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168 lines
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<document id="D0D1D3F779FC5F3C6561BB4EF4B3380E" ID-DOI="10.11646/zootaxa.5437.4.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="10985633" ID-ZooBank="7114B49F-4A44-4165-9FC6-8A52A6E898F9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713347432768" checkinUser="plazi" docAuthor="Kasparyan, Dmitry R. & Khalaim, Andrey I." docDate="2024" docId="03A03624FFF4B10D4DA4AA90FEC7FAFB" docLanguage="en" docName="zootaxa.5437.4.9.pdf" docOrigin="Zootaxa 5437 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5437.4.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glodianus Cameron 1902" docType="treatment" docVersion="2" lastPageNumber="572" masterDocId="FF994E5CFFF5B10C4D33A81EFFA1FF95" masterDocTitle="Two new species of Glodianus Cameron and Hylophasma Townes (Hymenoptera: Ichneumonidae: Cryptinae) from Mexico" masterLastPageNumber="581" masterPageNumber="571" pageNumber="572" updateTime="1713348150872" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="E9A5E9EB27288AD30FE9BD30E3E312F1">
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<mods:title id="09174E6B4FAFBEC6CD771498A971D59F">Two new species of Glodianus Cameron and Hylophasma Townes (Hymenoptera: Ichneumonidae: Cryptinae) from Mexico</mods:title>
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</mods:titleInfo>
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<mods:name id="0FF35142802CF9AB2BBFAD6839FD959E" type="personal">
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<mods:role id="7728EBB360B6CD6C66C40A7B3DB1250D">
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<mods:roleTerm id="7010D01790052DE74AFD1AF686ED9BBF">Author</mods:roleTerm>
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<mods:namePart id="F0BF418DF1F61C634A193DE199FD5CBA">Kasparyan, Dmitry R.</mods:namePart>
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<mods:affiliation id="D2558A89852869CDD4774C9E1D246808">Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia</mods:affiliation>
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<mods:name id="48394C86A65FB58B8A49101390F37125" type="personal">
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<mods:roleTerm id="06B533907536EE23A514223B4C699BFF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7559EDD1EB25969E2C3C734E01A0D17D">Khalaim, Andrey I.</mods:namePart>
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<mods:nameIdentifier id="02B7ABD0D194604FC6E51635C16F0207" type="ORCID">0000-0003-1802-2649</mods:nameIdentifier>
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<mods:affiliation id="767C2DF65D4DDF2D1856406B1F1B7D29">Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia</mods:affiliation>
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<mods:nameIdentifier id="25AD2B7EF56B9BD9C7C07BC77C28CF61" type="email">akhalaim@gmail.com</mods:nameIdentifier>
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<mods:title id="16296248A0518948E5FA6DA71BAEC4A1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="251E9BBCB8413DA4B7BE784C1674594A">
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<mods:date id="6B582C4305F14686256F063936FE0703">2024</mods:date>
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<mods:number id="5A1882BE8ED617B721E4899613012B7E">2024-04-16</mods:number>
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<mods:number id="6C84BA4D4D923E24D4D1D0393603BBE2">5437</mods:number>
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<mods:number id="717F6E8C3F8B347338A7AA83AF4C42D8">4</mods:number>
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<mods:start id="A6784A830C952DD331B7B03185473166">571</mods:start>
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<mods:end id="71E39C9F6DE00DD45A1271AD97F1E8AC">581</mods:end>
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<mods:url id="1BADE5EE30D5948BAFEC35CC319BD29C">http://dx.doi.org/10.11646/zootaxa.5437.4.9</mods:url>
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<mods:classification id="BAB0941F82C85B9FD473A9B0210CC448">journal article</mods:classification>
