treatments-xml/data/03/EF/40/03EF403E007AFFDAFB9CDBAEFDA21892.xml

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<document id="0DB44EC3C9072498BE6F69CBA335EFA2" ID-CLB-Dataset="3413" ID-DOI="10.1206/0003-0082(2003)417&lt;0001:CSOACF&gt;2.0.CO;2" ID-GBIF-Dataset="0464179d-9549-4885-9ddd-32ffef32942e" ID-ISSN="0003-0082" ID-Zenodo-Dep="4734880" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620067893553" checkinUser="felipe" docAuthor="BAIN, RAOUL H., LATHROP, AMY, MURPHY, ROBERT W., ORLOV, NIKOLAI L. &amp; CUC, HO THU" docDate="2003" docId="03EF403E007AFFDAFB9CDBAEFDA21892" docLanguage="en" docName="AmMusNovit.2003.3417.1-60.pdf" docOrigin="American Museum Novitates 3417" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0082%282003%29417%3C0001%3ACSOACF%3E2.0.CO%3B2" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Rana megatympanum BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC 2003, new species" docType="treatment" docVersion="8" lastPageNumber="53" masterDocId="FFD63846004BFFEEF972DA67FF941E5F" masterDocTitle="Cryptic Species of a Cascade Frog from Southeast Asia: Taxonomic Revisions and Descriptions of Six New Species" masterLastPageNumber="60" masterPageNumber="1" pageNumber="50" updateTime="1698938121846" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E31FBADFC22C67C3856409BE895697EC">Cryptic Species of a Cascade Frog from Southeast Asia: Taxonomic Revisions and Descriptions of Six New Species</mods:title>
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<mods:namePart id="BF1E511F5BD90FCA40F3C982E4A357EB">BAIN, RAOUL H.</mods:namePart>
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<mods:namePart id="9584A8024FD48AA429FD2DC8081A9E33">LATHROP, AMY</mods:namePart>
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<mods:namePart id="595BE2BE403D074400FAF839A52E502D">ORLOV, NIKOLAI L.</mods:namePart>
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<treatment id="03EF403E007AFFDAFB9CDBAEFDA21892" ID-DOI="http://doi.org/10.5281/zenodo.5713834" ID-GBIF-Taxon="190301392" ID-Zenodo-Dep="5713834" LSID="urn:lsid:plazi:treatment:03EF403E007AFFDAFB9CDBAEFDA21892" httpUri="http://treatment.plazi.org/id/03EF403E007AFFDAFB9CDBAEFDA21892" lastPageId="52" lastPageNumber="53" pageId="49" pageNumber="50">
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<paragraph id="8BF9F128007AFFDFFB9CDBAEFB0C1FBF" blockId="49.[750,1176,457,481]" box="[750,1176,457,481]" pageId="49" pageNumber="50">
<heading id="D0B14644007AFFDFFB9CDBAEFB0C1FBF" box="[750,1176,457,481]" centered="true" fontSize="36" level="2" pageId="49" pageNumber="50" reason="8">
<emphasis id="B9322D3A007AFFDFFB9CDBAEFB941FBE" bold="true" box="[750,1024,457,481]" pageId="49" pageNumber="50">
<taxonomicName id="4C468AAB007AFFDFFB9CDBAEFC6D1FBE" ID-CoL="4RDXR" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[750,1017,457,481]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="megatympanum" status="sp. nov.">
