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<mods:title id="980C117BF0741F89D1C2CD7BD914E9BB">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<heading id="D089F966FFC6FFE0FF48FF5C3341FBBA" bold="true" box="[192,573,1074,1098]" fontSize="10" level="2" pageId="13" pageNumber="12" reason="3">
<taxonomicName id="4C7E3589FFC6FFE0FF48FF5C3341FBBA" ID-CoL="628LF" authority="Latreille, 1804" authorityName="Latreille" authorityYear="1804" box="[192,573,1074,1098]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Lucaninae">
<emphasis id="B90A9218FFC6FFE0FF48FF5C3341FBBA" bold="true" box="[192,573,1074,1098]" pageId="13" pageNumber="12">LUCANINAE Latreille, 1804</emphasis>
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<heading id="D089F966FFC6FFE0FF48FF123340FB64" bold="true" box="[192,572,1148,1172]" fontSize="10" level="2" pageId="13" pageNumber="12" reason="3">
<emphasis id="B90A9218FFC6FFE0FF48FF123340FB64" bold="true" box="[192,572,1148,1172]" pageId="13" pageNumber="12">
FIGULINI
<bibRefCitation id="EFEF33FBFFC6FFE0FEDCFF123340FB64" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[340,572,1148,1172]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="13" pageNumber="12" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister, 1847</bibRefCitation>
</emphasis>
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<subSubSection id="C3641D81FFC6FFE0FF60FFC23595FA56" pageId="13" pageNumber="12" type="description">
<paragraph id="8BC14E0AFFC6FFE0FF60FFC232FFFB31" blockId="13.[232,899,1195,1246]" box="[232,899,1195,1217]" pageId="13" pageNumber="12">
= Penichrolucanini Arrow, 1950 -
<bibRefCitation id="EFEF33FBFFC6FFE0FDD7FFC53271FB30" author="Paulsen" box="[607,781,1195,1217]" firstAuthor="Paulsen" pageId="13" pageNumber="12" pagination="1 - 13" refId="ref219175" refString="Paulsen, M. J. 2018. Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini). Insecta Mundi 0624: 1 - 13." type="book chapter" year="2018">Paulsen 2018: 2</bibRefCitation>
(syn. nov.)
</paragraph>
<paragraph id="8BC14E0AFFC6FFE0FF60FFA632FCFB2D" blockId="13.[232,899,1195,1246]" box="[232,896,1224,1246]" pageId="13" pageNumber="12">
= Brasilucanini
<bibRefCitation id="EFEF33FBFFC6FFE0FE14FFA6333BFB2D" author="Nikolajev" box="[412,583,1224,1246]" firstAuthor="Nikolajev" pageId="13" pageNumber="12" pagination="171 - 172" refId="ref217077" refString="Nikolajev, G. V. 1999. On the polyphyly of the subfamily Penichrolucaninae (Coleoptera, Lucanidae), with the erection of the new monotypic tribe Brasilucanini. Tethys Entomological Research 1: 171 - 172." type="journal article" year="1999">Nikolajev, 1999</bibRefCitation>
-
<bibRefCitation id="EFEF33FBFFC6FFE0FDD4FFA63276FB2D" author="Paulsen" box="[604,778,1224,1246]" firstAuthor="Paulsen" pageId="13" pageNumber="12" pagination="1 - 13" refId="ref219175" refString="Paulsen, M. J. 2018. Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini). Insecta Mundi 0624: 1 - 13." type="book chapter" year="2018">Paulsen 2018: 2</bibRefCitation>
(syn. nov.)
</paragraph>
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<emphasis id="B90A9218FFC6FFE0FF48FF803074FAF6" bold="true" box="[192,264,1262,1286]" pageId="13" pageNumber="12">Note.</emphasis>
In their description of the genus
<taxonomicName id="4C7E3589FFC6FFE0FD19FF8035BDFAF6" authority=", Vulcano and Pereira (1961: 473)" authorityName=", Vulcano and Pereira" authorityPageNumber="473" authorityYear="1961" box="[657,1217,1262,1286]" class="Insecta" family="Lucanidae" genus="Brasilucanus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B90A9218FFC6FFE0FD19FF80324FFAF6" box="[657,819,1262,1286]" italics="true" pageId="13" pageNumber="12">Brasilucanus</emphasis>
,
<bibRefCitation id="EFEF33FBFFC6FFE0FCC8FF8035BDFAF6" author="Vulcano and Pereira" box="[832,1217,1262,1286]" firstAuthor="Vulcano" pageId="13" pageNumber="12" pagination="471 - 480" refId="ref228907" refString="Vulcano, M. A., and F. S. Pereira. 1961. A subfamilia Penichrolucaninae (Col. Lucanidae) representada em America. Studia Entomologica 4 (1 - 4): 471 - 480." type="journal article" year="1961">Vulcano and Pereira (1961: 473)</bibRefCitation>
</taxonomicName>
place it within the tribe Penichrolucanini.
