treatments-xml/data/03/8E/87/038E879AFFF8D1221DB2FC8DFAC4E2EF.xml

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<mods:title id="B99451ECC898584B35EC4CFA46A93783">New species and records for the mayfly families Caenidae, Leptohyphidae and Coryphoridae (Ephemeroptera, Pannota) from Venezuelan Guayana’s Uplands</mods:title>
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<mods:namePart id="311B7F4EA2C0F04C7457D9B494C185F2">Dominguez, Eduardo</mods:namePart>
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<heading id="D0D081E0FFF8D12D1DB2FC8DFD54E026" bold="true" box="[151,710,769,795]" fontSize="11" level="1" pageId="7" pageNumber="46" reason="1">
<taxonomicName id="4C274D0FFFF8D12D1DB2FC8DFD54E026" ID-CoL="58N6D" authority="Dias, Salles &amp; Ferreira" authorityName="Dias, Salles &amp; Ferreira" box="[151,710,769,795]" class="Insecta" family="Leptohyphidae" genus="Tricorythopsis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="yucupe">
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<emphasis id="B953EA9EFFF8D12D1DB2FC8DFE33E026" bold="true" box="[151,417,769,795]" italics="true" pageId="7" pageNumber="46">Tricorythopsis yucupe</emphasis>
Dias, Salles &amp; Ferreira
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(
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)
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<taxonomicName id="4C274D0FFFF8D12D1DB2FCE0FD49E0BC" ID-CoL="58N6D" authority="Dias, Salles &amp; Ferreira, 2008: 238" authorityName="Dias, Salles &amp; Ferreira" authorityPageNumber="238" authorityYear="2008" box="[151,731,876,898]" class="Insecta" family="Leptohyphidae" genus="Tricorythopsis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="yucupe">
<emphasis id="B953EA9EFFF8D12D1DB2FCE0FEE5E0BF" box="[151,375,876,898]" italics="true" pageId="7" pageNumber="46">Tricorythopsis yucupe</emphasis>
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: 238
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.
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<subSubSection id="C33D6507FFF8D12D1DB2FC22FB41E586" pageId="7" pageNumber="46" type="materials_examined">
<paragraph id="8B98368CFFF8D12D1DB2FC22FE6DE746" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="46">
<emphasis id="B953EA9EFFF8D12D1DB2FC22FE94E0FA" bold="true" box="[151,262,942,967]" pageId="7" pageNumber="46">Material.</emphasis>
<collectingCountry id="F330761CFFF8D12D1C2BFC23FE16E0FA" box="[270,388,943,967]" name="Venezuela" pageId="7" pageNumber="46">Venezuela</collectingCountry>
, Estado Bolivar: 2 nymphs from Soroape, 
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– 
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, 
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, 
<date id="FF99104CFFF8D12D1830FC23FA05E0FA" box="[1301,1431,943,967]" pageId="7" pageNumber="46" value="2006-03-21">21/III/2006</date>
, EG, AMO and MEG cols.; 3 nymphs from Aponwao, 
<geoCoordinate id="EE13504BFFF8D12D1FD4FC58FCFFE0D6" box="[753,877,979,1003]" direction="north" orientation="latitude" pageId="7" pageNumber="46" precision="15" value="5.8505554">N 5° 51' 2&quot;</geoCoordinate>
– 
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, 
<quantity id="4CDF9B69FFF8D12D1916FC5FFB18E0D6" box="[1075,1162,979,1003]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="7" pageNumber="46" unit="m" value="1340.0">1340 m</quantity>
, 
<date id="FF99104CFFF8D12D19B0FC5FFA8BE0D6" box="[1173,1305,979,1003]" pageId="7" pageNumber="46" value="2007-04-27">27/IV/2007</date>
, EG, AMO, CA and MEG cols.; 1 nymph idem except date: 
<date id="FF99104CFFF8D12D1F90FC7BFCABE732" box="[693,825,1015,1039]" pageId="7" pageNumber="46" value="2006-03-18">18/III/2006</date>
; 2 Ƥ imagos and 2 nymphs from Mareman-Parú, 
<geoCoordinate id="EE13504BFFF8D12D1842FC74FF7AE70E" direction="north" orientation="latitude" pageId="7" pageNumber="46" precision="15" value="5.746945">N 5° 44' 49&quot;</geoCoordinate>
– 
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, 
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, 
<date id="FF99104CFFF8D12D1F32FB97FD09E70E" box="[535,667,1051,1075]" pageId="7" pageNumber="46" value="2007-06-27">27/VI/2007</date>
, EG, AMO, CA and MEG cols.; 1 pharate 3 subimago (slide 513) from Kamá, 
<geoCoordinate id="EE13504BFFF8D12D1C03FBCCFE23E76A" box="[294,433,1087,1111]" direction="north" orientation="latitude" pageId="7" pageNumber="46" precision="15" value="5.419722">N 5° 25' 11&quot;</geoCoordinate>
– 
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, 
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, 
<date id="FF99104CFFF8D12D1FF6FBB3FCC4E76A" box="[723,854,1087,1111]" pageId="7" pageNumber="46" value="2005-11-24">24/XI/2005</date>
, EG, AMO, CA and MEG cols. Four nymphs and 1 Ƥ imago in 
<collectionCode id="ED36AE49FFF8D12D1C39FBE8FEFEE746" box="[284,364,1124,1147]" pageId="7" pageNumber="46">MLBV</collectionCode>
, rest in 
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.
