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<document id= "855A5E7B108E0D8138A0E9EFE344ABBB" ID-DOI= "10.11646/phytotaxa.363.1.1" ID-ISSN= "1179-3163" ID-Zenodo-Dep= "13703414" IM.bibliography_approvedBy= "felipe" IM.illustrations_approvedBy= "felipe" IM.materialsCitations_approvedBy= "felipe" IM.metadata_approvedBy= "felipe" IM.tables_requiresApprovalFor= "GgImagineBatch" IM.taxonomicNames_approvedBy= "felipe" IM.treatmentCitations_approvedBy= "felipe" IM.treatments_approvedBy= "felipe" checkinTime= "1725581356715" checkinUser= "felipe" docAuthor= "Negaresh, Kazem & Rahiminejad, Mohammad Reza" docDate= "2018" docId= "03E887F4DF23FFFF19D439F5F592FC3C" docLanguage= "en" docName= "phytotaxa.363.1.1.pdf" docOrigin= "Phytotaxa 363 (1)" docSource= "http://dx.doi.org/10.11646/phytotaxa.363.1.1" docStyle= "DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph" docStyleId= "F08184CE06D8A97EA3E6DE35D99648B0" docStyleName= "Phytotaxa.2014-.monograph" docStyleVersion= "4" docTitle= "Centaurea haussknechtii Boissier 1875" docType= "treatment" docVersion= "2" lastPageNumber= "450" masterDocId= "FFD1FF8CDF12FFCC195C3E3CF14EFF83" masterDocTitle= "A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae)" masterLastPageNumber= "450" masterPageNumber= "448" pageNumber= "448" updateTime= "1725593506943" updateUser= "ExternalLinkService" >
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<mods:title id= "0D125CDEFA0BD6DF88283D79F8E2280C" > A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae)</mods:title>
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<mods:namePart id= "F82A932FFC47446EE147A32D952B730D" > Negaresh, Kazem</mods:namePart>
<mods:affiliation id= "344E2DF28EFC8374D3D6FC6EFF8BAA9F" > Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir)</mods:affiliation>
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<mods:namePart id= "8A9CECBD2652D6D37B38051A41420874" > Rahiminejad, Mohammad Reza</mods:namePart>
<mods:affiliation id= "E15A3D88BF081B602A04FBA1F2442D16" > Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir) & Department of Biology, University of Isfahan, Isfahan 81746 - 73441, Iran. & Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir)</mods:affiliation>
<mods:nameIdentifier id= "9D10F04329CF0BEADE989E375FAB156C" type= "email" > negaresh@ramin.ac.ir</mods:nameIdentifier>
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<subSubSection id= "C35B6569DF23FFFD19D439F5F26EF7A9" pageId= "49" type= "nomenclature" >
<paragraph id= "8BFE36E2DF23FFFD19D439F5F26EF7A9" blockId= "49.[136,1451,1991,2090]" pageId= "49" >
<emphasis id= "B935EAF0DF23FFFD19D439F5F1E2F861" bold= "true" box= "[136,172,1993,2018]" pageId= "49" > 15.</emphasis>
<taxonomicName id= "4C414D61DF23FFFD19EF39FBF386F861" ID-CoL= "S6NX" authority= "Boissier (1875: 670)" authorityName= "Boissier" authorityPageNumber= "670" authorityYear= "1875" box= "[179,712,1991,2018]" class= "Magnoliopsida" family= "Asteraceae" genus= "Centaurea" kingdom= "Plantae" order= "Asterales" pageId= "49" phylum= "Tracheophyta" rank= "species" species= "haussknechtii" >
<emphasis id= "B935EAF0DF23FFFD19EF39FBF09AF861" bold= "true" box= "[179,468,1991,2018]" italics= "true" pageId= "49" > Centaurea haussknechtii</emphasis>
<bibRefCitation id= "EFD04B13DF23FFFD188039F5F386F861" author= "Boissier, E." box= "[476,712,1993,2018]" pageId= "49" refId= "ref65414" refString= "Boissier, E. (1875) Flora Orientalis, vol. 3. H. Georg, Geneva & Basilea [Basel], 1033 pp." type= "book" year= "1875" > Boissier (1875: 670)</bibRefCitation>
</taxonomicName>
. Type:—
<collectingCountry id= "F3567672DF23FFFD1A7039F5F2D1F861" box= "[812,927,1993,2018]" name= "Turkey" pageId= "49" > TURKEY</collectingCountry>
