dorsal view. Abbreviations: go: genital opening; gp: glandular pouch; m; muscular strands; ps: penis sheath; v: vagina; 41: Diagram of the stimulatorlike structure extending into atrium towards distal penis, scale bar = 5 mm. Abbreviations: a: atrium; p: penis; st: stimulatorlike structure; v: vagina; 42: Pilaster running from top of flagellum to junction with epiphallus showing the dorsal groove, scale bar = 5 mm. Abbreviations: e: epiphallus; f: flagellum; g: groove; pi: pilaster." captionText-2="FIGURES 43 – 48: Isomeria awa n. sp. 43: central and first lateral teeth, scale bar = 10 m. 44: lateral teeth, rows 3 to 5, scale bar = 10 m; 45: Lateral teeth, rows 20 to 27, scale bar = 10 m. 46: Dorsal view of the jaw, scale bar = 100 m. 47: Dorsal view of a hooked denticle from the inner wall of penis, scale bar = 10 m. 48: Lateral view of a hooked denticle from the penis, scale bar = 10" httpUri-0="https://zenodo.org/record/172572/files/figure.png" httpUri-1="https://zenodo.org/record/172573/files/figure.png" httpUri-2="https://zenodo.org/record/172574/files/figure.png" pageId="16" pageNumber="17">Figures 36–48</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31743FEFFD6E376BA4047679D314AD5" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="8BB21075FFD6E376BA404767987E4A65" blockId="16.[264,1325,1078,1905]" pageId="16" pageNumber="17">
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<emphasis id="B979CC67FFD6E376BA4047679B134A4C" bold="true" box="[264,577,1078,1104]" pageId="16" pageNumber="17">
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<typeStatus id="54B6AED7FFD6E376BA40476798174A4C" box="[264,325,1078,1104]" pageId="16" pageNumber="17">Type</typeStatus>
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material:
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<typeStatus id="54B6AED7FFD6E376BA8E47679B6E4A4C" box="[454,572,1078,1104]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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AvH unnumbered.
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<typeStatus id="54B6AED7FFD6E376B86E47679AF14A4C" box="[806,931,1078,1104]" pageId="16" pageNumber="17" type="paratype">
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<emphasis id="B979CC67FFD6E376B86E47679AF14A4C" bold="true" box="[806,931,1078,1104]" pageId="16" pageNumber="17">Paratypes</emphasis>
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</typeStatus>
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:
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<collectionCode id="ED1C88B0FFD6E376B8F147669AA74A4D" box="[953,1013,1079,1105]" httpUri="http://grbio.org/cool/sggj-9jsz" name="Fundacion Miguel Lillo" pageId="16" pageNumber="17">FML</collectionCode>
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14796 A, from
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<typeStatus id="54B6AED7FFD6E376BFF847669DB14A4D" box="[1200,1251,1079,1105]" pageId="16" pageNumber="17">type</typeStatus>
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locality.
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</paragraph>
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<paragraph id="8BB21075FFD6E376BA7047D79D314AD5" blockId="16.[264,1325,1078,1905]" pageId="16" pageNumber="17">
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<emphasis id="B979CC67FFD6E376BA7047D798B94ABC" bold="true" box="[312,491,1158,1184]" pageId="16" pageNumber="17">
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<typeStatus id="54B6AED7FFD6E376BA7047D798274ABC" box="[312,373,1158,1184]" pageId="16" pageNumber="17">Type</typeStatus>
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Locality
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</emphasis>
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:
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<collectingCountry id="F31A50E5FFD6E376B94C47D79BF84ABC" box="[516,682,1158,1184]" name="Colombia" pageId="16" pageNumber="17">
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<emphasis id="B979CC67FFD6E376B94C47D79BF84ABC" bold="true" box="[516,682,1158,1184]" pageId="16" pageNumber="17">COLOMBIA</emphasis>
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</collectingCountry>
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,
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<emphasis id="B979CC67FFD6E376B98947D69AFB4ABC" box="[705,937,1159,1184]" italics="true" pageId="16" pageNumber="17">Nariño department</emphasis>
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, Reserva Natural La Planada, Nariño, 1730 mts,
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<geoCoordinate id="EE3976B2FFD6E376BAAB47FE9B164AD5" box="[483,580,1199,1225]" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="1.8333333">N 1° 50'</geoCoordinate>
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<geoCoordinate id="EE3976B2FFD6E376B90347FE9B934AD5" box="[587,705,1199,1225]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="-77.71667">W 77° 43'</geoCoordinate>
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Orejuella, J. E. & Cantillo, G. Coll.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31743FEFFD6E376BA7047879B294B75" pageId="16" pageNumber="17" type="etymology">
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<paragraph id="8BB21075FFD6E376BA7047879B294B75" blockId="16.[264,1325,1078,1905]" pageId="16" pageNumber="17">
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<emphasis id="B979CC67FFD6E376BA704787989A4AEC" bold="true" box="[312,456,1238,1264]" pageId="16" pageNumber="17">Etymology:</emphasis>
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the species name refers to the indigenous Inkal
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<taxonomicName id="4C0D6BF6FFD6E376BF6647869D304AED" box="[1070,1122,1239,1265]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="22" phylum="Mollusca" rank="species" species="awa">Awa</taxonomicName>
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community that lives close to the Reserva Natural La Planada in the region of the Cordillera Occidental, Ricaurte County, Nariño Department in
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<collectingCountry id="F31A50E5FFD6E376B9A246769A304B5D" box="[746,866,1319,1345]" name="Colombia" pageId="16" pageNumber="17">Colombia</collectingCountry>
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. The name
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<taxonomicName id="4C0D6BF6FFD6E376B8B846769D774B5D" box="[1008,1061,1319,1345]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="22" phylum="Mollusca" rank="species" species="awa">Awa</taxonomicName>
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means "people of the mountain". Noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31743FEFFD6E376BA7046279A12482D" pageId="16" pageNumber="17" type="diagnosis">
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<paragraph id="8BB21075FFD6E376BA7046279A12482D" blockId="16.[264,1325,1078,1905]" pageId="16" pageNumber="17">
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<emphasis id="B979CC67FFD6E376BA70462798E84B8C" bold="true" box="[312,442,1398,1424]" pageId="16" pageNumber="17">Diagnosis:</emphasis>
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shell body whorl with axial growth lines, small pustules all over surface. Irregular spots of dark brown coloration on dorsal shell surface. Parietal lamellae small, triangular, not merging with lip, small conical tooth on lower palatal peristomal lip. Umbilicus narrow, open. Vagina long with glandular pouch in distal portion. Presence of inner conical stimulatorlike structure in distal vagina.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31743FEFFD6E375BA70456F9861497F" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17" type="description">
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<paragraph id="8BB21075FFD6E376BA70456F9D78496D" blockId="16.[264,1325,1078,1905]" pageId="16" pageNumber="17">
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<emphasis id="B979CC67FFD6E376BA70456F98804844" bold="true" box="[312,466,1598,1624]" pageId="16" pageNumber="17">Description:</emphasis>
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<emphasis id="B979CC67FFD6E376BA92456E9B474844" box="[474,533,1599,1624]" italics="true" pageId="16" pageNumber="17">Shell</emphasis>
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(
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<figureCitation id="13360CF0FFD6E376B96C456E9BFD4845" box="[548,687,1599,1625]" captionStart="FIGURES 36 – 38" captionStartId="17.[264,383,603,627]" captionTargetBox="[271,1315,283,575]" captionTargetId="figure@17.[271,1317,281,575]" captionTargetPageId="17" captionText="FIGURES 36 – 38: Isomeria awa n. sp. 36: Holotype, dorsal view, scale bar = 1 cm; 37: lateral view, scale bar = 1 cm; 38: ventral view, scale bar = 1 cm." httpUri="https://zenodo.org/record/172572/files/figure.png" pageId="16" pageNumber="17">Figs 36–38</figureCitation>
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): Globose, 4 1/2 whorls, strong, body whorl periphery obtusely angulated; golden yellowish. Axial growth lines with small pustules all over shell surface. Irregular small spots of dark brown color on dorsal side of shell. Body whorl strongly descending behind aperture. Aperture subquadrate, chestnut coloration inside aperture. Peristomal lip white, strongly reflexed, parietal lip with prominent raised edge. Small, triangular parietal lamellae present, not merging with peristomal lip, lower palatal lip with small conical tooth. Suture shallow. Umbilicus narrow, open, with columnar groove noticeable, basal lip extension covering only a small portion.
