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<document id= "7F9EE3B154E5E15B4F708995A7F4874F" ID-CLB-Dataset= "298249" ID-DOI= "10.1653/024.098.0405" ID-GBIF-Dataset= "7d14bbc5-e5e9-464f-b58f-e35fe117cfa2" ID-ISSN= "1938-5102" ID-Zenodo-Dep= "11548347" IM.bibliography_approvedBy= "felipe" IM.illustrations_approvedBy= "felipe" IM.materialsCitations_approvedBy= "juliana" IM.metadata_approvedBy= "juliana" IM.taxonomicNames_approvedBy= "juliana" IM.treatments_approvedBy= "juliana" checkinTime= "1717620029050" checkinUser= "felipe" docAuthor= "Morón, Miguel Ángel, Lugo-García, Gabriel A. & Aragón-García, Agustín" docDate= "2015" docId= "03AE87EDFF9ECA5CFF31A7E9FDD7A145" docLanguage= "en" docName= "FlaEntomol.98.4.1036-1043.pdf" docOrigin= "Florida Entomologist 98 (4)" docSource= "http://dx.doi.org/10.1653/024.098.0405" docStyle= "DocumentStyle:C0CD22107794FB65089F701CBA118AA5.2:FlaEntomol.2015-.journal_article" docStyleId= "C0CD22107794FB65089F701CBA118AA5" docStyleName= "FlaEntomol.2015-.journal_article" docStyleVersion= "2" docTitle= "Phyllophaga collaroides Morón, Lugo-García & Aragón-García, 2015, sp. nov." docType= "treatment" docVersion= "3" lastPageNumber= "1043" masterDocId= "FF97FF95FF98CA54FFA1A135FFABA322" masterDocTitle= "Five new species of Phyllophaga (Coleoptera: Melolonthidae) of northwestern Mexico" masterLastPageNumber= "1043" masterPageNumber= "1036" pageNumber= "1041" updateTime= "1718028493070" updateUser= "ExternalLinkService" zenodo-license-document= "CC-BY-4.0" >
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<mods:title id= "63F99445D0DBA53B4D7EBF9E02B06915" > Five new species of Phyllophaga (Coleoptera: Melolonthidae) of northwestern Mexico</mods:title>
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<mods:namePart id= "213B7741F40F5F2544CF62F2DB34034C" > Morón, Miguel Ángel</mods:namePart>
<mods:affiliation id= "E8AAD9B98EE4CBC9C98B5FCBD01490B1" > Red de Biodiversidad y Sistemática, Instituto de Ecología, A. C. Apdo. Postal 63, Xalapa, Veracruz 91000, México</mods:affiliation>
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<mods:namePart id= "60C55B2F47D39528BFF40F013531D88D" > Lugo-García, Gabriel A.</mods:namePart>
<mods:affiliation id= "BEAC808E46C85FD75A168BB4FD3973CC" > Escuela Superior de Agricultura del Valle del Fuerte, Universidad Autónoma de Sinaloa, Calle 16 y Av. Japaraqui, El Estero, Juan José Rios, Ahome, Sinaloa 81110, México</mods:affiliation>
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<mods:namePart id= "F46AE88FDB556614D5CE2E264D55070B" > Aragón-García, Agustín</mods:namePart>
<mods:affiliation id= "491A35942EE6DAEF58E843C99E073972" > Centro de Agroecología, Instituto de Ciencias, Benemérita Universidad Autónoma de Puebla, 14 Sur 6301, San Manuel, Puebla, Puebla 72570, México</mods:affiliation>
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<treatment id= "03AE87EDFF9ECA5CFF31A7E9FDD7A145" ID-DOI= "http://doi.org/10.5281/zenodo.11548160" ID-GBIF-Taxon= "232723916" ID-Zenodo-Dep= "11548160" LSID= "urn:lsid:plazi:treatment:03AE87EDFF9ECA5CFF31A7E9FDD7A145" httpUri= "http://treatment.plazi.org/id/03AE87EDFF9ECA5CFF31A7E9FDD7A145" lastPageId= "8" lastPageNumber= "1043" pageId= "6" pageNumber= "1041" >
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<subSubSection id= "C31D6570FF9ECA52FF31A7E9FC85A5D6" box= "[144,814,1756,1781]" pageId= "6" pageNumber= "1041" type= "nomenclature" >
<paragraph id= "8BB836FBFF9ECA52FF31A7E9FC85A5D6" blockId= "6.[144,814,1756,1781]" box= "[144,814,1756,1781]" pageId= "6" pageNumber= "1041" >
<heading id= "D0F08197FF9ECA52FF31A7E9FC85A5D6" box= "[144,814,1756,1781]" centered= "true" fontSize= "10" level= "2" pageId= "6" pageNumber= "1041" reason= "3" >
<taxonomicName id= "4C074D78FF9ECA52FF31A7E9FE4DA5D7" ID-CoL= "4H8ZQ" authorityName= "Moron, Lugo-Garcia & Aragon-Garcia" authorityYear= "2015" box= "[144,486,1756,1781]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" isSensuStricto= "true" kingdom= "Animalia" order= "Coleoptera" pageId= "6" pageNumber= "1041" phylum= "Arthropoda" rank= "species" species= "collaroides" status= "sp. nov." >
<emphasis id= "B973EAE9FF9ECA52FF31A7E9FEB6A5D7" box= "[144,285,1756,1781]" italics= "true" pageId= "6" pageNumber= "1041" > Phyllophaga</emphasis>
(
<emphasis id= "B973EAE9FF9ECA52FE8AA7E9FEF5A5D7" box= "[299,350,1756,1781]" italics= "true" pageId= "6" pageNumber= "1041" > s.str.</emphasis>
)