. C6
<collectingRegion id= "4985F800DF23FFFD1A8839F5F504F861" box= "[980,1098,1993,2018]" country= "Turkey" name= "Gaziantep" pageId= "49" > Gaziantep</collectingRegion>
: in graminosis calcareis montis Ssoffdagh (So f Da.) Syriae borealis,
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,
<date id= "FFFF1022DF23FFFD1BC439D1F260F785" box= "[664,814,2029,2054]" pageId= "49" value= "1865-07-27" > 27 July 1865</date>
,
<emphasis id= "B935EAF0DF23FFFD1A6639D1F2B6F785" box= "[826,1016,2029,2054]" italics= "true" pageId= "49" > Haussknecht s.n.</emphasis>
(
<typeStatus id= "54FA8840DF23FFFD1D5439D1F53FF785" box= "[1032,1137,2029,2054]" pageId= "49" type= "lectotype" > lectotype</typeStatus>
G! designated by
<bibRefCitation id= "EFD04B13DF23FFFD1C1E39D1F1B2F7A9" author= "Wagenitz, G." pageId= "49" pagination= "469 - 494" refId= "ref69162" refString= "Wagenitz, G. (1960) Centaurea L. sect. Cynaropsis, eine neue Sektion der Gattung aus Vorderasien. Willdenowia 2: 469 - 494." type= "journal article" year= "1960" > Wagenitz 1960: 490</bibRefCitation>
,
<typeStatus id= "54FA8840DF23FFFD185B362DF0D7F7A9" box= "[263,409,2065,2090]" pageId= "49" type= "isolectotype" > isolectotypes</typeStatus>
G!, JE!: two sheets, W!). (
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<figureCitation id= "137A2A67DF23FFFD1B9A362DF259F7A9" box= "[710,791,2065,2090]" captionStart= "FIGURE 32" captionStartId= "50.[136,229,1953,1974]" captionTargetBox= "[196,1393,203,1924]" captionTargetId= "figure-17@50.[194,1393,190,1929]" captionTargetPageId= "50" captionText= "FIGURE 32. Centaurea haussknechtii Boissier (Haussknecht s.n., G). A. habit. B. close up of capitule. Scale B = 1 cm." figureDoi= "http://doi.org/10.5281/zenodo.13703492" httpUri= "https://zenodo.org/record/13703492/files/figure.png" pageId= "49" > Fig. 32</figureCitation>
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)
</paragraph>
</subSubSection>
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<caption id= "DF3E666ADF20FFFE19D4399DF447F835" ID-DOI= "http://doi.org/10.5281/zenodo.13703492" ID-Zenodo-Dep= "13703492" box= "[136,1289,1953,1974]" httpUri= "https://zenodo.org/record/13703492/files/figure.png" pageId= "50" startId= "50.[136,229,1953,1974]" targetBox= "[196,1393,203,1924]" targetPageId= "50" targetType= "figure" >
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<paragraph id= "8BFE36E2DF20FFFE19D4399DF447F835" blockId= "50.[136,1289,1953,1974]" box= "[136,1289,1953,1974]" pageId= "50" >
<emphasis id= "B935EAF0DF20FFFE19D4399DF047F835" bold= "true" box= "[136,265,1953,1974]" pageId= "50" > FIGURE 32.</emphasis>
<taxonomicName id= "4C414D61DF20FFFE1853399DF31FF835" authority= "Boissier" authorityName= "Boissier" authorityYear= "1875" box= "[271,593,1953,1974]" class= "Magnoliopsida" family= "Asteraceae" genus= "Centaurea" kingdom= "Plantae" order= "Asterales" pageId= "50" phylum= "Tracheophyta" rank= "species" species= "haussknechtii" >
<emphasis id= "B935EAF0DF20FFFE1853399DF0B3F835" box= "[271,509,1953,1974]" italics= "true" pageId= "50" > Centaurea haussknechtii</emphasis>
Boissier
</taxonomicName>
(
<emphasis id= "B935EAF0DF20FFFE1B03399DF24CF835" box= "[607,770,1953,1974]" italics= "true" pageId= "50" > Haussknecht s.n.</emphasis>
, G).
<emphasis id= "B935EAF0DF20FFFE1A6E399DF204F835" bold= "true" box= "[818,842,1953,1974]" pageId= "50" > A.</emphasis>
habit.
<emphasis id= "B935EAF0DF20FFFE1AD0399DF2ECF835" bold= "true" box= "[908,930,1953,1974]" pageId= "50" > B.</emphasis>
close up of capitule. Scale
<emphasis id= "B935EAF0DF20FFFE1DF5399DF5F7F835" bold= "true" box= "[1193,1209,1953,1974]" pageId= "50" > B</emphasis>
= 1 cm.
