treatments-xml/data/F6/51/DA/F651DA1CFFD8A56D57BCFA86E45D42E3.xml

<document id="03DFDCA170FAC413B6A40D31A56211C8" ID-CLB-Dataset="3899" ID-DOI="10.11646/zootaxa.5057.1.6" ID-GBIF-Dataset="3f50ca0f-843f-494f-aff8-defcc449ad44" ID-ISSN="1175-5326" ID-Zenodo-Dep="5705482" ID-ZooBank="4074E652-255A-44E9-B837-5CB26745D06F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634714061645" checkinUser="plazi" docAuthor="Sithole, Yonela, Heemstra, Elaine &amp; Mwale, Monica" docDate="2021" docId="F651DA1CFFD8A56D57BCFA86E45D42E3" docLanguage="en" docName="zootaxa.5057.1.6.pdf" docOrigin="Zootaxa 5057 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Serranus cabrilla" docType="treatment" docVersion="11" lastPageNumber="108" masterDocId="0A68A264FFD0A564572BFFE3E63A475A" masterDocTitle="Revalidation and redescription of Serranus knysnaensis Gilchrist, 1904 (Perciformes: Serranidae) with a new distribution record" masterLastPageNumber="113" masterPageNumber="99" pageNumber="107" updateTime="1699460442245" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8F6A06213884C64C08383CC0C984EA2B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="370C9EDB9FA638FB2DD70FD7DD2C76D6">
<mods:title id="4B2D771BD468B210D2856078FF6E9487">Revalidation and redescription of Serranus knysnaensis Gilchrist, 1904 (Perciformes: Serranidae) with a new distribution record</mods:title>
</mods:titleInfo>
<mods:name id="67195EB9DA3C76739EFC954D5B1BEAED" type="personal">
<mods:role id="92B1891F862D7F097313F0176B77D746">
<mods:roleTerm id="11A612BB128BE6F950E2321F80A1C9E8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="710896623D9FA445945594E7DE61A076">Sithole, Yonela</mods:namePart>
<mods:nameIdentifier id="FA193A8F0731A741CC75E293B45335D2" type="ORCID">0000-0003-4084-6695</mods:nameIdentifier>
<mods:affiliation id="CFBAFF840905E06E4DB5AEC85A9CB80C">Department of Ichthyology and Fisheries Science, Rhodes University, Makhanda, 6140. &amp; National Research Foundation-South African Institute for Aquatic Biodiversity, Makhanda, 6140.</mods:affiliation>
</mods:name>
<mods:name id="A1E360BE168537C15EEB4DEDB2A153DE" type="personal">
<mods:role id="10F58D42484C3370B4D82E2247666C89">
<mods:roleTerm id="90E093A6D83FE7AECA5DA247CD9CE604">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DA63DFF73B6FCC1285C404E8EF72FEE0">Heemstra, Elaine</mods:namePart>
<mods:nameIdentifier id="BEE352C5B6E83E7285BC55FBE4BED65D" type="ORCID">0000-0001-5871-6990</mods:nameIdentifier>
<mods:affiliation id="81A409FB452CCE8562EB1BA934110280">National Research Foundation-South African Institute for Aquatic Biodiversity, Makhanda, 6140. &amp; e. heemstra @ saiab. nrf. ac. za; https: // orcid. org / 0000 - 0001 - 5871 - 6990</mods:affiliation>
<mods:nameIdentifier id="AF5DAD5F020BE8902D345BA41DB88930" type="email">e.heemstra@saiab.nrf.ac.za</mods:nameIdentifier>
</mods:name>
<mods:name id="CB6CB1C97AF818E46E1BF7BC9DDF26A2" type="personal">
<mods:role id="10318F74322EED0203E595B3A73B0AAF">
<mods:roleTerm id="EE4759045B336AA73B517F8233F71DBE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ECA87A0D578B3008A81296970ED26D70">Mwale, Monica</mods:namePart>
<mods:nameIdentifier id="09FBA54BC2AC94115E6E89B2C2A4983B" type="ORCID">0000-0003-2180-8917</mods:nameIdentifier>
