<documentid="1C149F75B52FF598F7338ECCC075AC5D"ID-DOI="10.5852/ejt.2020.706"ID-GBIF-Dataset="269e39e4-19ff-4a70-9376-8b7eaf09f663"ID-ISSN="2118-9773"ID-Zenodo-Dep="4010954"ID-ZooBank="C4C4D130-1EA7-48AA-A664-391DBC59C484"IM.bibliography_approvedBy="juliana"IM.materialsCitations_approvedBy="juliana"IM.metadata_approvedBy="valdenar"IM.tables_approvedBy="valdenar"IM.taxonomicNames_approvedBy="juliana"IM.treatmentCitations_approvedBy="juliana"IM.treatments_approvedBy="juliana"checkinTime="1598981996449"checkinUser="valdenar"docAuthor="Kantor, Yuri I., Castelin, Magalie, Fedosov, Alexander & Bouchet, Philippe"docDate="2020"docId="03A7D504FFECFFD76791FCF5A559F972"docLanguage="en"docName="ejt-706_kantor_castelin_fedosov.pdf"docOrigin="European Journal of Taxonomy 706"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Amalda aureomarginata Kilburn & Bouchet 1988"docType="treatment"docVersion="11"lastPageNumber="27"masterDocId="FF9EAD7CFFFAFFCD667AFFC7A410FFC8"masterDocTitle="The Indo-Pacific Amalda (Neogastropoda, Olivoidea, Ancillariidae) revisited with molecular data, with special emphasis on New Caledonia"masterLastPageNumber="59"masterPageNumber="1"pageNumber="23"updateTime="1733512817148"updateUser="juliana"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="42119E8C772F065F29A7E92013E8B89B">The Indo-Pacific Amalda (Neogastropoda, Olivoidea, Ancillariidae) revisited with molecular data, with special emphasis on New Caledonia</mods:title>
<mods:affiliationid="EFC1629B13C621295C6036310EBC685B">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninski Prospect 33, 119071 Moscow.</mods:affiliation>
<mods:affiliationid="2F1BA9B9F1109120E36169BDC5762DA4">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninski Prospect 33, 119071 Moscow.</mods:affiliation>
<taxonomicNameid="4C0E1F91FFECFFDB6791FCF5A058FC85"authority="Kilburn & Bouchet, 1988"authorityName="Kilburn & Bouchet"authorityYear="1988"box="[491,1096,818,845]"class="Gastropoda"family="Olividae"genus="Amalda"kingdom="Animalia"order="Neogastropoda"pageId="22"pageNumber="23"phylum="Mollusca"rank="species"species="aureomarginata">
<bibRefCitationid="EF9F19E3FFECFFDB6566FCF5A058FC85"author="Kilburn R. N. & Bouchet P."box="[796,1096,818,845]"pageId="22"pageNumber="23"pagination="277 - 300"refId="ref31223"refString="Kilburn R. N. & Bouchet P. 1988. The genus Amalda in New Caledonia (Mollusca, Gastropoda, Olividae, Ancillinae). Bulletin du Museum national d'histoire naturelle, ser. 4, 10, section A 2: 277 - 300."type="journal article"year="1988">Kilburn & Bouchet, 1988</bibRefCitation>
<figureCitationid="13357897FFECFFDB64DDFC91A718FCB9"box="[679,776,854,881]"captionStart="Fig"captionStartId="23.[189,232,1754,1780]"captionTargetBox="[189,1398,533,1712]"captionTargetId="figure@23.[189,1398,533,1712]"captionTargetPageId="23"captionText="Fig. 7. Radulae of species of Amalda H. Adams & A. Adams, 1853. A–G. A. hilgendorfi (von Martens, 1897). A–B. MNHN IM-2019-621 (shell on Fig. 4D). C. MNHN IM-2009-11961 (shell on Fig. 4J). D. MNHN IM-2013-58045, Papua New Guinea. E. MNHN IM-2013-19909, Papua New Guinea. F. MNHN IM-2013-63430, southern New Caledonia. G. MNHN IM-2007-33312 (shell on Fig. 5F). H. A. aureomarginata Kilburn & Bouchet, 1988, MNHN IM-2013-63421, specimen not sequenced, southern New Caledonia. I–J. A. miriky sp. nov. I. Miriky, stn CP3210, specimen not sequenced, Madagascar. J. Miriky, stn DW3215 (shell on Fig. 10J). K–L. A. cacao sp. nov., holotype, MNHN IM- 2000-35296 (shell on Fig. 11A–E)."figureDoi="http://doi.org/10.5281/zenodo.4010968"httpUri="https://zenodo.org/record/4010968/files/figure.png"pageId="22"pageNumber="23">Figs 7H</figureCitation>
