317 lines
41 KiB
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317 lines
41 KiB
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<document ID-DOI="http://doi.org/10.5281/zenodo.5943533" ID-GBIF-Dataset="775fe351-8dad-4490-9d4d-34701f62b5c1" ID-GBIF-Taxon="151742853" ID-Zenodo-Dep="5943533" checkinTime="1548713912964" checkinUser="plazi" docAuthor="Evgeny Viktorovich Yana, Rolf Georg Beutel, John Francis Lawrence, Margarita Igorevna Yavorskaya, Thomas Hörnschemeyer, Hans Pohl, Dmitry Vladimirovich Vassilenko, Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" docDate="2019" docId="038E878E224F2213F26AFAE4FD9AFD15" docLanguage="en" docName="historicalBiology.08912963.2018.1561672.pdf" docOrigin="Historical Biology 1561672" docStyle="DocumentStyle{}" docTitle="Archaeomalthus Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko 2019, gen. nov." docType="treatment" docVersion="4" lastPageId="4" lastPageNumber="5" masterDocId="FFB7FFF6224D2217F362FFCBFFB0FFD3" masterDocTitle="Archaeomalthus - (Coleoptera, Archostemata) a ‘ ghost adult ’ of Micromalthidae from Upper Permian deposits of Siberia?" masterLastPageNumber="9" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547027548802" originalUpdateUser="plazi" pageId="2" pageNumber="3" updateTime="1643761532324" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Archaeomalthus - (Coleoptera, Archostemata) a ‘ ghost adult ’ of Micromalthidae from Upper Permian deposits of Siberia?</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Evgeny Viktorovich Yana</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rolf Georg Beutel</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>John Francis Lawrence</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Margarita Igorevna Yavorskaya</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Thomas Hörnschemeyer</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hans Pohl</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dmitry Vladimirovich Vassilenko</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Alexey Semenovich Bashkuev</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Alexander Georgievich Ponomarenko</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Historical Biology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2019-01-07</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1561672</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>9</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1080/08912963.2018.1561672</mods:identifier>
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<mods:identifier type="GBIF-Dataset">775fe351-8dad-4490-9d4d-34701f62b5c1</mods:identifier>
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<mods:identifier type="Zenodo-Dep">2536029</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5943533" ID-GBIF-Taxon="151742853" ID-Zenodo-Dep="5943533" LSID="urn:lsid:plazi:treatment:038E878E224F2213F26AFAE4FD9AFD15" httpUri="http://treatment.plazi.org/id/038E878E224F2213F26AFAE4FD9AFD15" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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<subSubSection pageId="2" pageNumber="3" type="nomenclature">
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<paragraph blockId="2.[148,778,1327,1415]" box="[264,662,1327,1351]" pageId="2" pageNumber="3">
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<heading box="[264,662,1327,1351]" centered="true" fontSize="10" level="4" pageId="2" pageNumber="3" reason="4">
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Genus
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<emphasis bold="true" box="[343,547,1327,1351]" pageId="2" pageNumber="3">
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†
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<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[357,547,1327,1351]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[357,547,1327,1351]" italics="true" pageId="2" pageNumber="3">Archaeomalthus</emphasis>
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</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[556,662,1327,1351]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph blockId="2.[148,778,1327,1415]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[148,302,1359,1383]" pageId="2" pageNumber="3">Type species</emphasis>
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.
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<taxonomicName authority="Yan et al." authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[316,733,1359,1383]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="synoriacos" status="sp. nov.">
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<emphasis box="[316,616,1359,1383]" italics="true" pageId="2" pageNumber="3">Archaeomalthus synoriacos</emphasis>
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Yan et al.