<mods:affiliation id="C58723D5EC8E886A5A9FFA74F2B6EE90">National Research Foundation-South African Institute for Aquatic Biodiversity, Makhanda, 6140. &amp; South African National Biodiversity Institute (SANBI), National Zoological Gardens, P. O. Box 754, Pretoria, South Africa 0001. m. mwale @ sanbi. org. za; https: // orcid. org / 000 - 0003 - 2180 - 8917</mods:affiliation>
<mods:nameIdentifier id="8497C7E37A5A928A14D624BF7AA08751" type="email">m.mwale@sanbi.org.za</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1F4CBF138519EC307B08E77918AAB495">text</mods:typeOfResource>
<mods:relatedItem id="FE1E6131F667BEF2CBEAC35811CE1D21" type="host">
<mods:titleInfo id="578EE1EC138979CE18AC81C232DE3DF9">
<mods:title id="CBD77D0B1F7372E48570BAEECE7C3EFD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D2834DFEBC8462C2BA845FCCDBCDD472">
<mods:date id="2C4984E23F26C9F25AD7C15C26295EE7">2021</mods:date>
<mods:detail id="D1A214537282FCA9BE3B4C87922899E7" type="pubDate">
<mods:number id="A4D144B268FB63D0A0625699D8CFF146">2021-10-20</mods:number>
</mods:detail>
<mods:detail id="8478D29ED2247DC4EFAEE3EFBDC10421" type="volume">
<mods:number id="39E15FD8F85B4C79693B4782B1B1F3C7">5057</mods:number>
</mods:detail>
<mods:detail id="249106D7F84A29F639C9D748067BDC26" type="issue">
<mods:number id="72DFD2ADFCEE6F3F6D60B6BB56A93A7B">1</mods:number>
</mods:detail>
<mods:extent id="2D8FB6204AEAF53191D0E5DE863C1652" unit="page">
<mods:start id="5AB35DFBAE3E39D9927D589DCF037826">99</mods:start>
<mods:end id="0260E40A6DB4E05A7E04D7659EE5181D">113</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EDE8F9493F94545E98AACF3FAAD236BE">journal article</mods:classification>
<mods:identifier id="909F9726DB7195A2D8B304E77D504096" type="CLB-Dataset">3899</mods:identifier>
<mods:identifier id="A77FA8F5AD1F9335AC04538EF3D23524" type="DOI">10.11646/zootaxa.5057.1.6</mods:identifier>
<mods:identifier id="C30D67E7DD29094FB0AA3D4B924DC895" type="GBIF-Dataset">3f50ca0f-843f-494f-aff8-defcc449ad44</mods:identifier>
<mods:identifier id="1D6155265B5887FEDC36187950C3DF0F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D43BD6E5D81223FC680512688AA64FA2" type="Zenodo-Dep">5705482</mods:identifier>
<mods:identifier id="DD747E90A1B75669BADB15370456D425" type="ZooBank">4074E652-255A-44E9-B837-5CB26745D06F</mods:identifier>
</mods:mods>
<treatment id="F651DA1CFFD8A56D57BCFA86E45D42E3" ID-DOI="http://doi.org/10.5281/zenodo.5705494" ID-GBIF-Taxon="189313329" ID-Zenodo-Dep="5705494" LSID="urn:lsid:plazi:treatment:F651DA1CFFD8A56D57BCFA86E45D42E3" httpUri="http://treatment.plazi.org/id/F651DA1CFFD8A56D57BCFA86E45D42E3" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107">
<subSubSection id="36E23881FFD8A56C57BCFA86E7A342FE" pageId="8" pageNumber="107" type="nomenclature">
<paragraph id="7E476B0AFFD8A56C57BCFA86E4784225" blockId="8.[151,578,1381,1480]" box="[151,578,1381,1408]" pageId="8" pageNumber="107">
<heading id="250FDC66FFD8A56C57BCFA86E4784225" bold="true" box="[151,578,1381,1408]" fontSize="11" level="1" pageId="8" pageNumber="107" reason="1">
<taxonomicName id="B9F81089FFD8A56C57BCFA86E4784225" ID-CoL="4WWXZ" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,578,1381,1408]" class="Actinopterygii" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="107" phylum="Chordata" rank="species" species="cabrilla">
