treatments-xml/data/03/AA/10/03AA10085657FFE7C1D8FC8FFC50FD39.xml

<document id="981E3A9E2BA128FCE9FFAB993AECEEC1" ID-DOI="10.11646/zootaxa.3664.2.1" ID-GBIF-Dataset="a50b71f5-48a6-464f-aea1-f29bbc8ee322" ID-ISSN="1175-5326" ID-Zenodo-Dep="220503" ID-ZooBank="105D178E-6DBF-401D-85BB-CDB041B67E91" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460615203542" checkinUser="plazi" docAuthor="Shi, Hongliang, Sciaky, Riccardo, Liang, Hongbin &amp; Zhou, Hongzhang" docDate="2013" docId="03AA10085657FFE7C1D8FC8FFC50FD39" docLanguage="en" docName="zt03664p135.pdf" docOrigin="Zootaxa 3664 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Wraseiellus) diversus Fairmaire 1886" docType="treatment" docVersion="9" lastPageNumber="115" masterDocId="FF936870565BFFE9C14FFF95FFA9FF9E" masterDocTitle="A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions" masterLastPageNumber="135" masterPageNumber="101" pageNumber="113" updateTime="1698330049886" updateUser="plazi">
<mods:mods id="8788EC29F0AF51E1EFD40566D694583B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0DD4825BF2E4F2A3BE2608F46958D0E7">
<mods:title id="9281648B5F44C0A95509F3767CE580D4">A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions</mods:title>
</mods:titleInfo>
<mods:name id="9BF3D94DEAA38022CC5CA39CD85B5599" type="personal">
<mods:role id="6B30AEBD29AAB79A91DC48E57467FFF9">
<mods:roleTerm id="E359923CF060C391E2786E6D9945E7E1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0D148EB697E0D470767FF7580603A55C">Shi, Hongliang</mods:namePart>
</mods:name>
<mods:name id="AF2C38AF742A8B86D5BC7752DF8A1B31" type="personal">
<mods:role id="8711E7C857321DE1ADB06DDEAD2D662B">
<mods:roleTerm id="A50AFD6FB99C1E3ED1604F5AD7DC7257">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="512FA0E304CE51553F79F2EEF8352DB3">Sciaky, Riccardo</mods:namePart>
</mods:name>
<mods:name id="A79388CF85453A33A8BEF0DDBEFEFE0D" type="personal">
<mods:role id="CB28631ABF495E8B75FBB8E56ADC5622">
<mods:roleTerm id="28422BB43B879D0A3A00AC9F8D46042C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C0CE4B7564155225D14C561258081A72">Liang, Hongbin</mods:namePart>
</mods:name>
<mods:name id="E59BA866B5EC71F32E74E6F4CBD4208C" type="personal">
<mods:role id="1145E8FD5DEE3009F7738938E7BB9272">
<mods:roleTerm id="D2AF0C2253EB4860A568ADEF6884907E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D60E201329437EBCCD6F73A0BE3C09F9">Zhou, Hongzhang</mods:namePart>
</mods:name>
<mods:typeOfResource id="24F5526F0C26742C969E71909D8699EC">text</mods:typeOfResource>
<mods:relatedItem id="A6E9CC08F381848A2653B1AAF96BC436" type="host">
<mods:titleInfo id="C56ECB3B8317F81AF95566F941BC3F26">
<mods:title id="D78937F9FA6377A6B57CFAE479547C2C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A22D7CB76C345BC4FCE35FA316B9B046">
<mods:date id="CC1DC7F0902D221F30D2329EA8EF334D">2013</mods:date>
<mods:detail id="9D21F9FBBA880DE4DFE61D9CE9842B41" type="volume">
<mods:number id="C1AC6B4BE9DEBAC0D027120E3C191F38">3664</mods:number>
</mods:detail>
<mods:detail id="1543676E65BD3405330A6DC69B043B96" type="issue">
<mods:number id="A74AB15E1C55F3D996A0D4EF97BBC7A0">2</mods:number>
</mods:detail>
<mods:extent id="470CA5EDC898089C945C911A65288AA8" unit="page">
<mods:start id="E5179967ED6B80DC133A91A6D929A553">101</mods:start>
<mods:end id="0146DDBCDDF3287B3038BDEEF1BDB1B7">135</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2E1A76BB5DFF93F0855E25ECB1B7D364">journal article</mods:classification>
<mods:identifier id="6B2FE23AE11F70B694CBC24617F9BE7C" type="DOI">10.11646/zootaxa.3664.2.1</mods:identifier>
<mods:identifier id="B9F1823911F7145E928CE0FC8B7DA37F" type="GBIF-Dataset">a50b71f5-48a6-464f-aea1-f29bbc8ee322</mods:identifier>
<mods:identifier id="D8EC275216C34DDAA525101D55C05B6D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="834BF206FB388EE3FA195443AC512659" type="Zenodo-Dep">220503</mods:identifier>
<mods:identifier id="EB5C41BDC3279D9DB8DEBCBC5992CCA1" type="ZooBank">105D178E-6DBF-401D-85BB-CDB041B67E91</mods:identifier>
</mods:mods>
<treatment id="03AA10085657FFE7C1D8FC8FFC50FD39" ID-DOI="http://doi.org/10.5281/zenodo.6154511" ID-GBIF-Taxon="119568518" ID-Zenodo-Dep="6154511" LSID="urn:lsid:plazi:treatment:03AA10085657FFE7C1D8FC8FFC50FD39" httpUri="http://treatment.plazi.org/id/03AA10085657FFE7C1D8FC8FFC50FD39" lastPageId="14" lastPageNumber="115" pageId="12" pageNumber="113">
<subSubSection id="C319F2955657FFE5C1D8FC8FFBEBFC46" pageId="12" pageNumber="113" type="nomenclature">
<paragraph id="8BBCA11E5657FFE5C1D8FC8FFC89FCAA" blockId="12.[151,988,794,853]" box="[151,800,794,820]" pageId="12" pageNumber="113">
<heading id="D0F416725657FFE5C1D8FC8FFC89FCAA" bold="true" box="[151,800,794,820]" fontSize="11" level="1" pageId="12" pageNumber="113" reason="1">
<taxonomicName id="4C03DA9D5657FFE5C1D8FC8FFC89FCAA" ID-CoL="4Q4HN" authority="Fairmaire, 1886" authorityName="Fairmaire" authorityYear="1886" box="[151,800,794,820]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="diversus" subGenus="Wraseiellus">
<emphasis id="B9777D0C5657FFE5C1D8FC8FFC89FCAA" bold="true" box="[151,800,794,820]" pageId="12" pageNumber="113">
<emphasis id="B9777D0C5657FFE5C1D8FC8FFE84FCAA" bold="true" box="[151,301,794,820]" italics="true" pageId="12" pageNumber="113">Pterostichus</emphasis>
(
<emphasis id="B9777D0C5657FFE5C072FC8FFE61FCAA" bold="true" box="[317,456,794,820]" italics="true" pageId="12" pageNumber="113">Wraseiellus</emphasis>
) 
<emphasis id="B9777D0C5657FFE5C095FC8FFD94FCAA" bold="true" box="[474,573,794,820]" italics="true" pageId="12" pageNumber="113">diversus</emphasis>
(Fairmaire, 1886)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBCA11E5657FFE5C1D8FCA8FC75FCCB" blockId="12.[151,988,794,853]" box="[151,988,829,853]" pageId="12" pageNumber="113">
(
<figureCitation id="1338BD9B5657FFE5C1D1FCA8FE9CFCCB" box="[158,309,829,853]" captionStart="FIGURES 7 – 12" captionStartId="22.[151,264,1765,1788]" captionTargetBox="[167,1429,198,1733]" captionTargetId="figure@22.[151,1436,183,1744]" captionTargetPageId="22" captionText="FIGURES 7 – 12. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 7, Pt. crassiapex n. sp. holotype; 8, Pt. near crassiapex n. sp., a female from Liziping; 9, Pt. diversus (Fairmaire), a female syntype; 10, Pt. diversus (Fairmaire), a male from Yulongshan; 11, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 12, Pt. stictopleurus cangshanensis n. ssp., holotype." httpUri="https://zenodo.org/record/220505/files/figure.png" pageId="12" pageNumber="113">Figures 9, 10</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C00EFCA8FEC9FCCB" box="[321,352,829,853]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="12" pageNumber="113">24</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C024FCA8FE1AFCCB" box="[363,435,829,853]" captionStart="FIGURES 29 – 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 – 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="12" pageNumber="113">33, 34</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C0F0FCA8FDA1FCCB" box="[447,520,829,853]" captionStart="FIGURES 41 – 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 – 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 – 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 – 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="12" pageNumber="113">41, 42</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C35CFCA8FD98FCCB" box="[531,561,829,853]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="12" pageNumber="113">61</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C372FCA8FD73FCCB" box="[573,730,829,853]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="12" pageNumber="113">74, 75, 76, 77</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C3AAFCA8FCAAFCCB" box="[741,771,829,853]" captionStart="FIGURES 85 – 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 – 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="12" pageNumber="113">87</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C240FCA8FC84FCCB" box="[783,813,829,853]" captionStart="FIGURES 90 – 93" captionStartId="31.[151,264,1640,1663]" captionTargetBox="[161,1418,216,1605]" captionTargetId="figure@31.[151,1436,193,1619]" captionTargetPageId="31" captionText="FIGURES 90 – 93. Left stylomere of ovipositor of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, ventral view; B, dorsal view: 90, Pt. diversus (Fairmaire), a female from Yulongshan; 91, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 92, Pt. pseudodiversus n. sp., a paratype from Luguhu. 93, Pt. andrewesi Jedlička, a female from Jinfoshan. Abbreviations: es, ensiform setae; ns, nematiform setae; s 1, stylomere 1; s 2, stylomere 2." httpUri="https://zenodo.org/record/220514/files/figure.png" pageId="12" pageNumber="113">90</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C276FCA8FCCCFCCB" box="[825,869,829,853]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="12" pageNumber="113">102</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C23EFCA8FC7DFCCB" box="[881,980,829,853]" captionStart="FIGURES 107 – 109" captionStartId="32.[151,264,1625,1648]" captionText="FIGURES 107 – 109. Elytral microsculpture of Pterostichus (Wraseiellus) spp., sub-basal area of left elytron, interval 3 – 4: scale bar = 0.2 mm; 107, Pt. crassiapex n. sp., male (transverse); 108, Pt. diversus (Fairmaire), male (isodiametric, normal); 109, Pt. diversus (Fairmaire), female (isodiametric, extraordinary strong)." pageId="12" pageNumber="113">108, 109</figureCitation>
