treatments-xml/data/A4/80/54/A4805418925C8B55EDE0F4AC4D0BBAC5.xml

<document id="84FBF6A2625CA8F02F7FC539E3898720" ID-CLB-Dataset="35011" ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1453" ID-GBIF-Dataset="51bb4676-675d-429f-86ed-819a712454bc" ID-PMC="PMC3260773" ID-Pensoft-Pub="1313-2970-130-421" ID-PubMed="22259291" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-421" ModsDocOrigin="ZooKeys 130" ModsDocTitle="New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae)" checkinTime="1451249849988" checkinUser="pensoft" docAuthor="Perrichot, Vincent, Ortega-Blanco, Jaime, McKellar, Ryan C., Delclos, Xavier, Azar, Dany, Nel, Andre, Tafforeau, Paul &amp; Engel, Michael S." docDate="2011" docId="A4805418925C8B55EDE0F4AC4D0BBAC5" docLanguage="en" docName="ZooKeys 130: 421-453" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1453" docTitle="Iberomaimetsha nihtmara Ortega-Blanco, Delclos &amp; Engel, sp. n." docType="treatment" docVersion="6" lastPageNumber="437" masterDocId="7C0CFFB3434B4C34FFA1FFD2FFEEFFD8" masterDocTitle="New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae)" masterLastPageNumber="453" masterPageNumber="421" pageNumber="433" updateTime="1732746649147" updateUser="ExternalLinkService">
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<mods:title id="8C5B5EF1A576343164A0E92FDD5454CB">New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae)</mods:title>
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<mods:namePart id="C1FD8E4B857689DA399E4472C2989095">Perrichot, Vincent</mods:namePart>
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<mods:namePart id="4326AD884106439E6CD3E69144EF7FA5">Ortega-Blanco, Jaime</mods:namePart>
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<mods:namePart id="907D08F20C547343309D43AF72FBF727">McKellar, Ryan C.</mods:namePart>
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<mods:namePart id="18B70A0FE818FA796F36C0EEBD8917C9">Delclos, Xavier</mods:namePart>
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<mods:namePart id="044DDE6B2194D7B35F80BDDA35B04E33">Azar, Dany</mods:namePart>
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<mods:namePart id="C22D4B5FFB97E806EADB9AD083FA187E">Nel, Andre</mods:namePart>
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<mods:namePart id="D56C2EE2007D9DFC63F93C6B187F10B3">Tafforeau, Paul</mods:namePart>
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<mods:namePart id="63F62D497D332D730EA98BEB1409BC43">Engel, Michael S.</mods:namePart>
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<treatment id="A4805418925C8B55EDE0F4AC4D0BBAC5" ID-GBIF-Taxon="152030999" LSID="urn:lsid:zoobank.org:act:8E3C73D0-4CE1-4910-9DE6-45A0D3BC2F1D" httpUri="http://treatment.plazi.org/id/A4805418925C8B55EDE0F4AC4D0BBAC5" lastPageId="16" lastPageNumber="437" pageId="12" pageNumber="433" scope_family="Maimetshidae">
<subSubSection id="19E6D9DE038558E4CC5052377169C9C6" pageId="12" pageNumber="433" type="nomenclature">
<paragraph id="247136867DB597DE2370F984D3310A9C" pageId="12" pageNumber="433">
<taxonomicName id="E62F24B08A84A78962C245D12E86038F" LSID="urn:lsid:zoobank.org:act:8E3C73D0-4CE1-4910-9DE6-45A0D3BC2F1D" authority="Ortega-Blanco, Delclos &amp; Engel" genus="Iberomaimetsha" lsidName="Iberomaimetsha nihtmara" pageId="12" pageNumber="433" rank="species" species="nihtmara">
Iberomaimetsha nihtmara Ortega-Blanco, 
<normalizedToken id="39F357703270F06DB8F90837B035DB9E" originalValue="Delclòs">Delclos</normalizedToken>
&amp; Engel
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<taxonomicNameLabel id="8933AB06E3C7E9277DE49EAD246101D5" pageId="12" pageNumber="433">sp. n.</taxonomicNameLabel>
Figs 810
</paragraph>
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<subSubSection id="2A0A0F385A064A4CE5211D23CAC1CE4F" pageId="12" pageNumber="433" type="type material">
<paragraph id="0B4B4711F05E2C1F9363D5DAA34AD301" pageId="12" pageNumber="433">Type material.</paragraph>
<paragraph id="127BF7D09520C489A91DB6C5B3F101DD" pageId="12" pageNumber="433">
Holotype MCNA 13049 (?); paratypes MCNA 8758(♀?), MCNA 8790 (♂), MCNA 9373 (?), MCNA 9918 (♀), and MCNA 10732 (?). All specimens deposited in the Museo de Ciencias Naturales de 
<normalizedToken id="782E0D70F018E2D6D026E301FAD68BEC" originalValue="Álava">Alava</normalizedToken>
, Vitoria-Gasteiz, Spain.