<bibRefCitation id="EFEF33FBFFC6FFE0FE57FE60339EFAD6" author="Nikolajev" box="[479,738,1294,1318]" firstAuthor="Nikolajev" pageId="13" pageNumber="12" pagination="171 - 172" refId="ref217077" refString="Nikolajev, G. V. 1999. On the polyphyly of the subfamily Penichrolucaninae (Coleoptera, Lucanidae), with the erection of the new monotypic tribe Brasilucanini. Tethys Entomological Research 1: 171 - 172." type="journal article" year="1999">Nikolajev (1999: 171)</bibRefCitation>
places it within the new monogeneric tribe Brasilucanini. Both these tribes are treated as synonyms of the subfamily
<taxonomicName id="4C7E3589FFC6FFE0FC06FE403573FAB6" authorityName="Latreille" authorityYear="1804" box="[910,1039,1326,1350]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Lucaninae">Lucaninae</taxonomicName>
by
<bibRefCitation id="EFEF33FBFFC6FFE0FBB1FE40346DFAB6" author="Smith" box="[1081,1297,1326,1350]" firstAuthor="Smith" pageId="13" pageNumber="12" pagination="144 - 204" refId="ref225329" refString="Smith, A. B. T. 2006. A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification. Coleopterists Society Monograph 5: 144 - 204." type="journal article" year="2006">Smith (2006: 155)</bibRefCitation>
and as synonyms of the tribe
<taxonomicName id="4C7E3589FFC6FFE0FE17FE203370FA96" authorityName="Latreille" authorityYear="1804" box="[415,524,1358,1382]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Lucanini">Lucanini</taxonomicName>
by
<bibRefCitation id="EFEF33FBFFC6FFE0FDBDFE20320EFA96" author="Bouchard, P. &amp; Y. Bousquet &amp; A. E. Davies &amp; M. A. Alonso-Zarazaga &amp; J. F. Lawrence &amp; C. H. C. Lyal &amp; A. F. Newton &amp; C. A. M. Reid &amp; M. Schmitt &amp; S. A. Slipinski &amp; A. B. T. Smith" box="[565,882,1358,1382]" journalOrPublisher="ZooKeys" pageId="13" pageNumber="12" pagination="1 - 972" part="88" refId="ref194757" refString="Bouchard, P., Y. Bousquet, A. E. Davies, M. A. Alonso-Zarazaga, J. F. Lawrence, C. H. C. Lyal, A. F. Newton, C. A. M. Reid, M. Schmitt, S. A. Slipinski, and A. B. T. Smith. 2011. Familygroup names in Coleoptera (Insecta). ZooKeys 88: 1 - 972." title="Familygroup names in Coleoptera (Insecta)" type="journal article" year="2011">Bouchard et al. (2011: 233</bibRefCitation>
, 235). The genus is again placed in the revived Penichrolucanini by
<bibRefCitation id="EFEF33FBFFC6FFE0FE34FE003232FA76" author="Bartolozzi, L. &amp; M. Zilioli" box="[444,846,1390,1414]" journalOrPublisher="Onychium" pageId="13" pageNumber="12" pagination="123 - 128" part="12" refId="ref193271" refString="Bartolozzi, L., and M. Zilioli. 2016. Remarks on the genus Penichrolucanus Deyrolle, 1863 (Coleoptera: Lucanidae). Onychium 12: 123 - 128." title="Remarks on the genus Penichrolucanus Deyrolle, 1863 (Coleoptera: Lucanidae)" type="journal article" year="2016">Bartolozzi and Zilioli (2016: 123)</bibRefCitation>
.
<bibRefCitation id="EFEF33FBFFC6FFE0FCD1FE003551FA76" author="Paulsen" box="[857,1069,1390,1414]" firstAuthor="Paulsen" pageId="13" pageNumber="12" pagination="1 - 13" refId="ref219175" refString="Paulsen, M. J. 2018. Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini). Insecta Mundi 0624: 1 - 13." type="book chapter" year="2018">Paulsen (2018: 2)</bibRefCitation>
considers Penichrolucanini as well as Brasilucanini to be synonyms of the tribe Figulini. We follow the latter opinion.
</paragraph>
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