,
<figureCitationid="13357897FFECFFDB656EFC90A731FCB9"box="[788,801,855,881]"captionStart="Fig"captionStartId="24.[189,232,1824,1850]"captionTargetBox="[189,1398,287,1781]"captionTargetId="figure@24.[189,1398,287,1781]"captionTargetPageId="24"captionText="Fig. 8. A–H.Amalda aureomarginata Kilburn& Bouchet,1988, southernNew Caledonia.A–C. Holotype, MNHN IM-2000-1345, SL 34.1 mm. D. MNHN IM-2007-43644, SL 26.6 mm. E. MNHN IM-2007- 43655, SL 31.5 mm. F. MNHN IM-2007-43551, SL 30.5 mm. G–H. MNHN IM-2007-43654, SL 28.2 mm. I–J. A. hilgendorfi (von Martens, 1897), MNHN IM-2009-11968, northern New Caledonia, SL 31.6 mm. Shells at the same scale; H, J = enlarged upper part of the shell. Arrow on H points at dark brown line, bordering secondary spire callus."figureDoi="http://doi.org/10.5281/zenodo.4010970"httpUri="https://zenodo.org/record/4010970/files/figure.png"pageId="22"pageNumber="23">8</figureCitation>
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<taxonomicNameid="4C0E1F91FFECFFDB66C7FC5EA763FC7B"authority="Kilburn & Bouchet, 1988: 282"authorityName="Kilburn & Bouchet"authorityPageNumber="282"authorityYear="1988"box="[189,883,921,947]"class="Gastropoda"family="Olividae"genus="Amalda"kingdom="Animalia"order="Neogastropoda"pageId="22"pageNumber="23"phylum="Mollusca"rank="species"species="aureomarginata">
<bibRefCitationid="EF9F19E3FFECFFDB678AFC5EA763FC7B"author="Kilburn R. N. & Bouchet P."box="[496,883,921,947]"pageId="22"pageNumber="23"pagination="277 - 300"refId="ref31223"refString="Kilburn R. N. & Bouchet P. 1988. The genus Amalda in New Caledonia (Mollusca, Gastropoda, Olividae, Ancillinae). Bulletin du Museum national d'histoire naturelle, ser. 4, 10, section A 2: 277 - 300."type="journal article"year="1988">Kilburn & Bouchet, 1988: 282</bibRefCitation>
<taxonomicNameid="4C0E1F91FFECFFDB66C7FC38A5CCFBD1"authorityName="Kilburn & Bouchet"authorityYear="1988"box="[189,476,1023,1049]"class="Gastropoda"family="Olividae"genus="Amalda"kingdom="Animalia"order="Neogastropoda"pageId="22"pageNumber="23"phylum="Mollusca"rank="species"species="aureomarginata">
<bibRefCitationid="EF9F19E3FFECFFDB6783FC38A6B6FBD1"author="Gratecap D."box="[505,678,1023,1049]"pageId="22"pageNumber="23"pagination="13 - 20"refId="ref30817"refString="Gratecap D. 2014. Le genre Amalda en Nouvelle-Caledonie. Xenophora 146: 13 - 20."type="journal article"year="2014">Gratecap 2014</bibRefCitation>
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) (
<tableCitationid="C68C51A9FFECFFDB64FCFA89A6CEFAA0"box="[646,734,1358,1384]"captionStart="Table 1"captionStartId="2.[180,249,549,577]"captionTargetBox="[180,1931,635,1722]"captionTargetPageId="3"captionText="Table 1 (continued on next six pages). List of sequenced specimens analyzed, including voucher registration and GenBank accession numbers. Original sequences highlighted in bold typeface. The BOLD ID refers to the The Barcode of Life Data System."pageId="22"pageNumber="23">Table 1</tableCitation>