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<subSubSection id="BA092979668E1739D54EDE81915D695A" pageId="12" pageNumber="433" type="type locality">
<paragraph id="F5F083C5FA87019E0B803B2DF38696FE" pageId="12" pageNumber="433">Type locality.</paragraph>
<paragraph id="CC53E21E25617B35F83FCED4FC13E4B2" pageId="12" pageNumber="433">
<normalizedToken id="BDE52FF963CB854C7A5A9530D193C0CE" originalValue="Peñacerrada">Penacerrada</normalizedToken>
I, Moraza, Burgos Province, Basque-Cantabrian Basin, Spain.
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<subSubSection id="870B2BBD7500C72CB8FC266F03630DF4" pageId="12" pageNumber="433" type="stratigraphic horizon">
<paragraph id="A3259CCED51B945B9C6BD079014DA9D2" pageId="12" pageNumber="433">Stratigraphic horizon.</paragraph>
<paragraph id="E35CEDAD11B4488AF051E013E69866C3" pageId="12" pageNumber="433">Escucha Formation, Lower Cretaceous, Lower Albian.</paragraph>
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<subSubSection id="2A7B053E2CC9F41521C7C9C67C1D2551" pageId="12" pageNumber="433" type="diagnosis">
<paragraph id="02C4234CBC36AADCB4F526CC9CAB4E8F" pageId="12" pageNumber="433">Diagnosis.</paragraph>
<paragraph id="F739B008C4CA3F89D9362BFBDE900373" pageId="12" pageNumber="433">
Mandibles distinctly large, exserted, asymmetrical, right 4-toothed, left 3-toothed, arched inwards (teeth pointing toward teeth of opposite mandible); basal flagellomeres (from F1 to F3-4) distinctly larger than apical flagellomeres (F1-F4 about 2 
<normalizedToken id="7A1F34C66BB27D64F8A498456A279719" originalValue="–1.5×">-1.5x</normalizedToken>
the length of F7-F14); protibial distal margin with a comb of stiff setae; cell [1M] square or rhomboidal, distinctly smaller than [2Cu] which has a short but distinct connection between 1M+Cu and M (then, [2Cu] composed by five aristae); 2Rs+M longer than 1m-cu and longer than cell [1Rs]; apicalmost sector of Cu arising from almost apical posterior margin of [2Cu]; metasoma moderately short, as long as mesosoma.
</paragraph>
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<subSubSection id="23E71D94C7919E2F9EC0E13401E301F2" lastPageId="16" lastPageNumber="437" pageId="12" pageNumber="433" type="description">
<paragraph id="A21988A3F655433312B06679B8795D00" pageId="12" pageNumber="433">Description.</paragraph>
<paragraph id="B58D0536C44E8B95C696D3849C926457" lastPageId="15" lastPageNumber="436" pageId="12" pageNumber="433">
Body length (from paratypes, as incomplete metasoma in holotype) around 2.55-2.61 mm. Head rounded in anterior view, length 0.41 mm, width 0.68 mm with a short occipital carina and apparently some polygonal dorsal sculpture; compound eyes not bulging, slightly oval, separated dorsally by 0.39 mm; toruli about at same distance to inner margin of compound eyes and each other; antennae 2.54 mm in length, inserted on frons below midpoint of compound eyes; scape and pedicel short and globular, flagellomeres cylindrical; lengths of antennomeres as follows (all in mm): scape 0.12, pedicel 0.09, flagellomeres 0.29, 0.28, 0.27, 0.23, 0.20, 0.18, 0.15, 0.14, 0.12, 0.12, 0.11, 0.11, 0.10, 0.12; malar space short, length approximately 0.11 mm. Mesosoma (dorsal view) 0.95 mm in length; pronotum not distinct dorsally; mesoscutal length 0.27 mm, with 
<pageBreakToken id="DF91C45C4809DB80886E7F016B3A591E" pageId="13" pageNumber="434" start="start">parallel</pageBreakToken>
notauli and longitudinal medial line, all three present as linear series of grooves more than well-delimited lines (Fig. 9); axillae wide and long, well distinct, separating completely mesoscutellum and mesoscutum; mesoscutellum distinctly convex, length 0.22 mm; propodeum 0.29 mm in length, areolate (seen in paratype MCNA 8790). Legs distinctly elongate; division of trochantellus not visible; tarsi with no plantar lobes visible; leg measurements (all in mm): profemur 0.82, protibia 0.46, protarsomeres 
<normalizedToken id="6495891530A9DD65F1304F70B6B6501D" originalValue="I–V">I-V</normalizedToken>
0.34/0.15/0.10/0.09/0.10; mid-legs very poorly preserved but apparently with mesofemur 0.68, mesotibia?, meso 