</paragraph>
<paragraph id="8B98368CFFF8D12D1DE0FB0BFCF6E682" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="46">
Female imago (
<figureCitation id="131C2A09FFF8D12D1C51FB0BFE5AE7A2" box="[372,456,1159,1183]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 18</figureCitation>
). Length (mm): body, 2.7–2.8; forewing, 3.1–3.2; caudal filaments, 0.6. General coloration yellowish brown shaded gray. Head yellowish except occiput and ecdysial suture whitish, shaded gray on hind margin behind eyes. Antennae yellowish translucent. Thorax. Pronotum hyaline except lateral regions yellowish, shaded gray almost completely. Mesonotum yellowish brown shaded widely with gray; membranous filaments short and wide, medially curved. Thoracic sterna paler. Legs. Forelegs absent as usual in the genus, middle and hind legs yellowish with gray shading on femora (except small pale marks) and subapical band on tibiae. Wings. Membrane hyaline, longitudinal veins blackish. Abdomen (
<figureCitation id="131C2A09FFF8D12D1E15FAD3FC11E64A" box="[816,899,1375,1399]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 18</figureCitation>
). Segments yellowish white shaded widely with gray dorsally, except on anterior and posterior margins of all terga, on lateral zones of terga 1–6, and pale anterolateral marks on tergum 7. Caudal filaments yellowish translucent.
</paragraph>
<paragraph id="8B98368CFFF8D12D1DE0FA47FEA0E572" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="46">
Male subimago, pharate. Body length, 
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. General coloration as female, except head paler, and abdominal terga 1–7 not shaded gray on lateral zones. Genitalia (
<figureCitation id="131C2A09FFF8D12D1FD9FA63FC15E53A" box="[764,903,1519,1543]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Figs. 20–21</figureCitation>
): styliger plate with long projections forming a columnar base for each forceps (sp in 
<figureCitation id="131C2A09FFF8D12D1F60F99FFD0BE516" box="[581,665,1555,1579]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 20</figureCitation>
); penes slender, medially narrow with a moderately expanded apical part (
<figureCitation id="131C2A09FFF8D12D1DF7F9BBFEB7E572" box="[210,293,1591,1615]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8B98368CFFF8D12D1DE0F9D7FB41E586" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="46">
Nymphs (
<figureCitation id="131C2A09FFF8D12D1C10F9D7FE18E54E" box="[309,394,1627,1651]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 19</figureCitation>
). Length (mm): body, 2.2 (male)–2.7 (female); terminal filaments, 1.8 (male)–2.3 (female); cerci, 1.9 (male)–2.5 (female). As in the original description except that abdominal dorsal tubercles (
<figureCitation id="131C2A09FFF8D12D1833F9F3FAFBE5AA" box="[1302,1385,1663,1687]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 19</figureCitation>
) are present on segments 6 to 9, much larger on 7–8 (original description reported them in terga 7–9).