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</paragraph>
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<caption id="DF7240FDFFD7E377BA40410A9A724C89" httpUri="https://zenodo.org/record/172572/files/figure.png" pageId="17" pageNumber="18" targetBox="[271,1315,283,575]" targetPageId="17">
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<paragraph id="8BB21075FFD7E377BA40410A9A724C89" blockId="17.[264,1323,603,661]" pageId="17" pageNumber="18">
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<emphasis id="B979CC67FFD7E377BA40410A98954C6F" bold="true" box="[264,455,603,627]" pageId="17" pageNumber="18">FIGURES 36–38</emphasis>
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:
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<taxonomicName id="4C0D6BF6FFD7E377BA9D410D9B354C6F" box="[469,615,604,627]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="awa" status="sp. nov.">
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<emphasis id="B979CC67FFD7E377BA9D410D9B354C6F" box="[469,615,604,627]" italics="true" pageId="17" pageNumber="18">Isomeria awa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24A711CFFD7E377B925410D9BF44C6F" box="[621,678,604,627]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
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36: Holotype, dorsal view, scale bar= 1 cm; 37: lateral view, scale bar= 1 cm; 38: ventral view, scale bar= 1cm.
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</paragraph>
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</caption>
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<paragraph id="8BB21075FFD7E377BA7041989B534D47" blockId="17.[264,1324,713,1899]" pageId="17" pageNumber="18">
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<typeStatus id="54B6AED7FFD7E377BA70419898F74CFF" box="[312,421,713,739]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
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shell measurements: D major= 39.7; D minor= 34.4; H= 21.5; D ap= 19.7; Hap= 16.3.
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<typeStatus id="54B6AED7FFD7E377BAD641A09B424D17" box="[414,528,753,779]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
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shell measurements (n= 3): D major= 38.8–39.4 (0 = 38.6); D minor= 33.2–34.2 (0 = 33.4); H=
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<date id="FFB336B5FFD7E377B9DF40489BAC4D2F" box="[663,766,793,819]" pageId="17" pageNumber="18" value="1921-05-21">21–21–5</date>
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(0 = 21.3); D ap= 19.3–20.1 (0 = 19.6): H ap= 16.2–16.9 (0 = 16.4).
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</paragraph>
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<paragraph id="8BB21075FFD7E377BA7040389B044A57" blockId="17.[264,1324,713,1899]" pageId="17" pageNumber="18">
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<emphasis id="B979CC67FFD7E377BA70403898BA4D9E" box="[312,488,873,898]" italics="true" pageId="17" pageNumber="18">Pallial Organs</emphasis>
|
|||
|
|
: Pulmonary roof black due to dense dark pigmentation. Thin grayish minor veins weakly marked, distinguishable mostly between rectum and kidney. Main pulmonary vein running from pericardial cavity to mantle collar, not branched. Kidney narrow, long, occupying about 70% of pulmonary roof length. Secondary ureter closes from top of pulmonary roof to mantle collar. Ureteric interramus triangular, deeply excavated, overlapped by mantle collar.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BB21075FFD7E374BA7047089BEF4F42" blockId="17.[264,1324,713,1899]" lastBlockId="18.[264,1322,284,350]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
|||
|
|
<emphasis id="B979CC67FFD7E377BA7047089B614A6E" box="[312,563,1113,1138]" italics="true" pageId="17" pageNumber="18">Reproductive system</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13360CF0FFD7E377B90447089B8C4A6F" box="[588,734,1113,1139]" captionStart="FIGURES 39 – 42" captionStartId="18.[264,382,1580,1604]" captionTargetBox="[279,1297,402,1535]" captionTargetId="figure@18.[255,1333,390,1570]" captionTargetPageId="18" captionText="FIGURES 39 – 42: Isomeria awa n. sp. 39: Genital system, ventral view, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; bcd: duct of bursa copulatrix; e: epiphallus; f: flagellum; fo: free oviduct; gp: glandular pouch; hd: hermaphroditic duct; pr: penis retractor muscle; ps: penis sheath; s: spermoviduct; v: vagina; vd: vas deferens; 40: Detail of distal genital system in dorsal view. Abbreviations: go: genital opening; gp: glandular pouch; m; muscular strands; ps: penis sheath; v: vagina; 41: Diagram of the stimulatorlike structure extending into atrium towards distal penis, scale bar = 5 mm. Abbreviations: a: atrium; p: penis; st: stimulatorlike structure; v: vagina; 42: Pilaster running from top of flagellum to junction with epiphallus showing the dorsal groove, scale bar = 5 mm. Abbreviations: e: epiphallus; f: flagellum; g: groove; pi: pilaster." httpUri="https://zenodo.org/record/172573/files/figure.png" pageId="17" pageNumber="18">Figs 39–42</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13360CF0FFD7E377B9BC47089A1E4A6F" box="[756,844,1113,1139]" captionStart="FIGURES 43 – 48" captionStartId="19.[264,383,1729,1753]" captionTargetBox="[271,1315,514,1704]" captionTargetId="figure@19.[271,1317,514,1704]" captionTargetPageId="19" captionText="FIGURES 43 – 48: Isomeria awa n. sp. 43: central and first lateral teeth, scale bar = 10 m. 44: lateral teeth, rows 3 to 5, scale bar = 10 m; 45: Lateral teeth, rows 20 to 27, scale bar = 10 m. 46: Dorsal view of the jaw, scale bar = 100 m. 47: Dorsal view of a hooked denticle from the inner wall of penis, scale bar = 10 m. 48: Lateral view of a hooked denticle from the penis, scale bar = 10" httpUri="https://zenodo.org/record/172574/files/figure.png" pageId="17" pageNumber="18">47, 48</figureCitation>
|
|||
|
|
): Ovotestis formed by long, apically branched acini embedded in digestive gland. Hermaphroditic duct long, convoluted; distal portion of hermaphroditic duct reflexed over itself before inserting in distal portion of albumen gland. Fertilization pouchspermathecal complex (FPSC) not distinguishable. Duct of bursa copulatrix thicker than free oviduct, 1 1/2 times as long as spermoviduct length (
|
|||
|
|