<emphasis id="4C8CB718FFD8A56C57BCFA86E4784225" bold="true" box="[151,578,1381,1408]" pageId="8" pageNumber="107">
<emphasis id="4C8CB718FFD8A56C57BCFA86E75042DA" bold="true" box="[151,362,1381,1408]" italics="true" pageId="8" pageNumber="107">Serranus cabrilla</emphasis>
(
<bibRefCitation id="1A6916FBFFD8A56C5650FA86E4004225" author="Linnaeus, C." box="[379,570,1381,1407]" pageId="8" pageNumber="107" refId="ref10124" refString="Linnaeus, C. (1758) Systema Naturae. 10 th Edition. Impensis Direct. Laurentii Salvii, Holmiae, ii + 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="7E476B0AFFD8A56C57BCFA69E7A342FE" blockId="8.[151,578,1381,1480]" box="[151,409,1418,1444]" pageId="8" pageNumber="107">English name: Comber</paragraph>
</subSubSection>
<subSubSection id="36E23881FFD8A56C57BCFA4DE79E4292" box="[151,420,1453,1480]" pageId="8" pageNumber="107" type="description">
<paragraph id="7E476B0AFFD8A56C57BCFA4DE79E4292" blockId="8.[151,578,1381,1480]" box="[151,420,1453,1480]" pageId="8" pageNumber="107">
<figureCitation id="E6C3778FFFD8A56C57BCFA4DE7254292" box="[151,287,1454,1480]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="7.[151,250,1609,1633]" captionStartId-1="9.[151,250,1260,1284]" captionTargetBox-0="[163,1423,185,1581]" captionTargetBox-1="[157,1431,633,1237]" captionTargetId-0="figure-17@7.[151,1435,181,1585]" captionTargetId-1="figure-233@9.[157,1431,633,1237]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="FIGURE 3. A maximum likelihood tree based on 92 COI mtDNA sequences (597 bp) of 17 Serranus species based on the General Time Reversible model. All sequences were downloaded from BOLD and GenBank (Table 1). The bottom scale bar represents branch lengths measured in the number of substitutions per site. Study species clusters are also highlighted: Serranus cabrilla (SC), S. knysnaensis (SK) and S. novemcinctus (SN)." captionText-1="FIGURE 4. Serranus cabrilla, 95 mm SL from Gulf of Suez, Egypt, photograph by John E. Randall." figureDoi-0="http://doi.org/10.5281/zenodo.5705492" figureDoi-1="http://doi.org/10.5281/zenodo.5705496" httpUri-0="https://zenodo.org/record/5705492/files/figure.png" httpUri-1="https://zenodo.org/record/5705496/files/figure.png" pageId="8" pageNumber="107">Figures 3–4</figureCitation>
; 
<tableCitation id="337A5EB1FFD8A56C5601FA4DE79E4292" box="[298,420,1453,1480]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStart-4="TABLE 5" captionStartId-0="2.[151,239,440,464]" captionStartId-1="6.[151,239,151,175]" captionStartId-2="8.[151,239,151,175]" captionStartId-3="11.[151,239,699,725]" captionStartId-4="12.[151,239,557,583]" captionTargetBox-1="[166,1407,263,652]" captionTargetBox-4="[165,2019,670,1490]" captionTargetPageId-0="2" captionTargetPageId-1="6" captionTargetPageId-2="8" captionTargetPageId-3="11" captionText-0="TABLE 1. List of mtDNA COI sequences of Serranus knysnaensis, S. cabrilla, S. novemcinctus and other additional Serranus species downloaded from GenBank &amp; BOLD (see accession numbers) with localities. Accession numbers in bold represent tissue samples derived from voucher specimens that were examined morphologically." captionText-1="TABLE 2. Frequency distribution of the number of circumpeduncular scales, the number of lower