<emphasis id= "B973EAE9FF9ECA52FECDA7E9FE4DA5D7" box= "[364,486,1756,1781]" italics= "true" pageId= "6" pageNumber= "1041" > collaroides</emphasis>
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<emphasis id= "B973EAE9FF9ECA52FE4DA7EBFDEEA5D7" bold= "true" box= "[492,581,1758,1781]" pageId= "6" pageNumber= "1041" >
<taxonomicNameLabel id= "A2405792FF9ECA52FE4DA7EBFDEEA5D7" box= "[492,581,1758,1781]" pageId= "6" pageNumber= "1041" rank= "species" > sp. nov.</taxonomicNameLabel>
</emphasis>
(Figs. 21– –
</heading>
</paragraph>
</subSubSection>
<subSubSection id= "C31D6570FF9ECA53FF31A62DFBD6A3D6" lastPageId= "7" lastPageNumber= "1042" pageId= "6" pageNumber= "1041" type= "description" >
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<materialsCitation id= "3B6F3CA6FF9ECA52FF31A62DFEECA40C" box= "[144,327,1816,1838]" collectionCode= "MALE" pageId= "6" pageNumber= "1041" specimenCount= "1" typeStatus= "holotype" >
<collectionCode id= "ED16AE3EFF9ECA52FF31A62DFF65A40C" box= "[144,206,1816,1838]" pageId= "6" pageNumber= "1041" > MALE</collectionCode>
<typeStatus id= "54BC8859FF9ECA52FF75A62DFEECA40C" box= "[212,327,1816,1838]" pageId= "6" pageNumber= "1041" type= "holotype" > HOLOTYPE</typeStatus>
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</heading>
</paragraph>
<paragraph id= "8BB836FBFF9ECA52FF18A676FAEAA585" blockId= "6.[144,825,1859,2026]" lastBlockId= "6.[856,1537,1625,2026]" pageId= "6" pageNumber= "1041" >
Total body length
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, humeral width
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. Head and pronotum glossy reddish brown, elytra glossy pale brownish yellow (Figs. 21– × ×
</paragraph>
<paragraph id= "8BB836FBFF9ECA52FCDDA784FC67A4C8" blockId= "6.[856,1537,1625,2026]" pageId= "6" pageNumber= "1041" > Pronotum wider than long (2:1) and 2.0× × × × </paragraph>
<paragraph id= "8BB836FBFF9FCA53FF25A1F6FE57A2FD" blockId= "7.[96,777,195,655]" pageId= "7" pageNumber= "1042" > Propygidium densely covered with short yellow setae. Pygidium convex (Fig. 21), glossy, with sparse short setae throughout; apical margin widely rounded, with 10 short setae. Pterosternum with long, abundant yellowish setae. Second to fifh abdominal sternites convex, with many short setae; posterior border of 5th sternite slightly depressed; anal plate large, shallowly excavated, finely rugo-punctate with some erect setae, posterior margin widely curved, clearly notched at middle, with few short setae. Protibia as long as protarsus (1:1). Metatibia shorter than metatarsus (1.0:1.1). Tarsal claws with large tooth at middle of ventral border (Fig. 33).</paragraph>
<paragraph id= "8BB836FBFF9FCA53FF25A0DDFCAEA1AD" blockId= "7.[96,777,195,655]" pageId= "7" pageNumber= "1042" > Parameres symmetrical, long, fused dorsally, strongly curved downward with large tooth-like projection on the middle of distal surface; phallobase long, convex; sclerotized support of aedeagus widened apically and curved upward, and with sclerotized accessories on the dorsum of pre-apical membranous sac (Figs. 34– </paragraph>
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FEMALE
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</heading>
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<paragraph id= "8BB836FBFF9FCA53FF25A3C2FDD8A0FA" blockId= "7.[96,778,759,984]" pageId= "7" pageNumber= "1042" >
Similar to male,except:Body length 19.0 mm.Humeral width
<quantity id= "4CFF9B1EFF9FCA53FD1AA3C2FCAEA029" box= "[699,773,759,779]" metricMagnitude= "-3" metricUnit= "m" metricValue= "7.4" pageId= "7" pageNumber= "1042" unit= "mm" value= "7.4" > 7.4 mm</quantity>
. Antennal club 0.9×
</paragraph>
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</paragraph>
<paragraph id= "8BB836FBFF9FCA53FF25A575FE46A78E" blockId= "7.[96,776,1088,1196]" pageId= "7" pageNumber= "1042" >
Male: The only other male examined has a body length of
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and humeral width of
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. Dorsal color slightly darker than
<typeStatus id= "54BC8859FF9FCA53FD74A568FF24A7AD" pageId= "7" pageNumber= "1042" type= "holotype" > holotype</typeStatus>
. Punctation of head is slightly denser, and vestiture of pronotum and abdomen is longer than in
<typeStatus id= "54BC8859FF9FCA53FE2FA5ADFE43A78E" box= "[398,488,1176,1196]" pageId= "7" pageNumber= "1042" type= "holotype" > holotype</typeStatus>
.