<figureCitation id="13360CF0FFD7E377BA2E467098914B27" box="[358,451,1313,1339]" captionStart="FIGURES 39 – 42" captionStartId="18.[264,382,1580,1604]" captionTargetBox="[279,1297,402,1535]" captionTargetId="figure@18.[255,1333,390,1570]" captionTargetPageId="18" captionText="FIGURES 39 – 42: Isomeria awa n. sp. 39: Genital system, ventral view, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; bcd: duct of bursa copulatrix; e: epiphallus; f: flagellum; fo: free oviduct; gp: glandular pouch; hd: hermaphroditic duct; pr: penis retractor muscle; ps: penis sheath; s: spermoviduct; v: vagina; vd: vas deferens; 40: Detail of distal genital system in dorsal view. Abbreviations: go: genital opening; gp: glandular pouch; m; muscular strands; ps: penis sheath; v: vagina; 41: Diagram of the stimulatorlike structure extending into atrium towards distal penis, scale bar = 5 mm. Abbreviations: a: atrium; p: penis; st: stimulatorlike structure; v: vagina; 42: Pilaster running from top of flagellum to junction with epiphallus showing the dorsal groove, scale bar = 5 mm. Abbreviations: e: epiphallus; f: flagellum; g: groove; pi: pilaster." httpUri="https://zenodo.org/record/172573/files/figure.png" pageId="17" pageNumber="18">Fig. 39</figureCitation>
|
|||
|
|
). Sac oval, voluminous, not reflexed over its duct. Free oviduct short, slightly thinner than vagina. Vagina long, distal glandular pouch close to penioviducal angle (
|
|||
|
|
<figureCitation id="13360CF0FFD7E377BA10462098B64B97" box="[344,484,1393,1419]" captionStart="FIGURES 39 – 42" captionStartId="18.[264,382,1580,1604]" captionTargetBox="[279,1297,402,1535]" captionTargetId="figure@18.[255,1333,390,1570]" captionTargetPageId="18" captionText="FIGURES 39 – 42: Isomeria awa n. sp. 39: Genital system, ventral view, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; bcd: duct of bursa copulatrix; e: epiphallus; f: flagellum; fo: free oviduct; gp: glandular pouch; hd: hermaphroditic duct; pr: penis retractor muscle; ps: penis sheath; s: spermoviduct; v: vagina; vd: vas deferens; 40: Detail of distal genital system in dorsal view. Abbreviations: go: genital opening; gp: glandular pouch; m; muscular strands; ps: penis sheath; v: vagina; 41: Diagram of the stimulatorlike structure extending into atrium towards distal penis, scale bar = 5 mm. Abbreviations: a: atrium; p: penis; st: stimulatorlike structure; v: vagina; 42: Pilaster running from top of flagellum to junction with epiphallus showing the dorsal groove, scale bar = 5 mm. Abbreviations: e: epiphallus; f: flagellum; g: groove; pi: pilaster." httpUri="https://zenodo.org/record/172573/files/figure.png" pageId="17" pageNumber="18">Figs 39, 40</figureCitation>
|
|||
|
|
). Inner structure of proximal portion of vagina consisting of longitudinal, parallel, deep cords with scattered white hooked denticles. Conical stimulatorlike structure, present in distal portion of vagina (
|
|||
|
|
<figureCitation id="13360CF0FFD7E377B86346909AD64BC7" box="[811,900,1473,1499]" captionStart="FIGURES 39 – 42" captionStartId="18.[264,382,1580,1604]" captionTargetBox="[279,1297,402,1535]" captionTargetId="figure@18.[255,1333,390,1570]" captionTargetPageId="18" captionText="FIGURES 39 – 42: Isomeria awa n. sp. 39: Genital system, ventral view, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; bcd: duct of bursa copulatrix; e: epiphallus; f: flagellum; fo: free oviduct; gp: glandular pouch; hd: hermaphroditic duct; pr: penis retractor muscle; ps: penis sheath; s: spermoviduct; v: vagina; vd: vas deferens; 40: Detail of distal genital system in dorsal view. Abbreviations: go: genital opening; gp: glandular pouch; m; muscular strands; ps: penis sheath; v: vagina; 41: Diagram of the stimulatorlike structure extending into atrium towards distal penis, scale bar = 5 mm. Abbreviations: a: atrium; p: penis; st: stimulatorlike structure; v: vagina; 42: Pilaster running from top of flagellum to junction with epiphallus showing the dorsal groove, scale bar = 5 mm. Abbreviations: e: epiphallus; f: flagellum; g: groove; pi: pilaster." httpUri="https://zenodo.org/record/172573/files/figure.png" pageId="17" pageNumber="18">Fig. 41</figureCitation>
|
|||
|
|
), attached through base to vagina's wall near vaginaatrium junction, raised free into vagina's lumen. Portion of stimulatorlike structure extending into atrium towards distal penis, ending into two fingerlike branches (
|
|||
|
|
<figureCitation id="13360CF0FFD7E377BACA4568988E484F" box="[386,476,1593,1619]" captionStart="FIGURES 39 – 42" captionStartId="18.[264,382,1580,1604]" captionTargetBox="[279,1297,402,1535]" captionTargetId="figure@18.[255,1333,390,1570]" captionTargetPageId="18" captionText="FIGURES 39 – 42: Isomeria awa n. sp. 39: Genital system, ventral view, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; bcd: duct of bursa copulatrix; e: epiphallus; f: flagellum; fo: free oviduct; gp: glandular pouch; hd: hermaphroditic duct; pr: penis retractor muscle; ps: penis sheath; s: spermoviduct; v: vagina; vd: vas deferens; 40: Detail of distal genital system in dorsal view. Abbreviations: go: genital opening; gp: glandular pouch; m; muscular strands; ps: penis sheath; v: vagina; 41: Diagram of the stimulatorlike structure extending into atrium towards distal penis, scale bar = 5 mm. Abbreviations: a: atrium; p: penis; st: stimulatorlike structure; v: vagina; 42: Pilaster running from top of flagellum to junction with epiphallus showing the dorsal groove, scale bar = 5 mm. Abbreviations: e: epiphallus; f: flagellum; g: groove; pi: pilaster." httpUri="https://zenodo.org/record/172573/files/figure.png" pageId="17" pageNumber="18">Fig. 41</figureCitation>
|
|||
|
|
). Penial complex formed by flagellum, epiphallus, penis (
|
|||
|
|
<figureCitation id="13360CF0FFD7E377BFD945689C4C484F" box="[1169,1310,1593,1619]" captionStart="FIGURES 39 – 42" captionStartId="18.[264,382,1580,1604]" captionTargetBox="[279,1297,402,1535]" captionTargetId="figure@18.[255,1333,390,1570]" captionTargetPageId="18" captionText="FIGURES 39 – 42: Isomeria awa n. sp. 39: Genital system, ventral view, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; bcd: duct of bursa copulatrix; e: epiphallus; f: flagellum; fo: free oviduct; gp: glandular pouch; hd: hermaphroditic duct; pr: penis retractor muscle; ps: penis sheath; s: spermoviduct; v: vagina; vd: vas deferens; 40: Detail of distal genital system in dorsal view. Abbreviations: go: genital opening; gp: glandular pouch; m; muscular strands; ps: penis sheath; v: vagina; 41: Diagram of the stimulatorlike structure extending into atrium towards distal penis, scale bar = 5 mm. Abbreviations: a: atrium; p: penis; st: stimulatorlike structure; v: vagina; 42: Pilaster running from top of flagellum to junction with epiphallus showing the dorsal groove, scale bar = 5 mm. Abbreviations: e: epiphallus; f: flagellum; g: groove; pi: pilaster." httpUri="https://zenodo.org/record/172573/files/figure.png" pageId="17" pageNumber="18">Figs 39, 40</figureCitation>
|
|||
|
|
). Flagellum short, uniform diameter, tip folded. Internally, longitudinal pilaster with open dorsal groove running from top to junction with epiphallus, ending at point of insertion of vas deferens or slightly below (