<pageBreakToken id="3D1680192381D83C5010BA51D75ECC57" pageId="14" pageNumber="435" start="start">tarsomeres</pageBreakToken>
0.24/0.12/0.10/?/?; metafemur 1.0, metatibia 0.83, metatarsomeres 0.46/0.20/0.15/0.09/0.12. Forewing hyaline, length 2.1 mm, maximum width 0.80 mm, densely covered by microtrichia; all veins tubular except for nebulous 2rs-m and 2m-cu, thus, only cells [2Rs] and [2M] pseudo-opened apically (Figs 9, 10)
<pageBreakToken id="D4BFDA51E7703266A7124EEA8A90C42C" pageId="15" pageNumber="436" start="start">;</pageBreakToken>
1rs-m not aligned with r-rs; 2m-cu curved S-like; veins M and Cu reaching wing margin (although M somewhat nebulous), with apicalmost sector of Cu slightly angled posteriorly; cell [1R1] longer than both cells [1Rs]+[2Rs]; cell [1Rs] almost 
<normalizedToken id="D0A2E3B50FD8BED9097BC29D5069D903" originalValue="¾">3/4</normalizedToken>
the surface of cell [1M] and almost half of [2Rs]; vein r-rs emerging at two-thirds from base of pterostigma; cell [2R1] short, with vein Rs slightly anteriorly bent distad 2rs-m. Hind wing difficult to observe, hyaline and densely covered by microtrichia, length approximately 1.22 mm, maximum width approximately 0.41 mm; three strong hamuli present on costal vein in distal half of wing. Metasoma (checked on paratype MCNA 8790) pedunculate, short and slightly flattened, length 0.98 mm, greatest height 0.41 mm; sterna hard, convex; no apparent metasomal armature on sternum II or III.
</paragraph>
<paragraph id="B9D99BD126D1BD8910234A442889050C" pageId="16" pageNumber="437">
<pageBreakToken id="2953006AAF92D20032D3E55D6A581B9B" pageId="16" pageNumber="437" start="start">Male</pageBreakToken>
: Parameres spatulate, with apparently three short stiff setae apically (Fig. 10).
</paragraph>
<caption id="05997D1A9D843633D0D394D3A6420476" pageId="16" pageNumber="437">
<paragraph id="5679AD553C760C25C9A78E8D59608ECB" pageId="16" pageNumber="437">
Figure 8. Photomicrographs of holotype of 
<taxonomicName id="B1E65452EE88805AB1D151FD87E8D6C9" genus="Iberomaimetsha" lsidName="Iberomaimetsha nihtmara" pageId="16" pageNumber="437" rank="species" species="nihtmara">Iberomaimetsha nihtmara</taxonomicName>
Ortega-Blanco, 
<normalizedToken id="3F4E2384C847D8E9DD7333A5E68C1F7C" originalValue="Delclòs">Delclos</normalizedToken>
, and Engel sp. n. (MCNA 13049) A Dorsal aspect B Ventral aspect.
</paragraph>
</caption>
<caption id="B2C29F7E8BB13707C26AFFF3092C74DE" pageId="16" pageNumber="437">
<paragraph id="E28C16F2EF7CB4A526318F0514553ECF" pageId="16" pageNumber="437">
Figure 9. Habitus diagram of holotype of 
<taxonomicName id="BB10E90BA9F03EC8E90D2989A2C2B79C" genus="Iberomaimetsha" lsidName="Iberomaimetsha nihtmara" pageId="16" pageNumber="437" rank="species" species="nihtmara">Iberomaimetsha nihtmara</taxonomicName>
Ortega-Blanco, 
<normalizedToken id="8E4971632590F92E465B96281337281E" originalValue="Delclòs">Delclos</normalizedToken>
, and Engel sp. n. (MCNA 13049), with inset of facial aspect.
</paragraph>
</caption>
<caption id="A4FD7E0A08BBF2590F0A42287D2C8B40" pageId="16" pageNumber="437">
<paragraph id="187C80D4ED800C1797BB2BFF0872B342" pageId="16" pageNumber="437">
Figure 10. Habitus diagram of male paratype of 
<taxonomicName id="44B6C9B92668A0757325435A271F211A" genus="Iberomaimetsha" lsidName="Iberomaimetsha nihtmara" pageId="16" pageNumber="437" rank="species" species="nihtmara">Iberomaimetsha nihtmara</taxonomicName>
Ortega-Blanco, 
<normalizedToken id="EE0F129040E378DA9FD21DD245823864" originalValue="Delclòs">Delclos</normalizedToken>
, and Engel sp. n. (MCNA 8790), with inset of facial aspect.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E347D85D3DDA3FC9AD07F3FBC4A30B67" pageId="16" pageNumber="437" type="etymology">
<paragraph id="1CB1C9A8F67343CB53A82A00DF3C3AFE" pageId="16" pageNumber="437">Etymology.</paragraph>
<paragraph id="A2BBF6D3E829C372DF11FA50259AF4CC" pageId="16" pageNumber="437">The specific epithet is a noun in apposition and is the Anglo-Saxon word for nightmare, in reference to the terrifying appearance of the species in frontal view and with its enlarged mandibles.</paragraph>
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</treatment>
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