</paragraph>
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Female: The second female examined has a body length of
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and humeral width of
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.
</paragraph>
</subSubSection>
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<typeStatus id= "54BC8859FF9FCA53FCEDA065FC1CA246" box= "[844,951,336,356]" pageId= "7" pageNumber= "1042" type= "holotype" > HOLOTYPE</typeStatus>
:
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—
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:
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,
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,
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,
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<collectingDate id= "EFFDE9D3FF9FCA53FA2EA065FCF7A2A3" pageId= "7" pageNumber= "1042" value= "2009-07-09" > 09-VII-2009</collectingDate>
</date>
,
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,
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(
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)
</materialsCitation>
.
<materialsCitation id= "3B6F3CA6FF9FCA53FBF5A058FB1BA2BD" collectingDate= "2009-07-20" collectionCode= "IEXA" collectorName= "Graciela" country= "Mexico" elevation= "1488" location= "Los Tascates" municipality= "Choix" pageId= "7" pageNumber= "1042" specimenCount= "1" stateProvince= "Sinaloa" typeStatus= "allotype" >
<typeStatus id= "54BC8859FF9FCA53FBF5A058FB12A2A3" box= "[1108,1209,365,385]" pageId= "7" pageNumber= "1042" type= "allotype" > ALLOTYPE</typeStatus>
:
<specimenCount id= "9D01FD72FF9FCA53FB61A058FAA9A2A3" box= "[1216,1282,365,385]" pageId= "7" pageNumber= "1042" type= "female" > female</specimenCount>
—same data as holotype except
<date id= "FFB9103BFF9FCA53FC01A0BEFBBFA2BD" box= "[928,1044,395,415]" pageId= "7" pageNumber= "1042" value= "2009-07-20" >
<collectingDate id= "EFFDE9D3FF9FCA53FC01A0BEFBBFA2BD" box= "[928,1044,395,415]" pageId= "7" pageNumber= "1042" value= "2009-07-20" > 20-VII-2009</collectingDate>
</date>
,
<collectorName id= "26F2532DFF9FCA53FBBCA0BEFBC7A2BD" box= "[1053,1132,395,415]" pageId= "7" pageNumber= "1042" > Graciela</collectorName>
(
<collectionCode id= "ED16AE3EFF9FCA53FBD6A0BEFB00A2BD" box= "[1143,1195,395,415]" pageId= "7" pageNumber= "1042" > IEXA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id= "3B6F3CA6FF9FCA53FB18A0BEFB69A29E" collectingDate= "2009-07-09" collectionCode= "MXAL" collectorName= "Graciela" country= "Mexico" elevation= "1488" location= "Los Tascates" municipality= "Choix" pageId= "7" pageNumber= "1042" specimenCount= "1" specimenCount-female= "1" stateProvince= "Sinaloa" typeStatus= "paratype" >
<typeStatus id= "54BC8859FF9FCA53FB18A0BEFA82A2BD" box= "[1209,1321,395,415]" pageId= "7" pageNumber= "1042" type= "paratype" > PARATYPES</typeStatus>
:
<specimenCount id= "9D01FD72FF9FCA53FA90A0BEFA2EA2BD" box= "[1329,1413,395,415]" count= "1" pageId= "7" pageNumber= "1042" type= "female" > 1 female</specimenCount>
—same data as holotype except
<collectorName id= "26F2532DFF9FCA53FB81A09DFBC4A29E" box= "[1056,1135,424,444]" pageId= "7" pageNumber= "1042" > Graciela</collectorName>
(
<collectionCode id= "ED16AE3EFF9FCA53FBDDA09DFB17A29E" box= "[1148,1212,424,444]" pageId= "7" pageNumber= "1042" > MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id= "3B6F3CA6FF9FCA53FB6CA09DFA6BA2FB" collectingDate= "2009-07-09" collectionCode= "MXAL" collectorName= "G. A. Lugo" country= "Mexico" elevation= "1488" location= "Los Tascates" pageId= "7" pageNumber= "1042" specimenCount= "1" specimenCount-male= "1" stateProvince= "Chihuahua" typeStatus= "paratype" >
<specimenCount id= "9D01FD72FF9FCA53FB6CA09DFAA4A29E" box= "[1229,1295,424,444]" count= "1" pageId= "7" pageNumber= "1042" type= "male" > 1 male</specimenCount>
—
<collectingCountry id= "F310766BFF9FCA53FA85A09DFAD1A29E" box= "[1316,1402,424,444]" name= "Mexico" pageId= "7" pageNumber= "1042" > MEXICO</collectingCountry>
:
<collectingRegion id= "49C3F819FF9FCA53FA24A09DFCFBA2FB" country= "Mexico" name= "Chihuahua" pageId= "7" pageNumber= "1042" > Chihuahua</collectingRegion>