)
</paragraph>
<paragraph id="8BBCA11E5657FFE5C1D8FC10FBEBFC46" blockId="12.[151,1437,901,984]" pageId="12" pageNumber="113">
Fairmaire 1886: 311 (Original: 
<taxonomicName id="4C03DA9D5657FFE5C092FC13FD93FC05" box="[477,570,902,923]" class="Insecta" family="Carabidae" genus="Omaseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C092FC13FD93FC05" box="[477,570,902,923]" italics="true" pageId="12" pageNumber="113">Omaseus</emphasis>
</taxonomicName>
; a 
<typeStatus id="54B81FBC5657FFE5C312FC13FD05FC05" box="[605,684,902,923]" pageId="12" pageNumber="113" type="syntype">syntype</typeStatus>
deposited in ZRAS); Tschitschérine 1897: 297 (
<taxonomicName id="4C03DA9D5657FFE5C5EAFC13FB5CFC05" box="[1189,1269,902,923]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C5EAFC13FB5CFC05" box="[1189,1269,902,923]" italics="true" pageId="12" pageNumber="113">Feronia</emphasis>
</taxonomicName>
(
<taxonomicName id="4C03DA9D5657FFE5C449FC10FA2AFC05" box="[1286,1411,901,923]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C449FC10FA2AFC05" box="[1286,1411,901,923]" italics="true" pageId="12" pageNumber="113">Pterostichus</emphasis>
</taxonomicName>
)); Jedlička 1962: 234 (
<taxonomicName id="4C03DA9D5657FFE5C0D7FC31FDBFFC24" box="[408,534,932,954]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C0D7FC31FDBFFC24" box="[408,534,932,954]" italics="true" pageId="12" pageNumber="113">Pterostichus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C03DA9D5657FFE5C369FC31FD40FC24" box="[550,745,932,954]" class="Insecta" family="Carabidae" genus="Morphohaptoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C369FC31FD40FC24" box="[550,745,932,954]" italics="true" pageId="12" pageNumber="113">Morphohaptoderus</emphasis>
</taxonomicName>
)); Sciaky 1994: 4 (
<taxonomicName id="4C03DA9D5657FFE5C2FCFC31FB98FC24" box="[947,1073,932,954]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C2FCFC31FB98FC24" box="[947,1073,932,954]" italics="true" pageId="12" pageNumber="113">Pterostichus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C03DA9D5657FFE5C50EFC31FB72FC24" box="[1089,1243,932,954]" class="Insecta" family="Carabidae" genus="Neohaptoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C50EFC31FB72FC24" box="[1089,1243,932,954]" italics="true" pageId="12" pageNumber="113">Neohaptoderus</emphasis>
</taxonomicName>
)); Bousquet 2003: 510 (
<taxonomicName id="4C03DA9D5657FFE5C1B3FC57FED3FC46" box="[252,378,962,984]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C1B3FC57FED3FC46" box="[252,378,962,984]" italics="true" pageId="12" pageNumber="113">Pterostichus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C03DA9D5657FFE5C0C7FC57FD8BFC46" box="[392,546,962,984]" class="Insecta" family="Carabidae" genus="Neohaptoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C0C7FC57FD8BFC46" box="[392,546,962,984]" italics="true" pageId="12" pageNumber="113">Neohaptoderus</emphasis>
</taxonomicName>
)); Lorenz 2005: 278 (
<taxonomicName id="4C03DA9D5657FFE5C24BFC57FC28FC46" box="[772,897,962,984]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C24BFC57FC28FC46" box="[772,897,962,984]" italics="true" pageId="12" pageNumber="113">Pterostichus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C03DA9D5657FFE5C2DFFC57FB82FC46" box="[912,1067,962,984]" class="Insecta" family="Carabidae" genus="Neohaptoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5657FFE5C2DFFC57FB82FC46" box="[912,1067,962,984]" italics="true" pageId="12" pageNumber="113">Neohaptoderus</emphasis>
</taxonomicName>
)).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955657FFE4C1D8FB90FAB8FDD9" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113" type="materials_examined">
<paragraph id="8BBCA11E5657FFE5C1D8FB90FCE3FB17" blockId="12.[151,1436,1029,2036]" pageId="12" pageNumber="113">
<emphasis id="B9777D0C5657FFE5C1D8FB90FE87FB80" bold="true" box="[151,302,1029,1054]" pageId="12" pageNumber="113">
<typeStatus id="54B81FBC5657FFE5C1D8FB90FF79FB83" box="[151,208,1029,1053]" pageId="12" pageNumber="113">Type</typeStatus>
locality
</emphasis>
. Yunnan. The detailed 
<typeStatus id="54B81FBC5657FFE5C37BFB93FDCDFB80" box="[564,612,1030,1054]" pageId="12" pageNumber="113">type</typeStatus>
locality of this species was neither indicated in the original literature nor in the labels of the examined 
<typeStatus id="54B81FBC5657FFE5C090FBBFFD93FBDC" box="[479,570,1066,1090]" pageId="12" pageNumber="113" type="syntype">syntype</typeStatus>
. It was indicated in the preface of original literature that species from Yunnan in this paper were collected by Largeteau and Chasle (Fairmaire, 1886: 303). However, there is no way to infer the exactly 
<typeStatus id="54B81FBC5657FFE5C1A0FBE7FEB6FB14" box="[239,287,1138,1162]" pageId="12" pageNumber="113">type</typeStatus>
locality of this species from their journey record.
</paragraph>
<paragraph id="8BBCA11E5657FFE5C188FB00FDBEFA1B" blockId="12.[151,1436,1029,2036]" pageId="12" pageNumber="113">
There are two other 
<taxonomicName id="4C03DA9D5657FFE5C087FB00FDFCFB33" box="[456,597,1173,1197]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
species (
<taxonomicName id="4C03DA9D5657FFE5C382FB00FC73FB33" box="[717,986,1173,1197]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="simillimus">
<emphasis id="B9777D0C5657FFE5C382FB00FC73FB33" box="[717,986,1173,1197]" italics="true" pageId="12" pageNumber="113">Pterostichus simillimus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C03DA9D5657FFE5C2BFFB00FB4EFB33" box="[1008,1255,1173,1197]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="curtatus">
<emphasis id="B9777D0C5657FFE5C2BFFB00FB4EFB33" box="[1008,1255,1173,1197]" italics="true" pageId="12" pageNumber="113">Pterostichus curtatus</emphasis>
</taxonomicName>
) from Yunnan described in the same paper together with 
<taxonomicName id="4C03DA9D5657FFE5C3CEFB2FFC52FB4F" authority="Fairmaire. We" authorityName="Fairmaire. We" box="[641,1019,1209,1233]" class="Insecta" family="Carabidae" genus="Omaseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5657FFE5C3CEFB2FFCE7FB4F" box="[641,846,1209,1233]" italics="true" pageId="12" pageNumber="113">Omaseus diversus</emphasis>
Fairmaire. We
</taxonomicName>
have examined more complete 
<typeStatus id="54B81FBC5657FFE5C424FB2FFA32FB4C" box="[1387,1435,1210,1234]" pageId="12" pageNumber="113">type</typeStatus>
series of the other two, and confirmed that all these three species are distributed and abundant in Yulongshan Mountain according to our recent expedition results and examined materials from different collections. Moreover, except 
<taxonomicName id="4C03DA9D5657FFE5C1A9FAB0FE63FAA3" box="[230,458,1317,1341]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="diverus">
<emphasis id="B9777D0C5657FFE5C1A9FAB0FE63FAA3" box="[230,458,1317,1341]" italics="true" pageId="12" pageNumber="113">Pterostichus diverus</emphasis>
</taxonomicName>
, the other two species are rediscovered only from Yulongshan Mountain. So, according the above evidence, we suppose that the 
<typeStatus id="54B81FBC5657FFE5C312FADFFD24FAFC" box="[605,653,1354,1378]" pageId="12" pageNumber="113">type</typeStatus>
locality of this species is probably not far from Yulongshan mountain (
<geoCoordinate id="EE37C7D95657FFE5C1D0FAF8FF56FA1B" box="[159,255,1389,1413]" direction="north" orientation="latitude" pageId="12" pageNumber="113" precision="555" value="27.17">N27.17º</geoCoordinate>
, 
<geoCoordinate id="EE37C7D95657FFE5C045FAF8FEDAFA1B" box="[266,371,1389,1413]" direction="east" orientation="longitude" pageId="12" pageNumber="113" precision="555" value="100.26">E100.26º</geoCoordinate>
) from Lijiang.