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</taxonomicName>
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,
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<taxonomicNameLabel pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="diagnosis">
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<paragraph blockId="2.[128,797,1475,1724]" box="[128,238,1475,1500]" pageId="2" pageNumber="3">
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<heading bold="true" box="[128,238,1475,1500]" fontSize="10" level="3" pageId="2" pageNumber="3" reason="6">
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<emphasis bold="true" box="[128,238,1475,1500]" italics="true" pageId="2" pageNumber="3">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="2.[128,797,1475,1724]" pageId="2" pageNumber="3">
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Antennal sockets visible from above; propleuron exposed, narrow, not reaching anterior pronotal margin; procoxae slightly widened, almost contiguous medially; metanotum with metascutum delimited by very distinct longitudinal bulges (alacrista) (
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<figureCitation box="[337,438,1636,1660]" captionStart="Figure 1" captionStartId="2.[128,183,987,1006]" captionTargetBox="[302,1325,140,960]" captionTargetId="figure@2.[301,1325,139,962]" captionTargetPageId="2" captionText="Figure 1. Archaeomalthus synoriacos sp. nov., Habitus. (a) Scanning electron micrograph of the holotype 5381/32. (b) Interpretative drawing." httpUri="https://zenodo.org/record/2536031/files/figure.png" pageId="2" pageNumber="3">Figures 1</figureCitation>
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(b) and 2(i,k,l)); anterior intercoxal process of metaventrite strongly developed, completely separating mesocoxae. Abdomen with seven visible sternites.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="etymology">
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<paragraph blockId="2.[128,797,1768,1889]" box="[128,351,1768,1793]" pageId="2" pageNumber="3">
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<heading bold="true" box="[128,351,1768,1793]" fontSize="10" level="3" pageId="2" pageNumber="3" reason="6">
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<emphasis bold="true" box="[128,351,1768,1793]" italics="true" pageId="2" pageNumber="3">Derivation of name</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="2.[128,797,1768,1889]" pageId="2" pageNumber="3">
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Generic name from Greek ‘archaios’ – ‘ancient’ and
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<taxonomicName authority="LeConte, 1878" authorityName="LeConte" authorityYear="1878" box="[128,483,1833,1857]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis box="[128,277,1833,1857]" italics="true" pageId="2" pageNumber="3">Micromalthus</emphasis>
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LeConte, 1878
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</taxonomicName>
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, type genus of family
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<taxonomicName authorityName=", Barber" authorityYear="1913" box="[128,304,1865,1889]" class="Insecta" family="Micromalthidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Micromalthidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
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<paragraph blockId="2.[128,797,1933,2022]" box="[128,256,1933,1958]" pageId="2" pageNumber="3">
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<heading bold="true" box="[128,256,1933,1958]" fontSize="10" level="3" pageId="2" pageNumber="3" reason="6">
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<emphasis bold="true" box="[128,256,1933,1958]" italics="true" pageId="2" pageNumber="3">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="2.[128,797,1933,2022]" lastBlockId="2.[829,1498,1066,2018]" pageId="2" pageNumber="3">Body elongated, slender, probably slightly flattened, with distinct shoulder region, slightly rounded laterally posterior to prothorax. Total length ca. 1.18 mm, maximum width at posterior metathoracic region ca. 0.4 mm.</paragraph>
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<paragraph blockId="2.[829,1498,1066,2018]" pageId="2" pageNumber="3">Cuticular surface largely smooth on all body regions, without recognizable scales, punctures or tubercles.</paragraph>
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<paragraph blockId="2.[829,1498,1066,2018]" pageId="2" pageNumber="3">
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Head large, as long as pronotum, rectangular, slightly narrower than anterior pronotal margin; with widely separated well-developed compound eyes, inner margins of eyes are finely bordered; ocelli not recognizable, probably absent. Labrum not articulated, apparently fused with clypeus. Concave transverse line present on anterior head capsule, possibly indicating clypeofrontal strengthening ridge. Antennae inserted dorsally but still widely separated, articulatory area close to anteromesal edge of eyes. Mandibles sickle-shaped, with distal halves curved inwards. Maxillae not recognizable. Proximal elements of labium not visible as separate structures, submentum apparently fused with gula and ventral wall of head capsule (
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<figureCitation box="[1220,1316,1578,1603]" captionStart="Figure 2" captionStartId="3.[128,183,1045,1064]" captionTargetBox="[253,1373,156,1020]" captionTargetId="figure@3.[253,1373,139,1020]" captionTargetPageId="3" captionText="Figure 2. Details of the organization in extinct and recent Micromalthidae. (a) Archaeomalthus synoriacos sp. nov. head, scanning electron micrograph. (b) A. synoriacos sp. nov. head interpretative drawing. (c) Micromalthus debilis, head dorsal aspect, scanning electron micrograph. (d) M. debilis interpretative drawing of head dorsal aspect. (e) M. debilis, head ventral aspect, scanning electron micrograph. (f) M. debilis interpretative drawing of head ventral aspect. (g) A. synoriacos sp. nov. pronotum, scanning electron micrograph. (h) A. synoriacos sp. nov. pronotum interpretative drawing. (i) A. synoriacos sp. nov. metathorax, scanning electron micrograph. (j) A. synoriacos sp. nov. metathorax interpretative drawing. (k) M. debilis, thoracic dorsal aspect, scanning electron micrograph. (l) M. debilis interpretative drawing of thoracic dorsal aspect. (m) M. debilis prothoracic lateral aspect, scanning electron micrograph. (n) M. debilis photograph of thoracic ventrall aspect. (m) M. debilis interpretative drawing of thoracic ventral aspect." httpUri="https://zenodo.org/record/2536033/files/figure.png" pageId="2" pageNumber="3">Figure 2</figureCitation>
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(a,b)). Temporal region posterior to eyes very long, more than half as long as the length of head; narrowed neck region absent. Dorsal protuberances missing. Gular plate present, large.
