195 lines
18 KiB
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195 lines
18 KiB
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<document ID-DOI="10.5252/g2016n4a4" ID-ISSN="1638-9395" ID-Zenodo-Dep="5208463" ID-ZooBank="urn:lsid:zoobank.org:pub:CFFB0AA0-D396-40EB-BE75-D2E417257B87" approvalRequired="27" approvalRequired_for_textStreams="26" approvalRequired_for_treatments="1" checkinTime="1629146399156" checkinUser="felipe" docAuthor="Devillez, Julien, Charbonnier, Sylvain, Hyžný, Matúš & Leroy, Lucien" docDate="2016" docId="E32D87EF8209FE09FE8FFABAFA7E0E58" docLanguage="en" docName="Geodiversitas.38.4.515-541.pdf" docOrigin="Geodiversitas 38 (4)" docSource="http://dx.doi.org/10.5252/g2016n4a4" docStyle="DocumentStyle:4DA100580BA89F561A6D37CADC50A8FD.2:Geodiversitas.2015-2017.journal_article.type2" docStyleId="4DA100580BA89F561A6D37CADC50A8FD" docStyleName="Geodiversitas.2015-2017.journal_article.type2" docStyleVersion="2" docTitle="Enoploclytia gigantea Devillez & Charbonnier & Hyžný & Leroy 2016, n. sp." docType="treatment" docUuid="622C486B-B2A1-454F-B540-C6B50A17880C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="531" masterDocId="1F14FF978219FE19FFBEFFC9FFCC0A76" masterDocTitle="Review of the Early Cretaceous erymid lobsters (Crustacea: Decapoda) from the Western Tethys" masterLastPageNumber="541" masterPageNumber="515" pageNumber="531" updateTime="1675446794094" updateUser="tatiana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Early Cretaceous erymid lobsters (Crustacea: Decapoda) from the Western Tethys</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Devillez, Julien</mods:namePart>
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<mods:affiliation>Muséum national d’Histoire naturelle, Paris Centre de Recherche sur la Paléobiodiversité et les Paléoenvironnements (CR 2 P, UMR 7207), Sorbonne Universités, MNHN, UPMC, CNRS, 57 rue Cuvier F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:nameIdentifier type="email">julien.devillez@edu.mnhn.fr</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Charbonnier, Sylvain</mods:namePart>
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<mods:affiliation>Muséum national d’Histoire naturelle, Paris Centre de Recherche sur la Paléobiodiversité et les Paléoenvironnements (CR 2 P, UMR 7207), Sorbonne Universités, MNHN, UPMC, CNRS, 57 rue Cuvier F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:nameIdentifier type="email">sylvain.charbonnier@mnhn.fr</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hyžný, Matúš</mods:namePart>
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<mods:affiliation>Department of Geology and Palaeontology, Faculty of Natural Sciences, Comenius University, Mlynská dolina G 1, 842 15 Bratislava (Slovakia); Geologisch-paläontologische Abteilung, Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna (Austria)</mods:affiliation>
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<mods:nameIdentifier type="email">hyzny.matus@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Leroy, Lucien</mods:namePart>
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<mods:affiliation>Hameau de Taloire, F- 04120 Castellane (France)</mods:affiliation>
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<mods:nameIdentifier type="email">lucien.leroy@orange.fr</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-12-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>38</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>515</mods:start>
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<mods:end>541</mods:end>
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</mods:extent>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.5252/g2016n4a4</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5252/g2016n4a4</mods:identifier>
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<mods:identifier type="ISSN">1638-9395</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5208463</mods:identifier>
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<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:CFFB0AA0-D396-40EB-BE75-D2E417257B87</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5208494" ID-Zenodo-Dep="5208494" LSID="urn:lsid:zoobank.org:act:622C486B-B2A1-454F-B540-C6B50A17880C" httpUri="http://treatment.plazi.org/id/E32D87EF8209FE09FE8FFABAFA7E0E58" lastPageNumber="531" pageId="16" pageNumber="531">
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<subSubSection box="[305,603,1395,1422]" pageId="16" pageNumber="531" type="nomenclature">
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<paragraph blockId="16.[305,603,1395,1454]" box="[305,603,1395,1422]" pageId="16" pageNumber="531">
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<heading box="[305,603,1395,1422]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="531" reason="2">