</paragraph>
<paragraph id="8BBCA11E5657FFE5C188FA07FB04F986" blockId="12.[151,1436,1029,2036]" pageId="12" pageNumber="113">
<emphasis id="B9777D0C5657FFE5C188FA07FE50FA34" bold="true" box="[199,505,1425,1450]" pageId="12" pageNumber="113">
<typeStatus id="54B81FBC5657FFE5C188FA07FEADFA34" box="[199,260,1426,1450]" pageId="12" pageNumber="113">Type</typeStatus>
materials examined
</emphasis>
. 
<emphasis id="B9777D0C5657FFE5C345FA04FDC3FA34" bold="true" box="[522,618,1425,1450]" pageId="12" pageNumber="113">
<typeStatus id="54B81FBC5657FFE5C345FA04FDC3FA34" box="[522,618,1425,1450]" pageId="12" pageNumber="113" type="syntype">Syntype</typeStatus>
</emphasis>
of 
<taxonomicName id="4C03DA9D5657FFE5C3DBFA07FC75FA37" authority="Fairmaire" authorityName="Fairmaire" box="[660,988,1425,1449]" class="Insecta" family="Carabidae" genus="Omaseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5657FFE5C3DBFA07FCC8FA37" box="[660,865,1425,1449]" italics="true" pageId="12" pageNumber="113">Omaseus diversus</emphasis>
Fairmaire
</taxonomicName>
: 
<specimenCount id="9D056A975657FFE5C2A6FA04FBE4FA37" box="[1001,1101,1425,1449]" pageId="12" pageNumber="113" type="female">1 female</specimenCount>
(ZRAS), body length = 
<quantity id="4CFB0CFB5657FFE5C424FA04FF6CFA53" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="12" pageNumber="113" unit="mm" value="14.2">14.2 mm</quantity>
, pinned, &quot; 
<taxonomicName id="4C03DA9D5657FFE5C070FA20FE35FA53" box="[319,412,1461,1485]" class="Insecta" family="Carabidae" genus="Omaseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5657FFE5C070FA20FE35FA53" box="[319,412,1461,1485]" italics="true" pageId="12" pageNumber="113">diversus</emphasis>
</taxonomicName>
/ Ƥ 
<emphasis id="B9777D0C5657FFE5C094FA23FD87FA53" box="[475,558,1462,1485]" italics="true" pageId="12" pageNumber="113">yunnan</emphasis>
&quot; [handwritten by Fairmaire]; &quot; 
<emphasis id="B9777D0C5657FFE5C2D5FA23FC5DFA53" box="[922,1012,1462,1485]" italics="true" pageId="12" pageNumber="113">yunnan.</emphasis>
/ 
<emphasis id="B9777D0C5657FFE5C55DFA20FB66FA53" box="[1042,1231,1461,1485]" italics="true" pageId="12" pageNumber="113">Coll. Fairmaire.</emphasis>
&quot; [handwritten by Semenov]; &quot; 
<taxonomicName id="4C03DA9D5657FFE5C06CFA4CFDABFA6F" authority="Fairmaire." authorityName="Fairmaire." box="[291,514,1497,1521]" class="Insecta" family="Carabidae" genus="Omaseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5657FFE5C06CFA4CFDABFA6F" box="[291,514,1497,1521]" italics="true" pageId="12" pageNumber="113">diversus Fairmaire.</emphasis>
</taxonomicName>
/ Ƥ 
<emphasis id="B9777D0C5657FFE5C37AFA4FFD77FA6F" box="[565,734,1497,1521]" italics="true" pageId="12" pageNumber="113">
typ. (
<taxonomicName id="4C03DA9D5657FFE5C321FA4CFD7EFA6F" box="[622,727,1497,1521]" class="Insecta" family="Carabidae" genus="Omaseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">Omaseus</taxonomicName>
)
</emphasis>
/ 
<emphasis id="B9777D0C5657FFE5C3BAFA4FFC19FA6F" box="[757,944,1497,1521]" italics="true" pageId="12" pageNumber="113">L. Fairmaire det</emphasis>
&quot; [handwritten probably by Tschitschérine]; &quot;Zoological Institute / Russian Academy / of Sciences / St. Petersburg&quot; [yellow label] [
<figureCitation id="1338BD9B5657FFE5C511FA68FB22F995" box="[1118,1163,1533,1547]" captionStart="FIGURES 7 – 12" captionStartId="22.[151,264,1765,1788]" captionTargetBox="[167,1429,198,1733]" captionTargetId="figure@22.[151,1436,183,1744]" captionTargetPageId="22" captionText="FIGURES 7 – 12. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 7, Pt. crassiapex n. sp. holotype; 8, Pt. near crassiapex n. sp., a female from Liziping; 9, Pt. diversus (Fairmaire), a female syntype; 10, Pt. diversus (Fairmaire), a male from Yulongshan; 11, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 12, Pt. stictopleurus cangshanensis n. ssp., holotype." httpUri="https://zenodo.org/record/220505/files/figure.png" pageId="12" pageNumber="113">Figs. 9</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C5DEFA68FB0BF995" box="[1169,1186,1533,1547]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="12" pageNumber="113">24</figureCitation>
].
</paragraph>
<paragraph id="8BBCA11E5657FFE5C188F9B1FAF0F88A" blockId="12.[151,1436,1029,2036]" pageId="12" pageNumber="113">
<emphasis id="B9777D0C5657FFE5C188F9B1FED7F9A2" bold="true" box="[199,382,1572,1596]" pageId="12" pageNumber="113">
Notes on 
<typeStatus id="54B81FBC5657FFE5C00FF9B1FED7F9A2" box="[320,382,1572,1596]" pageId="12" pageNumber="113">types</typeStatus>
</emphasis>
. This species was described on an uncertain number of specimens, but both sexes were mentioned in the original description. Later, Tschitschérine (1897) redescribed this species on a male belonging to the 
<typeStatus id="54B81FBC5657FFE5C18EF9F8FF58F91B" box="[193,241,1645,1669]" pageId="12" pageNumber="113">type</typeStatus>
series. The collection of Fairmaire should be in MNHN, but we failed to find any specimen belonging to the 
<typeStatus id="54B81FBC5657FFE5C18FF904FF59F937" box="[192,240,1681,1705]" pageId="12" pageNumber="113">type</typeStatus>
series of this species after a carefully searching among the collection of MNHN. The studied female by us was the only 
<typeStatus id="54B81FBC5657FFE5C07FF920FE2EF953" box="[304,391,1717,1741]" pageId="12" pageNumber="113" type="syntype">syntype</typeStatus>
found in the collection of ZRAS (
<figureCitation id="1338BD9B5657FFE5C25EF921FCCBF952" box="[785,866,1716,1740]" captionStart="FIGURES 7 – 12" captionStartId="22.[151,264,1765,1788]" captionTargetBox="[167,1429,198,1733]" captionTargetId="figure@22.[151,1436,183,1744]" captionTargetPageId="22" captionText="FIGURES 7 – 12. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 7, Pt. crassiapex n. sp. holotype; 8, Pt. near crassiapex n. sp., a female from Liziping; 9, Pt. diversus (Fairmaire), a female syntype; 10, Pt. diversus (Fairmaire), a male from Yulongshan; 11, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 12, Pt. stictopleurus cangshanensis n. ssp., holotype." httpUri="https://zenodo.org/record/220505/files/figure.png" pageId="12" pageNumber="113">Figs. 9</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C23FF921FC39F952" box="[880,912,1716,1740]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="12" pageNumber="113">24</figureCitation>
). This specimen could be exchanged from the collection of Fairmaire through Tschitschérine. But we don’t think it is appropriate to designate a 
<typeStatus id="54B81FBC5657FFE5C5A9F94DFAE5F96E" box="[1254,1356,1752,1776]" pageId="12" pageNumber="113" type="lectotype">lectotype</typeStatus>
before a male 
<typeStatus id="54B81FBC5657FFE5C1A7F968FE96F88B" box="[232,319,1789,1813]" pageId="12" pageNumber="113" type="syntype">syntype</typeStatus>
were found out, which should exist according to Fairmaire (1886) and Tschitschérine (1897).