</paragraph>
</caption>
<subSubSection id= "C35B6569DF21FFFF19D43E9CF44FFC1F" pageId= "51" type= "description" >
<paragraph id= "8BFE36E2DF21FFFF19D43E9CF2D8FD6F" blockId= "51.[136,1452,160,959]" pageId= "51" >
Biennial herbs, with long and stout taproot, whole plant usually grayish or green-yellowish,
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tall; collar of fibrous petiolar remains present at stem base. Stem erect, simple,
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in diam. at base, cylindrical, sparsely leafy throughout, lower part purplish and loosely covered with hirsute-articulate hairs, others ± densely covered with white-tomentose hairs. Leaves firm or rigid (on drying), usually divided, densely covered with white-tomentose hairs, sometimes basal and lower ones sparsely covered with hirsute-articulate hairs, denser along midrib and veins, and mixed with tomentose hairs. Basal and lower cauline leaves petiolate, (7–
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long, lyrate; terminal segments larger, oblong, triangular-ovate or lanceolate, 4– × –
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, entire or shallowly serrulate, obtuse or truncate at base, acute or rounded at apex; lateral segments smaller, 1– – ×
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, very shortly decurrent, sometimes not decurrent, entire, acuminate or mucronate at apex. Capitula few, 4 to 7, arranged in a spike or raceme, often sessile or subsessile, sometimes with
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peduncles. Involucres ovoid, truncate at base, (33– – ×
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. Phyllaries multiseriate, yellow-greenish, imbricate, coriaceous-scarious, glabrous.Appendages rigid, totally concealing phyllaries, not decurrent, lanceolate, straw-coloured or pale brown, 3– – –
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long; spines narrowly triangular, (7–
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long, strongly longer than nearest cilia. Flowers pink-purple; central florets hermaphroditic, limb ca.
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long, 5-lobed, lobes ca.
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long; peripheral florets sterile, slightly shorter than central ones, conspicuous, finely dissected, not radiant, 5-lobed, limb lobes linear. Achenes and pappus unknown.
</paragraph>
<paragraph id= "8BFE36E2DF21FFFF19E13CCAF5F1FCD6" blockId= "51.[136,1452,160,959]" pageId= "51" >
<emphasis id= "B935EAF0DF21FFFF19E13CCAF3C5FC8C" bold= "true" box= "[189,651,758,783]" pageId= "51" > Taxonomic and distribution remarks:</emphasis>
—
<taxonomicName id= "4C414D61DF21FFFF1BFB3CCAF2F1FC8C" authorityName= "Boissier" authorityYear= "1875" box= "[679,959,758,783]" class= "Magnoliopsida" family= "Asteraceae" genus= "Centaurea" kingdom= "Plantae" order= "Asterales" pageId= "51" phylum= "Tracheophyta" rank= "species" species= "haussknechtii" >
<emphasis id= "B935EAF0DF21FFFF1BFB3CCAF2F1FC8C" box= "[679,959,758,783]" italics= "true" pageId= "51" > Centaurea haussknechtii</emphasis>
</taxonomicName>
is a rare endemic of
<collectingRegion id= "4985F800DF21FFFF1D983CCAF4E9FC8C" box= "[1220,1447,758,783]" country= "Turkey" name= "Gaziantep" pageId= "51" > Gaziantep Province</collectingRegion>
, S
<collectingCountry id= "F3567672DF21FFFF19FD3D25F1BFFCB1" box= "[161,241,793,818]" name= "Turkey" pageId= "51" > Turkey</collectingCountry>
(
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<figureCitation id= "137A2A67DF21FFFF185F3D25F014FCB1" box= "[259,346,793,818]" captionStart= "FIGURE 30" captionStartId= "47.[136,229,1389,1410]" captionTargetBox= "[166,1381,628,1339]" captionTargetId= "figure-193@47.[151,1436,605,1363]" captionTargetPageId= "47" captionText= "FIGURE 30. Distribution of Centaurea gudrunensis (circles), C. haradjianii (squares), and C. haussknechtii (triangles)." figureDoi= "http://doi.org/10.5281/zenodo.13703484" httpUri= "https://zenodo.org/record/13703484/files/figure.png" pageId= "51" > Fig. 30</figureCitation>
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).
<taxonomicName id= "4C414D61DF21FFFF182C3D25F3C8FCB1" authorityName= "Boissier" authorityYear= "1875" box= "[368,646,793,818]" class= "Magnoliopsida" family= "Asteraceae" genus= "Centaurea" kingdom= "Plantae" order= "Asterales" pageId= "51" phylum= "Tracheophyta" rank= "species" species= "haussknechtii" >
<emphasis id= "B935EAF0DF21FFFF182C3D25F3C8FCB1" box= "[368,646,793,818]" italics= "true" pageId= "51" > Centaurea haussknechtii</emphasis>
</taxonomicName>
is an Irano-Turanian element, and grows on limestone, grassland and calcareous mountains, at elevations of
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. It closely resembles
<taxonomicName id= "4C414D61DF21FFFF1A9B3D00F574FCD6" authorityName= "Post" authorityYear= "1888" box= "[967,1082,828,853]" class= "Magnoliopsida" family= "Asteraceae" genus= "Centaurea" kingdom= "Plantae" order= "Asterales" pageId= "51" phylum= "Tracheophyta" rank= "species" species= "doddsii" >
<emphasis id= "B935EAF0DF21FFFF1A9B3D00F574FCD6" box= "[967,1082,828,853]" italics= "true" pageId= "51" > C. doddsii</emphasis>
</taxonomicName>
in its habit.