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</paragraph>
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<paragraph blockId="2.[829,1498,1066,2018]" pageId="2" pageNumber="3">Pronotum almost quadrangular, as long as wide, with distinct lateral carina; anterior and posterior angles distinct, triangular; anterior pronotal margin at the same level as the anterior prosternal edge. Prosternal intercoxal process not visible, apparently missing; wide membranous area present posterior to procoxae. Large spiracle recognizable in prothoracic-mesothoracic intersegmental region.</paragraph>
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<paragraph blockId="2.[829,1498,1066,2018]" lastBlockId="3.[128,797,1294,1926]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">Elytral bases distinctly wider than maximum pronotal width; pronoto-elytral angle, i.e. humeral bulges, very distinct, rounded. The shape of elytra roughly quadrangular, with truncated apices. Mesal base enclosed by the semioval scutellar shield. Window punctures and other patterns of surface sculpture absent, degree of sclerotization uniform.</paragraph>
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<caption httpUri="https://zenodo.org/record/2536033/files/figure.png" pageId="3" pageNumber="4" startId="3.[128,183,1045,1064]" targetBox="[253,1373,156,1020]" targetPageId="3">
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<paragraph blockId="3.[128,1499,1045,1209]" pageId="3" pageNumber="4">
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<emphasis bold="true" box="[128,209,1045,1064]" pageId="3" pageNumber="4">Figure 2.</emphasis>
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Details of the organization in extinct and recent
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<taxonomicName authorityName=", Barber" authorityYear="1913" box="[658,790,1046,1065]" class="Insecta" family="Micromalthidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Micromalthidae</taxonomicName>
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. (a)
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<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[833,1058,1045,1065]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="synoriacos" status="sp. nov.">
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<emphasis box="[833,1058,1045,1065]" italics="true" pageId="3" pageNumber="4">Archaeomalthus synoriacos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1068,1138,1046,1065]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
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head, scanning electron micrograph. (b)
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<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[128,233,1069,1089]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="synoriacos" status="sp. nov.">
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<emphasis box="[128,233,1069,1089]" italics="true" pageId="3" pageNumber="4">A. synoriacos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[239,305,1070,1089]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
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head interpretative drawing. (c)
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<taxonomicName box="[578,743,1069,1089]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="debilis">
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<emphasis box="[578,743,1069,1089]" italics="true" pageId="3" pageNumber="4">Micromalthus debilis</emphasis>
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</taxonomicName>
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, head dorsal aspect, scanning electron micrograph. (d)
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<taxonomicName box="[1207,1284,1069,1089]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="debilis">
|
|||
|
|
<emphasis box="[1207,1284,1069,1089]" italics="true" pageId="3" pageNumber="4">M. debilis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
interpretative drawing of head dorsal aspect. (e)
|
|||
|
|
<taxonomicName box="[322,398,1093,1113]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="debilis">
|
|||