, Morelos,
<location id= "8ED86020FF9FCA53FC15A0F0FB84A2FB" LSID= "urn:lsid:plazi:treatment:03AE87EDFF9ECA5CFF31A7E9FDD7A145:8ED86020FF9FCA53FC15A0F0FB84A2FB" box= "[948,1071,453,473]" country= "Mexico" name= "Los Tascates" pageId= "7" pageNumber= "1042" stateProvince= "Chihuahua" > Los Tascates</location>
,
<date id= "FFB9103BFF9FCA53FB97A0F0FB00A2FB" box= "[1078,1195,453,473]" pageId= "7" pageNumber= "1042" value= "2009-07-09" >
<collectingDate id= "EFFDE9D3FF9FCA53FB97A0F0FB00A2FB" box= "[1078,1195,453,473]" pageId= "7" pageNumber= "1042" value= "2009-07-09" > 09-VII-2009</collectingDate>
</date>
,
<quantity id= "4CFF9B1EFF9FCA53FB12A0F0FAAEA2FB" box= "[1203,1285,453,473]" metricMagnitude= "3" metricUnit= "m" metricValue= "1.488" pageId= "7" pageNumber= "1042" unit= "m" value= "1488.0" >
<elevation id= "002AD1C8FF9FCA53FB12A0F0FAAEA2FB" box= "[1203,1285,453,473]" metricMagnitude= "3" metricUnit= "m" metricValue= "1.488" pageId= "7" pageNumber= "1042" unit= "m" value= "1488.0" > 1,488 m</elevation>
</quantity>
,
<collectorName id= "26F2532DFF9FCA53FAAFA0F0FADBA2FB" box= "[1294,1392,453,473]" pageId= "7" pageNumber= "1042" > G. A. Lugo</collectorName>
(
<collectionCode id= "ED16AE3EFF9FCA53FADAA0F0FA10A2FB" box= "[1403,1467,453,473]" pageId= "7" pageNumber= "1042" > MXAL</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id= "8BB836FBFF9FCA53FC89A33FFC62A102" blockId= "7.[808,969,522,544]" box= "[808,969,522,544]" pageId= "7" pageNumber= "1042" >
<heading id= "D0F08197FF9FCA53FC89A33FFC62A102" box= "[808,969,522,544]" fontSize= "10" level= "3" pageId= "7" pageNumber= "1042" reason= "9" >
<collectionCode id= "ED16AE3EFF9FCA53FC89A33FFCF6A102" box= "[808,861,522,544]" pageId= "7" pageNumber= "1042" > TYPE</collectionCode>
<collectionCode id= "ED16AE3EFF9FCA53FCC2A33FFC62A102" box= "[867,969,522,544]" pageId= "7" pageNumber= "1042" > LOCALITY</collectionCode>
</heading>
</paragraph>
<paragraph id= "8BB836FBFF9FCA53FCEDA300FA7AA16B" blockId= "7.[844,1489,565,585]" box= "[844,1489,565,585]" pageId= "7" pageNumber= "1042" >
<materialsCitation id= "3B6F3CA6FF9FCA53FCEDA300FA66A16B" box= "[844,1485,565,585]" country= "Mexico" latitude= "26.645945" location= "Los Tascates" longLatPrecision= "1" longitude= "-107.85244" municipality= "Choix" pageId= "7" pageNumber= "1042" specimenCount= "1" stateProvince= "Sinaloa" >
<location id= "8ED86020FF9FCA53FCEDA300FC6EA16B" LSID= "urn:lsid:plazi:treatment:03AE87EDFF9ECA5CFF31A7E9FDD7A145:8ED86020FF9FCA53FCEDA300FC6EA16B" box= "[844,965,565,585]" country= "Mexico" latitude= "26.645945" longLatPrecision= "1" longitude= "-107.85244" municipality= "Choix" name= "Los Tascates" pageId= "7" pageNumber= "1042" stateProvince= "Sinaloa" > Los Tascates</location>
,
<collectingMunicipality id= "6BDCAC81FF9FCA53FC6DA300FBAEA16B" box= "[972,1029,565,585]" pageId= "7" pageNumber= "1042" > Choix</collectingMunicipality>
,
<collectingRegion id= "49C3F819FF9FCA53FBADA300FBFDA16B" box= "[1036,1110,565,585]" country= "Mexico" name= "Sinaloa" pageId= "7" pageNumber= "1042" > Sinaloa</collectingRegion>
,
<collectingCountry id= "F310766BFF9FCA53FBFCA300FB0FA16B" box= "[1117,1188,565,585]" name= "Mexico" pageId= "7" pageNumber= "1042" > Mexico</collectingCountry>
(
<geoCoordinate id= "EE33503CFF9FCA53FB0CA300FA9FA16B" box= "[1197,1332,565,585]" degrees= "26" direction= "north" minutes= "38" orientation= "latitude" pageId= "7" pageNumber= "1042" precision= "1" seconds= "45.4" value= "26.645945" > 26°38'45.4" N</geoCoordinate>
,
<geoCoordinate id= "EE33503CFF9FCA53FA9AA300FA6CA16B" box= "[1339,1479,565,585]" degrees= "107" direction= "west" minutes= "51" orientation= "longitude" pageId= "7" pageNumber= "1042" precision= "1" seconds= "8.8" value= "-107.85244" > 107°51'8.8" W</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id= "C31D6570FF9FCA53FC89A34FFB84A04B" pageId= "7" pageNumber= "1042" type= "biology_ecology" >
<paragraph id= "8BB836FBFF9FCA53FC89A34FFC41A1B2" blockId= "7.[808,1002,634,656]" box= "[808,1002,634,656]" pageId= "7" pageNumber= "1042" >