</paragraph>
</subSubSection>
<subSubSection id="C33D6507FFF8D1221DE0F94AFAC4E2EF" lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46" type="discussion">
<paragraph id="8B98368CFFF8D1221DE0F94AFF57E202" blockId="7.[151,1437,942,2011]" lastBlockId="8.[151,1436,151,467]" lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46">
<emphasis id="B953EA9EFFF8D12D1DE0F94AFED8E5E2" bold="true" box="[197,330,1734,1759]" pageId="7" pageNumber="46">Discussion.</emphasis>
The nymphs are easily distinguished from other species of the genus by their large size and other characters listed in 
<bibRefCitation id="EFB64B7DFFF8D12D1C5EF967FDDFE43E" box="[379,589,1771,1795]" pageId="7" pageNumber="49" refString="Dias, L. G., Salles, F. F. &amp; Ferreira, P. S. F. (2008) New species of Tricorythopsis Traver (Ephemeroptera: Leptohyphidae) from northern Brazil. Studies on Neotropical Fauna and Environment, 43, 237 - 241." type="journal article">
Dias 
<emphasis id="B953EA9EFFF8D12D1C9CF960FE67E43E" box="[441,501,1771,1795]" italics="true" pageId="7" pageNumber="46">et al.</emphasis>
(2008)
</bibRefCitation>
, mainly the mesonotal paired keels on hind margin (mk in 
<figureCitation id="131C2A09FFF8D12D182EF967FAF2E43E" box="[1291,1376,1771,1795]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 19</figureCitation>
), the abdominal dorsal tubercles (at in 
<figureCitation id="131C2A09FFF8D12D1F36F883FDF4E41A" box="[531,614,1807,1831]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 19</figureCitation>
), the operculate gills with 3 dark marks (
<figureCitation id="131C2A09FFF8D12D1918F883FB00E41A" box="[1085,1170,1807,1831]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 22</figureCitation>
), the pale head without gray shading, contrary to the rest of body widely shaded with gray and black. The adults can be distinguished from the other species by the following combination of characters: 1) large size (body length&gt; 
<quantity id="4CDF9B69FFF8D12D1984F8DBFB6CE452" box="[1185,1278,1879,1903]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="7" pageNumber="46" unit="mm" value="2.7">2.7 mm</quantity>
); 2) abdomen shaded with gray almost completely except at lateral margins of terga 1–6 and pale anterior marks at each side of tergum 7 (
<figureCitation id="131C2A09FFF8D12D1C2CF813FECEE48A" box="[265,348,1951,1975]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 18</figureCitation>
); 3) middle and hind femora shaded widely with gray or black; 4) penes slender, medially narrow but widening in distal portion (
<figureCitation id="131C2A09FFF8D12D1CCBF84FFDD3E4E6" box="[494,577,1987,2011]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="7" pageNumber="46">Fig. 21</figureCitation>
). The combination of mesonotal keel, abdominal dorsal tubercles and projections on middle and hind coxae is also present in 
<taxonomicName id="4C274D0FFFF7D1221FE3FF15FC14E392" authority="Allen" authorityName="Allen" box="[710,902,151,176]" class="Insecta" family="Leptohyphidae" genus="Tricorythopsis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="gibbus">
<emphasis id="B953EA9EFFF7D1221FE3FF15FCBFE38D" box="[710,813,152,176]" italics="true" pageId="8" pageNumber="47">T. gibbus</emphasis>
(Allen)
</taxonomicName>
and 
<taxonomicName id="4C274D0FFFF7D1221E98FF15FAC3E392" authority="Dias &amp; Salles. The" authorityName="Dias &amp; Salles. The" box="[957,1361,151,176]" class="Insecta" family="Leptohyphidae" genus="Tricorythopsis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudogibbus">
<emphasis id="B953EA9EFFF7D1221E98FF15FBE6E38D" box="[957,1140,152,176]" italics="true" pageId="8" pageNumber="47">T. pseudogibbus</emphasis>
Dias &amp; Salles. The
</taxonomicName>
adults also share some features with 
<taxonomicName id="4C274D0FFFF7D1221CC0FF31FDD8E3E9" box="[485,586,188,212]" class="Insecta" family="Leptohyphidae" genus="Tricorythopsis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="gibbus">
<emphasis id="B953EA9EFFF7D1221CC0FF31FDD8E3E9" box="[485,586,188,212]" italics="true" pageId="8" pageNumber="47">T. gibbus</emphasis>
</taxonomicName>
(those of 
<taxonomicName id="4C274D0FFFF7D1221F9FFF31FCFDE3E9" box="[698,879,188,212]" class="Insecta" family="Leptohyphidae" genus="Tricorythopsis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudogibbus">