|
|||
|
|
<figureCitation id="13360CF0FFD7E377B9C345E09BBA48D7" box="[651,744,1713,1739]" captionStart="FIGURES 39 – 42" captionStartId="18.[264,382,1580,1604]" captionTargetBox="[279,1297,402,1535]" captionTargetId="figure@18.[255,1333,390,1570]" captionTargetPageId="18" captionText="FIGURES 39 – 42: Isomeria awa n. sp. 39: Genital system, ventral view, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; bcd: duct of bursa copulatrix; e: epiphallus; f: flagellum; fo: free oviduct; gp: glandular pouch; hd: hermaphroditic duct; pr: penis retractor muscle; ps: penis sheath; s: spermoviduct; v: vagina; vd: vas deferens; 40: Detail of distal genital system in dorsal view. Abbreviations: go: genital opening; gp: glandular pouch; m; muscular strands; ps: penis sheath; v: vagina; 41: Diagram of the stimulatorlike structure extending into atrium towards distal penis, scale bar = 5 mm. Abbreviations: a: atrium; p: penis; st: stimulatorlike structure; v: vagina; 42: Pilaster running from top of flagellum to junction with epiphallus showing the dorsal groove, scale bar = 5 mm. Abbreviations: e: epiphallus; f: flagellum; g: groove; pi: pilaster." httpUri="https://zenodo.org/record/172573/files/figure.png" pageId="17" pageNumber="18">Fig. 42</figureCitation>
|
|||
|
|
). Pilaster dorsal groove partially overlapped by extension of lateral wall, forming flap. Epiphallus longer, thinner than penis, distally reflexed over penis, overlapped by penis sheath. Penis long. Penial retractor inserting in distal epiphallus. Thick muscular strands attached penis to penis sheath (
|
|||
|
|
<figureCitation id="13360CF0FFD7E377BF2C44789DEC495F" box="[1124,1214,1833,1859]" captionStart="FIGURES 39 – 42" captionStartId="18.[264,382,1580,1604]" captionTargetBox="[279,1297,402,1535]" captionTargetId="figure@18.[255,1333,390,1570]" captionTargetPageId="18" captionText="FIGURES 39 – 42: Isomeria awa n. sp. 39: Genital system, ventral view, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; bcd: duct of bursa copulatrix; e: epiphallus; f: flagellum; fo: free oviduct; gp: glandular pouch; hd: hermaphroditic duct; pr: penis retractor muscle; ps: penis sheath; s: spermoviduct; v: vagina; vd: vas deferens; 40: Detail of distal genital system in dorsal view. Abbreviations: go: genital opening; gp: glandular pouch; m; muscular strands; ps: penis sheath; v: vagina; 41: Diagram of the stimulatorlike structure extending into atrium towards distal penis, scale bar = 5 mm. Abbreviations: a: atrium; p: penis; st: stimulatorlike structure; v: vagina; 42: Pilaster running from top of flagellum to junction with epiphallus showing the dorsal groove, scale bar = 5 mm. Abbreviations: e: epiphallus; f: flagellum; g: groove; pi: pilaster." httpUri="https://zenodo.org/record/172573/files/figure.png" pageId="17" pageNumber="18">Fig. 40</figureCitation>
|
|||
|
|
). Interior wall of proximal penis with thick, deep pilasters bearing hooked denticles regularly distributed, with oval bases of
|
|||
|
|
<quantity id="4CF5BD90FFD4E374B907424D9BB44F2A" box="[591,742,284,310]" metricMagnitude="2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="18" pageNumber="19" unit="m" value="190.0" valueMax="200.0" valueMin="180.0">180–200 m</quantity>
|
|||
|
|
of length (
|
|||
|
|
<figureCitation id="13360CF0FFD4E374B82F424D9AA24F2A" box="[871,1008,284,310]" captionStart="FIGURES 43 – 48" captionStartId="19.[264,383,1729,1753]" captionTargetBox="[271,1315,514,1704]" captionTargetId="figure@19.[271,1317,514,1704]" captionTargetPageId="19" captionText="FIGURES 43 – 48: Isomeria awa n. sp. 43: central and first lateral teeth, scale bar = 10 m. 44: lateral teeth, rows 3 to 5, scale bar = 10 m; 45: Lateral teeth, rows 20 to 27, scale bar = 10 m. 46: Dorsal view of the jaw, scale bar = 100 m. 47: Dorsal view of a hooked denticle from the inner wall of penis, scale bar = 10 m. 48: Lateral view of a hooked denticle from the penis, scale bar = 10" httpUri="https://zenodo.org/record/172574/files/figure.png" pageId="18" pageNumber="19">Figs 47, 48</figureCitation>
|
|||
|
|
). Denticles more abundant in penis than in vagina. Verge absent.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF7240FDFFD4E374BA40457D9D9C4945" httpUri="https://zenodo.org/record/172573/files/figure.png" pageId="18" pageNumber="19" targetBox="[279,1297,402,1535]" targetPageId="18">
|
|||
|
|
<paragraph id="8BB21075FFD4E374BA40457D9D9C4945" blockId="18.[264,1326,1580,1881]" pageId="18" pageNumber="19">
|
|||
|
|
<emphasis id="B979CC67FFD4E374BA40457D989A4858" bold="true" box="[264,456,1580,1604]" pageId="18" pageNumber="19">FIGURES 39–42</emphasis>
|
|||
|
|
:
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD4E374BA9D457C9B344858" box="[469,614,1581,1604]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="awa" status="sp. nov.">
|
|||
|
|
<emphasis id="B979CC67FFD4E374BA9D457C9B344858" box="[469,614,1581,1604]" italics="true" pageId="18" pageNumber="19">Isomeria awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A24A711CFFD4E374B924457C9BF74858" box="[620,677,1581,1604]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
39: Genital system, ventral view, scale bar= 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; bcd: duct of bursa copulatrix; e: epiphallus; f: flagellum; fo: free oviduct; gp: glandular pouch; hd: hermaphroditic duct; pr: penis retractor muscle; ps: penis sheath; s: spermoviduct; v: vagina; vd: vas deferens; 40: Detail of distal genital system in dorsal view. Abbreviations: go: genital opening; gp: glandular pouch; m; muscular strands; ps: penis sheath; v: vagina; 41: Diagram of the stimulatorlike structure extending into atrium towards distal penis, scale bar= 5 mm. Abbreviations: a: atrium; p: penis; st: stimulatorlike structure; v: vagina; 42: Pilaster running from top of flagellum to junction with epiphallus showing the dorsal groove, scale bar= 5 mm. Abbreviations: e: epiphallus; f: flagellum; g: groove; pi: pilaster.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BB21075FFD5E375BA70424D9D5C4FB2" blockId="19.[264,1323,283,430]" pageId="19" pageNumber="20">
|
|||
|
|
<emphasis id="B979CC67FFD5E375BA70424D9B524F29" box="[312,512,284,309]" italics="true" pageId="19" pageNumber="20">Digestive system</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13360CF0FFD5E375B95B424D9BCF4F2A" box="[531,669,284,310]" captionStart="FIGURES 43 – 48" captionStartId="19.[264,383,1729,1753]" captionTargetBox="[271,1315,514,1704]" captionTargetId="figure@19.[271,1317,514,1704]" captionTargetPageId="19" captionText="FIGURES 43 – 48: Isomeria awa n. sp. 43: central and first lateral teeth, scale bar = 10 m. 44: lateral teeth, rows 3 to 5, scale bar = 10 m; 45: Lateral teeth, rows 20 to 27, scale bar = 10 m. 46: Dorsal view of the jaw, scale bar = 100 m. 47: Dorsal view of a hooked denticle from the inner wall of penis, scale bar = 10 m. 48: Lateral view of a hooked denticle from the penis, scale bar = 10" httpUri="https://zenodo.org/record/172574/files/figure.png" pageId="19" pageNumber="20">Figs 43–46</figureCitation>