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<taxonomicName authority="Devillez & Charbonnier & Hyžný & Leroy, 2016" authorityName="Devillez & Charbonnier & Hyžný & Leroy" authorityYear="2016" box="[305,537,1395,1422]" class="Malacostraca" family="Erymidae" genus="Enoploclytia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="gigantea" status="n. sp.">
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<emphasis bold="true" box="[305,537,1395,1422]" italics="true" pageId="16" pageNumber="531">Enoploclytia gigantea</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[544,603,1396,1422]" pageId="16" pageNumber="531" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[390,518,1427,1454]" pageId="16" pageNumber="531" type="description">
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<paragraph blockId="16.[305,603,1395,1454]" box="[390,518,1427,1454]" pageId="16" pageNumber="531">
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(
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<figureCitation box="[399,461,1427,1453]" captionStart="FIG" captionStartId="18.[132,143,1894,1912]" captionTargetBox="[132,1455,216,1848]" captionTargetId="figure-30@18.[698,1267,770,1326]" captionTargetPageId="18" captionText="FIG. 8. — New species of Early Cretaceous Enoploclytia M’Coy, 1849: A-D, holotype MNHN.F.B14557 of Enoploclytia augustobonae n. sp. from the Barremian of Amance, France: carapace in right lateral view (A), and line drawing (B), dorsal view (C) showing the deviation of median line in cephalic region (intercalated plate) and line drawing (D); E-H, holotype of Enoploclytia gigantea n. sp. from the Albian of Fort Worth, USA: original figures of carapace by Rathbun (1935: pl. 4; figs 1, 2), left lateral view (E), right lateral view (F), and line drawings (G, H). Abbreviations: a, branchiocardiac groove; b, antennal groove; b1, hepatic groove; c, postcervical groove; d, gastro-orbital groove; e1e, cervical groove; i, inferior groove; ω, attachment site of mandibular muscle. Photographs: L. Cazes (A, C). Line drawings: J. Devillez. Scale bars: A-D, 10 mm; E-H, 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5208479" httpUri="https://zenodo.org/record/5208479/files/figure.png" pageId="16" pageNumber="531">Fig. 8</figureCitation>
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E-H)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="531" type="materials_examined">
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<paragraph blockId="16.[131,775,1484,1535]" pageId="16" pageNumber="531">
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<materialsCitation location="Bureau of Economic Geology" pageId="16" pageNumber="531" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
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TYPE MATERIAL. —
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<typeStatus box="[322,415,1484,1508]" pageId="16" pageNumber="531">Holotype</typeStatus>
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n°201 stored in the collections of the
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<location LSID="urn:lsid:plazi:treatment:E32D87EF8209FE09FE8FFABAFA7E0E58:6E5B60228209FE09FF3DFA2EFE6A0F88" box="[131,422,1510,1534]" name="Bureau of Economic Geology" pageId="16" pageNumber="531" stateProvince="Texas">Bureau of Economic Geology</location>
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,
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<collectingRegion box="[430,483,1511,1534]" country="United States of America" name="Texas" pageId="16" pageNumber="531">Texas</collectingRegion>
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(after
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<bibRefCitation author="RATHBUN M. J." box="[549,692,1510,1534]" pageId="16" pageNumber="531" pagination="1 - 160" refId="ref22066" refString="RATHBUN M. J. 1935. - Fossil Crustacea of the Atlantic and Gulf Coastal Plain. Geological Society of America Special Papers 2: 1 - 160. https: // doi. org / 10.1130 / spe 2 - p 1" type="journal article" year="1935">Rathbun 1935</bibRefCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="16.[132,773,1558,1582]" box="[132,773,1558,1582]" pageId="16" pageNumber="531">
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<typeStatus box="[132,181,1559,1582]" pageId="16" pageNumber="531">TYPE</typeStatus>
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LOCALITY. — Fort Worth,
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<collectingRegion box="[446,499,1559,1582]" country="United States of America" name="Texas" pageId="16" pageNumber="531">Texas</collectingRegion>
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(
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<collectingCountry box="[512,762,1558,1582]" name="United States of America" pageId="16" pageNumber="531">United States of America</collectingCountry>
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).