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<mods:identifier id="0E353A31BD0661625C8F14B663D7FA12" type="DOI">10.11646/zootaxa.5437.4.9</mods:identifier>
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<mods:identifier id="FBDF820223B6BC6F3E16C7B97997E439" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5C783EDE05E5B3D3ACF965045B50FDA0" type="Zenodo-Dep">10985633</mods:identifier>
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<treatment id="03A03624FFF4B10D4DA4AA90FEC7FAFB" ID-DOI="http://doi.org/10.5281/zenodo.10985653" ID-Zenodo-Dep="10985653" LSID="urn:lsid:plazi:treatment:03A03624FFF4B10D4DA4AA90FEC7FAFB" httpUri="http://treatment.plazi.org/id/03A03624FFF4B10D4DA4AA90FEC7FAFB" lastPageNumber="572" pageId="1" pageNumber="572">
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<subSubSection id="C313D4B9FFF4B10D4DA4AA90FD93FD3D" box="[151,562,654,681]" pageId="1" pageNumber="572" type="nomenclature">
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<paragraph id="8BB68732FFF4B10D4DA4AA90FD93FD3D" blockId="1.[151,562,654,681]" box="[151,562,654,681]" pageId="1" pageNumber="572">
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<heading id="D0FE305EFFF4B10D4DA4AA90FD93FD3D" bold="true" box="[151,562,654,681]" fontSize="11" level="1" pageId="1" pageNumber="572" reason="1">
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<emphasis id="B97D5B20FFF4B10D4DA4AA90FD93FD3D" bold="true" box="[151,562,654,681]" pageId="1" pageNumber="572">
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Genus
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<taxonomicName id="4C09FCB1FFF4B10D4DDDAA90FD93FD3D" ID-CoL="4P45" ID-ENA="540422" authority="Cameron, 1902" authorityName="Cameron" authorityYear="1902" box="[238,562,654,681]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="genus">
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<emphasis id="B97D5B20FFF4B10D4DDDAA90FECDFD3C" bold="true" box="[238,364,654,681]" italics="true" pageId="1" pageNumber="572">Glodianus</emphasis>
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<bibRefCitation id="EF98FAC3FFF4B10D4C40AA90FD93FD3D" author="Cameron, P." box="[371,562,654,680]" pageId="1" pageNumber="572" pagination="369 - 377" refId="ref6875" refString="Cameron, P. (1902) Descriptions of new genera and species of American Hymenoptera. Transactions of the American Entomological Society, 28, 369 - 377." type="journal article" year="1902">Cameron, 1902</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C313D4B9FFF4B10D4DA4AACDFE76FD7E" box="[151,471,723,747]" pageId="1" pageNumber="572" type="reference_group">
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<paragraph id="8BB68732FFF4B10D4DA4AACDFE76FD7E" blockId="1.[151,864,723,778]" box="[151,471,723,747]" pageId="1" pageNumber="572">
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<treatmentCitationGroup id="AB19A01CFFF4B10D4DA4AACDFE76FD7E" box="[151,471,723,747]" pageId="1" pageNumber="572">
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<treatmentCitation id="0AA8A123FFF4B10D4DA4AACDFE72FD7E" author="Cameron, P." box="[151,467,723,747]" page="369" pageId="1" pageNumber="572" year="1902">
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<taxonomicName id="4C09FCB1FFF4B10D4DA4AACDFE72FD7E" ID-CoL="4P45" authority="Cameron 1902: 369" authorityName="Cameron" authorityPageNumber="369" authorityYear="1902" box="[151,467,723,747]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="genus">
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<emphasis id="B97D5B20FFF4B10D4DA4AACDFEA0FD7E" box="[151,257,723,747]" italics="true" pageId="1" pageNumber="572">Glodianus</emphasis>
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<bibRefCitation id="EF98FAC3FFF4B10D4C34AACDFE72FD7E" author="Cameron, P." box="[263,467,723,747]" pageId="1" pageNumber="572" pagination="369 - 377" refId="ref6875" refString="Cameron, P. (1902) Descriptions of new genera and species of American Hymenoptera. Transactions of the American Entomological Society, 28, 369 - 377." type="journal article" year="1902">Cameron 1902: 369</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C313D4B9FFF4B10D4DA4AAECFEC7FAFB" pageId="1" pageNumber="572" type="materials_examined">