|
|||
|
|
): Jaw arched, incipient plaque in middle of jaw, fine transversal grooves more evident close to one edge (
|
|||
|
|
<figureCitation id="13360CF0FFD5E375B83242159A874F42" box="[890,981,324,350]" captionStart="FIGURES 43 – 48" captionStartId="19.[264,383,1729,1753]" captionTargetBox="[271,1315,514,1704]" captionTargetId="figure@19.[271,1317,514,1704]" captionTargetPageId="19" captionText="FIGURES 43 – 48: Isomeria awa n. sp. 43: central and first lateral teeth, scale bar = 10 m. 44: lateral teeth, rows 3 to 5, scale bar = 10 m; 45: Lateral teeth, rows 20 to 27, scale bar = 10 m. 46: Dorsal view of the jaw, scale bar = 100 m. 47: Dorsal view of a hooked denticle from the inner wall of penis, scale bar = 10 m. 48: Lateral view of a hooked denticle from the penis, scale bar = 10" httpUri="https://zenodo.org/record/172574/files/figure.png" pageId="19" pageNumber="20">Fig. 46</figureCitation>
|
|||
|
|
). Upper, lower cutting edges even. Central tooth of radula monocuspid (
|
|||
|
|
<figureCitation id="13360CF0FFD5E375B846423D9A3B4F9A" box="[782,873,364,390]" captionStart="FIGURES 43 – 48" captionStartId="19.[264,383,1729,1753]" captionTargetBox="[271,1315,514,1704]" captionTargetId="figure@19.[271,1317,514,1704]" captionTargetPageId="19" captionText="FIGURES 43 – 48: Isomeria awa n. sp. 43: central and first lateral teeth, scale bar = 10 m. 44: lateral teeth, rows 3 to 5, scale bar = 10 m; 45: Lateral teeth, rows 20 to 27, scale bar = 10 m. 46: Dorsal view of the jaw, scale bar = 100 m. 47: Dorsal view of a hooked denticle from the inner wall of penis, scale bar = 10 m. 48: Lateral view of a hooked denticle from the penis, scale bar = 10" httpUri="https://zenodo.org/record/172574/files/figure.png" pageId="19" pageNumber="20">Fig. 43</figureCitation>
|
|||
|
|
). First laterals monocuspid, changing into bicuspid (
|
|||
|
|
<figureCitation id="13360CF0FFD5E375BAF842C59B6B4FB2" box="[432,569,404,430]" captionStart="FIGURES 43 – 48" captionStartId="19.[264,383,1729,1753]" captionTargetBox="[271,1315,514,1704]" captionTargetId="figure@19.[271,1317,514,1704]" captionTargetPageId="19" captionText="FIGURES 43 – 48: Isomeria awa n. sp. 43: central and first lateral teeth, scale bar = 10 m. 44: lateral teeth, rows 3 to 5, scale bar = 10 m; 45: Lateral teeth, rows 20 to 27, scale bar = 10 m. 46: Dorsal view of the jaw, scale bar = 100 m. 47: Dorsal view of a hooked denticle from the inner wall of penis, scale bar = 10 m. 48: Lateral view of a hooked denticle from the penis, scale bar = 10" httpUri="https://zenodo.org/record/172574/files/figure.png" pageId="19" pageNumber="20">Figs 44, 45</figureCitation>
|
|||
|
|
). Condition of marginal teeth unknown.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF7240FDFFD5E375BA4045909C79495C" httpUri="https://zenodo.org/record/172574/files/figure.png" pageId="19" pageNumber="20" targetBox="[271,1315,514,1704]" targetPageId="19">
|
|||
|
|
<paragraph id="8BB21075FFD5E375BA4045909C79495C" blockId="19.[264,1323,1729,1891]" pageId="19" pageNumber="20">
|
|||
|
|
<emphasis id="B979CC67FFD5E375BA404590989A48C5" bold="true" box="[264,456,1729,1753]" pageId="19" pageNumber="20">FIGURES 43–48</emphasis>
|
|||
|
|
:
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD5E375BA9F45909B3B48C4" box="[471,617,1729,1752]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="awa" status="sp. nov.">
|
|||
|
|
<emphasis id="B979CC67FFD5E375BA9F45909B3B48C4" box="[471,617,1729,1752]" italics="true" pageId="19" pageNumber="20">Isomeria awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A24A711CFFD5E375B93945909BF848C4" box="[625,682,1729,1752]" pageId="19" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
43: central and first lateral teeth, scale bar= 10 m. 44: lateral teeth, rows 3 to 5, scale bar= 10 m; 45: Lateral teeth, rows 20 to 27, scale bar= 10 m. 46: Dorsal view of the jaw, scale bar= 100 m. 47: Dorsal view of a hooked denticle from the inner wall of penis, scale bar= 10 m. 48: Lateral view of a hooked denticle from the penis, scale bar= 10
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BB21075FFD5E375BA5F441D9861497F" blockId="19.[264,1323,1729,1891]" box="[279,307,1868,1891]" pageId="19" pageNumber="20">m.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C31743FEFFD2E372BA70424A9A424FCA" pageId="20" pageNumber="21" type="materials_examined">
|
|||
|
|
<paragraph id="8BB21075FFD2E372BA70424A9A424FCA" blockId="20.[264,1323,283,1870]" pageId="20" pageNumber="21">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA70424A9AF94F29" bold="true" box="[312,939,283,309]" pageId="20" pageNumber="21">
|
|||
|
|
Comparative Material examined:
|
|||
|
|
<collectingCountry id="F31A50E5FFD2E372B9B4424A9AF44F29" box="[764,934,283,309]" name="Colombia" pageId="20" pageNumber="21">COLOMBIA</collectingCountry>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
ZMZ 544602
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BF3A424D98FB4F42" authority="Koch, 1844" authorityName="Koch" authorityYear="1844" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oreas">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BF3A424D9C784F29" box="[1138,1322,284,309]" italics="true" pageId="20" pageNumber="21">Isomeria oreas</emphasis>
|
|||
|
|
(Koch, 1844)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, G. Schneider, 1912; ZMZ 509673
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B81C42159C744F42" authority="Mousson, 1869" authorityName="Mousson" authorityYear="1869" box="[852,1318,324,350]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="subelliptica">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B81C42159D1C4F41" box="[852,1102,324,349]" italics="true" pageId="20" pageNumber="21">Isomeria subelliptica</emphasis>
|
|||
|
|
(Mousson, 1869)
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<typeStatus id="54B6AED7FFD2E372BA40423D982A4F9A" box="[264,376,364,390]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
|
|||
|
|
, Bagua, Amazonas,
|
|||
|
|
<collectingCountry id="F31A50E5FFD2E372B93B423D9BEF4F9A" box="[627,701,364,390]" name="United Kingdom" pageId="20" pageNumber="21">Wallis</collectingCountry>
|
|||
|
|
leg., 1869.
|
|||
|
|
<emphasis id="B979CC67FFD2E372B815423A9AA94F99" bold="true" box="[861,1019,363,389]" pageId="20" pageNumber="21">
|
|||
|
|
<collectingCountry id="F31A50E5FFD2E372B815423A9AA44F99" box="[861,1014,363,389]" name="Ecuador" pageId="20" pageNumber="21">ECUADOR</collectingCountry>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