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</paragraph>
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<paragraph blockId="16.[132,518,1606,1630]" box="[132,518,1606,1630]" pageId="16" pageNumber="531">
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<typeStatus box="[132,181,1606,1629]" pageId="16" pageNumber="531">TYPE</typeStatus>
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AGE. — Albian, Early Cretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="531" type="etymology">
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<paragraph blockId="16.[132,776,1653,1704]" pageId="16" pageNumber="531">
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ETYMOLOGY. — The specific epithet refers to the Latin
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<emphasis box="[652,737,1654,1677]" italics="true" pageId="16" pageNumber="531">giganteus</emphasis>
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, -
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<emphasis box="[757,768,1654,1677]" italics="true" pageId="16" pageNumber="531">a</emphasis>
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, -
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<emphasis box="[140,171,1680,1703]" italics="true" pageId="16" pageNumber="531">um</emphasis>
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(= enormous) alluding to the exceptional size of the carapace.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="531" type="description">
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<paragraph blockId="16.[130,777,1746,2028]" box="[132,277,1746,1772]" pageId="16" pageNumber="531">DESCRIPTION</paragraph>
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<paragraph blockId="16.[130,777,1746,2028]" lastBlockId="16.[811,1457,215,560]" pageId="16" pageNumber="531">
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Carapace subrectangular in lateral view (
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<typeStatus box="[527,620,1778,1804]" pageId="16" pageNumber="531">holotype</typeStatus>
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:CL=
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<quantity box="[682,771,1778,1805]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" pageId="16" pageNumber="531" unit="mm" value="175.0">175 mm</quantity>
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,
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<collectingCountry box="[130,172,1810,1836]" name="Switzerland" pageId="16" pageNumber="531">CH</collectingCountry>
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=
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<quantity box="[195,288,1810,1836]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="16" pageNumber="531" unit="mm" value="110.0">110 mm</quantity>
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); rostrum not preserved; deep, sinuous cervical groove, joined to dorsal margin and to antennal groove; deep antennal groove, strongly curved and delimiting a slightly raised antennal lobe; long gastro-orbital groove, originating as a deep median inflexion of cervical groove, with two divergent branches delimiting two gastro-orbital lobes (flat upper lobe, slightly prominent lower lobe); deep, sinuous postcervical groove (strong sinuosity at carapace mid-height), joined to dorsal margin and to hepatic groove, with short ventral extension; short, shallow branchiocardiac groove, strongly inclined, joined to dorsal margin and not to postcervical groove; sinuous hepatic groove, joined to cervical groove; strongly inflated ω bulge; flat attachment site of adductor testis muscle (χ bulge); deep inferior groove; carapace covered with widely spaced tubercles; coarse tubercles in upper half of carapace and small tubercles in lower half of carapace; gastric region with a row of coarse tubercles parallel to intercalated plate; row of spines directed forward along dorsal margin.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="531" type="discussion">
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<paragraph blockId="16.[812,1458,598,1071]" box="[813,942,598,624]" pageId="16" pageNumber="531">DISCUSSION</paragraph>
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<paragraph blockId="16.[812,1458,598,1071]" pageId="16" pageNumber="531">
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The present carapace was initially identified as
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<taxonomicName authorityName="Whitfield" authorityYear="1883" class="Malacostraca" family="Erymidae" genus="Palaeastacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="walkeri">
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<emphasis italics="true" pageId="16" pageNumber="531">Palaeastacus walkeri</emphasis>
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</taxonomicName>
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byRathbun (1935), who was followed by
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<bibRefCitation author="STENZEL H. B." box="[1308,1447,661,688]" pageId="16" pageNumber="531" pagination="401 - 476" refId="ref22965" refString="STENZEL H. B. 1945. - Decapod Crustacea from the Cretaceous of Texas. Texas University Publications 4401: 401 - 476." type="journal article" year="1945">Stenzel (1945</bibRefCitation>
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; see discussion about
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<taxonomicName authorityName="Bell" authorityYear="1850" box="[1044,1169,693,719]" class="Malacostraca" family="Erymidae" genus="Palaeastacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="531" phylum="Arthropoda" rank="genus">
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<emphasis box="[1044,1169,693,719]" italics="true" pageId="16" pageNumber="531">Palaeastacus</emphasis>
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</taxonomicName>
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). Our examination of the
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<typeStatus box="[814,860,726,752]" pageId="16" pageNumber="531">type</typeStatus>
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material of
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<taxonomicName authorityName="Whitfield" authorityYear="1883" box="[988,1089,725,751]" class="Malacostraca" family="Erymidae" genus="Palaeastacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="walkeri">
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<emphasis box="[988,1089,725,751]" italics="true" pageId="16" pageNumber="531">P. walkeri</emphasis>
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</taxonomicName>
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leads us to a different conclusion. Indeed, this carapace shows groove pattern typical of
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<taxonomicName authorityName="M'Coy" authorityYear="1849" class="Malacostraca" family="Erymidae" genus="Enoploclytia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="531" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="16" pageNumber="531">Enoploclytia</emphasis>
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</taxonomicName>
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(long gastro-orbital groove, postcervical groove joined to hepatic groove, short branchiocardiac groove not joined to postcervical nor to hepatic grooves).