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<paragraph id="8BB68732FFF4B10D4DA4AAECFCC1FC9F" blockId="1.[151,864,723,778]" box="[151,864,754,778]" pageId="1" pageNumber="572">
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<typeStatus id="54B23990FFF4B10D4DA4AAECFF68FC9F" box="[151,201,754,778]" pageId="1" pageNumber="572">Type</typeStatus>
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species:
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<taxonomicName id="4C09FCB1FFF4B10D4C15AAECFD68FC9F" authority="Cameron, 1902" authorityName="Cameron" authorityYear="1902" box="[294,713,754,778]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="species" species="bombycivorus">
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<emphasis id="B97D5B20FFF4B10D4C15AAECFD85FC9F" box="[294,548,754,778]" italics="true" pageId="1" pageNumber="572">Glodianus bombycivorus</emphasis>
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<bibRefCitation id="EF98FAC3FFF4B10D4F18AAECFD68FC9F" author="Cameron, P." box="[555,713,754,778]" pageId="1" pageNumber="572" pagination="369 - 377" refId="ref6875" refString="Cameron, P. (1902) Descriptions of new genera and species of American Hymenoptera. Transactions of the American Entomological Society, 28, 369 - 377." type="journal article" year="1902">Cameron, 1902</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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<paragraph id="8BB68732FFF4B10D4DA4AB26FC1BFC77" blockId="1.[151,1437,824,1390]" pageId="1" pageNumber="572">
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The genus
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<taxonomicName id="4C09FCB1FFF4B10D4C24AB26FE2DFCC7" authorityName="Cameron" authorityYear="1902" box="[279,396,824,850]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="genus">
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<emphasis id="B97D5B20FFF4B10D4C24AB26FE2DFCC7" box="[279,396,824,850]" italics="true" pageId="1" pageNumber="572">Glodianus</emphasis>
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</taxonomicName>
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belongs to the small subtribe
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<taxonomicName id="4C09FCB1FFF4B10D4FDFAB26FCC9FCC7" box="[748,872,824,850]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="subTribe" subTribe="Glodianina">Glodianina</taxonomicName>
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comprising three exclusively Neotropical genera (
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<bibRefCitation id="EF98FAC3FFF4B10D4DACAB42FED3FCE3" author="Townes, H. K." box="[159,370,860,886]" pageId="1" pageNumber="572" pagination="1 - 537" refId="ref7309" refString="Townes, H. K. (1970) The genera of Ichneumonidae. Part 2. Memoirs of the American Entomological Institute, 12, 1 - 537." type="journal article" year="1970">Townes 1970: 232</bibRefCitation>
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,
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<bibRefCitation id="EF98FAC3FFF4B10D4C4EAB43FDBEFCE3" author="Yu, D. S. K. & Achterberg, C. van & Horstmann, K." box="[381,543,860,887]" pageId="1" pageNumber="572" refId="ref7497" refString="Yu, D. S. K., Achterberg, C. van & Horstmann, K. (2016) Taxapad 2016, Ichneumonoidea 2015. Nepean, Ontario. [database on flash-drive]" type="book" year="2016">
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Yu
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<emphasis id="B97D5B20FFF4B10D4C90AB43FE75FCE3" box="[419,468,860,886]" italics="true" pageId="1" pageNumber="572">et al</emphasis>
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. 2016
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</bibRefCitation>
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). All species of the subtribe are characterized by a small, rectangular or slightly trapeziform areolet in the fore wing (
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<figureCitation id="13329BB7FFF4B10D4F0FAB9EFD20FC0E" box="[572,641,896,923]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="1" pageNumber="572">Fig. 5</figureCitation>
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), polished first metasomal tergite (