MUC s/n,
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BFC0423D986D4FB1" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="globosa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BFC0423D986D4FB1" italics="true" pageId="20" pageNumber="21">Isomeria globosa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA0642C59B644FB1" box="[334,566,404,429]" italics="true" pageId="20" pageNumber="21">Pichincha Province</emphasis>
|
|||
|
|
. Puerto Quito, and
|
|||
|
|
<date id="FFB336B5FFD2E372B85442C59AC54FB2" box="[796,919,404,430]" pageId="20" pageNumber="21" value="1983-12-05">5/12/1983</date>
|
|||
|
|
.M. Ituwalde. MUC s/n,
|
|||
|
|
<emphasis id="B979CC67FFD2E372BFFC42C5986C4FC9" italics="true" pageId="20" pageNumber="21">Pichincha Prov</emphasis>
|
|||
|
|
., Puerto Quito,
|
|||
|
|
<date id="FFB336B5FFD2E372BAB142ED9B214FCA" box="[505,627,444,470]" pageId="20" pageNumber="21" value="1984-01-16">16/1/1984</date>
|
|||
|
|
. Leg. V. Zak.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C31743FEFFD2E373BA7042B29B124F42" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
|
|||
|
|
<paragraph id="8BB21075FFD2E372BA7042B29AFA4AD1" blockId="20.[264,1323,283,1870]" pageId="20" pageNumber="21">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA7042B298E14FE1" bold="true" box="[312,435,483,509]" pageId="20" pageNumber="21">Remarks:</emphasis>
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA8942B59B3A4FE1" box="[449,616,484,509]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA8942B59B3A4FE1" box="[449,616,484,509]" italics="true" pageId="20" pageNumber="21">Isomeria awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
belongs to the
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B87E42B59ABF4FE1" box="[822,1005,484,509]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oreas">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B87E42B59ABF4FE1" box="[822,1005,484,509]" italics="true" pageId="20" pageNumber="21">Isomeria oreas</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group of species (
|
|||
|
|
<bibRefCitation id="EF9C6D84FFD2E372BF9542B598154C3A" author="Solem" pageId="20" pageNumber="23" refString="Solem, A. (1966) The neotropical land snail genera Labyrinthus and Isomeria (Pulmonata, Camaenidae). Fieldiana, Zoology, 50, 1 - 226." type="journal article" year="1966">Solem 1966</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA16415D9B504C39" box="[350,514,524,549]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA16415D9B504C39" box="[350,514,524,549]" italics="true" pageId="20" pageNumber="21">Isomeria awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is most similar in shell morphology to
|
|||
|
|
<emphasis id="B979CC67FFD2E372B8A2415D9D1E4C39" box="[1002,1100,524,549]" italics="true" pageId="20" pageNumber="21">
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B8A2415D9D1A4C39" box="[1002,1096,524,549]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oreas">I. oreas</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
differing in being smaller (adult specimens of
|
|||
|
|
<emphasis id="B979CC67FFD2E372B91941659BE34C51" box="[593,689,564,589]" italics="true" pageId="20" pageNumber="21">
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B91941659BFE4C51" box="[593,684,564,589]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oreas">I. oreas</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
mean diameter = 57.3 mm;
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B8B141659D104C51" box="[1017,1090,564,589]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B8B141659D104C51" box="[1017,1090,564,589]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
mean diameter=
|
|||
|
|
<quantity id="4CF5BD90FFD2E372BE45416598684C6A" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="20" pageNumber="21" unit="mm" value="39.0">39 mm</quantity>
|
|||
|
|
);
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA07410D98F54C69" box="[335,423,604,629]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oreas">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA07410D98F54C69" box="[335,423,604,629]" italics="true" pageId="20" pageNumber="21">I. oreas</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has fine radial striae and malleations on the surface of the body whorl, while
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA4041D598034C81" box="[264,337,644,669]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA4041D598034C81" box="[264,337,644,669]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has axial growth lines with small pustules all over shell surface; the shell coloration in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA6F41FD98D34CD9" box="[295,385,684,709]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oreas">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA6F41FD98D34CD9" box="[295,385,684,709]" italics="true" pageId="20" pageNumber="21">I. oreas</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is darker than in
|
|||
|
|
<emphasis id="B979CC67FFD2E372B91B41FD9BF64CD9" box="[595,676,684,709]" italics="true" pageId="20" pageNumber="21">
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B91B41FD9BF24CD9" box="[595,672,684,709]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">I. awa</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
without the characteristics irregular spots present in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BE5141FD986A4CF1" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BE5141FD986A4CF1" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
shells. The number of shell whorls is lower in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B81241859AF14CF1" box="[858,931,724,749]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B81241859AF14CF1" box="[858,931,724,749]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(41/2 whorls) than in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BFED41859DAF4CF1" box="[1189,1277,724,749]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oreas">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BFED41859DAF4CF1" box="[1189,1277,724,749]" italics="true" pageId="20" pageNumber="21">I. oreas</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(5– 5 1/2 whorls). Comparison based on anatomical characters is not possible because the anatomy of
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BADB407598B94D21" box="[403,491,804,829]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oreas">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BADB407598B94D21" box="[403,491,804,829]" italics="true" pageId="20" pageNumber="21">I. oreas</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is unknown.