</paragraph>
<paragraph id="8BBCA11E5657FFE4C188F8B6FAB8FDD9" blockId="12.[151,1436,1029,2036]" lastBlockId="13.[151,1438,151,2023]" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113">
<emphasis id="B9777D0C5657FFE5C188F8B6FD83F8A5" bold="true" box="[199,554,1826,1851]" pageId="12" pageNumber="113">Non-type materials examined</emphasis>
(total 
<specimenCount id="9D056A975657FFE5C338F8B6FCA6F8A5" box="[631,783,1827,1851]" pageId="12" pageNumber="113" type="generic">30 specimens</specimenCount>
). 
<specimenCount id="9D056A975657FFE5C266F8B6FCDFF8A5" box="[809,886,1827,1851]" pageId="12" pageNumber="113" type="male">1 male</specimenCount>
(MNHN), &quot;Yunnan, R.P.Delavay&quot; [
<figureCitation id="1338BD9B5657FFE5C443F8B5FA93F8B0" box="[1292,1338,1824,1838]" captionStart="FIGURES 13 – 18" captionStartId="23.[151,264,1749,1772]" captionTargetBox="[158,1427,204,1715]" captionTargetId="figure@23.[151,1436,193,1728]" captionTargetPageId="23" captionText="FIGURES 13 – 18. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 13, Pt. pseudodiversus n. sp., holotype; 14, Pt. pseudodiversus n. sp., a female paratype from Muli; 15, Pt. pseudodiversus n. sp., a male paratype from Deqin; 16, Pt. pseudodiversus n. sp., a male from Weixi; 17, Pt. kambaiti (Andrewes), holotype [photoed by Dr. J. Bergsten]; 18, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220506/files/figure.png" pageId="12" pageNumber="113">Fig. 17</figureCitation>
]. 
<specimenCount id="9D056A975657FFE5C400F8B6FA32F8A5" box="[1359,1435,1827,1851]" pageId="12" pageNumber="113" type="male">1 male</specimenCount>
(MNHN), &quot;Yunnan, Ta-pin-tze, R.P.Delavay&quot; [
<figureCitation id="1338BD9B5657FFE5C3FAF8D3FD42F8CA" box="[693,747,1862,1876]" captionStart="FIGURES 29 – 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 – 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="12" pageNumber="113">Figs. 34</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C3BBF8D3FCACF8CA" box="[756,773,1862,1876]" captionStart="FIGURES 41 – 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 – 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 – 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 – 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="12" pageNumber="113">42</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C242F8D3FCB7F8CA" box="[781,798,1862,1876]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="12" pageNumber="113">76</figureCitation>
]. 
<specimenCount id="9D056A975657FFE5C27CF8DCFC39F8FF" box="[819,912,1865,1889]" pageId="12" pageNumber="113" type="male">3 males</specimenCount>
, 
<specimenCount id="9D056A975657FFE5C2D0F8DCFBA6F8FF" box="[927,1039,1865,1889]" pageId="12" pageNumber="113" type="female">2 females</specimenCount>
(NHMB), &quot;Yunnan 
<quantity id="4CFB0CFB5657FFE5C44EF8DCFA31F8FF" box="[1281,1432,1865,1889]" metricMagnitude="3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="12" pageNumber="113" unit="m" value="3750.0" valueMax="4000.0" valueMin="3500.0">3500–4000m</quantity>
, 27.10 
<geoCoordinate id="EE37C7D95657FFE5C199F8F8FEEDF81B" box="[214,324,1901,1925]" direction="north" orientation="latitude" pageId="12" pageNumber="113" precision="555" value="100.13">N 100.13</geoCoordinate>
E, YULONGSHAN mts., vít Kubáň leg. 
<date id="FFBD87DE5657FFE5C26BF8F8FC61F81B" box="[804,968,1901,1925]" pageId="12" pageNumber="113" value="1993-06-16" valueMax="1993-06-19" valueMin="1993-06-16">16–19/6/1993</date>
.&quot;. 
<specimenCount id="9D056A975657FFE5C2ABF8F8FB9CF81B" box="[996,1077,1901,1925]" pageId="12" pageNumber="113" type="male">1 male</specimenCount>
, 
<specimenCount id="9D056A975657FFE5C50AF8F8FB1CF81B" box="[1093,1205,1901,1925]" pageId="12" pageNumber="113" type="female">2 females</specimenCount>
(NHMB), &quot;Yunnan 
<quantity id="4CFB0CFB5657FFE5C1D8F804FE84F837" box="[151,301,1937,1961]" metricMagnitude="3" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.6" pageId="12" pageNumber="113" unit="m" value="3950.0" valueMax="4300.0" valueMin="3600.0">3600–4300m</quantity>
, 27.02 
<geoCoordinate id="EE37C7D95657FFE5C034F804FE41F837" box="[379,488,1937,1961]" direction="north" orientation="latitude" pageId="12" pageNumber="113" precision="555" value="100.11">N 100.11</geoCoordinate>
E, YULONGSHAN mts., vít Kubáň leg. 
<date id="FFBD87DE5657FFE5C289F804FB94F837" box="[966,1085,1937,1961]" pageId="12" pageNumber="113" value="1993-06-22">22/6/1993</date>
&quot;. 
<specimenCount id="9D056A975657FFE5C51BF804FB18F837" box="[1108,1201,1937,1961]" pageId="12" pageNumber="113" type="male">2 males</specimenCount>
, 
<specimenCount id="9D056A975657FFE5C5F0F804FA8DF837" box="[1215,1316,1937,1961]" pageId="12" pageNumber="113" type="female">1 female</specimenCount>
(NHMB), &quot;Yunnan 
<quantity id="4CFB0CFB5657FFE5C049F82DFE35F84E" box="[262,412,1976,2000]" metricMagnitude="3" metricUnit="m" metricValue="3.85" metricValueMax="4.1" metricValueMin="3.6" pageId="12" pageNumber="113" unit="m" value="3850.0" valueMax="4100.0" valueMin="3600.0">3600–4100m</quantity>
, 27.02 
<geoCoordinate id="EE37C7D95657FFE5C0A4F82DFDFEF84E" box="[491,599,1976,2000]" direction="north" orientation="latitude" pageId="12" pageNumber="113" precision="555" value="100.11">N 100.11</geoCoordinate>
E, YULONGSHAN mts., vít Kubáň leg. 
<date id="FFBD87DE5657FFE5C575F82DFB07F84E" box="[1082,1198,1976,2000]" pageId="12" pageNumber="113" value="1993-05-27">27/5/1993</date>
&quot; [
<figureCitation id="1338BD9B5657FFE5C589F820FB5FF85D" box="[1222,1270,1973,1987]" captionStart="FIGURES 85 – 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 – 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="12" pageNumber="113">Fig. 87</figureCitation>
, 
<figureCitation id="1338BD9B5657FFE5C5B0F820FABEF85D" box="[1279,1303,1973,1987]" captionStart="FIGURES 107 – 109" captionStartId="32.[151,264,1625,1648]" captionText="FIGURES 107 – 109. Elytral microsculpture of Pterostichus (Wraseiellus) spp., sub-basal area of left elytron, interval 3 – 4: scale bar = 0.2 mm; 107, Pt. crassiapex n. sp., male (transverse); 108, Pt. diversus (Fairmaire), male (isodiametric, normal); 109, Pt. diversus (Fairmaire), female (isodiametric, extraordinary strong)." pageId="12" pageNumber="113">109</figureCitation>
]. 
<specimenCount id="9D056A975657FFE5C462F82DFAD7F84E" box="[1325,1406,1976,2000]" pageId="12" pageNumber="113" type="male">1 male</specimenCount>
, 
<specimenCount id="9D056A975657FFE5C4C1F82DFF4AF86A" pageId="12" pageNumber="113" type="female">1 female</specimenCount>
(NHMB), &quot;YUNNAN, YULONG Mts., 
<date id="FFBD87DE5657FFE5C3F2F849FCDEF86A" box="[701,887,2012,2036]" pageId="12" pageNumber="113" value="1993-06-20" valueMax="1993-06-21" valueMin="1993-06-20">20–21 Jun 1993</date>
, 27.07 
<geoCoordinate id="EE37C7D95657FFE5C28AF849FB9BF86A" box="[965,1074,2012,2036]" direction="north" orientation="latitude" pageId="12" pageNumber="113" precision="555" value="100.13">N 100.13</geoCoordinate>
E, Bolm lgt., 
<quantity id="4CFB0CFB5657FFE5C581F849FA8AF86A" box="[1230,1315,2012,2036]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="113" unit="m" value="3400.0">3400m</quantity>
&quot;. 