limb and total gill rakers (both including rudiments) for specimens of Serranus cabrilla (SC), S. knysnaensis (SK), and S. novemcinctus (SN)." captionText-2="TABLE 3. Minimum and maximum proportional measurements (expressed as percentage of Standard length) for Serranus cabrilla, S. knysnaensis and S. novemcinctus." captionText-3="TABLE 4. The genetic distance estimated using the Kimura 2-parameter model from the number of base substitutions per site from averaging over all sequence pairs between species (below diagonal) and standard error estimate(s) are shown above the diagonal. Values were estimated in MEGA7." captionText-4="TABLE 5. Summary statistics reported by the species delimitation plugin for mtDNA COI gene in each putative species using pairwise interspecific genetic distances between Serranus species (distances between S. cabrilla, S. knysnaensis and S. novemcinctus are highlighted in bold). The barcoding gap analysis results for two relative interspecific gap" httpUri-0="http://table.plazi.org/id/2A873B82FFD2A56657BCFE5BE2FD4542" httpUri-1="http://table.plazi.org/id/2A873B82FFD6A56257BCFF74E6E847AD" httpUri-2="http://table.plazi.org/id/2A873B82FFD8A56C57BCFF74E4404789" httpUri-3="http://table.plazi.org/id/2A873B82FFDBA56F57BCFD58E39345A3" httpUri-4="http://table.plazi.org/id/2A873B82FFDCA56857BCFDCEE50145D5" pageId="8" pageNumber="107" tableUuid-0="2A873B82FFD2A56657BCFE5BE2FD4542" tableUuid-1="2A873B82FFD6A56257BCFF74E6E847AD" tableUuid-2="2A873B82FFD8A56C57BCFF74E4404789" tableUuid-3="2A873B82FFDBA56F57BCFD58E39345A3" tableUuid-4="2A873B82FFDCA56857BCFDCEE50145D5">Tables 1–5</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="36E23881FFD8A56C57BCFA0EE4DC41C4" pageId="8" pageNumber="107" type="reference_group">
<paragraph id="7E476B0AFFD8A56C57BCFA0EE59A415F" blockId="8.[151,1437,1517,1694]" box="[151,928,1517,1541]" pageId="8" pageNumber="107">
<treatmentCitationGroup id="5EE84C24FFD8A56C57BCFA0EE59A415F" box="[151,928,1517,1541]" pageId="8" pageNumber="107">
<treatmentCitation id="FF594D1BFFD8A56C57BCFA0EE43B415F" ID-DOI="http://doi.org/10.5281/zenodo.3917826" ID-Zenodo-Dep="3917826" LSID="urn:lsid:zoobank.org:act:90A4514A-3F79-467E-9875-AAD956D03EC3" author="Linnaeus, C." box="[151,513,1517,1541]" httpUri="http://treatment.plazi.org/id/7D0C6CCCBA4669CF607492FB71F84EB0" page="294" pageId="8" pageNumber="107" year="1758">
<taxonomicName id="B9F81089FFD8A56C57BCFA0EE43B415F" authority=": Linnaeus 1758: 294" authorityName=": Linnaeus" authorityPageNumber="294" authorityYear="1758" box="[151,513,1517,1541]" class="Actinopterygii" family="Percidae" genus="Perca" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="107" phylum="Chordata" rank="species" species="cabrilla">
<emphasis id="4C8CB718FFD8A56C57BCFA0EE708415F" box="[151,306,1517,1541]" italics="true" pageId="8" pageNumber="107">Perca cabrilla:</emphasis>
<bibRefCitation id="1A6916FBFFD8A56C5613FA0EE43B415F" author="Linnaeus, C." box="[312,513,1517,1541]" pageId="8" pageNumber="107" refId="ref10124" refString="Linnaeus, C. (1758) Systema Naturae. 10 th Edition. Impensis Direct. Laurentii Salvii, Holmiae, ii + 824 pp." type="book" year="1758">Linnaeus 1758: 294</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(no locality given [Mediterranean Sea]).