<emphasis id="B9322D3A007AFFDFFB9CDBAEFC6D1FBE" bold="true" box="[750,1017,457,481]" italics="true" pageId="49" pageNumber="50">Rana megatympanum</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2019041007AFFDFFD78DBAEFB0C1FBF" box="[1034,1176,457,480]" pageId="49" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BF9F128007AFFDFFB9FDB92FBC51C19" blockId="49.[749,1176,501,583]" pageId="49" pageNumber="50">
(Previously referred to as species 7, Large)
<figureCitation id="137DEDAD007AFFDFFA47D854FC501C19" box="[821,964,563,582]" captionStart="Fig" captionStartId="33.[149,187,1380,1401]" captionTargetBox="[131,1668,442,1357]" captionTargetId="figure-5@33.[131,1668,439,1358]" captionTargetPageId="33" captionText="Fig. 12. Heads of members of the Rana chloronota complex in lateral and dorsal views: (A, B) Rana chloronota, male (ROM 26402), and (C, D) female (ROM 26406); paratypes of (E, F) Rana bacboensis, new species (ROM 29529); (G, H) Rana daorum, new species (ROM 38538); (I, J) Rana hmongorum, new species (ROM 39891); (K, L) Rana morafkai, new species (ROM 39906); (M, N) Rana banaorum, new species (ROM39941); and (O, P) Rana megatympanum, new species (ROM 26398)." figureDoi="http://doi.org/10.5281/zenodo.4734907" httpUri="https://zenodo.org/record/4734907/files/figure.png" pageId="49" pageNumber="50">Figures 12O, P</figureCitation>
,
<figureCitation id="137DEDAD007AFFDFFAA3D854FC6D1C19" box="[977,1017,563,582]" captionStart="Fig" captionStartId="34.[110,148,1407,1428]" captionTargetBox="[92,1628,413,1387]" captionTargetId="figure-5@34.[92,1629,412,1387]" captionTargetPageId="34" captionText="Fig. 13. Hands and feet of members of the Rana chloronota complex: (A) Rana chloronota (ROM 26402); paratypes of (B) Rana bacboensis, new species (ROM 29529); (C) Rana daorum, new species (ROM 38538); (D) Rana hmongorum, new species (ROM 26379); (E) Rana morafkai, new species (ROM 39906); (F) Rana banaorum, new species (ROM 39941); and (G) Rana megatympanum, new species (ROM 26398). Scale equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4734911" httpUri="https://zenodo.org/record/4734911/files/figure.png" pageId="49" pageNumber="50">13G</figureCitation>
,
<figureCitation id="137DEDAD007AFFDFFD78D854FBC51C19" box="[1034,1105,563,582]" captionStart="Fig" captionStartId="41.[149,187,1715,1736]" captionTargetBox="[123,1227,187,1693]" captionTargetId="figure-10@41.[123,1227,187,1693]" captionTargetPageId="41" captionText="Fig. 14. Members of the Rana chloronota complex from Vietnam. A. Paratype of R. hmongorum, new species, male from Lao Cai Province. (B) Female, brown (night) phase, (C) female, green (day) phase, and (D) male of R. morafkai, new species, from Gia Lai Province. (E) Female and (F) male of R. banaorum, new species, from Gia Lai Province. (G, H) female of R. megatympanum new species." figureDoi="http://doi.org/10.5281/zenodo.4734915" httpUri="https://zenodo.org/record/4734915/files/figure.png" pageId="49" pageNumber="50">14G, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35CA2A3007AFFDFFBAAD83BFBE51B4A" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="8BF9F128007AFFDFFBAAD83BFC041D24" blockId="49.[699,1228,604,1741]" pageId="49" pageNumber="50">