<specimenCount id="9D056A975657FFE5C477F849FA35F86A" box="[1336,1436,2012,2036]" pageId="12" pageNumber="113" type="female">1 female</specimenCount>
(NHMB), &quot;YUNNAN, 
<date id="FFBD87DE5656FFE4C0EAFF02FDEAFF31" box="[421,579,151,176]" pageId="13" pageNumber="114" value="1993-05-27">27 May 1993</date>
, YULONG MTS. 
<quantity id="4CFB0CFB5656FFE4C250FF02FCC6FF31" box="[799,879,151,175]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="114" unit="m" value="4000.0">4000m</quantity>
, 27.02 
<geoCoordinate id="EE37C7D95656FFE4C2F3FF02FB83FF31" box="[956,1066,151,175]" direction="north" orientation="latitude" pageId="13" pageNumber="114" precision="555" value="100.11">N 100.11</geoCoordinate>
E, Bolm lgt.&quot;. 
<specimenCount id="9D056A975656FFE4C59AFF02FA8AFF31" box="[1237,1315,151,175]" pageId="13" pageNumber="114" type="male">1 male</specimenCount>
(NHMB), &quot;YUNNAN, 
<date id="FFBD87DE5656FFE4C060FF29FE05FF4A" box="[303,428,188,212]" pageId="13" pageNumber="114" value="1995-07-09">9.Jul 1995</date>
, YULONGSHAN 
<quantity id="4CFB0CFB5656FFE4C3C2FF29FD77FF4A" box="[653,734,188,212]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="13" pageNumber="114" unit="m" value="3300.0">3300m</quantity>
, 27.07 
<geoCoordinate id="EE37C7D95656FFE4C262FF29FC34FF4A" box="[813,925,188,212]" direction="north" orientation="latitude" pageId="13" pageNumber="114" precision="555" value="100.14">N 100.14</geoCoordinate>
E, BOLM lgt.&quot;. 
<specimenCount id="9D056A975656FFE4C52EFF29FB61FF4A" box="[1121,1224,188,212]" pageId="13" pageNumber="114" type="female">1 female</specimenCount>
(CRS), &quot; 
<collectingCountry id="F314E18E5656FFE4C475FF29FA31FF4A" box="[1338,1432,188,212]" name="China" pageId="13" pageNumber="114">CHINA</collectingCountry>
, YUNNAN, LIJIANG area, YULONGSHAN mts.-E slopes, 27º09'–12'/100º13'–16', 
<quantity id="4CFB0CFB5656FFE4C511FF4AFB58FF69" box="[1118,1265,223,247]" metricMagnitude="3" metricUnit="m" metricValue="3.45" metricValueMax="3.9" metricValueMin="3.0" pageId="13" pageNumber="114" unit="m" value="3450.0" valueMax="3900.0" valueMin="3000.0">3000–3900m</quantity>
18– 
<date id="FFBD87DE5656FFE4C46CFF4AFA31FF69" box="[1315,1432,223,247]" pageId="13" pageNumber="114" value="1990-07-25">25.7.1990</date>
, coll.D.Král&quot;. 
<specimenCount id="9D056A975656FFE4C062FE91FED1FE82" box="[301,376,260,284]" pageId="13" pageNumber="114" type="male">1 male</specimenCount>
(CRS), &quot;YUNNAN 
<quantity id="4CFB0CFB5656FFE4C310FE91FD19FE82" box="[607,688,260,284]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="114" unit="m" value="3000.0">3000m</quantity>
, 27.05 
<geoCoordinate id="EE37C7D95656FFE4C3B6FE91FCCBFE82" box="[761,866,260,284]" direction="north" orientation="latitude" pageId="13" pageNumber="114" precision="555" value="100.15">N 100.15</geoCoordinate>
E, YULONGSHAN mts., Ganhaizi pass, vít Kubáň leg. 
<date id="FFBD87DE5656FFE4C188FEB2FEBDFEA1" box="[199,276,295,319]" pageId="13" pageNumber="114" value="1992-07-04">4/7/92</date>
&quot;. 
<specimenCount id="9D056A975656FFE4C068FEB2FE29FEA1" box="[295,384,295,319]" pageId="13" pageNumber="114" type="male">2 males</specimenCount>
, 
<specimenCount id="9D056A975656FFE4C0C4FEB2FE42FEA1" box="[395,491,295,319]" pageId="13" pageNumber="114" type="female">1 female</specimenCount>
(CRS), &quot; 
<collectingCountry id="F314E18E5656FFE4C31FFEB2FD07FEA1" box="[592,686,295,319]" name="China" pageId="13" pageNumber="114">CHINA</collectingCountry>
, Yunnan prov., 27º06 N 100º15 E, 
<quantity id="4CFB0CFB5656FFE4C579FEB2FB65FEA1" box="[1078,1228,295,319]" metricMagnitude="3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="13" pageNumber="114" unit="m" value="3250.0" valueMax="3500.0" valueMin="3000.0">3000–3500m</quantity>
, Yulongshan mts., GANHAIZI pass, vít Kubáň leg. 
<date id="FFBD87DE5656FFE4C347FEDAFD14FEF9" box="[520,701,335,359]" pageId="13" pageNumber="114" value="1990-07-18" valueMax="1990-07-23" valueMin="1990-07-18">18–23.VII.1990</date>
&quot; [
<figureCitation id="1338BD9B5656FFE4C39EFED9FD56FEC4" box="[721,767,332,346]" captionStart="FIGURES 7 – 12" captionStartId="22.[151,264,1765,1788]" captionTargetBox="[167,1429,198,1733]" captionTargetId="figure@22.[151,1436,183,1744]" captionTargetPageId="22" captionText="FIGURES 7 – 12. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 7, Pt. crassiapex n. sp. holotype; 8, Pt. near crassiapex n. sp., a female from Liziping; 9, Pt. diversus (Fairmaire), a female syntype; 10, Pt. diversus (Fairmaire), a male from Yulongshan; 11, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 12, Pt. stictopleurus cangshanensis n. ssp., holotype." httpUri="https://zenodo.org/record/220505/files/figure.png" pageId="13" pageNumber="114">Fig. 10</figureCitation>
, 
<figureCitation id="1338BD9B5656FFE4C24BFED9FCB5FEC4" box="[772,796,332,346]" captionStart="FIGURES 107 – 109" captionStartId="32.[151,264,1625,1648]" captionText="FIGURES 107 – 109. Elytral microsculpture of Pterostichus (Wraseiellus) spp., sub-basal area of left elytron, interval 3 – 4: scale bar = 0.2 mm; 107, Pt. crassiapex n. sp., male (transverse); 108, Pt. diversus (Fairmaire), male (isodiametric, normal); 109, Pt. diversus (Fairmaire), female (isodiametric, extraordinary strong)." pageId="13" pageNumber="114">108</figureCitation>
]. 
<specimenCount id="9D056A975656FFE4C261FEDAFC27FEF9" box="[814,910,335,359]" pageId="13" pageNumber="114" type="female">1 female</specimenCount>
(CRS), &quot; 
<collectingCountry id="F314E18E5656FFE4C2BCFEDAFBF8FEF9" box="[1011,1105,335,359]" name="China" pageId="13" pageNumber="114">CHINA</collectingCountry>
, YUNNAN prov., HEISHUI- 35KM n Lijiang, 18.6.– 
<date id="FFBD87DE5656FFE4C0EBFEE7FDA2FE14" box="[420,523,370,394]" pageId="13" pageNumber="114" value="1993-07-04">4.7.1993</date>
, 27,13N; 100,19E, lgt.S.Becvar&quot;. 
<specimenCount id="9D056A975656FFE4C2DEFEE7FC48FE14" box="[913,993,370,394]" pageId="13" pageNumber="114" type="male">1 male</specimenCount>
, 
<specimenCount id="9D056A975656FFE4C2A1FEE7FBF8FE14" box="[1006,1105,370,394]" pageId="13" pageNumber="114" type="female">1 female</specimenCount>
(CSF), &quot; 
<collectingCountry id="F314E18E5656FFE4C5F8FEE7FABCFE14" box="[1207,1301,370,394]" name="China" pageId="13" pageNumber="114">CHINA</collectingCountry>
, YUNNAN prov., HEISHUI-35KM n Lijiang, 18.6.– 
<date id="FFBD87DE5656FFE4C3CAFE0CFD44FE2F" box="[645,749,409,433]" pageId="13" pageNumber="114" value="1993-07-04">4.7.1993</date>
, 27,13N; 100,19E, lgt.S.Becvar&quot; [
<figureCitation id="1338BD9B5656FFE4C5CBFE03FB11FE3A" box="[1156,1208,406,420]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="13" pageNumber="114">Fig. 75</figureCitation>
]. 