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B9CE40759D194D22" authority="Mousson, 1869" authorityName="Mousson" authorityYear="1869" box="[646,1099,804,830]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="subelliptica">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B9CE40759A2C4D21" box="[646,894,804,829]" italics="true" pageId="20" pageNumber="21">Isomeria subelliptica</emphasis>
|
|||
|
|
(Mousson, 1869)
|
|||
|
|
</taxonomicName>
|
|||
|
|
is similar to
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BFAA40759C794D21" box="[1250,1323,804,829]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BFAA40759C794D21" box="[1250,1323,804,829]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in that they both possess a small parietal tooth and open umbilicus, but
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BF1A401D9DCE4D79" box="[1106,1180,844,869]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BF1A401D9DCE4D79" box="[1106,1180,844,869]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has a larger shell [mean diameter of
|
|||
|
|
<quantity id="4CF5BD90FFD2E372B97E40259BC24D92" box="[566,656,884,910]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="20" pageNumber="21" unit="mm" value="39.0">39 mm</quantity>
|
|||
|
|
, while
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B9A540259AC14D91" box="[749,915,884,909]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="subelliptica">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B9A540259AC14D91" box="[749,915,884,909]" italics="true" pageId="20" pageNumber="21">I. subelliptica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
shell mean diameter is 30.9 mm (
|
|||
|
|
<bibRefCitation id="EF9C6D84FFD2E372BA5840CD98F64DAA" author="Solem" box="[272,420,924,950]" pageId="20" pageNumber="23" refString="Solem, A. (1966) The neotropical land snail genera Labyrinthus and Isomeria (Pulmonata, Camaenidae). Fieldiana, Zoology, 50, 1 - 226." type="journal article" year="1966">Solem 1966</bibRefCitation>
|
|||
|
|
)], the umbilicus is narrower than in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B80240CD9AB84DA9" box="[842,1002,924,949]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="subelliptica">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B80240CD9AB84DA9" box="[842,1002,924,949]" italics="true" pageId="20" pageNumber="21">I. subelliptica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Both species also differ in shell color,
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BAD9409598894DC1" box="[401,475,964,989]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BAD9409598894DC1" box="[401,475,964,989]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has a golden yellowish coloration with darker irregular spots while in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BE51409598C34A19" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="subelliptica">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BE51409598C34A19" italics="true" pageId="20" pageNumber="21">I. subelliptica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
the shell possess an even dark brown coloration.
|
|||
|
|
<collectingCountry id="F31A50E5FFD2E372B8B840BD9D424A1A" box="[1008,1040,1004,1030]" name="American Samoa" pageId="20" pageNumber="21">As</collectingCountry>
|
|||
|
|
in the case of
|
|||
|
|
<emphasis id="B979CC67FFD2E372BF8040BD9C784A19" box="[1224,1322,1004,1029]" italics="true" pageId="20" pageNumber="21">
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BF8040BD9C744A19" box="[1224,1318,1004,1029]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oreas">I. oreas</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
anatomical comparisons of
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B91A47459BCF4A31" box="[594,669,1044,1069]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B91A47459BCF4A31" box="[594,669,1044,1069]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
with
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B9AC47459AD54A31" box="[740,903,1044,1069]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="subelliptica">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B9AC47459AD54A31" box="[740,903,1044,1069]" italics="true" pageId="20" pageNumber="21">I. subelliptica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are not possible due to the lack of information on the later.