<materialsCitation id="3B2EFB75007AFFDFFBAAD83BFC041D24" ID-GBIF-Occurrence="3408947338" accessionNumber="ROM 39684" collectingDate="1994-10-24" collectingDateMax="1994-10-29" collectingDateMin="1994-10-24" collectorName="I. S. Darevsky &amp; L. A. Lowcock &amp; R. W. Murphy &amp; N. L. Orlov" country="Vietnam" county="Con Cuong District" location="Khe Moi River" municipality="Con Cuong village" pageId="49" pageNumber="50" specimenCount="2" stateProvince="Nghe An" typeStatus="holotype">
<typeStatus id="54FD4F8A007AFFDFFBAAD83BFCDA1C2D" box="[728,846,604,627]" pageId="49" pageNumber="50" type="holotype">HOLOTYPE</typeStatus>
: (ROM field no. 12999) ROM 39684, a gravid adult female from
<location id="8E99A7F3007AFFDFFD12D81EFD681CF2" LSID="urn:lsid:plazi:treatment:03EF403E007AFFDAFB9CDBAEFDA21892:8E99A7F3007AFFDFFD12D81EFD681CF2" country="Vietnam" county="Con Cuong District" municipality="Con Cuong village" name="Khe Moi River" pageId="49" pageNumber="50" stateProvince="Nghe An">Khe Moi River</location>
, approximately
<quantity id="4CBE5CCD007AFFDFFABBD8F1FB881CF2" box="[969,1052,662,685]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="49" pageNumber="50" unit="km" value="24.0">24 km</quantity>
west of
<collectingMunicipality id="6B9D6B52007AFFDFFDE8D8F1FCCB1C94" pageId="49" pageNumber="50">Con Cuong village</collectingMunicipality>
(by road),
<collectingCounty id="629889A4007AFFDFFA93D8D3FB531C94" box="[993,1223,692,715]" pageId="49" pageNumber="50">Con Cuong District</collectingCounty>
,
<collectingRegion id="49823FCA007AFFDFFBC9D8B6FC3E1CB7" box="[699,938,721,744]" country="Vietnam" name="Nghe An" pageId="49" pageNumber="50">Nghe An Province</collectingRegion>
,
<collectingCountry id="F351B1B8007AFFDFFACDD8B6FBB61CB7" box="[959,1058,721,744]" name="Vietnam" pageId="49" pageNumber="50">Vietnam</collectingCountry>
(18°56̍30̎N, 104°48̍35̎E) found between
<date id="FFF8D7E8007AFFDFFD69D889FCAE1D7C" pageId="49" pageNumber="50" value="1994-10-24" valueMax="1994-10-29" valueMin="1994-10-24">
<collectingDate id="EFBC2E00007AFFDFFD69D889FCAE1D7C" pageId="49" pageNumber="50" value="1994-10-24" valueMax="1994-10-29" valueMin="1994-10-24">24 and 29 October 1994</collectingDate>
</date>
by
<collectorName id="26B394FE007AFFDFFA18D96BFB991D7C" box="[874,1037,780,803]" pageId="49" pageNumber="50">I.S. Darevsky</collectorName>
,
<collectorName id="26B394FE007AFFDFFD68D96BFB531D7C" box="[1050,1223,780,803]" pageId="49" pageNumber="50">L.A. Lowcock</collectorName>
,
<collectorName id="26B394FE007AFFDFFBC9D94EFCC91D1F" box="[699,861,809,832]" pageId="49" pageNumber="50">R.W. Murphy</collectorName>
, and
<collectorName id="26B394FE007AFFDFFAECD94EFBC81D1F" box="[926,1116,809,832]" pageId="49" pageNumber="50">N.L. Orlov. The</collectorName>
<typeStatus id="54FD4F8A007AFFDFFD17D94EFB5F1D1F" box="[1125,1227,809,832]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
had leg and liver tissue removed shortly after it was euthanised.