<locationDeviationid="9A037D30FFECFFDB6734F9AEA72DF94B"box="[334,829,1639,1667]"pageId="22"pageNumber="23">between Île des Pins and Grand Récif Sud</locationDeviation>
<locationDeviationid="9A037D30FFECFFDB67E0F94AA78AF96F"box="[410,922,1675,1703]"pageId="22"pageNumber="23">between Île des Pins and Grand Récif Sud</locationDeviation>
<locationDeviationid="9A037D30FFECFFDB6787F976A7FFF903"box="[509,1007,1711,1739]"pageId="22"pageNumber="23">between Île des Pins and Grand Récif Sud</locationDeviation>
<locationDeviationid="9A037D30FFECFFDB643FF912A028F927"box="[581,1080,1747,1775]"pageId="22"pageNumber="23">between Île des Pins and Grand Récif Sud</locationDeviation>
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ca 30 dd; S of
<locationid="8ED132C9FFECFFDB6215F8F8A0ECF893"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFECFFDB6215F8F8A0ECF893"box="[1135,1276,1855,1883]"latitude="-22.8"longLatPrecision="1258"longitude="167.25"name="Ile des Pins"pageId="22"pageNumber="23">
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3 dd; S of
<locationid="8ED132C9FFECFFDB6599F8A4A062F8B7"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFECFFDB6599F8A4A062F8B7"box="[995,1138,1891,1919]"latitude="-22.866667"longLatPrecision="1258"longitude="167.26666"name="Ile des Pins"pageId="22"pageNumber="23">
<materialsCitationid="3B666E4FFFECFFDB64E5F84EA5BEF80F"ID-GBIF-Occurrence="2854824358"elevation="431"latitude="-22.833334"location="Ile des Pins"longLatPrecision="1258"longitude="167.45"pageId="22"pageNumber="23"specimenCount="1">
4 dd; S of
<locationid="8ED132C9FFECFFDB6560F840A7B5F86B"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFECFFDB6560F840A7B5F86B"box="[794,933,1927,1955]"latitude="-22.833334"longLatPrecision="1258"longitude="167.45"name="Ile des Pins"pageId="22"pageNumber="23">
<materialsCitationid="3B666E4FFFEDFFDA6713FEE8A53CFEA4"ID-GBIF-Occurrence="2854824423"collectingDate="1978"collectingDateMax="1979"collectingDateMin="1978"collectorName="Vauban"elevation="395"latitude="-22.783333"location="W of Ile des Pins"longLatPrecision="1258"longitude="167.2"pageId="23"pageNumber="24"specimenCount="1">
4 dd;
<locationid="8ED132C9FFEDFFDA67C9FEE8A680FE80"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFEDFFDA67C9FEE8A680FE80"box="[435,656,301,329]"latitude="-22.783333"longLatPrecision="1258"longitude="167.2"name="W of Ile des Pins"pageId="23"pageNumber="24">
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25 dd; SW of
<locationid="8ED132C9FFEDFFDA6719FEB4A5FFFE47"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFEDFFDA6719FEB4A5FFFE47"box="[355,495,371,399]"latitude="-22.95"longLatPrecision="1258"longitude="167.26666"name="Ile des Pins"pageId="23"pageNumber="24">
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20 dd; SW of
<locationid="8ED132C9FFEDFFDA6776FE51A583FE7A"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFEDFFDA6776FE51A583FE7A"box="[268,403,406,434]"latitude="-22.9"longLatPrecision="1258"longitude="167.21666"name="Ile des Pins"pageId="23"pageNumber="24">
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<locationid="8ED132C9FFEDFFDA66A6FE7FA576FE1C"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFEDFFDA66A6FE7FA576FE1C"box="[220,358,440,468]"latitude="-22.883333"longLatPrecision="1258"longitude="167.21666"name="Ile des Pins"pageId="23"pageNumber="24">