<specimenCount id="9D056A975656FFE4C59CFE0CFA8EFE2F" box="[1235,1319,409,433]" pageId="13" pageNumber="114" type="male">1 male</specimenCount>
(CCCC), &quot;Yunnan, Lijiang, Yulongxueshan, Zhu Xiaoyu leg., 2009. 
<date id="FFBD87DE5656FFE4C216FE28FC31FE4B" box="[857,920,445,469]" pageId="13" pageNumber="114" value="1917-06">VI.17</date>
, JC09DJ0757&quot;. 
<specimenCount id="9D056A975656FFE4C512FE28FB06FE4B" box="[1117,1199,445,469]" pageId="13" pageNumber="114" type="male">1 male</specimenCount>
(CCCC), &quot;Yunnan, Lijiang, Yulongxueshan, Zhu Xiaoyu leg., 2009. 
<date id="FFBD87DE5656FFE4C3F4FE74FD52FE67" box="[699,763,481,505]" pageId="13" pageNumber="114" value="1917-06">VI.17</date>
, JC09DJ0761&quot;. 
<specimenCount id="9D056A975656FFE4C2FEFE74FC56FE67" box="[945,1023,481,505]" pageId="13" pageNumber="114" type="male">1 male</specimenCount>
, 
<specimenCount id="9D056A975656FFE4C545FE74FBC2FE67" box="[1034,1131,481,505]" pageId="13" pageNumber="114" type="female">1 female</specimenCount>
(IZAS), &quot;Yunnan, Lijiang, Yulongxueshan cons., 
<quantity id="4CFB0CFB5656FFE4C0D9FD9DFE4EFDBE" box="[406,487,520,544]" metricMagnitude="3" metricUnit="m" metricValue="2.98" pageId="13" pageNumber="114" unit="m" value="2980.0">2980m</quantity>
, pine forest, by pitfall trap, 
<date id="FFBD87DE5656FFE4C26FFD9DFC6BFDBE" box="[800,962,520,544]" pageId="13" pageNumber="114" value="2000-08-01" valueMax="2000-08-03" valueMin="2000-08-01">2000 VIII 1–3</date>
, Yu Xiaodong, Zhou Hongzhang leg.&quot; [
<figureCitation id="1338BD9B5656FFE4C433FD90FF01FDA4" captionStart="FIGURES 29 – 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 – 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="13" pageNumber="114">Figs. 33</figureCitation>
, 
<figureCitation id="1338BD9B5656FFE4C1E1FDB9FF16FDA4" box="[174,191,556,570]" captionStart="FIGURES 41 – 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 – 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 – 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 – 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="13" pageNumber="114">41</figureCitation>
, 
<figureCitation id="1338BD9B5656FFE4C18AFDB9FF7FFDA4" box="[197,214,556,570]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="13" pageNumber="114">61</figureCitation>
, 
<figureCitation id="1338BD9B5656FFE4C193FDB9FF44FDA4" box="[220,237,556,570]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="13" pageNumber="114">74</figureCitation>
, 
<figureCitation id="1338BD9B5656FFE4C1BCFDB9FEADFDA4" box="[243,260,556,570]" captionStart="FIGURES 90 – 93" captionStartId="31.[151,264,1640,1663]" captionTargetBox="[161,1418,216,1605]" captionTargetId="figure@31.[151,1436,193,1619]" captionTargetPageId="31" captionText="FIGURES 90 – 93. Left stylomere of ovipositor of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, ventral view; B, dorsal view: 90, Pt. diversus (Fairmaire), a female from Yulongshan; 91, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 92, Pt. pseudodiversus n. sp., a paratype from Luguhu. 93, Pt. andrewesi Jedlička, a female from Jinfoshan. Abbreviations: es, ensiform setae; ns, nematiform setae; s 1, stylomere 1; s 2, stylomere 2." httpUri="https://zenodo.org/record/220514/files/figure.png" pageId="13" pageNumber="114">90</figureCitation>
, 
<figureCitation id="1338BD9B5656FFE4C045FDB9FE8BFDA4" box="[266,290,556,570]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="13" pageNumber="114">102</figureCitation>
]. 
<specimenCount id="9D056A975656FFE4C07AFDBAFE3CFDD9" box="[309,405,559,583]" pageId="13" pageNumber="114" type="female">1 female</specimenCount>
(SNUM), &quot;Mt. Yulong, Lijiang, Yunnan Prov., 
<date id="FFBD87DE5656FFE4C2FDFDBAFBE5FDD9" box="[946,1100,559,583]" pageId="13" pageNumber="114" value="2003-07-11">11–VII–2003</date>
, Li-zhen Li leg.&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C319F2955656FFE4C188FDC7FC34FB85" pageId="13" pageNumber="114" type="diagnosis">
<paragraph id="8BBCA11E5656FFE4C188FDC7FC34FB85" blockId="13.[151,1438,151,2023]" pageId="13" pageNumber="114">
<emphasis id="B9777D0C5656FFE4C188FDC7FEE8FDF5" bold="true" box="[199,321,594,619]" pageId="13" pageNumber="114">Diagnosis.</emphasis>
Most species of 
<taxonomicName id="4C03DA9D5656FFE4C342FDC6FCABFDF5" box="[525,770,595,619]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5656FFE4C342FDC6FCABFDF5" box="[525,770,595,619]" italics="true" pageId="13" pageNumber="114">Pterostichus diversus</emphasis>
</taxonomicName>
species group are very difficult to distinguish from each other by external characters, except the subspecies 
<taxonomicName id="4C03DA9D5656FFE4C3B5FDE2FB7AFD11" box="[762,1235,631,655]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" status="subsp. nov." subSpecies="cangshanensis">
<emphasis id="B9777D0C5656FFE4C3B5FDE2FB7AFD11" box="[762,1235,631,655]" italics="true" pageId="13" pageNumber="114">Pterostichus stictopleurus cangshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A244C0775656FFE4C590FDEDFA85FD11" box="[1247,1324,632,655]" pageId="13" pageNumber="114" rank="subSpecies">n. ssp.</taxonomicNameLabel>
which is different by its 5th tarsomere setose beneath. From other allied species of this species group, 
<taxonomicName id="4C03DA9D5656FFE4C58CFD09FAE0FD2D" box="[1219,1353,667,691]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5656FFE4C58CFD09FAE0FD2D" box="[1219,1353,667,691]" italics="true" pageId="13" pageNumber="114">Pt. diversus</emphasis>
</taxonomicName>
can be easily distinguished by male genital characters: (1) apical lamella of median lobe of aedeagus long and capitate, apex slightly truncate (
<figureCitation id="1338BD9B5656FFE4C0D5FD76FD8CFD65" box="[410,549,739,763]" captionStart="FIGURES 41 – 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 – 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 – 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 – 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="13" pageNumber="114">Figs. 41, 42</figureCitation>
); (2) right paramere long and wide, apex usually truncate, not narrowed (
<figureCitation id="1338BD9B5656FFE4C42CFD76FF1CFC81" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="13" pageNumber="114">Figs. 74</figureCitation>
B–77B). But it is easier to distinguish this species from others by their different localities except from 
<taxonomicName id="4C03DA9D5656FFE4C433FC9DFE69FCDD" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" subSpecies="stictopleurus">
<emphasis id="B9777D0C5656FFE4C433FC9DFE69FCDD" italics="true" pageId="13" pageNumber="114">Pt. stictopleurus stictopleurus</emphasis>
</taxonomicName>
, which is sympatric with 
<taxonomicName id="4C03DA9D5656FFE4C3ACFCB9FCCEFCDD" box="[739,871,811,835]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5656FFE4C3ACFCB9FCCEFCDD" box="[739,871,811,835]" italics="true" pageId="13" pageNumber="114">Pt. diversus</emphasis>
</taxonomicName>
at Jizushan Mt. (Dali). Except for their difference on male genitalia, 
<taxonomicName id="4C03DA9D5656FFE4C029FCC5FE43FCF9" box="[358,490,847,871]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5656FFE4C029FCC5FE43FCF9" box="[358,490,847,871]" italics="true" pageId="13" pageNumber="114">Pt. diversus</emphasis>
</taxonomicName>
is different from 
<taxonomicName id="4C03DA9D5656FFE4C3FFFCC5FBA9FCF9" box="[688,1024,847,871]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" subSpecies="stictopleurus">
<emphasis id="B9777D0C5656FFE4C3FFFCC5FBA9FCF9" box="[688,1024,847,871]" italics="true" pageId="13" pageNumber="114">Pt. stictopleurus stictopleurus</emphasis>
</taxonomicName>
also in: (1) area between outer basal foveal groove and lateral margin of pronotum strongly convex, forming a distinct ridge (
<figureCitation id="1338BD9B5656FFE4C5DEFCE6FB5BFC15" box="[1169,1266,883,907]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="13" pageNumber="114">Fig. 102</figureCitation>
) (only slightly convex not forming a distinct ridge in 
<taxonomicName id="4C03DA9D5656FFE4C30DFC0DFC39FC31" box="[578,912,919,943]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" subSpecies="stictopleurus">
<emphasis id="B9777D0C5656FFE4C30DFC0DFC39FC31" box="[578,912,919,943]" italics="true" pageId="13" pageNumber="114">Pt. stictopleurus stictopleurus</emphasis>