</treatmentCitationGroup>
</paragraph>
<paragraph id="7E476B0AFFD8A56C57BCF9EFE454417E" blockId="8.[151,1437,1517,1694]" box="[151,622,1547,1572]" pageId="8" pageNumber="107">
<treatmentCitationGroup id="5EE84C24FFD8A56C57BCF9EFE454417E" box="[151,622,1547,1572]" pageId="8" pageNumber="107">
<treatmentCitation id="FF594D1BFFD8A56C57BCF9EFE454417E" author="Fowler, H. W." box="[151,622,1547,1572]" page="1291" pageId="8" pageNumber="107" year="1936">
<taxonomicName id="B9F81089FFD8A56C57BCF9EFE454417E" ID-CoL="4CP2P" authority=": Fowler 1936: 1291" authorityName="Fowler" authorityPageNumber="1291" authorityYear="1936" box="[151,622,1547,1572]" class="Actinopterygii" family="Serranidae" genus="Paracentropristis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="107" phylum="Chordata" rank="species" species="cabrilla">
<emphasis id="4C8CB718FFD8A56C57BCF9EFE7A54179" box="[151,415,1547,1571]" italics="true" pageId="8" pageNumber="107">Paracentropristis cabrilla</emphasis>
: 
<bibRefCitation id="1A6916FBFFD8A56C5686F9E8E454417E" author="Fowler, H. W." box="[429,622,1547,1572]" pageId="8" pageNumber="107" pagination="607 - 1493" refId="ref9379" refString="Fowler, H. W. (1936) The marine fishes of West Africa based on the collection of the American Museum Congo expedition, 1909 - 1915. Part II. Bulletin of the American Museum of Natural History, 70, 607 - 1493." type="journal article" year="1936">Fowler 1936: 1291</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="7E476B0AFFD8A56C57BCF9C9E4DC41C4" blockId="8.[151,1437,1517,1694]" pageId="8" pageNumber="107">
<treatmentCitationGroup id="5EE84C24FFD8A56C57BCF9C9E4DC41C4" pageId="8" pageNumber="107">
<treatmentCitation id="FF594D1BFFD8A56C57BCF9C9E43A4118" author="Poll, M." box="[151,512,1578,1602]" page="72" pageId="8" pageNumber="107" year="1954">
<taxonomicName id="B9F81089FFD8A56C57BCF9C9E43A4118" ID-CoL="4WWXZ" authority=": Poll 1954: 69 - 72" authorityName=": Poll" authorityPageNumber="69 - 72" authorityYear="1954" box="[151,512,1578,1602]" class="Actinopterygii" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="107" phylum="Chordata" rank="species" species="cabrilla">
<emphasis id="4C8CB718FFD8A56C57BCF9C9E76B4118" box="[151,337,1578,1602]" italics="true" pageId="8" pageNumber="107">Serranus cabrilla:</emphasis>
<bibRefCitation id="1A6916FBFFD8A56C567EF9C9E43A4118" author="Poll, M." box="[341,512,1578,1602]" pageId="8" pageNumber="107" pagination="1 - 390" refId="ref10256" refString="Poll, M. (1954) Poissons IV. Teleosteens Acanthopterygiens (premiere partie). Resultats Scientifiques. Expedition Oceanographique Belge dans les Eaux Cotieres Africaines de l'Atlantique Sud (1948 - 1949), Bruxelles, 4, 1 - 390." type="journal article" year="1954">Poll 1954:69−72</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, Fig. 19 (
<collectingCountry id="06EF2B9AFFD8A56C5577F9C9E49B4118" box="[604,673,1578,1602]" name="Gabon" pageId="8" pageNumber="107">Gabon</collectingCountry>
to s. 
<collectingCountry id="06EF2B9AFFD8A56C55F8F9C9E5184118" box="[723,802,1578,1602]" name="Angola" pageId="8" pageNumber="107">Angola</collectingCountry>
); 
<treatmentCitation id="FF594D1BFFD8A56C5419F9C9E2BD4118" author="Robins, C. R. &amp; Starck II, W. A." box="[818,1159,1578,1602]" page="261" pageId="8" pageNumber="107" year="1961">
<bibRefCitation id="1A6916FBFFD8A56C5419F9C9E2BD4118" author="Robins, C. R. &amp; Starck II, W. A." box="[818,1159,1578,1602]" pageId="8" pageNumber="107" pagination="259 - 314" refId="ref10425" refString="Robins, C. R. &amp;. Starck II, W. A. (1961) Materials for a revision of Serranus and related fish genera. Proceedings of the Academy of Natural Sciences of Philadelphia, 113, 259 - 314." type="journal article" year="1961">Robins and Starck 1961: 261−262</bibRefCitation>
</treatmentCitation>
(figs. 1a, 7c, 8a, 8c; + 
<collectingCountry id="06EF2B9AFFD8A56C5249F9C9E3A74118" box="[1378,1437,1578,1602]" name="Spain" pageId="8" pageNumber="107">Spain</collectingCountry>
and 
<collectingCountry id="06EF2B9AFFD8A56C57DEF9AAE712413B" box="[245,296,1609,1633]" name="Italy" pageId="8" pageNumber="107">Italy</collectingCountry>
); 
<treatmentCitation id="FF594D1BFFD8A56C5610F9AAE404413B" author="Smith, J. L. B. &amp; Smith, M. M." box="[315,574,1609,1633]" page="88" pageId="8" pageNumber="107" year="1966">