<heading id= "D0F08197FF9FCA53FC89A34FFC41A1B2" box= "[808,1002,634,656]" fontSize= "10" level= "3" pageId= "7" pageNumber= "1042" reason= "9" >
<collectionCode id= "ED16AE3EFF9FCA53FC89A34FFC07A1B2" box= "[808,940,634,656]" pageId= "7" pageNumber= "1042" > BIOLOGICAL</collectionCode>
<collectionCode id= "ED16AE3EFF9FCA53FC13A34FFC41A1B2" box= "[946,1002,634,656]" pageId= "7" pageNumber= "1042" > DATA</collectionCode>
</heading>
</paragraph>
<paragraph id= "8BB836FBFF9FCA53FCEDA390FB84A04B" blockId= "7.[808,1489,677,873]" pageId= "7" pageNumber= "1042" >
Specimens of
<taxonomicName id= "4C074D78FF9FCA53FC77A390FBFEA19B" authorityName= "Moron, Lugo-Garcia & Aragon-Garcia" authorityYear= "2015" box= "[982,1109,677,697]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "collaroides" >
<emphasis id= "B973EAE9FF9FCA53FC77A390FBFEA19B" box= "[982,1109,677,697]" italics= "true" pageId= "7" pageNumber= "1042" > P. collaroides</emphasis>
</taxonomicName>
were collected in a pine–
<quantity id= "4CFF9B1EFF9FCA53FC89A3F6FCD3A1F5" box= "[808,888,707,727]" metricMagnitude= "3" metricUnit= "m" metricValue= "1.488" pageId= "7" pageNumber= "1042" unit= "m" value= "1488.0" > 1,488 m</quantity>
elevation using a black-light trap and white-fluorescent–
<specimenCount id= "9D01FD72FF9FCA53FC3BA3D5FBB0A1D6" box= "[922,1051,736,756]" count= "4" pageId= "7" pageNumber= "1042" type= "generic" > 4 specimens</specimenCount>
). Other species of
<taxonomicName id= "4C074D78FF9FCA53FB76A3D5FAE4A1D6" authorityName= "Harris" authorityYear= "1826" box= "[1239,1359,736,756]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "genus" >
<emphasis id= "B973EAE9FF9FCA53FB76A3D5FAE4A1D6" box= "[1239,1359,736,756]" italics= "true" pageId= "7" pageNumber= "1042" > Phyllophaga</emphasis>
</taxonomicName>
flying at the same time and place were
<taxonomicName id= "4C074D78FF9FCA53FB90A3C8FAD1A033" authority= "(Blanchard)" baseAuthorityName= "Blanchard" box= "[1073,1402,765,785]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "obsoleta" subGenus= "Phytalus" >
<emphasis id= "B973EAE9FF9FCA53FB90A3C8FBEFA033" box= "[1073,1092,765,785]" italics= "true" pageId= "7" pageNumber= "1042" > P.</emphasis>
(
<emphasis id= "B973EAE9FF9FCA53FBEFA3C8FB0BA033" box= "[1102,1184,765,785]" italics= "true" pageId= "7" pageNumber= "1042" > Phytalus</emphasis>
)
<emphasis id= "B973EAE9FF9FCA53FB0FA3C8FAAAA033" box= "[1198,1281,765,785]" italics= "true" pageId= "7" pageNumber= "1042" > obsoleta</emphasis>
(Blanchard)
</taxonomicName>
,
<taxonomicName id= "4C074D78FF9FCA53FA22A3C8FB91A00D" authority= "(Blanchard)" baseAuthorityName= "Blanchard" baseAuthorityYear= "1850" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "ravida" subGenus= "Phyllophaga" >
<emphasis id= "B973EAE9FF9FCA53FA22A3C8FA3CA033" box= "[1411,1431,765,785]" italics= "true" pageId= "7" pageNumber= "1042" > P.</emphasis>
(
<emphasis id= "B973EAE9FF9FCA53FA01A3C8FCDCA00D" italics= "true" pageId= "7" pageNumber= "1042" > Phyllophaga</emphasis>
)
<emphasis id= "B973EAE9FF9FCA53FC25A22EFC6AA00D" box= "[900,961,795,815]" italics= "true" pageId= "7" pageNumber= "1042" > ravida</emphasis>
(Blanchard)
</taxonomicName>
,
<taxonomicName id= "4C074D78FF9FCA53FBE3A22EFB56A00D" authority= "(Horn)" baseAuthorityName= "Horn" baseAuthorityYear= "1887" box= "[1090,1277,795,815]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "vetula" subGenus= "Phyllophaga" >
<emphasis id= "B973EAE9FF9FCA53FBE3A22EFBFEA00D" box= "[1090,1109,795,815]" italics= "true" pageId= "7" pageNumber= "1042" > P.</emphasis>
(
<emphasis id= "B973EAE9FF9FCA53FBFCA22EFBC5A00D" box= "[1117,1134,795,815]" italics= "true" pageId= "7" pageNumber= "1042" > P.</emphasis>
)
<emphasis id= "B973EAE9FF9FCA53FBD8A22EFB1EA00D" box= "[1145,1205,795,815]" italics= "true" pageId= "7" pageNumber= "1042" > vetula</emphasis>
(Horn)
</taxonomicName>
,
<taxonomicName id= "4C074D78FF9FCA53FAA4A22EFA65A00D" authority= "(Moser)" baseAuthorityName= "Moser" baseAuthorityYear= "1918" box= "[1285,1486,795,815]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "opaca" subGenus= "Phyllophaga" >