|
|
<emphasis box="[322,398,1093,1113]" italics="true" pageId="3" pageNumber="4">M. debilis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, head ventral aspect, scanning electron micrograph. (f)
|
|||
|
|
<taxonomicName box="[865,941,1093,1113]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="debilis">
|
|||
|
|
<emphasis box="[865,941,1093,1113]" italics="true" pageId="3" pageNumber="4">M. debilis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
interpretative drawing of head ventral aspect. (g)
|
|||
|
|
<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[1363,1468,1093,1113]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="synoriacos" status="sp. nov.">
|
|||
|
|
<emphasis box="[1363,1468,1093,1113]" italics="true" pageId="3" pageNumber="4">A. synoriacos</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
pronotum, scanning electron micrograph. (h)
|
|||
|
|
<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[558,664,1117,1137]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="synoriacos" status="sp. nov.">
|
|||
|
|
<emphasis box="[558,664,1117,1137]" italics="true" pageId="3" pageNumber="4">A. synoriacos</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[671,738,1118,1137]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
pronotum interpretative drawing. (i)
|
|||
|
|
<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[1056,1162,1117,1137]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="synoriacos" status="sp. nov.">
|
|||
|
|
<emphasis box="[1056,1162,1117,1137]" italics="true" pageId="3" pageNumber="4">A. synoriacos</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[1169,1236,1118,1137]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
metathorax, scanning electron micrograph. (j)
|
|||
|
|
<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[266,377,1141,1161]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="synoriacos" status="sp. nov.">
|
|||
|
|
<emphasis box="[266,377,1141,1161]" italics="true" pageId="3" pageNumber="4">A. synoriacos</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[387,458,1142,1161]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
metathorax interpretative drawing. (k)
|
|||
|
|
<taxonomicName box="[812,893,1141,1161]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="debilis">
|
|||
|
|
<emphasis box="[812,893,1141,1161]" italics="true" pageId="3" pageNumber="4">M. debilis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, thoracic dorsal aspect, scanning electron micrograph. (l)
|
|||
|
|
<taxonomicName box="[1416,1498,1141,1161]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="debilis">
|
|||
|
|
<emphasis box="[1416,1498,1141,1161]" italics="true" pageId="3" pageNumber="4">M. debilis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
interpretative drawing of thoracic dorsal aspect. (m)
|
|||
|
|
<taxonomicName box="[590,670,1165,1185]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="debilis">
|
|||
|
|
<emphasis box="[590,670,1165,1185]" italics="true" pageId="3" pageNumber="4">M. debilis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
prothoracic lateral aspect, scanning electron micrograph. (n)
|
|||
|
|
<taxonomicName box="[1210,1289,1165,1185]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="debilis">
|
|||
|
|
<emphasis box="[1210,1289,1165,1185]" italics="true" pageId="3" pageNumber="4">M. debilis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
photograph of thoracic ventrall aspect. (m)
|
|||
|
|
<taxonomicName box="[297,375,1189,1209]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="debilis">
|
|||
|
|
<emphasis box="[297,375,1189,1209]" italics="true" pageId="3" pageNumber="4">M. debilis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
interpretative drawing of thoracic ventral aspect.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph blockId="3.[128,797,1294,1926]" pageId="3" pageNumber="4">Metanotum with distinct longitudinal median groove delimited with strongly developed alacrista; additional longitudinal bulges visible laterally. Mesocoxae widely separated, more or less rounded, nearly triangular, not transverse. Mesocoxal cavities laterally closed by mesepimeron and metanepisternum.</paragraph>