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4 dd; SW of
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<locationid="8ED132C9FFE3FFD46350FECEA4E3FE8F"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFE3FFD46350FECEA4E3FE8F"latitude="-22.85"longLatPrecision="1258"longitude="167.23334"name="Ile des Pins"pageId="25"pageNumber="26">
<materialsCitationid="3B666E4FFFE3FFD46255FEE9A740FEA2"ID-GBIF-Occurrence="2854824371"elevation="273"latitude="-22.716667"location="Ile des Pins"longLatPrecision="1258"longitude="167.26666"pageId="25"pageNumber="26"specimenCount="1">
1 dd,
<locationid="8ED132C9FFE3FFD46208FEEBA0EDFE8F"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFE3FFD46208FEEBA0EDFE8F"box="[1138,1277,300,328]"latitude="-22.716667"longLatPrecision="1258"longitude="167.26666"name="Ile des Pins"pageId="25"pageNumber="26">
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30 dd;
<locationid="8ED132C9FFE3FFD465CCFE89A02EFEA1"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFE3FFD465CCFE89A02EFEA1"box="[950,1086,334,362]"latitude="-22.866667"longLatPrecision="1258"longitude="167.21666"name="Ile des Pins"pageId="25"pageNumber="26">
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<locationid="8ED132C9FFE0FFD76512FECEA7E2FEED"LSID="urn:lsid:plazi:treatment:03A7D504FFECFFD76791FCF5A559F972:8ED132C9FFE0FFD76512FECEA7E2FEED"box="[872,1010,265,293]"latitude="-22.8"longLatPrecision="1258"longitude="167.25"name="Ile des Pins"pageId="26"pageNumber="27">
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, MNHN IM-2013-63421, specimen not sequenced, southern New Caledonia.
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<taxonomicNameid="4C0E1F91FFE3FFD462B6F84CA5F0F800"authority="Kilburn & Bouchet, 1988"authorityName="Kilburn & Bouchet"authorityYear="1988"class="Gastropoda"family="Olividae"genus="Amalda"kingdom="Animalia"order="Neogastropoda"pageId="25"pageNumber="26"phylum="Mollusca"rank="species"species="fuscolingua">
<bibRefCitationid="EF9F19E3FFE3FFD466C7F86AA5F0F800"author="Kilburn R. N. & Bouchet P."box="[189,480,1965,1992]"pageId="25"pageNumber="26"pagination="277 - 300"refId="ref31223"refString="Kilburn R. N. & Bouchet P. 1988. The genus Amalda in New Caledonia (Mollusca, Gastropoda, Olividae, Ancillinae). Bulletin du Museum national d'histoire naturelle, ser. 4, 10, section A 2: 277 - 300."type="journal article"year="1988">Kilburn & Bouchet, 1988</bibRefCitation>
(mean 0.91, n =5). Primary spire callus thick, covering all except protoconch and one or two teleoconch whorls, secondary callus forming fairly thin, linguiform pad, extending to second last suture. Plication plate with 4–7 ridges, sometimes bifid. Olivoid groove rather shallow, labral denticle very weak. Upper anterior band weakly convex, nearly flat, may be slightly thickened at rear edge of anterior band. Lower anterior band flat, smooth. Colour light yellowish-brown with paler olivoid band, spire callus with orange-yellow lower band, bordered abapically by narrow light line and creamy upper band with purple tint. Adapically primary callus with narrow but distinct brown borderline. Parietal callus thick, typically purplish-pink. Upper anterior band yellowish orange, lower anterior band and plication plate white.