<emphasis id= "B973EAE9FF9FCA53FAA4A22EFAB3A00D" box= "[1285,1304,795,815]" italics= "true" pageId= "7" pageNumber= "1042" > P.</emphasis>
(
<emphasis id= "B973EAE9FF9FCA53FA80A22EFA99A00D" box= "[1313,1330,795,815]" italics= "true" pageId= "7" pageNumber= "1042" > P.</emphasis>
)
<emphasis id= "B973EAE9FF9FCA53FA9DA22EFADCA00D" box= "[1340,1399,795,815]" italics= "true" pageId= "7" pageNumber= "1042" > opaca</emphasis>
(Moser)
</taxonomicName>
,
<taxonomicName id= "4C074D78FF9FCA53FC89A20DFB9AA06E" authority= "(Burmeister)" baseAuthorityName= "Burmeister" baseAuthorityYear= "1855" box= "[808,1073,824,844]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "setifera" subGenus= "Phyllophaga" >
<emphasis id= "B973EAE9FF9FCA53FC89A20DFC90A06E" box= "[808,827,824,844]" italics= "true" pageId= "7" pageNumber= "1042" > P.</emphasis>
(
<emphasis id= "B973EAE9FF9FCA53FCE7A20DFCFCA06E" box= "[838,855,824,844]" italics= "true" pageId= "7" pageNumber= "1042" > P.</emphasis>
)
<emphasis id= "B973EAE9FF9FCA53FCC5A20DFC06A06E" box= "[868,941,824,844]" italics= "true" pageId= "7" pageNumber= "1042" > setifera</emphasis>
(Burmeister)
</taxonomicName>
,
<taxonomicName id= "4C074D78FF9FCA53FB9AA20DFB43A06E" authorityName= "Moron, Lugo-Garcia & Aragon-Garcia" authorityYear= "2015" box= "[1083,1256,824,844]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "reyesolivasi" status= "sp. nov." subGenus= "Listrochelus" >
<emphasis id= "B973EAE9FF9FCA53FB9AA20DFBE4A06E" box= "[1083,1103,824,844]" italics= "true" pageId= "7" pageNumber= "1042" > P.</emphasis>
(
<emphasis id= "B973EAE9FF9FCA53FBF8A20DFBC1A06E" box= "[1113,1130,824,844]" italics= "true" pageId= "7" pageNumber= "1042" > L.</emphasis>
)
<emphasis id= "B973EAE9FF9FCA53FBD9A20DFB43A06E" box= "[1144,1256,824,844]" italics= "true" pageId= "7" pageNumber= "1042" > reyesolivasi</emphasis>
</taxonomicName>
<emphasis id= "B973EAE9FF9FCA53FB4EA20DFA96A06E" bold= "true" box= "[1263,1341,824,844]" pageId= "7" pageNumber= "1042" >
<taxonomicNameLabel id= "A2405792FF9FCA53FB4EA20DFA96A06E" box= "[1263,1341,824,844]" pageId= "7" pageNumber= "1042" rank= "species" > sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id= "4C074D78FF9FCA53FAD7A20DFC76A04B" authorityName= "Moron, Lugo-Garcia & Aragon-Garcia" authorityYear= "2015" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "tascatensis" status= "sp. nov." subGenus= "Listrochelus" >
<emphasis id= "B973EAE9FF9FCA53FAD7A20DFA22A06E" box= "[1398,1417,824,844]" italics= "true" pageId= "7" pageNumber= "1042" > P.</emphasis>
(
<emphasis id= "B973EAE9FF9FCA53FA32A20DFCCEA04B" italics= "true" pageId= "7" pageNumber= "1042" > Listrochelus</emphasis>
)
<emphasis id= "B973EAE9FF9FCA53FCD3A260FC76A04B" box= "[882,989,853,873]" italics= "true" pageId= "7" pageNumber= "1042" > tascatensis</emphasis>
</taxonomicName>
<emphasis id= "B973EAE9FF9FCA53FC43A260FB84A04B" bold= "true" box= "[994,1071,853,873]" pageId= "7" pageNumber= "1042" >
<taxonomicNameLabel id= "A2405792FF9FCA53FC43A260FB84A04B" box= "[994,1071,853,873]" pageId= "7" pageNumber= "1042" rank= "species" > sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id= "C31D6570FF9FCA5CFC89A294FED6A2FD" lastPageId= "8" lastPageNumber= "1043" pageId= "7" pageNumber= "1042" type= "discussion" >
<paragraph id= "8BB836FBFF9FCA53FC89A294FB8BA095" blockId= "7.[808,1056,929,951]" box= "[808,1056,929,951]" pageId= "7" pageNumber= "1042" >
<heading id= "D0F08197FF9FCA53FC89A294FB8BA095" box= "[808,1056,929,951]" fontSize= "10" level= "3" pageId= "7" pageNumber= "1042" reason= "9" > TAXONOMIC REMARKS</heading>
</paragraph>
<paragraph id= "8BB836FBFF9FCA5CFCEDA2F9FED6A2FD" blockId= "7.[808,1488,972,1197]" lastBlockId= "8.[144,825,195,479]" lastPageId= "8" lastPageNumber= "1043" pageId= "7" pageNumber= "1042" >
<taxonomicName id= "4C074D78FF9FCA53FCEDA2F9FBEFA0C2" authorityName= "Moron, Lugo-Garcia & Aragon-Garcia" authorityYear= "2015" box= "[844,1092,972,992]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "collaroides" >
<emphasis id= "B973EAE9FF9FCA53FCEDA2F9FBEFA0C2" box= "[844,1092,972,992]" italics= "true" pageId= "7" pageNumber= "1042" > Phyllophaga collaroides</emphasis>
</taxonomicName>