|
|||
|
|
<paragraph blockId="3.[128,797,1294,1926]" pageId="3" pageNumber="4">Metaventrite large, distinctly narrowed anteriorly, with large triangular, apically rounded anterior intercoxal process; separating mesocoxae completely. Transverse suture of metaventrite not recognizable, probably absent. Metatrochantin not exposed. Metacoxae short, transverse, reaching the lateral edge of ventrite, without recognizable metacoxal plates.</paragraph>
|
|||
|
|
<paragraph blockId="3.[128,797,1294,1926]" pageId="3" pageNumber="4">Abdomen of parabolic shape, evenly narrowing posteriorly, with rounded apex, as long as meso- and metaventrite combined; at least basal sternites separated by externally visible intersegmental membranes. Four distal abdominal tergites not covered by the elytra.</paragraph>
|
|||
|
|
<paragraph blockId="3.[128,315,1965,2022]" box="[128,315,1965,1990]" pageId="3" pageNumber="4">
|
|||
|
|
<heading bold="true" box="[128,315,1965,1990]" fontSize="10" level="3" pageId="3" pageNumber="4" reason="6">
|
|||
|
|
<emphasis bold="true" box="[128,315,1965,1990]" italics="true" pageId="3" pageNumber="4">Species included</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="3.[128,315,1965,2022]" box="[128,253,1998,2022]" pageId="3" pageNumber="4">Monotypic.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="3" pageNumber="4" type="distribution">
|
|||
|
|
<paragraph blockId="3.[829,1497,1293,1382]" box="[829,955,1293,1318]" pageId="3" pageNumber="4">
|
|||
|
|
<heading bold="true" box="[829,955,1293,1318]" fontSize="10" level="3" pageId="3" pageNumber="4" reason="6">
|
|||
|
|
<emphasis bold="true" box="[829,955,1293,1318]" italics="true" pageId="3" pageNumber="4">Occurrence</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="3.[829,1497,1293,1382]" pageId="3" pageNumber="4">Babiy Kamen`locality, Tom`river, Kuznetsk Basin, S. Siberian, in the upper part of the Maltsevo Formation.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="discussion">
|
|||
|
|
<paragraph blockId="3.[829,1498,1421,2023]" box="[829,926,1421,1446]" pageId="3" pageNumber="4">
|
|||
|
|
<heading bold="true" box="[829,926,1421,1446]" fontSize="10" level="3" pageId="3" pageNumber="4" reason="6">
|
|||
|
|
<emphasis bold="true" box="[829,926,1421,1446]" italics="true" pageId="3" pageNumber="4">Remarks</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="3.[829,1498,1421,2023]" pageId="3" pageNumber="4">
|
|||
|
|
Generally, size alone does not justify a placement of a beetle in a specific supraspecific taxon. However, after taking into consideration all the characters mentioned in the Diagonisis and Description, a length of only slightly more than 1 mm should be seen as an additional argument for keeping
|
|||
|
|
<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[829,1002,1614,1638]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis box="[829,1002,1614,1638]" italics="true" pageId="3" pageNumber="4">Archaeomalthus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in
|
|||
|
|
<taxonomicName authorityName=", Barber" authorityYear="1913" box="[1043,1219,1614,1638]" class="Insecta" family="Micromalthidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Micromalthidae</taxonomicName>
|
|||
|
|
. All other archostematan beetles with the exception of the strongly flattened
|
|||
|
|
<emphasis box="[829,1041,1678,1702]" italics="true" pageId="3" pageNumber="4">
|
|||
|
|
<taxonomicName box="[829,966,1678,1702]" class="Insecta" family="Crowsoniellidae" genus="Crowsoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Crowsoniella</taxonomicName>
|
|||
|
|
relicta
|
|||
|
|
</emphasis>
|
|||
|
|
(Pace 1975) are much larger.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="3.[829,1498,1421,2023]" lastBlockId="4.[128,797,142,710]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
|||
|
|
The degree of cuticular sclerotization can rarely be adequately estimated on fossil imprints. Therefore, it cannot be used as a solid diagnostic character, even though some surface features such as tubercles, large punctures, etc., can be used for species identification and also in a phylogenetic context (e.g.