<bibRefCitation id="1A6916FBFFD8A56C5610F9AAE404413B" author="Smith, J. L. B. &amp; Smith, M. M." box="[315,574,1609,1633]" pageId="8" pageNumber="107" refId="ref10497" refString="Smith, J. L. B. &amp; Smith, M. M. (1966) Fishes of the Tsitsikama Coastal National Park. Swan Press Ltd, Johannesburg, 161 pp." type="book" year="1966">Smith &amp; Smith 1966: 88</bibRefCitation>
</treatmentCitation>
(in part, not painting; Atlantic species from Mediterranean southwards, colonising 
<collectingCountry id="06EF2B9AFFD8A56C57ECF984E77D4125" box="[199,327,1639,1663]" name="South Africa" pageId="8" pageNumber="107">South Africa</collectingCountry>
to Natal); 
<treatmentCitation id="FF594D1BFFD8A56C569BF98BE4D24125" author="Heemstra, P. C. &amp; Randall, J. E." box="[432,744,1639,1664]" page="536" pageId="8" pageNumber="107" year="1986">
<bibRefCitation id="1A6916FBFFD8A56C569BF98BE4D24125" author="Heemstra, P. C. &amp; Randall, J. E." box="[432,744,1639,1664]" pageId="8" pageNumber="107" pagination="509 - 537" refId="ref9648" refString="Heemstra, P. C. &amp; Randall, J. E. (1986) Family Serranidae. In: Smith, M. M. &amp; Heemstra, P. C. (eds), Smiths' Sea Fishes. MacMillan, Johannesburg, pp. 509 - 537." type="book chapter" year="1986">Heemstra &amp; Randall 1986: 536</bibRefCitation>
</treatmentCitation>
(in part, not Figure or Plate; Mediterranean, eastern Atlantic, English Channel to 
<collectingCountry id="06EF2B9AFFD8A56C5617F965E78541C4" box="[316,447,1670,1694]" name="South Africa" pageId="8" pageNumber="107">South Africa</collectingCountry>
, and round coast to Durban).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="36E23881FFD8A56C57BCF924E3A64073" pageId="8" pageNumber="107" type="diagnosis">
<paragraph id="7E476B0AFFD8A56C57BCF924E3A64073" blockId="8.[151,1437,1735,2013]" pageId="8" pageNumber="107">
<emphasis id="4C8CB718FFD8A56C57BCF924E72841BB" bold="true" box="[151,274,1735,1761]" pageId="8" pageNumber="107">Diagnosis.</emphasis>
Dorsal-fin X, 13–14; pectoral-fin rays 14–16; gill rakers 7–8+14–16 (22–24 total); circumpeduncular scales 34–38; lateral-line scales 70–77; scales from dorsal fin to lateral line 9–12; scales from the anal-fin origin to lateral line 22–25. Caudal fin emarginate to slightly forked; cheek, nape, opercle, pectoral-fin base and chest scaly.
</paragraph>
</subSubSection>
<subSubSection id="36E23881FFD8A56D57ECF8D0E45D42E3" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107" type="description">
<paragraph id="7E476B0AFFD8A56D57ECF8D0E3B9478F" blockId="8.[151,1437,1735,2013]" lastBlockId="9.[151,1437,151,610]" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107">
<emphasis id="4C8CB718FFD8A56C57ECF8D0E7634017" bold="true" box="[199,345,1843,1869]" pageId="8" pageNumber="107">Description.</emphasis>
Body elongate and slightly compressed; body depth less than head length, about 
<quantity id="B900C6EFFFD8A56C53D2F8D0E3564017" box="[1273,1388,1843,1870]" metricMagnitude="-1" metricUnit="m" metricValue="1.0921999999999998" metricValueMax="1.3208" metricValueMin="0.8635999999999999" pageId="8" pageNumber="107" unit="in" value="4.3" valueMax="5.2" valueMin="3.4">3.4–5.2 in</quantity>
SL; head length 
<quantity id="B900C6EFFFD8A56C560AF8B4E7A9402B" box="[289,403,1879,1905]" metricMagnitude="-2" metricUnit="m" metricValue="6.858" metricValueMax="7.111999999999999" metricValueMin="6.604" pageId="8" pageNumber="107" unit="in" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8 in</quantity>
SL. Maxilla extending to below middle of the eye; lower jaw projecting beyond upper jaw; opercle with three flat spines, upper two spines well developed. Snout length greater than eye diameter, snout length 
<quantity id="B900C6EFFFD8A56C57BCF87CE72C40E3" box="[151,278,1951,1977]" metricMagnitude="-1" metricUnit="m" metricValue="2.5654000000000003" metricValueMax="2.8956000000000004" metricValueMin="2.2352000000000003" pageId="8" pageNumber="107" unit="in" value="10.1" valueMax="11.4" valueMin="8.8">8.8–11.4 in</quantity>
SL, eye diameter 10.0– 
<quantity id="B900C6EFFFD8A56C550FF87CE44B40E3" box="[548,625,1951,1977]" metricMagnitude="-1" metricUnit="m" metricValue="3.5306" pageId="8" pageNumber="107" unit="in" value="13.9">13.9 in</quantity>
SL. Snout, interorbital and maxilla naked. Small scales present at bases of dorsal, anal and pectoral fins; caudal and pelvic fins with or without some small scales basally. Preopercle margin coarsely serrate with developed spines near angle. Premaxillary teeth in several irregular rows, outer teeth enlarged, with large canines anteriorly. Vomer more or less V-shaped, with small teeth. Palatines with patch of small teeth.