is part of the “blanchardi” species group (sensu
<bibRefCitation id= "EF964B0AFF9FCA53FC18A2DCFB94A0DF" author= "Moron MA" box= "[953,1087,1001,1021]" pageId= "7" pageNumber= "1042" refId= "ref7626" refString= "Moron MA. 1986. El genero Phyllophaga en Mexico. Morfologia, Distribucion y Sistematica Supraespecifica (Insecta: Coleoptera). Instituto de Ecologia, A. C. Mexico." type= "book" year= "1986" > Morón 1986</bibRefCitation>
). This group is one of the largest and most complex within the subgenus
<taxonomicName id= "4C074D78FF9FCA53FB32A533FABBA738" authorityName= "Harris" authorityYear= "1826" box= "[1171,1296,1030,1050]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "genus" >
<emphasis id= "B973EAE9FF9FCA53FB32A533FABBA738" box= "[1171,1296,1030,1050]" italics= "true" pageId= "7" pageNumber= "1042" > Phyllophaga</emphasis>
</taxonomicName>
(
<emphasis id= "B973EAE9FF9FCA53FABEA533FAE1A738" box= "[1311,1354,1030,1050]" italics= "true" pageId= "7" pageNumber= "1042" > s.str</emphasis>
.), with some undescribed Mexican species and subspecies, and frequently represented by large numbers in light-trap samples. The group is currently under study by the senior author.
<taxonomicName id= "4C074D78FF9FCA53FB03A56BFA38A750" authorityName= "Moron, Lugo-Garcia & Aragon-Garcia" authorityYear= "2015" box= "[1186,1427,1118,1138]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "collaroides" >
<emphasis id= "B973EAE9FF9FCA53FB03A56BFA38A750" box= "[1186,1427,1118,1138]" italics= "true" pageId= "7" pageNumber= "1042" > Phyllophaga collaroides</emphasis>
</taxonomicName>
looks quite different from
<taxonomicName id= "4C074D78FF9FCA53FC5CA549FB1DA7B2" authority= "(Moser)" baseAuthorityName= "Moser" baseAuthorityYear= "1921" box= "[1021,1206,1148,1168]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "collaris" >
<emphasis id= "B973EAE9FF9FCA53FC5CA549FBF6A7B2" box= "[1021,1117,1148,1168]" italics= "true" pageId= "7" pageNumber= "1042" > P. collaris</emphasis>
(Moser)
</taxonomicName>
externally, but the form of parameres of the 2 species is very similar. Shape of the clypeus, proportions of the antennomeres, form of the tarsal claws, and punctation of the elytra are similar between the two, but in
<taxonomicName id= "4C074D78FF90CA5CFD02A1D5FCABA3D6" baseAuthorityName= "Moser" baseAuthorityYear= "1921" box= "[675,768,224,244]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "8" pageNumber= "1043" phylum= "Arthropoda" rank= "species" species= "collaris" >
<emphasis id= "B973EAE9FF90CA5CFD02A1D5FCABA3D6" box= "[675,768,224,244]" italics= "true" pageId= "8" pageNumber= "1043" > P. collaris</emphasis>
</taxonomicName>
body size is much smaller (
<quantity id= "4CFF9B1EFF90CA5CFEC7A1C8FDABA233" box= "[358,512,253,273]" metricMagnitude= "-2" metricUnit= "m" metricValue= "1.1" metricValueMax= "1.15" metricValueMin= "1.05" pageId= "8" pageNumber= "1043" unit= "mm" value= "11.0" valueMax= "11.5" valueMin= "10.5" > 10.5– </quantity>
); the abdominal sternites have few setae; the pygidial surface is glabrous and coarsely rugose; the male anal plate is moderately expanded; and the female pygidium is uniformly convex. In
<taxonomicName id= "4C074D78FF90CA5CFEDEA060FDA8A24B" authorityName= "Moron, Lugo-Garcia & Aragon-Garcia" authorityYear= "2015" box= "[383,515,341,361]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "8" pageNumber= "1043" phylum= "Arthropoda" rank= "species" species= "collaroides" >
<emphasis id= "B973EAE9FF90CA5CFEDEA060FDA8A24B" box= "[383,515,341,361]" italics= "true" pageId= "8" pageNumber= "1043" > P. collaroides</emphasis>
</taxonomicName>
body size is larger (
<quantity id= "4CFF9B1EFF90CA5CFD70A060FF17A2A5" metricMagnitude= "-2" metricUnit= "m" metricValue= "1.64" metricValueMax= "1.73" metricValueMin= "1.55" pageId= "8" pageNumber= "1043" unit= "mm" value= "16.4" valueMax= "17.3" valueMin= "15.5" > 15.5– </quantity>
); the abdominal sternites are covered with many short setae; the pygidial surface is densely punctate and setiferous; the male anal plate is widely expanded; and the female pygidium has a large pre-apical prominence.