<figureCitationid="13357897FFE0FFD76757FC6DA59DFC0C"box="[301,397,938,964]"captionStart="Fig"captionStartId="23.[189,232,1754,1780]"captionTargetBox="[189,1398,533,1712]"captionTargetId="figure@23.[189,1398,533,1712]"captionTargetPageId="23"captionText="Fig. 7. Radulae of species of Amalda H. Adams & A. Adams, 1853. A–G. A. hilgendorfi (von Martens, 1897). A–B. MNHN IM-2019-621 (shell on Fig. 4D). C. MNHN IM-2009-11961 (shell on Fig. 4J). D. MNHN IM-2013-58045, Papua New Guinea. E. MNHN IM-2013-19909, Papua New Guinea. F. MNHN IM-2013-63430, southern New Caledonia. G. MNHN IM-2007-33312 (shell on Fig. 5F). H. A. aureomarginata Kilburn & Bouchet, 1988, MNHN IM-2013-63421, specimen not sequenced, southern New Caledonia. I–J. A. miriky sp. nov. I. Miriky, stn CP3210, specimen not sequenced, Madagascar. J. Miriky, stn DW3215 (shell on Fig. 10J). K–L. A. cacao sp. nov., holotype, MNHN IM- 2000-35296 (shell on Fig. 11A–E)."figureDoi="http://doi.org/10.5281/zenodo.4010968"httpUri="https://zenodo.org/record/4010968/files/figure.png"pageId="26"pageNumber="27">Fig. 7H</figureCitation>
). With simple rather thin hook-shaped lateral teeth and tricuspid central teeth with central cusp smallest. Outer lower edges of outer cusps serrated.
<bibRefCitationid="EF9F19E3FFE0FFD764EBFB82A7D5FB97"author="Kilburn R. N. & Bouchet P."box="[657,965,1093,1119]"pageId="26"pageNumber="27"pagination="277 - 300"refId="ref31223"refString="Kilburn R. N. & Bouchet P. 1988. The genus Amalda in New Caledonia (Mollusca, Gastropoda, Olividae, Ancillinae). Bulletin du Museum national d'histoire naturelle, ser. 4, 10, section A 2: 277 - 300."type="journal article"year="1988">Kilburn & Bouchet (1988</bibRefCitation>
: figs 52–56) is rather similar to ours, but the lateral cusps of the central tooth with serrated inner edges as well.
<bibRefCitationid="EF9F19E3FFE0FFD766C7FB68A5ECFB01"author="Kilburn R. N. & Bouchet P."box="[189,508,1199,1225]"pageId="26"pageNumber="27"pagination="277 - 300"refId="ref31223"refString="Kilburn R. N. & Bouchet P. 1988. The genus Amalda in New Caledonia (Mollusca, Gastropoda, Olividae, Ancillinae). Bulletin du Museum national d'histoire naturelle, ser. 4, 10, section A 2: 277 - 300."type="journal article"year="1988">Kilburn & Bouchet (1988)</bibRefCitation>
form, the latter two characterized by a broader oval shell and details of colouration. A molecular analysis, however, revealed that the specimens identified as these two latter forms are in fact subadults of
<figureCitationid="13357897FFE0FFD7653AFAE6A782FAF3"box="[832,914,1313,1339]"captionStart="Fig"captionStartId="24.[189,232,1824,1850]"captionTargetBox="[189,1398,287,1781]"captionTargetId="figure@24.[189,1398,287,1781]"captionTargetPageId="24"captionText="Fig. 8. A–H.Amalda aureomarginata Kilburn& Bouchet,1988, southernNew Caledonia.A–C. Holotype, MNHN IM-2000-1345, SL 34.1 mm. D. MNHN IM-2007-43644, SL 26.6 mm. E. MNHN IM-2007- 43655, SL 31.5 mm. F. MNHN IM-2007-43551, SL 30.5 mm. G–H. MNHN IM-2007-43654, SL 28.2 mm. I–J. A. hilgendorfi (von Martens, 1897), MNHN IM-2009-11968, northern New Caledonia, SL 31.6 mm. Shells at the same scale; H, J = enlarged upper part of the shell. Arrow on H points at dark brown line, bordering secondary spire callus."figureDoi="http://doi.org/10.5281/zenodo.4010970"httpUri="https://zenodo.org/record/4010970/files/figure.png"pageId="26"pageNumber="27">Fig. 8I</figureCitation>