</materialsCitation>
</paragraph>
<paragraph id="8BF9F128007AFFDFFBAAD9E6FBE51B4A" blockId="49.[699,1228,604,1741]" pageId="49" pageNumber="50">
<materialsCitation id="3B2EFB75007AFFDFFBAAD9E6FC901A52" ID-GBIF-Occurrence="3408947310" accessionNumber="ROM 39263, ROM 39237, ROM 26398" collectingDate="1994-10-24" collectingDateMax="1996-05-27" collectingDateMin="1994-10-24" collectorName="L. A. Lowcock &amp; R. W. Murphy &amp; N. L. Orlov" country="Vietnam" county="Con Cuong District" location="Eight" municipality="Na Hang Nature Reserve" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Tuyen Quang" typeStatus="paratype">
<typeStatus id="54FD4F8A007AFFDFFBAAD9E6FCC21DC8" box="[728,854,897,920]" pageId="49" pageNumber="50" type="paratype">PARATYPES</typeStatus>
:
<location id="8E99A7F3007AFFDFFA1FD9E6FC3F1DC7" LSID="urn:lsid:plazi:treatment:03EF403E007AFFDAFB9CDBAEFDA21892:8E99A7F3007AFFDFFA1FD9E6FC3F1DC7" box="[877,939,897,920]" country="Vietnam" county="Con Cuong District" municipality="Na Hang Nature Reserve" name="Eight" pageId="49" pageNumber="50" stateProvince="Tuyen Quang">Eight</location>
females (ROM 39263, 3968539691) collected with holotype between
<date id="FFF8D7E8007AFFDFFA79D9DBFBA51D8C" box="[779,1073,956,979]" pageId="49" pageNumber="50" value="1994-10-24" valueMax="1994-10-29" valueMin="1994-10-24">
<collectingDate id="EFBC2E00007AFFDFFA79D9DBFBA51D8C" box="[779,1073,956,979]" pageId="49" pageNumber="50" value="1994-10-24" valueMax="1994-10-29" valueMin="1994-10-24">24 and 29 October 1994</collectingDate>
</date>
by
<collectorName id="26B394FE007AFFDFFD17D9DBFC941DAF" pageId="49" pageNumber="50">I.S. Darevsky</collectorName>
,
<collectorName id="26B394FE007AFFDFFA78D9BEFC271DAF" box="[778,947,985,1008]" pageId="49" pageNumber="50">L.A. Lowcock</collectorName>
,
<collectorName id="26B394FE007AFFDFFACCD9BEFBC81DAF" box="[958,1116,985,1008]" pageId="49" pageNumber="50">R.W. Murphy</collectorName>
, and
<collectorName id="26B394FE007AFFDFFDE5D9BEFC901A52" pageId="49" pageNumber="50">N.L. Orlov.</collectorName>
</materialsCitation>
<materialsCitation id="3B2EFB75007AFFDFFA79D991FB891AFF" ID-GBIF-Occurrence="3408947322" accessionNumber="ROM 39263, ROM 39237, ROM 26398" collectingDate="1994-10-24" collectingDateMax="1996-05-27" collectingDateMin="1994-10-24" collectorName="A. Lathrop &amp; R. W. Murphy " country="Vietnam" county="Con Cuong District" location="Eight" municipality="Na Hang Nature Reserve" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Tuyen Quang" typeStatus="paratype">
Four males (ROM 3923739240) from
<collectingMunicipality id="6B9D6B52007AFFDFFBC9DE73FC4A1A74" box="[699,990,1044,1067]" pageId="49" pageNumber="50">Na Hang Nature Reserve</collectingMunicipality>
,
<collectingRegion id="49823FCA007AFFDFFA98DE73FD651A17" country="Vietnam" name="Tuyen Quang" pageId="49" pageNumber="50">Tuyen Quang Province</collectingRegion>
,
<collectingCountry id="F351B1B8007AFFDFFA71DE56FCFF1A17" box="[771,875,1073,1096]" name="Vietnam" pageId="49" pageNumber="50">Vietnam</collectingCountry>
(22°21̍54̎N, 105°25̍40̎E) found along waterfalls between
<date id="FFF8D7E8007AFFDFFD38DE29FCA01ADC" pageId="49" pageNumber="50" value="1996-05-26" valueMax="1996-05-27" valueMin="1996-05-26">
<collectingDate id="EFBC2E00007AFFDFFD38DE29FCA01ADC" pageId="49" pageNumber="50" value="1996-05-26" valueMax="1996-05-27" valueMin="1996-05-26">26 and 27 May 1996</collectingDate>
</date>
by
<collectorName id="26B394FE007AFFDFFA1ADE0BFC7E1ADC" box="[872,1002,1132,1155]" pageId="49" pageNumber="50">A. Lathrop</collectorName>
and
<collectorName id="26B394FE007AFFDFFD59DE0BFB5F1ADC" box="[1067,1227,1132,1155]" pageId="49" pageNumber="50">R.W. Murphy</collectorName>
between 193 and 0200 hours.