</paragraph>
<paragraph id="7E476B0AFFD9A56D57ECFF3CE487461B" blockId="9.[151,1437,151,610]" pageId="9" pageNumber="108">The first two dorsal-fin spines shortest, the sixth spine usually longest. No distinct notch between dorsal spines and soft rays but soft rays slightly elevated. Pectoral-fin origin slightly behind pelvic-fin origin, distal tip of pectoral fin reaching anus, slightly behind anal-fin origin.</paragraph>
<paragraph id="7E476B0AFFD9A56D57ECFEA8E4784538" blockId="9.[151,1437,151,610]" pageId="9" pageNumber="108">
<emphasis id="4C8CB718FFD9A56D57ECFEA8E787463E" bold="true" box="[199,445,330,357]" pageId="9" pageNumber="108">
Fresh Color (
<figureCitation id="E6C3778FFFD9A56D564CFEA9E795463E" box="[359,431,330,356]" captionStart="FIGURE 4" captionStartId="9.[151,250,1260,1284]" captionTargetBox="[157,1431,633,1237]" captionTargetId="figure-233@9.[157,1431,633,1237]" captionTargetPageId="9" captionText="FIGURE 4. Serranus cabrilla, 95 mm SL from Gulf of Suez, Egypt, photograph by John E. Randall." figureDoi="http://doi.org/10.5281/zenodo.5705496" httpUri="https://zenodo.org/record/5705496/files/figure.png" pageId="9" pageNumber="108">Fig. 4</figureCitation>
).
</emphasis>
Body light brown to reddish with eight or nine dark vertical bars dorsolaterally; two or three white horizontal bands from head to tail; ventral part of head and belly mostly white; head with two or three orange-brownish stripes below and behind eye; caudal and soft dorsal fins with small bluish dots; tips of caudal fin reddish or dark brown; pelvic and pectoral fins pale. Juveniles with white midlateral stripe bordered by two thick black stripes, the upper stripe running from tip of snout through middle of eye to upper margin of operculum onto trunk above midlateral line to upper half of caudal peduncle, the lower stripe from base of pectoral fin to lower half of caudal peduncle; the dark stripes often partially broken into dark blotches (
<bibRefCitation id="1A6916FBFFD9A56D532DFDC0E3284567" author="Iwamoto, T. &amp; Wirtz, P." box="[1030,1298,547,573]" pageId="9" pageNumber="108" pagination="1 - 39" refId="ref9863" refString="Iwamoto, T. &amp; Wirtz, P. (2018) A synopsis of the Eastern and Central Atlantic combers of the genus Serranus (Teleostei: Perciformes: Serranidae). Proceedings of the California Academy of Sciences, 65, 1 - 39." type="journal article" year="2018">Iwamoto &amp; Wirtz 2018</bibRefCitation>
; no juvenile specimens available for examination).