|
|||
|
|
<bibRefCitation author="Beutel RG & Ge SQ & Hornschemeyer T." box="[1232,1491,1870,1894]" journalOrPublisher="Cladistics." pageId="3" pageNumber="4" pagination="270 - 298" part="24" refId="ref5728" refString="Beutel RG, Ge SQ, Hornschemeyer T. 2008. On the head morphology of Tetraphalerus, the phylogeny of Archostemata and the basal branching events in Coleoptera. Cladistics. 24: 270 - 298." title="On the head morphology of Tetraphalerus, the phylogeny of Archostemata and the basal branching events in Coleoptera" type="journal article" year="2008">Beutel et al. 2008</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation author="Hornschemeyer T." box="[829,1067,1902,1926]" journalOrPublisher="Syst Entomol." pageId="3" pageNumber="4" pagination="533 - 558" part="32" refId="ref6169" refString="Hornschemeyer T. 2009. The species-level phylogeny of archostematan beetles - where do Micromalthus debilis and Crowsoniella relicta belong? Syst Entomol. 32: 533 - 558." title="The species-level phylogeny of archostematan beetles - where do Micromalthus debilis and Crowsoniella relicta belong?" type="journal article" year="2009">Hörnschemeyer 2009</bibRefCitation>
|
|||
|
|
). In the case of
|
|||
|
|
<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[1268,1444,1902,1926]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis box="[1268,1444,1902,1926]" italics="true" pageId="3" pageNumber="4">Archaeomalthus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the visibility of metathoracic fine details (including metapostnotum) on mostly ventral imprint and elytra (
|
|||
|
|
<figureCitation box="[1344,1437,1966,1990]" captionStart="Figure 2" captionStartId="3.[128,183,1045,1064]" captionTargetBox="[253,1373,156,1020]" captionTargetId="figure@3.[253,1373,139,1020]" captionTargetPageId="3" captionText="Figure 2. Details of the organization in extinct and recent Micromalthidae. (a) Archaeomalthus synoriacos sp. nov. head, scanning electron micrograph. (b) A. synoriacos sp. nov. head interpretative drawing. (c) Micromalthus debilis, head dorsal aspect, scanning electron micrograph. (d) M. debilis interpretative drawing of head dorsal aspect. (e) M. debilis, head ventral aspect, scanning electron micrograph. (f) M. debilis interpretative drawing of head ventral aspect. (g) A. synoriacos sp. nov. pronotum, scanning electron micrograph. (h) A. synoriacos sp. nov. pronotum interpretative drawing. (i) A. synoriacos sp. nov. metathorax, scanning electron micrograph. (j) A. synoriacos sp. nov. metathorax interpretative drawing. (k) M. debilis, thoracic dorsal aspect, scanning electron micrograph. (l) M. debilis interpretative drawing of thoracic dorsal aspect. (m) M. debilis prothoracic lateral aspect, scanning electron micrograph. (n) M. debilis photograph of thoracic ventrall aspect. (m) M. debilis interpretative drawing of thoracic ventral aspect." httpUri="https://zenodo.org/record/2536033/files/figure.png" pageId="3" pageNumber="4">Figure 2</figureCitation>
|
|||
|
|
(i,k)), preserved only as faint lines, suggests that
|
|||
|
|
<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[1321,1497,1998,2022]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis box="[1321,1497,1998,2022]" italics="true" pageId="3" pageNumber="4">Archaeomalthus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
was overall weakly sclerotized. Elytra of most fossilized beetles, in contrast, are hard structures overlapping the body and usually retaining many structural details (
|
|||
|
|
<bibRefCitation author="Ponomarenko AG" box="[135,355,238,262]" journalOrPublisher="Tr Palaeontol Inst Akad Ski SSSR" pageId="4" pageNumber="5" pagination="1 - 233" refId="ref7091" refString="Ponomarenko AG. 1969. Historical development of archostematan beetles. Tr Palaeontol Inst Akad Ski SSSR. 1 - 233." title="Historical development of archostematan beetles" type="book chapter" year="1969">Ponomarenko 1969</bibRefCitation>
|
|||
|
|
). Beetle fossils previously known from Babiy Kamen`locality,
|
|||
|
|
<taxonomicName authority="Ponomarenko et Volkov, 2013" authorityName="Ponomarenko et Volkov" authorityYear="2013" class="Insecta" family="Carabidae" genus="Petrodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[429,578,270,294]" italics="true" pageId="4" pageNumber="5">Petrodromeus</emphasis>
|
|||
|
|
<bibRefCitation author="Ponomarenko AG & Volkov AN" journalOrPublisher="Paleontol J." pageId="4" pageNumber="5" pagination="601 - 606" part="47" refId="ref7167" refString="Ponomarenko AG, Volkov AN. 2013. Ademosynoides asiaticus Martynov, 1936, the earlies known member of an extant beetle family (Insecta, Coleoptera, Trachypachidae). Paleontol J. 47: 601 - 606." title="Ademosynoides asiaticus Martynov, 1936, the earlies known member of an extant beetle family (Insecta, Coleoptera, Trachypachidae)" type="journal article" year="2013">Ponomarenko et Volkov, 2013</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName authority="Ponomarenko et Volkov 2013" authorityName="Ponomarenko et Volkov" authorityYear="2013" class="Insecta" family="Carabidae" genus="Permunda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[368,481,302,326]" italics="true" pageId="4" pageNumber="5">Permunda</emphasis>
