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<document id="1E1FF152C2A2D3063F2B2370478A46FE" ID-CLB-Dataset="199060" ID-DOI="10.11646/zootaxa.5213.5.3" ID-GBIF-Dataset="c73ab242-8a3e-4f76-81b4-fd9eada9f0e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7381670" ID-ZooBank="449781B5-94E0-4B6C-9F6B-D0711FC08BB2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669810183154" checkinUser="plazi" docAuthor="Allsopp, Peter G." docDate="2022" docId="03AE87B9FFB5827BFF6772551621FB8A" docLanguage="en" docName="zootaxa.5213.5.3.pdf" docOrigin="Zootaxa 5213 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidiota ciliata Britton 1978" docType="treatment" docVersion="7" lastPageNumber="518" masterDocId="FF97FFC1FFB0827EFFF0714A1538FFF7" masterDocTitle="Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae): reclassification of eight species to Antitrogus Burmeister, 1855 and Barryfilius new genus" masterLastPageNumber="545" masterPageNumber="513" pageNumber="518" updateTime="1699412597972" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="92984FF548194F6130AF063481F4D02B">Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae): reclassification of eight species to Antitrogus Burmeister, 1855 and Barryfilius new genus</mods:title>
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<mods:namePart id="4924742A9DC5A15C8286DAAA8A8E6046">Allsopp, Peter G.</mods:namePart>
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<mods:date id="0BAB934C03AA11FB0A1586B35B83491E">2022</mods:date>
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<treatment id="03AE87B9FFB5827BFF6772551621FB8A" ID-DOI="http://doi.org/10.5281/zenodo.7383617" ID-GBIF-Taxon="205354366" ID-Zenodo-Dep="7383617" LSID="urn:lsid:plazi:treatment:03AE87B9FFB5827BFF6772551621FB8A" httpUri="http://treatment.plazi.org/id/03AE87B9FFB5827BFF6772551621FB8A" lastPageNumber="518" pageId="5" pageNumber="518">
<subSubSection id="C31D6524FFB5827BFF6772551789FCCE" box="[151,689,799,826]" pageId="5" pageNumber="518" type="nomenclature">
<paragraph id="8BB836AFFFB5827BFF6772551789FCCE" blockId="5.[151,689,799,826]" box="[151,689,799,826]" pageId="5" pageNumber="518">
<heading id="D0F081C3FFB5827BFF6772551789FCCE" bold="true" box="[151,689,799,826]" fontSize="11" level="1" pageId="5" pageNumber="518" reason="1">
<emphasis id="B973EABDFFB5827BFF6772551789FCCE" bold="true" box="[151,689,799,826]" pageId="5" pageNumber="518">
Placement of
<taxonomicName id="4C074D2CFFB5827BFEB172551789FCCE" authority="Britton, 1978" authorityName="Britton" authorityYear="1978" box="[321,689,799,826]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="ciliata">
<emphasis id="B973EABDFFB5827BFEB17255173DFCCD" bold="true" box="[321,517,799,826]" italics="true" pageId="5" pageNumber="518">Lepidiota ciliata</emphasis>
<bibRefCitation id="EF964B5EFFB5827BFDFC72551789FCCE" author="Britton, E. B." box="[524,689,799,825]" pageId="5" pageNumber="518" pagination="1 - 150" refId="ref32215" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton, 1978</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D6524FFB5827BFF6772221621FB8A" pageId="5" pageNumber="518" type="description">
<paragraph id="8BB836AFFFB5827BFF6772221024FBE6" blockId="5.[151,1436,871,1150]" pageId="5" pageNumber="518">
In
<bibRefCitation id="EF964B5EFFB5827BFF45722D145FFC76" author="Britton, E. B." box="[181,359,871,897]" pageId="5" pageNumber="518" pagination="1 - 150" refId="ref32215" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton’s (1978)</bibRefCitation>
key to the Australian
<taxonomicName id="4C074D2CFFB5827BFDAA722D1618FC76" box="[602,800,871,897]" pageId="5" pageNumber="518">
<emphasis id="B973EABDFFB5827BFDAA722D17FDFC76" box="[602,709,871,897]" italics="true" pageId="5" pageNumber="518">Lepidiota</emphasis>
species
</taxonomicName>
,
<taxonomicName id="4C074D2CFFB5827BFCDA72221111FC76" authority="Britton, 1978" authorityName="Britton" authorityYear="1978" box="[810,1065,871,897]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="ciliata">
<emphasis id="B973EABDFFB5827BFCDA722216B6FC76" box="[810,910,871,897]" italics="true" pageId="5" pageNumber="518">L. ciliata</emphasis>
<bibRefCitation id="EF964B5EFFB5827BFC64722D1111FC76" author="Britton, E. B." box="[916,1065,871,897]" pageId="5" pageNumber="518" pagination="1 - 150" refId="ref32215" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton, 1978</bibRefCitation>
</taxonomicName>
falls as an isolated species late in his key.
<bibRefCitation id="EF964B5EFFB5827BFF0572C114AEFC52" author="Britton, E. B." box="[245,406,907,933]" pageId="5" pageNumber="518" pagination="1 - 150" refId="ref32215" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton (1978)</bibRefCitation>
noted his difficulty with the placement of
<taxonomicName id="4C074D2CFFB5827BFC8872C616E7FC52" authorityName="Britton" authorityYear="1978" box="[888,991,907,933]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="ciliata">
<emphasis id="B973EABDFFB5827BFC8872C616E7FC52" box="[888,991,907,933]" italics="true" pageId="5" pageNumber="518">L. ciliata</emphasis>
</taxonomicName>
, remarking “This species has also been included in the key to
<taxonomicName id="4C074D2CFFB5827BFE6B72E51770FC3D" box="[411,584,943,970]" class="Insecta" family="Melolonthidae" genus="Antitrogus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B973EABDFFB5827BFE6B72E51728FC3E" box="[411,528,943,969]" italics="true" pageId="5" pageNumber="518">Antitrogus</emphasis>
spp.