</paragraph>
<caption id="2A873B82FFD9A56D57BCFB0FE2A3425E" ID-DOI="http://doi.org/10.5281/zenodo.5705496" ID-Zenodo-Dep="5705496" box="[151,1177,1260,1285]" httpUri="https://zenodo.org/record/5705496/files/figure.png" pageId="9" pageNumber="108" startId="9.[151,250,1260,1284]" targetBox="[157,1431,633,1237]" targetPageId="9">
<paragraph id="7E476B0AFFD9A56D57BCFB0FE2A3425E" blockId="9.[151,1177,1260,1285]" box="[151,1177,1260,1285]" pageId="9" pageNumber="108">
<emphasis id="4C8CB718FFD9A56D57BCFB0FE729425E" bold="true" box="[151,275,1260,1284]" pageId="9" pageNumber="108">FIGURE 4.</emphasis>
<taxonomicName id="B9F81089FFD9A56D5632FB0EE7F1425E" authorityName=": Poll" authorityYear="1954" box="[281,459,1260,1284]" class="Actinopterygii" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="108" phylum="Chordata" rank="species" species="cabrilla">
<emphasis id="4C8CB718FFD9A56D5632FB0EE7F1425E" box="[281,459,1260,1284]" italics="true" pageId="9" pageNumber="108">Serranus cabrilla</emphasis>
</taxonomicName>
, 95 mm SL from Gulf of Suez, Egypt, photograph by John E. Randall.
</paragraph>
</caption>
<paragraph id="7E476B0AFFD9A56D57ECFAD1E5A64217" blockId="9.[151,1436,1330,1465]" box="[199,924,1330,1357]" pageId="9" pageNumber="108">
<emphasis id="4C8CB718FFD9A56D57ECFAD1E7F44216" bold="true" box="[199,462,1330,1356]" pageId="9" pageNumber="108">Color in preservation.</emphasis>
Specimens lose all color in preservation.
</paragraph>
<paragraph id="7E476B0AFFD9A56D57ECFAB5E45D42E3" blockId="9.[151,1436,1330,1465]" pageId="9" pageNumber="108">
<emphasis id="4C8CB718FFD9A56D57ECFAB5E75B422A" bold="true" box="[199,353,1366,1392]" pageId="9" pageNumber="108">Distribution.</emphasis>
From the British Isles to 
<collectingCountry id="06EF2B9AFFD9A56D55B8FAB4E4D2422B" box="[659,744,1367,1393]" name="Angola" pageId="9" pageNumber="108">Angola</collectingCountry>
, including Azores, Madeira, Canary Islands, 
<collectingCountry id="06EF2B9AFFD9A56D53DDFAB4E3A6422B" box="[1270,1436,1367,1393]" name="Sao Tome and Principe" pageId="9" pageNumber="108">São Tomé and</collectingCountry>
Príncipe, and 
<collectingCountry id="06EF2B9AFFD9A56D5613FA98E78142CF" box="[312,443,1403,1429]" name="Cape Verde" pageId="9" pageNumber="108">Cape Verde</collectingCountry>
Islands, also through Mediterranean and Black Sea, and in the Red Sea (
<bibRefCitation id="1A6916FBFFD9A56D5224FA98E3A642CF" author="Heemstra, P. C. &amp; Anderson, W. D." box="[1295,1436,1403,1429]" pageId="9" pageNumber="108" pagination="2365 - 2413" refId="ref9517" refString="Heemstra, P. C. &amp; Anderson, W. D. (2016) Serranidae. Groupers (seabasses, hinds, creolefish, combers, anthiines, soapfish). In: Carpenter, K. E. &amp; De Angelis, N. (Eds.), The Living Marine Resources of the Eastern Central Atlantic. 4. Bony Fishes. Part 2 (Perciformes to Tetraodontiformes) and sea turtles. FAO species Identification Guide for Fishery Purposes. FAO, Rome, pp. 2365 - 2413." type="book chapter" year="2016">Heemstra &amp;</bibRefCitation>
Anderson 2016; 
<bibRefCitation id="1A6916FBFFD9A56D5679FA7CE46142E3" author="Iwamoto, T. &amp; Wirtz, P." box="[338,603,1439,1465]" pageId="9" pageNumber="108" pagination="1 - 39" refId="ref9863" refString="Iwamoto, T. &amp; Wirtz, P. (2018) A synopsis of the Eastern and Central Atlantic combers of the genus Serranus (Teleostei: Perciformes: Serranidae). Proceedings of the California Academy of Sciences, 65, 1 - 39." type="journal article" year="2018">Iwamoto &amp; Wirtz 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>