</taxonomicName>
(
<figureCitation id="1338BD9B5656FFE4C2D1FC02FC56FC31" box="[926,1023,919,943]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="13" pageNumber="114">Fig. 104</figureCitation>
)); (2) lateral margins slightly sinuate just before hind angles, hind angles obtuse but slightly pointed (
<figureCitation id="1338BD9B5656FFE4C2CFFC2EFC4BFC4D" box="[896,994,955,979]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="13" pageNumber="114">Fig. 102</figureCitation>
) (not sinuate before hind angles, hind angles not pointed in 
<taxonomicName id="4C03DA9D5656FFE4C0C8FC75FD7FFC69" box="[391,726,991,1015]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" subSpecies="stictopleurus">
<emphasis id="B9777D0C5656FFE4C0C8FC75FD7FFC69" box="[391,726,991,1015]" italics="true" pageId="13" pageNumber="114">Pt. stictopleurus stictopleurus</emphasis>
</taxonomicName>
(
<figureCitation id="1338BD9B5656FFE4C3AAFC4AFCEFFC69" box="[741,838,991,1015]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="13" pageNumber="114">Fig. 104</figureCitation>
)); (3) pronotum widest a little before middle (
<figureCitation id="1338BD9B5656FFE4C407FC4AFF1DFB85" captionStart="FIGURES 7 – 12" captionStartId="22.[151,264,1765,1788]" captionTargetBox="[167,1429,198,1733]" captionTargetId="figure@22.[151,1436,183,1744]" captionTargetPageId="22" captionText="FIGURES 7 – 12. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 7, Pt. crassiapex n. sp. holotype; 8, Pt. near crassiapex n. sp., a female from Liziping; 9, Pt. diversus (Fairmaire), a female syntype; 10, Pt. diversus (Fairmaire), a male from Yulongshan; 11, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 12, Pt. stictopleurus cangshanensis n. ssp., holotype." httpUri="https://zenodo.org/record/220505/files/figure.png" pageId="13" pageNumber="114">Figs. 9, 10</figureCitation>
) (widest in the middle in 
<taxonomicName id="4C03DA9D5656FFE4C09BFB91FC8DFB85" box="[468,804,1027,1051]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" subSpecies="stictopleurus">
<emphasis id="B9777D0C5656FFE4C09BFB91FC8DFB85" box="[468,804,1027,1051]" italics="true" pageId="13" pageNumber="114">Pt. stictopleurus stictopleurus</emphasis>
</taxonomicName>
(
<figureCitation id="1338BD9B5656FFE4C27CFB96FC2AFB85" box="[819,899,1027,1051]" captionStart="FIGURES 7 – 12" captionStartId="22.[151,264,1765,1788]" captionTargetBox="[167,1429,198,1733]" captionTargetId="figure@22.[151,1436,183,1744]" captionTargetPageId="22" captionText="FIGURES 7 – 12. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 7, Pt. crassiapex n. sp. holotype; 8, Pt. near crassiapex n. sp., a female from Liziping; 9, Pt. diversus (Fairmaire), a female syntype; 10, Pt. diversus (Fairmaire), a male from Yulongshan; 11, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 12, Pt. stictopleurus cangshanensis n. ssp., holotype." httpUri="https://zenodo.org/record/220505/files/figure.png" pageId="13" pageNumber="114">Fig. 11</figureCitation>
)).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955656FFE4C188FBB3FDDDF879" pageId="13" pageNumber="114" type="description">
<paragraph id="8BBCA11E5656FFE4C188FBB3FDDDF879" blockId="13.[151,1438,151,2023]" pageId="13" pageNumber="114">
<emphasis id="B9777D0C5656FFE4C188FBB3FEF8FBA1" bold="true" box="[199,337,1062,1087]" pageId="13" pageNumber="114">Description</emphasis>
. Body length 
<quantity id="4CFB0CFB5656FFE4C0BCFBB2FD33FBA1" box="[499,666,1063,1087]" metricMagnitude="-2" metricUnit="m" metricValue="1.365" metricValueMax="1.49" metricValueMin="1.24" pageId="13" pageNumber="114" unit="mm" value="13.65" valueMax="14.9" valueMin="12.4">12.4–14.9 mm</quantity>
; dorsal side black, shining in males; tibiae dark brown, tarsi brown; ventral side black or dark brown. Males with elytral microsculpture isodiametric. 
<emphasis id="B9777D0C5656FFE4C533FBDEFB15FBFD" bold="true" box="[1148,1212,1099,1123]" pageId="13" pageNumber="114">Head</emphasis>
. Frons and vertex impunctate, sometimes few punctures present in frontal furrows; antennae reaching elytra basal eighth; 3rd antennomere usually glabrous except apical setae, but with few additional setae in some specimens; males with terminal labial palpomere similar to that of females, slightly expanded and truncate. 
<emphasis id="B9777D0C5656FFE4C51CFB22FB64FB51" bold="true" box="[1107,1229,1207,1231]" pageId="13" pageNumber="114">Pronotum</emphasis>
rounded, widest a little before middle, PW/PL = 1.31–1.42; lateral margins fully rounded near the middle, slightly sinuate just before hind angles; hind angles slightly pointed; one mid-lateral seta present at about anterior third of lateral margins; basal fovea moderately deep, faintly defined; inner groove of basal fovea subparallel to median line, slightly curved to the outside posteriorly, ended at a distance from the posterior margin of pronotum; outer groove of basal fovea about half length as the inner one, as deep as the inner one, reaching the basal margin of pronotum; ridge between outer groove and lateral margin distinct; area between inner and outer basal foveal grooves convex and slightly rugose; basal fovea area densely punctate, some sparser punctures usually present in the area between two basal foveae. 
<emphasis id="B9777D0C5656FFE4C070FA42FE25FA71" bold="true" box="[319,396,1495,1519]" pageId="13" pageNumber="114">Elytra</emphasis>
with base slightly depressed between 3rd and 5th intervals; elytral shoulders slightly narrowed, humeral teeth small but distinct; intervals feebly convex; striae shallow, with very faint punctures; scutellar striae long, usually incomplete, located between 1st stria and elytral suture; umbilical pore series on 9th interval sparse in the middle, composed of 16–19 pores (6, 2–4, 7–9). 
<emphasis id="B9777D0C5656FFE4C2EEF9D6FB98F9C5" bold="true" box="[929,1073,1602,1627]" pageId="13" pageNumber="114">Ventral side</emphasis>
. Proepisterna and metaepisterna densely punctate; terminal sternum of males without special structure. 
<emphasis id="B9777D0C5656FFE4C287F9F2FBA9F9E1" bold="true" box="[968,1024,1639,1663]" pageId="13" pageNumber="114">Legs</emphasis>
. 5th tarsomere usually without seta beneath, rarely one pair of very fine setae present. 
<emphasis id="B9777D0C5656FFE4C3A4F91EFC31F93D" bold="true" box="[747,920,1674,1699]" pageId="13" pageNumber="114">Male genitalia</emphasis>
. Median lobe of male genitalia bent at a 90 degrees angle at about basal two-fifths; ventral margin almost straight in the middle, apical part not bent to the ventral side (
<figureCitation id="1338BD9B5656FFE4C065F946FE2FF975" box="[298,390,1747,1771]" captionStart="FIGURES 29 – 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 – 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="13" pageNumber="114">Figs. 33</figureCitation>
A, 34A); apical orifice large, slightly turned to the left side, apical lamella twisted to the left (
<figureCitation id="1338BD9B5656FFE4C1D0F962FEA8F891" box="[159,257,1783,1807]" captionStart="FIGURES 29 – 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 – 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="13" pageNumber="114">Figs. 33</figureCitation>
B, 34B); apical lamella laminar, about twice longer than its basal width, apex capitate and rounded, slightly bent to the left (
<figureCitation id="1338BD9B5656FFE4C0E1F88EFD95F8AD" box="[430,572,1819,1843]" captionStart="FIGURES 41 – 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 – 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 – 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 – 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="13" pageNumber="114">Figs. 41, 42</figureCitation>
); apex of apical lamella gradually turned to the dorsal side (
<figureCitation id="1338BD9B5656FFE4C5BDF88EFAEEF8AD" box="[1266,1351,1819,1843]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="13" pageNumber="114">Fig. 61</figureCitation>
). Right paramere laminar, elongate and curved, right and left margin subparallel, not narrowed apically, apex truncate or slightly pointed (
<figureCitation id="1338BD9B5656FFE4C019F8F6FE1AF8E5" box="[342,435,1891,1915]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="13" pageNumber="114">Figs. 74</figureCitation>
B–77B). 