</paragraph>
</subSubSection>
2024-07-12 22:07:22 +02:00
<caption id= "DF786673FF9FCA53FFCDA6AFFC17A4FE" ID-DOI= "http://doi.org/10.5281/zenodo.11548353" ID-Zenodo-Dep= "11548353" httpUri= "https://zenodo.org/record/11548353/files/figure.png" pageId= "7" pageNumber= "1042" startId= "7.[108,145,1946,1964]" targetBox= "[96,1481,1242,1909]" targetPageId= "7" targetType= "figure" >
2024-06-21 12:22:17 +02:00
<paragraph id= "8BB836FBFF9FCA53FFCDA6AFFC17A4FE" blockId= "7.[96,1490,1946,2012]" pageId= "7" pageNumber= "1042" >
<emphasis id= "B973EAE9FF9FCA53FFCDA6AFFEC0A48E" bold= "true" box= "[108,363,1946,1964]" pageId= "7" pageNumber= "1042" > Figs. 33– – </emphasis>
Habitus and morphological characters of
<taxonomicName id= "4C074D78FF9FCA53FD6EA6AFFC30A48E" authorityName= "Moron, Lugo-Garcia & Aragon-Garcia" authorityYear= "2015" box= "[719,923,1946,1964]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "7" pageNumber= "1042" phylum= "Arthropoda" rank= "species" species= "collaroides" status= "sp. nov." >
<emphasis id= "B973EAE9FF9FCA53FD6EA6AFFC30A48E" box= "[719,923,1946,1964]" italics= "true" pageId= "7" pageNumber= "1042" > Phyllophaga collaroides</emphasis>
</taxonomicName>
,
<emphasis id= "B973EAE9FF9FCA53FC05A6AFFBA4A48E" bold= "true" box= "[932,1039,1946,1964]" pageId= "7" pageNumber= "1042" >
<taxonomicNameLabel id= "A2405792FF9FCA53FC05A6AFFBA4A48E" box= "[932,1039,1946,1964]" pageId= "7" pageNumber= "1042" rank= "species" > new species</taxonomicNameLabel>
</emphasis>
.
<emphasis id= "B973EAE9FF9FCA53FBB9A6AFFBFFA48E" bold= "true" box= "[1048,1108,1946,1964]" pageId= "7" pageNumber= "1042" > 21– </emphasis>
Habitus of males: 21. lateral view, 22. dorsal view, scale bar = 2 mm.
<emphasis id= "B973EAE9FF9FCA53FE8FA687FEC1A4E6" bold= "true" box= "[302,362,1970,1988]" pageId= "7" pageNumber= "1042" > 33– </emphasis>
Morphological characters: 33. male tarsal claw; 34–
</paragraph>
</caption>
<subSubSection id= "C31D6570FF90CA5CFF31A33EFDD7A145" pageId= "8" pageNumber= "1043" type= "etymology" >
<paragraph id= "8BB836FBFF90CA5CFF31A33EFEB2A103" blockId= "8.[144,281,523,545]" box= "[144,281,523,545]" pageId= "8" pageNumber= "1043" >
<heading id= "D0F08197FF90CA5CFF31A33EFEB2A103" box= "[144,281,523,545]" fontSize= "10" level= "3" pageId= "8" pageNumber= "1043" reason= "9" > ETYMOLOGY</heading>
</paragraph>
<paragraph id= "8BB836FBFF90CA5CFF15A303FDD7A145" blockId= "8.[144,824,566,615]" pageId= "8" pageNumber= "1043" >
The Latinized name of the new species references the genital similarity between it and
<taxonomicName id= "4C074D78FF90CA5CFEC1A366FDD3A145" authority= "(Moser)" baseAuthorityName= "Moser" baseAuthorityYear= "1921" box= "[352,632,595,615]" class= "Insecta" family= "Scarabaeidae" genus= "Phyllophaga" kingdom= "Animalia" order= "Coleoptera" pageId= "8" pageNumber= "1043" phylum= "Arthropoda" rank= "species" species= "collaris" >
<emphasis id= "B973EAE9FF90CA5CFEC1A366FD89A145" box= "[352,546,595,615]" italics= "true" pageId= "8" pageNumber= "1043" > Phyllophaga collaris</emphasis>
(Moser)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