<emphasis id="B953EA9EFFF7D1221F9FFF31FCFDE3E9" box="[698,879,188,212]" italics="true" pageId="8" pageNumber="47">T. pseudogibbus</emphasis>
</taxonomicName>
are unknown): the widely pigmented femora, and the relatively great expansion of the distal lobes of the penes (
<figureCitation id="131C2A09FFF7D1221E7EFF53FC22E3CA" box="[859,944,223,247]" captionStart="FIGURES 14 – 22" captionStartId="8.[151,264,1849,1872]" captionTargetBox="[192,1399,522,1828]" captionTargetId="figure@8.[187,1399,522,1828]" captionTargetPageId="8" captionText="FIGURES 14 – 22. Macunahyphes pemonensis sp. nov.; 14, male imago; 15, female imago; 16, male fore wing. Macunahyphes incognitus: 17, male fore wing. Tricorythopsis yucupe: 18, female imago; 19, nymph, l. v. (at = abdominal tubercle, mk = mesonotal keel); 20, styliger and forceps of pharate male adult (sp = styliger projections, f 1 = forceps segment 1, f 2 = forceps segment 2); 21, penes of pharate male; 22, operculate gill, d. v. (tl = transverse line)." httpUri="https://zenodo.org/record/206968/files/figure.png" pageId="8" pageNumber="47">Fig. 21</figureCitation>
). Nevertheless, the strong narrowing of the penes on the medial portion, somewhat resembling those of 
<taxonomicName id="4C274D0FFFF7D1221E18FE89FC6FE221" authority="Traver" authorityName="Traver" box="[829,1021,261,284]" class="Insecta" family="Leptohyphidae" genus="Tricorythopsis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="artigas">
<emphasis id="B953EA9EFFF7D1221E18FE89FC3AE221" box="[829,936,261,284]" italics="true" pageId="8" pageNumber="47">T. artigas</emphasis>
Traver
</taxonomicName>
, distinguishes 
<taxonomicName id="4C274D0FFFF7D1221987FE89FA9FE221" box="[1186,1293,261,284]" class="Insecta" family="Leptohyphidae" genus="Tricorythopsis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="yucupe">
<emphasis id="B953EA9EFFF7D1221987FE89FA9FE221" box="[1186,1293,261,284]" italics="true" pageId="8" pageNumber="47">T. yucupe</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C274D0FFFF7D1221876FE89FF2CE27D" class="Insecta" family="Leptohyphidae" genus="Tricorythopsis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="gibbus">
<emphasis id="B953EA9EFFF7D1221876FE89FF2CE27D" italics="true" pageId="8" pageNumber="47">T. gibbus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B98368CFFF7D1221DE0FEC0FAC4E2EF" blockId="8.[151,1436,151,467]" pageId="8" pageNumber="47">
<emphasis id="B953EA9EFFF7D1221DE0FEC0FEB8E259" bold="true" box="[197,298,332,356]" pageId="8" pageNumber="47">Ecology.</emphasis>
Imagos were collected along the river margins, at night, during the rainy season (
<date id="FF99104CFFF7D1221993FEC0FAFFE259" box="[1206,1389,332,356]" pageId="8" pageNumber="47" value="2005-11">November 2005</date>
and 
<date id="FF99104CFFF7D1221DB2FEFCFE99E2BA" box="[151,267,367,391]" pageId="8" pageNumber="47" value="2007-06">June 2007</date>
) and dry season (
<date id="FF99104CFFF7D1221CE8FEE3FDC9E2BA" box="[461,603,367,391]" pageId="8" pageNumber="47" value="2006-03">March 2006</date>
), using light traps, whereas, nymphs were collected within the rivers using a D-net. All the sampled rivers (
<figureCitation id="131C2A09FFF7D1221F27FE18FD1EE291" box="[514,652,404,428]" captionStart="FIGURES 23 – 26" captionStartId="9.[151,264,1742,1765]" captionTargetBox="[209,1371,181,1723]" captionTargetId="figure@9.[209,1372,181,1723]" captionTargetPageId="9" captionText="FIGURES 23 – 26. Study sites: 23, Kaku Parú; 24, Kavanayén; 25, Mareman Parú; 26, Aponwao." httpUri="https://zenodo.org/record/206969/files/figure.png" pageId="8" pageNumber="47">Figs. 25–26</figureCitation>
), flowing through the upland savanna ecosystem (
<quantity id="4CDF9B69FFF7D122199AFE18FAC3E291" box="[1215,1361,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.1375" metricValueMax="1.34" metricValueMin="0.9349999999999999" pageId="8" pageNumber="47" unit="m" value="1137.5" valueMax="1340.0" valueMin="935.0">935–1340 m</quantity>
), were large (
<quantity id="4CDF9B69FFF7D1221DF8FE36FEDDE2EF" box="[221,335,442,466]" metricMagnitude="1" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="8" pageNumber="47" unit="m" value="65.0" valueMax="100.0" valueMin="30.0">30–100 m</quantity>
wide;&gt; 4th order), non shaded, and bed-rock bottom rivers, with very acidic waters (pH=5).
</paragraph>
</subSubSection>
</treatment>
</document>