<emphasis id="B9777D0C5656FFE4C357F8F6FD74F8E5" bold="true" box="[536,733,1890,1915]" pageId="13" pageNumber="114">Female genitalia</emphasis>
. Spermatheca (
<figureCitation id="1338BD9B5656FFE4C2C3F8F6FC74F8E5" box="[908,989,1891,1915]" captionStart="FIGURES 85 – 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 – 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="13" pageNumber="114">Fig. 87</figureCitation>
) with the seminal canal about ten times long as the receptaculum; spermathecal gland strongly expanded; the seminal canal inserted at the base of common oviduct, base of seminal canal sclerotized (as in 
<figureCitation id="1338BD9B5656FFE4C3F6F83EFCA5F85D" box="[697,780,1963,1987]" captionStart="FIGURES 85 – 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 – 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="13" pageNumber="114">Fig. 85</figureCitation>
). Stylomere 2 with two ensiform setae at the outer margin and three or four at inner margin (
<figureCitation id="1338BD9B5656FFE4C35CF85AFDC1F879" box="[531,616,1999,2023]" captionStart="FIGURES 90 – 93" captionStartId="31.[151,264,1640,1663]" captionTargetBox="[161,1418,216,1605]" captionTargetId="figure@31.[151,1436,193,1619]" captionTargetPageId="31" captionText="FIGURES 90 – 93. Left stylomere of ovipositor of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, ventral view; B, dorsal view: 90, Pt. diversus (Fairmaire), a female from Yulongshan; 91, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 92, Pt. pseudodiversus n. sp., a paratype from Luguhu. 93, Pt. andrewesi Jedlička, a female from Jinfoshan. Abbreviations: es, ensiform setae; ns, nematiform setae; s 1, stylomere 1; s 2, stylomere 2." httpUri="https://zenodo.org/record/220514/files/figure.png" pageId="13" pageNumber="114">Fig. 90</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955655FFE7C188FF02FB62FE82" pageId="14" pageNumber="115" type="distribution">
<paragraph id="8BBCA11E5655FFE7C188FF02FB62FE82" blockId="14.[151,1436,151,680]" pageId="14" pageNumber="115">
<emphasis id="B9777D0C5655FFE7C188FF02FEF0FF2E" bold="true" box="[199,345,151,176]" pageId="14" pageNumber="115">Distribution</emphasis>
. Yulongxueshan, Lijiang (Yunnan); Ta-pin-tze, Dali (Yunnan). The locality &quot;Ta-pin-tze&quot; is only recorded by a male collected about a hundred years ago. Ta-pin-tze (= Dapingdi, 
<geoCoordinate id="EE37C7D95655FFE7C508FF29FB0CFF4A" box="[1095,1189,188,212]" direction="north" orientation="latitude" pageId="14" pageNumber="115" precision="555" value="25.91">N25.91º</geoCoordinate>
, 
<geoCoordinate id="EE37C7D95655FFE7C5FCFF29FAB5FF4A" box="[1203,1308,188,212]" direction="east" orientation="longitude" pageId="14" pageNumber="115" precision="555" value="100.24">E100.24º</geoCoordinate>
) is a small village on the ancient way to the peak of Jizushan Mt., Dali District. All the specimens collected in recent years were from Yulongxueshan (=Yulongshan, 
<geoCoordinate id="EE37C7D95655FFE7C338FE91FD7CFE82" box="[631,725,260,284]" direction="north" orientation="latitude" pageId="14" pageNumber="115" precision="555" value="27.12">N27.12º</geoCoordinate>
, 
<geoCoordinate id="EE37C7D95655FFE7C3AFFE91FCE2FE82" box="[736,843,260,284]" direction="east" orientation="longitude" pageId="14" pageNumber="115" precision="555" value="100.2">E100.20º</geoCoordinate>
), Lijiang District (
<figureCitation id="1338BD9B5655FFE7C555FE91FB17FE82" box="[1050,1214,260,284]" captionStart="FIGURES 110 – 111" captionStartId="33.[151,264,1885,1908]" captionTargetBox="[224,1312,207,1827]" captionTargetId="figure@33.[224,1343,193,1844]" captionTargetPageId="33" captionText="FIGURES 110 – 111. Distribution maps for species of subgenus Pterostichus (Wraseiellus): - Pt. andrewesi Jedlička; « Pt. meyeri Jedlička; ˑ Pt. crassiapex n. sp.; ¿ Pt. comatus n. sp.; ● Pt. diversus (Fairmaire); Ο Pt. pseudodiversus n. sp.; ˔ Pt. stictopleurus stictopleurus (Fairmaire); r Pt. stictopleurus cangshanensis n. ssp.; ․ Pt. kambaiti (Andrewes). 111. shows an enlarged scale of the area in figure 110 marked by the dotted line square." httpUri="https://zenodo.org/record/220516/files/figure.png" pageId="14" pageNumber="115">Figs. 110, 111</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955655FFE7C188FEB2FBFAFE51" pageId="14" pageNumber="115" type="description">
<paragraph id="8BBCA11E5655FFE7C188FEB2FBFAFE51" blockId="14.[151,1436,151,680]" pageId="14" pageNumber="115">
<emphasis id="B9777D0C5655FFE7C188FEB2FE93FEDE" bold="true" box="[199,314,295,320]" pageId="14" pageNumber="115">Variation</emphasis>
. A large series of specimens of this species were studied in the present work, so singular individual variations in some characters were discovered: (1) additional setae on 3rd antennomere present in few individuals; (2) elytral 3rd interval may have one additional pore after the posterior pore or the position of pores may change; (3) in one specimen, 5th pro- and mesotarsomeres with a pair of very fine setae beneath; (4) apex of right paramere may vary in different individuals, being truncate or slightly pointed (
<figureCitation id="1338BD9B5655FFE7C2D7FE22FC5CFE51" box="[920,1013,439,463]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="14" pageNumber="115">Figs. 74</figureCitation>
B–77B).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955655FFE7C188FE49FC50FD39" pageId="14" pageNumber="115" type="discussion">
<paragraph id="8BBCA11E5655FFE7C188FE49FBD9FDA2" blockId="14.[151,1436,151,680]" pageId="14" pageNumber="115">
<emphasis id="B9777D0C5655FFE7C188FE49FE9DFE6A" bold="true" box="[199,308,476,500]" pageId="14" pageNumber="115">Remarks</emphasis>
. It is curious that the male 
<typeStatus id="54B81FBC5655FFE7C331FE48FD7CFE6B" box="[638,725,477,501]" pageId="14" pageNumber="115" type="syntype">syntype</typeStatus>
studied by Tschitschérine has two pairs of fine setae on 5th tarsomeres: &quot; 
<emphasis id="B9777D0C5655FFE7C07DFD95FB7FFD86" box="[306,1238,512,536]" italics="true" pageId="14" pageNumber="115">le 5e de tous porte en dessous 2 cils très fins de chaque côté, à peu près au milieu.</emphasis>
&quot; (Tschitschérine, 1897). But, the female 
<typeStatus id="54B81FBC5655FFE7C0D6FDB0FE59FDA3" box="[409,496,549,573]" pageId="14" pageNumber="115" type="syntype">syntype</typeStatus>
and other specimens examined by us without such setae.
</paragraph>
<paragraph id="8BBCA11E5655FFE7C188FDDDFC50FD39" blockId="14.[151,1436,151,680]" pageId="14" pageNumber="115">
In the present paper, we recognized this species based on a female 
<typeStatus id="54B81FBC5655FFE7C2FEFDDDFBA1FDFE" box="[945,1032,584,608]" pageId="14" pageNumber="115" type="syntype">syntype</typeStatus>
and other examined specimens. We are confident that these materials from Yulongshan Mountain are identical with the 
<typeStatus id="54B81FBC5655FFE7C529FDF8FB14FD1B" box="[1126,1213,621,645]" pageId="14" pageNumber="115" type="syntype">syntype</typeStatus>
for their accordant external characters and our speculation on the 
<typeStatus id="54B81FBC5655FFE7C3D0FD05FD66FD36" box="[671,719,656,680]" pageId="14" pageNumber="115">type</typeStatus>
locality mentioned above.
</paragraph>
</subSubSection>
</treatment>
</document>