). They differ from
<taxonomicNameid="4C0E1F91FFE0FFD7620EFAE6A141FAF3"authorityName="Kilburn & Bouchet"authorityYear="1988"box="[1140,1361,1313,1339]"class="Gastropoda"family="Olividae"genus="Amalda"kingdom="Animalia"order="Neogastropoda"pageId="26"pageNumber="27"phylum="Mollusca"rank="species"species="aureomarginata">
by the absence of a characteristic reddish-purple or purplish-pink upper part of the parietal callus and the median part of the primary spire callus. Another stable difference is that in
<taxonomicNameid="4C0E1F91FFE0FFD7625AFAAAA0EBFA4F"authorityName="Kilburn & Bouchet"authorityYear="1988"box="[1056,1275,1389,1415]"class="Gastropoda"family="Olividae"genus="Amalda"kingdom="Animalia"order="Neogastropoda"pageId="26"pageNumber="27"phylum="Mollusca"rank="species"species="aureomarginata">
, the upper limit of the secondary callus is bordered by a narrow distinct brown band (arrow on
<figureCitationid="13357897FFE0FFD762C2FA54A107FA65"box="[1208,1303,1427,1453]"captionStart="Fig"captionStartId="24.[189,232,1824,1850]"captionTargetBox="[189,1398,287,1781]"captionTargetId="figure@24.[189,1398,287,1781]"captionTargetPageId="24"captionText="Fig. 8. A–H.Amalda aureomarginata Kilburn& Bouchet,1988, southernNew Caledonia.A–C. Holotype, MNHN IM-2000-1345, SL 34.1 mm. D. MNHN IM-2007-43644, SL 26.6 mm. E. MNHN IM-2007- 43655, SL 31.5 mm. F. MNHN IM-2007-43551, SL 30.5 mm. G–H. MNHN IM-2007-43654, SL 28.2 mm. I–J. A. hilgendorfi (von Martens, 1897), MNHN IM-2009-11968, northern New Caledonia, SL 31.6 mm. Shells at the same scale; H, J = enlarged upper part of the shell. Arrow on H points at dark brown line, bordering secondary spire callus."figureDoi="http://doi.org/10.5281/zenodo.4010970"httpUri="https://zenodo.org/record/4010970/files/figure.png"pageId="26"pageNumber="27">Fig. 8H</figureCitation>
<taxonomicNameid="4C0E1F91FFE0FFD7656CFA7EA070FA1A"authorityName="Kilburn & Bouchet"authorityYear="1988"box="[790,1120,1464,1491]"class="Gastropoda"family="Olividae"genus="Amalda"kingdom="Animalia"order="Neogastropoda"pageId="26"pageNumber="27"phylum="Mollusca"rank="species"species="aureomarginata">
<bibRefCitationid="EF9F19E3FFE0FFD76724FA18A60EFA31"author="Gratecap D."box="[350,542,1503,1529]"pageId="26"pageNumber="27"pagination="13 - 20"refId="ref30817"refString="Gratecap D. 2014. Le genre Amalda en Nouvelle-Caledonie. Xenophora 146: 13 - 20."type="journal article"year="2014">Gratecap (2014)</bibRefCitation>
<figureCitationid="13357897FFE0FFD765B5F993A03CF9A6"box="[975,1068,1620,1646]"captionStart="Fig"captionStartId="25.[189,232,1930,1956]"captionTargetBox="[189,1398,406,1892]"captionTargetId="figure@25.[189,1398,406,1892]"captionTargetPageId="25"captionText="Fig. 9. Distribution map of Amalda aureomarginata Kilburn & Bouchet, 1988 (A) and A. fuscolingua Kilburn & Bouchet, 1988 (B) off southern New Caledonia. Red circles=the typelocalities; stars= indicate stations where the species were recovered."figureDoi="http://doi.org/10.5281/zenodo.4010972"httpUri="https://zenodo.org/record/4010972/files/figure.png"pageId="26"pageNumber="27">Fig. 9A</figureCitation>
), in a limited area off Île des Pins approximately 40 ×