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B90E476D9A4A4A49" box="[582,792,1084,1109]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="globosa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B90E476D9A4A4A49" box="[582,792,1084,1109]" italics="true" pageId="20" pageNumber="21">Isomeria globosa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, whose anatomy was described by
|
|||
|
|
<bibRefCitation id="EF9C6D84FFD2E372BFA8476D980B4A62" author="Solem" pageId="20" pageNumber="23" refString="Solem, A. (1966) The neotropical land snail genera Labyrinthus and Isomeria (Pulmonata, Camaenidae). Fieldiana, Zoology, 50, 1 - 226." type="journal article" year="1966">Solem (1966)</bibRefCitation>
|
|||
|
|
, differs from
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BAB347359B174A61" box="[507,581,1124,1149]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BAB347359B174A61" box="[507,581,1124,1149]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in not having the vagina's stimulator apparatus. Hooked denticles in vagina and penis, and a glandular pouch are present in
|
|||
|
|
<emphasis id="B979CC67FFD2E372B8A047DD9D344AB9" box="[1000,1126,1164,1189]" italics="true" pageId="20" pageNumber="21">
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B8A047DD9D314AB9" box="[1000,1123,1164,1189]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="globosa">I. globosa</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
A penial appendix is lacking in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA8447E59B474AD1" box="[460,533,1204,1229]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA8447E59B474AD1" box="[460,533,1204,1229]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
but it was described in
|
|||
|
|
<emphasis id="B979CC67FFD2E372B86347E59AFA4AD1" box="[811,936,1204,1229]" italics="true" pageId="20" pageNumber="21">
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B86347E59AF64AD1" box="[811,932,1204,1229]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="globosa">I. globosa</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BB21075FFD2E373BA70478D9B124F42" blockId="20.[264,1323,283,1870]" lastBlockId="21.[264,1323,284,350]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
|
|||
|
|
Two structures in the genitalia of
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B984478D9A454AE9" box="[716,791,1244,1269]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B984478D9A454AE9" box="[716,791,1244,1269]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
probably represent unique characters within the genus, the presence of a stimulatorlike organ within the vagina and a glandular pouch externally visible in the distal portion of the vagina. This kind of stimulatorlike organ has never been reported before in any
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B98C46059A074B72" box="[708,853,1364,1390]" class="Gastropoda" family="Camaenidae" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="family">Camaenidae</taxonomicName>
|
|||
|
|
and resembles the ones described in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA40462D9B604B8A" authority="Wiktor 1987" authorityName="Wiktor" authorityYear="1987" box="[264,562,1404,1430]" class="Gastropoda" family="Milacidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="family">
|
|||
|
|
Milacidae (
|
|||
|
|
<bibRefCitation id="EF9C6D84FFD2E372BAC5462D9B7B4B8A" author="Wiktor" box="[397,553,1404,1430]" pageId="20" pageNumber="23" refString="Wiktor, A. (1987) Milacidae (Gastropoda, Pulmonata): systematic monograph. Annales Zoologici, 41, 153 - 319." type="journal article" year="1987">Wiktor 1987</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Other stimulatorlike appendages have been reported in certain
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA4046F59B204BA2" authority="Hausdorf 1998" authorityName="Hausdorf" authorityYear="1998" box="[264,626,1444,1470]" class="Gastropoda" family="Hygromiidae" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="family">
|
|||
|
|
Hygromiidae (
|
|||
|
|
<bibRefCitation id="EF9C6D84FFD2E372BAFB46F59B3B4BA2" author="Hausdorf" box="[435,617,1444,1470]" pageId="20" pageNumber="23" refString="Hausdorf, B. (1998) Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora). Journal of Molluscan Studies, 64, 35 - 66." type="journal article" year="1998">Hausdorf 1998</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
. However, several appendages have been called as "stimulators" (
|
|||
|
|
<bibRefCitation id="EF9C6D84FFD2E372BA20469D9B794BFA" author="Nordsieck" box="[360,555,1484,1510]" pageId="20" pageNumber="23" refString="Nordsieck, H. (1987) Revision des Systems der Helicoidea (Gastropoda: Stylommatophora). Archiv fur Molluskenkunde, 118, 9 - 50." type="journal article" year="1987">Nordsieck 1987</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EF9C6D84FFD2E372B973469D9BAB4BFA" author="Schileyko" box="[571,761,1484,1510]" pageId="20" pageNumber="23" refString="Schileyko, A. A. (1991) Taxonomic status, phylogenetic relations and system of the Helicoidea sensu lato (Pulmonata). Archiv fur Molluskenkunde, 120, 187 - 236." type="journal article" year="1991">Schileyko 1991</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EF9C6D84FFD2E372B841469D9AEF4BFA" author="Hausdorf" box="[777,957,1484,1510]" pageId="20" pageNumber="23" refString="Hausdorf, B. (1998) Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora). Journal of Molluscan Studies, 64, 35 - 66." type="journal article" year="1998">Hausdorf 1998</bibRefCitation>
|
|||
|
|
) so that the homology of this kind of structures can only be tested by phylogenetic analyses. The glandular pouch found in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA6E454D98224829" box="[294,368,1564,1589]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA6E454D98224829" box="[294,368,1564,1589]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is also present in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B90A454D9BE54829" box="[578,695,1564,1589]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="globosa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B90A454D9BE54829" box="[578,695,1564,1589]" italics="true" pageId="20" pageNumber="21">I. globosa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in which it has been described as a ring of low knobs around the vagina (
|
|||
|
|
<bibRefCitation id="EF9C6D84FFD2E372BABF45159BDE4842" author="Solem" box="[503,652,1604,1630]" pageId="20" pageNumber="23" refString="Solem, A. (1966) The neotropical land snail genera Labyrinthus and Isomeria (Pulmonata, Camaenidae). Fieldiana, Zoology, 50, 1 - 226." type="journal article" year="1966">Solem 1966</bibRefCitation>
|
|||
|
|
). A glandular pouch is characteristic of some species of
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA40453D9B19489A" authority="Montfort, 1810" authorityName="Montfort" authorityYear="1810" box="[264,587,1644,1670]" class="Gastropoda" family="Camaenidae" genus="Polydontes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA40453D98D84899" box="[264,394,1644,1669]" italics="true" pageId="20" pageNumber="21">Polydontes</emphasis>
|
|||
|
|
Montfort, 1810
|
|||
|
|
</taxonomicName>
|
|||
|
|
, also member of the
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B80D453D9A87489A" box="[837,981,1644,1670]" class="Gastropoda" family="Camaenidae" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="family">Camaenidae</taxonomicName>
|
|||
|
|
, however, this kind of structure appears in the penial complex but not in the vagina. Hooked denticles with oval bases are also present, both in vagina and penis, in some
|
|||
|
|
<emphasis id="B979CC67FFD2E372B82045ED9AAF48C9" box="[872,1021,1724,1749]" italics="true" pageId="20" pageNumber="21">
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B82045ED9AAB48C9" box="[872,1017,1724,1749]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">Labyrinthus</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
with similar morphology although they are smaller in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B91D45B59BCC48E1" box="[597,670,1764,1789]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B91D45B59BCC48E1" box="[597,670,1764,1789]" italics="true" pageId="20" pageNumber="21">I. awa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. These denticles are usually more abundant in the penis than in the vagina in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B944445D9BC84939" box="[524,666,1804,1829]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B944445D9BC84939" box="[524,666,1804,1829]" italics="true" pageId="20" pageNumber="21">Labyrinthus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as well as in
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372B80F445D9AFD4939" box="[839,943,1804,1829]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B979CC67FFD2E372B80F445D9AFD4939" box="[839,943,1804,1829]" italics="true" pageId="20" pageNumber="21">Isomeria</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Another character shared with
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BA40446598C44951" box="[264,406,1844,1869]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B979CC67FFD2E372BA40446598C44951" box="[264,406,1844,1869]" italics="true" pageId="20" pageNumber="21">Labyrinthus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is the presence of an internal flagellar pilaster, which in
|
|||
|
|
<emphasis id="B979CC67FFD2E372BF7444659DDC4951" box="[1084,1166,1844,1869]" italics="true" pageId="20" pageNumber="21">
|
|||
|
|
<taxonomicName id="4C0D6BF6FFD2E372BF7444659DD84951" box="[1084,1162,1844,1869]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="awa">I. awa</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
shows a distinct open dorsal groove. Marginal teeth were not observed in the radulas preparations because they were broken.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|