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</paragraph>
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<paragraph blockId="16.[812,1458,598,1071]" pageId="16" pageNumber="531">
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As for
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<taxonomicName authority="Devillez & Charbonnier & Hyžný & Leroy, 2016" authorityName="Devillez & Charbonnier & Hyžný & Leroy" authorityYear="2016" box="[900,1166,885,911]" class="Malacostraca" family="Erymidae" genus="Enoploclytia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="augustobonae" status="n. sp.">
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<emphasis box="[900,1166,885,911]" italics="true" pageId="16" pageNumber="531">Enoploclytia augustobonae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1173,1231,885,911]" pageId="16" pageNumber="531" rank="species">n. sp.</taxonomicNameLabel>
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,
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<taxonomicName authority="Devillez & Charbonnier & Hyžný & Leroy, 2016" authorityName="Devillez & Charbonnier & Hyžný & Leroy" authorityYear="2016" box="[1239,1454,885,911]" class="Malacostraca" family="Erymidae" genus="Enoploclytia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="gigantea" status="n. sp.">
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<emphasis box="[1239,1454,885,911]" italics="true" pageId="16" pageNumber="531">Enoploclytia gigantea</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[813,869,917,943]" pageId="16" pageNumber="531" rank="species">n. sp.</taxonomicNameLabel>
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(known by its carapace) cannot be compared to
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<taxonomicName authorityName="Stenzel" authorityYear="1945" class="Malacostraca" family="Erymidae" genus="Enoploclytia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="wintoni">
|
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|
<emphasis italics="true" pageId="16" pageNumber="531">Enoploclytia wintoni</emphasis>
|
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|
</taxonomicName>
|
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|
|
(only known by P1 chelae). For a comparison, see discussion on
|
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|
|
<taxonomicName authority="Devillez & Charbonnier & Hyžný & Leroy, 2016" authorityName="Devillez & Charbonnier & Hyžný & Leroy" authorityYear="2016" box="[1003,1171,980,1006]" class="Malacostraca" family="Erymidae" genus="Enoploclytia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="augustobonae" status="n. sp.">
|
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|
|
<emphasis box="[1003,1171,980,1006]" italics="true" pageId="16" pageNumber="531">E. augustobonae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[1178,1237,981,1007]" pageId="16" pageNumber="531" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="16.[812,1458,598,1071]" pageId="16" pageNumber="531">
|
|||
|
|
In conclusion,
|
|||
|
|
<taxonomicName authority="Devillez & Charbonnier & Hyžný & Leroy, 2016" authorityName="Devillez & Charbonnier & Hyžný & Leroy" authorityYear="2016" box="[1007,1234,1012,1038]" class="Malacostraca" family="Erymidae" genus="Enoploclytia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="gigantea" status="n. sp.">
|
|||
|
|
<emphasis box="[1007,1234,1012,1038]" italics="true" pageId="16" pageNumber="531">Enoploclytia gigantea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[1242,1304,1013,1039]" pageId="16" pageNumber="531" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
possesses the biggest carapace currently known among all erymid lobsters.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|