</taxonomicName>
to allow for the possibility of interpretation of the very elongate scales as setae [at couplet 19(18)] in the key to genera of
<taxonomicName id="4C074D2CFFB5827BFD5772991607FC1A" baseAuthorityName="sensu Britton" baseAuthorityYear="1978" box="[679,831,979,1005]" class="Insecta" family="Scarabaeidae" genus="Melolonthini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="genus">Melolonthini</taxonomicName>
”.
<bibRefCitation id="EF964B5EFFB5827BFCA27299112EFC1A" author="Weir, T. A. &amp; Lawrence, J. F. &amp; Lemann, C. &amp; Gunter, N. L." box="[850,1046,979,1005]" pageId="5" pageNumber="518" pagination="1819" refId="ref34705" refString="Weir, T. A., Lawrence, J. F., Lemann, C. &amp; Gunter, N. L. (2019) Scarabaeidae: Melolonthinae Leach, 1819. In: Slipinski, A. &amp; Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 467 - 507." type="book chapter" year="2019">
Weir
<emphasis id="B973EABDFFB5827BFC7E729E16FDFC1A" box="[910,965,979,1005]" italics="true" pageId="5" pageNumber="518">et al.</emphasis>
(2019)
</bibRefCitation>
grouped
<taxonomicName id="4C074D2CFFB5827BFB8D729E11DAFC1A" authorityName="Britton" authorityYear="1978" box="[1149,1250,979,1005]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="ciliata">
<emphasis id="B973EABDFFB5827BFB8D729E11DAFC1A" box="[1149,1250,979,1005]" italics="true" pageId="5" pageNumber="518">L. ciliata</emphasis>
</taxonomicName>
with
<taxonomicName id="4C074D2CFFB5827BFAEF729E1415FBE6" authority="Britton, 1978" authorityName="Britton" authorityYear="1978" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="quinaria">
<emphasis id="B973EABDFFB5827BFAEF729E10A4FC1A" box="[1311,1436,980,1005]" italics="true" pageId="5" pageNumber="518">L. quinaria</emphasis>
<bibRefCitation id="EF964B5EFFB5827BFF6772BD1415FBE6" author="Britton, E. B." box="[151,301,1015,1041]" pageId="5" pageNumber="518" pagination="1 - 150" refId="ref32215" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton, 1978</bibRefCitation>
</taxonomicName>
at their couplet 22, suggesting that they were distinct from mainstream
<taxonomicName id="4C074D2CFFB5827BFBA172BD1024FBE6" box="[1105,1308,1015,1041]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="undefined-i">
<emphasis id="B973EABDFFB5827BFBA172BD1184FBE6" box="[1105,1212,1015,1041]" italics="true" pageId="5" pageNumber="518">Lepidiota</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="8BB836AFFFB5827BFF3775561621FB8A" blockId="5.[151,1436,871,1150]" pageId="5" pageNumber="518">
I consider that the “very elongate scales” of
<bibRefCitation id="EF964B5EFFB5827BFD2675511643FBC2" author="Britton, E. B." box="[726,891,1051,1077]" pageId="5" pageNumber="518" pagination="1 - 150" refId="ref32215" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton (1978)</bibRefCitation>
should be considered setae (
<figureCitation id="133C2A2AFFB5827BFB3F75511062FBC2" box="[1231,1370,1051,1077]" captionStart="FIGURES 10–13" captionStartId="7.[151,264,1030,1054]" captionTargetBox="[151,1436,181,1007]" captionTargetId="figure-15@7.[151,1436,181,1007]" captionTargetPageId="7" captionText="FIGURES 10–13. Male of Antitrogus ciliatus (Britton, 1978). 10, Dorsal view, Mount Kaputar specimen; 11, lateral view, Mount Kaputar specimen; 12, head, frontal view; 13, aedeagus, ventral and lateral views. Figs. 10–11 by Greg Daniels; Fig. 12 reproduced from Weir et al. (2019) (CSIRO, used with permission); Fig. 13 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.7381680" httpUri="https://zenodo.org/record/7381680/files/figure.png" pageId="5" pageNumber="518">Figs. 10–12</figureCitation>
) and, also because
<taxonomicName id="4C074D2CFFB5827BFEDC750A14ABFBAE" authorityName="Britton" authorityYear="1978" box="[300,403,1087,1113]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="ciliata">
<emphasis id="B973EABDFFB5827BFEDC750A14ABFBAE" box="[300,403,1087,1113]" italics="true" pageId="5" pageNumber="518">L. ciliata</emphasis>
</taxonomicName>
has a five-lamellate club in both males and females, it should be placed in
<taxonomicName id="4C074D2CFFB5827BFB1775751064FBAE" authorityName=": Blackburn" authorityYear="1911" box="[1255,1372,1087,1113]" class="Insecta" family="Melolonthidae" genus="Antitrogus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="B973EABDFFB5827BFB1775751064FBAE" box="[1255,1372,1087,1113]" italics="true" pageId="5" pageNumber="518">Antitrogus</emphasis>
</taxonomicName>
. This resolves Britton’s dilemma and simplifies keys to genera.
</paragraph>
</subSubSection>
</treatment>
</document>
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