|
|||
|
|
<bibRefCitation author="Ponomarenko AG & Volkov AN" journalOrPublisher="Paleontol J." pageId="4" pageNumber="5" pagination="601 - 606" part="47" refId="ref7167" refString="Ponomarenko AG, Volkov AN. 2013. Ademosynoides asiaticus Martynov, 1936, the earlies known member of an extant beetle family (Insecta, Coleoptera, Trachypachidae). Paleontol J. 47: 601 - 606." title="Ademosynoides asiaticus Martynov, 1936, the earlies known member of an extant beetle family (Insecta, Coleoptera, Trachypachidae)" type="journal article" year="2013">Ponomarenko et Volkov 2013</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(Adephaga;
|
|||
|
|
<taxonomicName authorityName="C.G. Thomson" authorityYear="1857" box="[339,515,334,358]" class="Insecta" family="Trachypachidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Trachypachidae</taxonomicName>
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) both have finely preserved elytra with well-defined longitudinal striae (
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<bibRefCitation author="Ponomarenko AG & Volkov AN" box="[135,496,398,422]" journalOrPublisher="Paleontol J." pageId="4" pageNumber="5" pagination="601 - 606" part="47" refId="ref7167" refString="Ponomarenko AG, Volkov AN. 2013. Ademosynoides asiaticus Martynov, 1936, the earlies known member of an extant beetle family (Insecta, Coleoptera, Trachypachidae). Paleontol J. 47: 601 - 606." title="Ademosynoides asiaticus Martynov, 1936, the earlies known member of an extant beetle family (Insecta, Coleoptera, Trachypachidae)" type="journal article" year="2013">Ponomarenko and Volkov 2013</bibRefCitation>
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).
|
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</paragraph>
|
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<paragraph blockId="4.[128,797,142,710]" pageId="4" pageNumber="5">
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The abdomen of
|
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<taxonomicName authority="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko, 2019" authorityName="Evgeny Viktorovich Yana & Rolf Georg Beutel & John Francis Lawrence & Margarita Igorevna Yavorskaya & Thomas Hörnschemeyer & Hans Pohl & Dmitry Vladimirovich Vassilenko & Alexey Semenovich Bashkuev & Alexander Georgievich Ponomarenko" authorityYear="2019" box="[351,521,430,454]" class="Insecta" family="Micromalthidae" genus="Archaeomalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
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<emphasis box="[351,521,430,454]" italics="true" pageId="4" pageNumber="5">Archaeomalthus</emphasis>
|
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</taxonomicName>
|
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is distinctly shorter than the head and pronotum combined; however, this character is variable in the extant
|
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<taxonomicName authorityName="LeConte" authorityYear="1878" box="[366,512,494,518]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
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|
<emphasis box="[366,512,494,518]" italics="true" pageId="4" pageNumber="5">Micromalthus</emphasis>
|
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</taxonomicName>
|
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|
due to varying degrees of extension of the intersegmental membranes. Thus it is not used in the generic diagnosis. The abdominal segments are often more contracted in males than in females, possibly due to different physiological conditions (e.g. development stage of eggs). The length of the abdomen, especially of females, varies in
|
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<taxonomicName authority="Hornschemeyer et al. 2010" authorityName="Hornschemeyer et al." authorityYear="2010" box="[156,548,686,710]" class="Insecta" family="Micromalthidae" genus="Micromalthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="debilis">
|
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|
|
<emphasis box="[156,258,686,710]" italics="true" pageId="4" pageNumber="5">M. debilis</emphasis>
|
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|
<bibRefCitation author="Hornschemeyer T & Wedman S & Poinar G." box="[265,548,686,710]" journalOrPublisher="Zool JLinn Soc." pageId="4" pageNumber="5" pagination="300 - 311" part="158" refId="ref6273" refString="Hornschemeyer T, Wedman S, Poinar G. 2010. How long can insect species exist? Evidence from extant and fossil Micromalthus beetles (Insecta: Coleoptera). Zool JLinn Soc. 158: 300 - 311." title="How long can insect species exist? Evidence from extant and fossil Micromalthus beetles (Insecta: Coleoptera)" type="journal article" year="2010">Hörnschemeyer et al. 2010</bibRefCitation>
|
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</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
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</subSubSection>
|
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|
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</treatment>
|
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</document>
|