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<figureCitation id="13329BB7FFF4B10D493EAB9EFBD2FC0F" box="[1037,1139,896,922]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1792,1816]" captionTargetBox="[151,1435,181,1759]" captionTargetId="figure-26@4.[151,1435,181,1768]" captionTargetPageId="4" captionText="FIGURES 6–10. Glodianus rufithorax sp. nov., ♀, holotype: 6—posterior part of mesosoma and base of metasoma, lateral; 7—propodeum and first tergite, dorsal; 8—apex of metasoma with ovipositor, lateral; 9—apex of ovipositor, lateral; 10—apex of ovipositor, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10985639" httpUri="https://zenodo.org/record/10985639/files/figure.png" pageId="1" pageNumber="572">Figs 6, 7</figureCitation>
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) and ovipositor apex with oblique grooves on dorsal valve (
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<figureCitation id="13329BB7FFF4B10D4F35ABBAFDE8FC2B" box="[518,585,932,958]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1792,1816]" captionTargetBox="[151,1435,181,1759]" captionTargetId="figure-26@4.[151,1435,181,1768]" captionTargetPageId="4" captionText="FIGURES 6–10. Glodianus rufithorax sp. nov., ♀, holotype: 6—posterior part of mesosoma and base of metasoma, lateral; 7—propodeum and first tergite, dorsal; 8—apex of metasoma with ovipositor, lateral; 9—apex of ovipositor, lateral; 10—apex of ovipositor, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10985639" httpUri="https://zenodo.org/record/10985639/files/figure.png" pageId="1" pageNumber="572">Fig. 9</figureCitation>
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). Within the subtribe
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<taxonomicName id="4C09FCB1FFF4B10D4E04ABBAFC17FC2B" box="[823,950,932,958]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="subTribe" subTribe="Glodianina">Glodianina</taxonomicName>
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,
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<taxonomicName id="4C09FCB1FFF4B10D4EF2ABBAFB97FC2B" authorityName="Cameron" authorityYear="1902" box="[961,1078,932,958]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="genus">
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<emphasis id="B97D5B20FFF4B10D4EF2ABBAFB97FC2B" box="[961,1078,932,958]" italics="true" pageId="1" pageNumber="572">Glodianus</emphasis>
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</taxonomicName>
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is easily distinguished from two other genera by having a median compressed horn on the frons (
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<figureCitation id="13329BB7FFF4B10D4E54ABD6FC0DFC77" box="[871,940,968,994]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="1" pageNumber="572">Fig. 2</figureCitation>
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).
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</paragraph>
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<paragraph id="8BB68732FFF4B10D4DF4ABF2FF5EFBE7" blockId="1.[151,1437,824,1390]" pageId="1" pageNumber="572">
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<taxonomicName id="4C09FCB1FFF4B10D4DF4ABF2FE9DFB93" authorityName="Cameron" authorityYear="1902" box="[199,316,1004,1030]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="genus">
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<emphasis id="B97D5B20FFF4B10D4DF4ABF2FE9DFB93" box="[199,316,1004,1030]" italics="true" pageId="1" pageNumber="572">Glodianus</emphasis>
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</taxonomicName>
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comprises ten species distributed from
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<collectingCountry id="F31EC7A2FFF4B10D4E26ABF2FCCAFB93" box="[789,875,1004,1030]" name="Mexico" pageId="1" pageNumber="572">Mexico</collectingCountry>
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to
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<collectingCountry id="F31EC7A2FFF4B10D4EABABF2FBAAFB93" box="[920,1035,1004,1030]" name="Venezuela" pageId="1" pageNumber="572">Venezuela</collectingCountry>
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and