</paragraph>
<paragraph id= "8BFE36E2DF21FFFF19E13D5CF44FFC1F" blockId= "51.[136,1452,160,959]" pageId= "51" >
<emphasis id= "B935EAF0DF21FFFF19E13D5CF094FCFA" bold= "true" box= "[189,474,864,889]" pageId= "51" > Chromosome numbers:</emphasis>
—
<bibRefCitation id= "EFD04B13DF21FFFF18AA3D5CF399FCFA" author= "Askoy, H. & Berber, A. A. & Celik, S. & Baser, B. & Karaveliogullari, F. A." box= "[502,727,864,889]" pageId= "51" pagination= "285 - 294" refId= "ref65175" refString= "Askoy, H., Berber, A. A., Celik, S., Baser, B. & Karaveliogullari, F. A. (2016) Karyological study on 12 Centaurea (Asteraceae) species from Turkey. Caryologia 69 (4): 285 - 294. http: // doi. org / 10.1080 / 00087114.2016.1188357" type= "journal article" year= "2016" >
Askoy
<emphasis id= "B935EAF0DF21FFFF1B143D5CF3CDFCFA" box= "[584,643,864,889]" italics= "true" pageId= "51" > et al.</emphasis>
(2016)
</bibRefCitation>
reported the chromosome number 2
<emphasis id= "B935EAF0DF21FFFF1D253D5CF5C9FCFA" box= "[1145,1159,864,889]" italics= "true" pageId= "51" > n</emphasis>
= 2
<emphasis id= "B935EAF0DF21FFFF1DE43D5CF58AFCFA" box= "[1208,1220,864,889]" italics= "true" pageId= "51" > x</emphasis>
= 18 for
<taxonomicName id= "4C414D61DF21FFFF1C6A3D5CF06EFC1F" authorityName= "Boissier" authorityYear= "1875" class= "Magnoliopsida" family= "Asteraceae" genus= "Centaurea" kingdom= "Plantae" order= "Asterales" pageId= "51" phylum= "Tracheophyta" rank= "species" species= "haussknechtii" >
<emphasis id= "B935EAF0DF21FFFF1C6A3D5CF06EFC1F" italics= "true" pageId= "51" > Centaurea haussknechtii</emphasis>
</taxonomicName>
collected from
<collectingCountry id= "F3567672DF21FFFF188C3DBFF36FFC1F" box= "[464,545,899,924]" name= "Turkey" pageId= "51" > Turkey</collectingCountry>
, Siverek-Kahta (
<collectingRegion id= "4985F800DF21FFFF1B813DBFF217FC1F" box= "[733,857,899,924]" country= "Turkey" name= "Adiyaman" pageId= "51" > Adı </collectingRegion>
). This is the only count for this taxon.
</paragraph>
</subSubSection>
<subSubSection id= "C35B6569DF21FFFF19E13D9AF592FC3C" box= "[189,1244,934,959]" pageId= "51" type= "materials_examined" >
<paragraph id= "8BFE36E2DF21FFFF19E13D9AF592FC3C" blockId= "51.[136,1452,160,959]" box= "[189,1244,934,959]" pageId= "51" >
<emphasis id= "B935EAF0DF21FFFF19E13D9AF3F9FC3C" bold= "true" box= "[189,695,934,959]" pageId= "51" >
Selected specimens examined:—
<collectingCountry id= "F3567672DF21FFFF1B633D9AF3F9FC3C" box= "[575,695,934,959]" name= "Turkey" pageId= "51" > TURKEY</collectingCountry>
</emphasis>
. Taurus cataonicus, 1865,
<emphasis id= "B935EAF0DF21FFFF1ABD3D9AF5D0FC3C" box= "[993,1182,934,959]" italics= "true" pageId= "51" > Haussknecht s.n.</emphasis>
(
<collectionCode id= "ED50AE27DF21FFFF1DF33D9AF584FC3C" box= "[1199,1226,934,959]" country= "Switzerland" lsid= "urn:lsid:biocol.org:col:15706" name= "Conservatoire et Jardin botaniques de la Ville de Genève" pageId= "51" type= "Herbarium" > G</collectionCode>
!).
</paragraph>
</subSubSection>
</treatment>
</document>