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<collectingCountry id="F31EC7A2FFF4B10D4978ABF2FB33FB93" box="[1099,1170,1004,1030]" name="Brazil" pageId="1" pageNumber="572">Brazil</collectingCountry>
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, including one species described in this paper and another one resurrected. Seven species of
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<taxonomicName id="4C09FCB1FFF4B10D4E8AAC0EFB8FFBBF" authorityName="Cameron" authorityYear="1902" box="[953,1070,1040,1066]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="genus">
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<emphasis id="B97D5B20FFF4B10D4E8AAC0EFB8FFBBF" box="[953,1070,1040,1066]" italics="true" pageId="1" pageNumber="572">Glodianus</emphasis>
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</taxonomicName>
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|||
|
are restricted to South America and three species are known only from
|
|||
|
<collectingCountry id="F31EC7A2FFF4B10D4F66AC2AFD0FFBDB" box="[597,686,1076,1102]" name="Mexico" pageId="1" pageNumber="572">Mexico</collectingCountry>
|
|||
|
. Nothing is known about the distribution of any species in Central America.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8BB68732FFF4B10D4DF4AC62FEC7FAFB" blockId="1.[151,1437,824,1390]" pageId="1" pageNumber="572">
|
|||
|
An identification key to all known species of
|
|||
|
<taxonomicName id="4C09FCB1FFF4B10D4FD1AC62FCF6FB03" authorityName="Cameron" authorityYear="1902" box="[738,855,1148,1174]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B97D5B20FFF4B10D4FD1AC62FCF6FB03" box="[738,855,1148,1174]" italics="true" pageId="1" pageNumber="572">Glodianus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
is provided. However,
|
|||
|
<bibRefCitation id="EF98FAC3FFF4B10D4958AC63FAF2FB02" author="Townes, H. K." box="[1131,1363,1148,1175]" pageId="1" pageNumber="572" pagination="1 - 537" refId="ref7309" refString="Townes, H. K. (1970) The genera of Ichneumonidae. Part 2. Memoirs of the American Entomological Institute, 12, 1 - 537." type="journal article" year="1970">Townes (1970: 235)</bibRefCitation>
|
|||
|
noted that the genus is large; Central and South American faunas of
|
|||
|
<taxonomicName id="4C09FCB1FFF4B10D4E44ACBEFC4DFB2F" authorityName="Cameron" authorityYear="1902" box="[887,1004,1184,1210]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B97D5B20FFF4B10D4E44ACBEFC4DFB2F" box="[887,1004,1184,1210]" italics="true" pageId="1" pageNumber="572">Glodianus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
require revision, therefore additional comparison of species with their original descriptions or
|
|||
|
<typeStatus id="54B23990FFF4B10D4E38ACDBFCE7FB4A" box="[779,838,1221,1247]" pageId="1" pageNumber="572">types</typeStatus>
|
|||
|
is strongly recommended. Two species,
|
|||
|
<taxonomicName id="4C09FCB1FFF4B10D4836ACDAFF51FA97" authority="(Brulle)" baseAuthorityName="Brulle" baseAuthorityYear="1846" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="species" species="dimidiatus">
|
|||
|
<emphasis id="B97D5B20FFF4B10D4836ACDAFA3DFB4B" box="[1285,1436,1220,1246]" italics="true" pageId="1" pageNumber="572">G. dimidiatus</emphasis>
|
|||
|
(Brullé)
|
|||
|
</taxonomicName>
|
|||
|
and
|
|||
|
<taxonomicName id="4C09FCB1FFF4B10D4C1FACF6FDFFFA97" authority="Cameron" authorityName="Cameron" authorityYear="1911" box="[300,606,1256,1282]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="species" species="xanthostomus">
|
|||
|
<emphasis id="B97D5B20FFF4B10D4C1FACF6FE4AFA97" box="[300,491,1256,1282]" italics="true" pageId="1" pageNumber="572">G. xanthostomus</emphasis>
|
|||
|
Cameron
|
|||
|
</taxonomicName>
|
|||
|
, are known only from males, the
|
|||
|
<typeStatus id="54B23990FFF4B10D4ED4ACF7FBB6FA96" box="[999,1047,1257,1283]" pageId="1" pageNumber="572">type</typeStatus>
|
|||
|
of the former species is lost, and original descriptions of both species are brief and incomplete. Thus, these two species cannot be distinguished from each other until the South American fauna of the genus is revised, and therefore both species are placed in one couplet of the key.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|