</materialsCitation>
<materialsCitation id="3B2EFB75007AFFDFFD5ADEEEFBFA1B4A" ID-GBIF-Occurrence="3408947308" accessionNumber="ROM 39263, ROM 39237, ROM 26398" collectingDate="1994-10-24" collectingDateMax="1996-05-27" collectingDateMin="1994-10-24" collectorName="B. Hubley &amp; A. Lathrop &amp; R. W. Murph &amp; N. L. Orlov" country="Vietnam" county="Con Cuong District" location="Eight" municipality="Na Hang Nature Reserve" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Tuyen Quang" typeStatus="paratype">
ROM 26398 26400
<collectingCounty id="629889A4007AFFDFFA62DEC1FC6B1AE2" box="[784,1023,1190,1213]" pageId="49" pageNumber="50">Con Cuong District</collectingCounty>
,
<collectingRegion id="49823FCA007AFFDFFD62DEC1FD641A84" country="Vietnam" name="Nghe An" pageId="49" pageNumber="50">Nghe An Province</collectingRegion>
,
<collectingCountry id="F351B1B8007AFFDFFA70DEA3FCFC1A84" box="[770,872,1220,1243]" name="Vietnam" pageId="49" pageNumber="50">Vietnam</collectingCountry>
(18°56̍30̎N, 104°48̍35̎E), collected
<date id="FFF8D7E8007AFFDFFA5FDE86FC2A1AA7" box="[813,958,1249,1272]" pageId="49" pageNumber="50" value="1995-06-05">
<collectingDate id="EFBC2E00007AFFDFFA5FDE86FC2A1AA7" box="[813,958,1249,1272]" pageId="49" pageNumber="50" value="1995-06-05">5 June 1995</collectingDate>
</date>
by
<collectorName id="26B394FE007AFFDFFA80DE86FBF91AA7" box="[1010,1133,1249,1272]" pageId="49" pageNumber="50">B. Hubley</collectorName>
,
<collectorName id="26B394FE007AFFDFFD0FDE86FD6E1B4A" pageId="49" pageNumber="50">A. Lathrop</collectorName>
,
<collectorName id="26B394FE007AFFDFFA7ADE99FC3C1B4A" box="[776,936,1278,1301]" pageId="49" pageNumber="50">R.W. Murphy</collectorName>
, and
<collectorName id="26B394FE007AFFDFFA98DE99FBFA1B4A" box="[1002,1134,1278,1301]" pageId="49" pageNumber="50">N.L. Orlov</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35CA2A3007AFFDDFBAADF7BFF401D8C" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="8BF9F128007AFFDCFBAADF7BFF511CCF" blockId="49.[699,1228,604,1741]" lastBlockId="50.[124,653,193,1742]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
DIAGNOSIS:
<taxonomicName id="4C468AAB007AFFDFFA10DF7BFBF51B6C" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[866,1121,1308,1331]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A007AFFDFFA10DF7BFBF51B6C" box="[866,1121,1308,1331]" italics="true" pageId="49" pageNumber="50">Rana megatympanum</emphasis>
</taxonomicName>
, a member of the subgenus
<taxonomicName id="4C468AAB007AFFDFFAC0DF5EFBB21B0F" authorityName="Fei, Ye, and Huang" authorityYear="1990" box="[946,1062,1337,1360]" class="Amphibia" family="Ranidae" genus="Odorrana" kingdom="Animalia" order="Anura" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B9322D3A007AFFDFFAC0DF5EFBB21B0F" box="[946,1062,1337,1360]" italics="true" pageId="49" pageNumber="50">Odorrana</emphasis>
</taxonomicName>
(sensu
<bibRefCitation id="EFD78CD9007AFFDFFDF4DF5EFCB21B32" author="Fei, L." pageId="49" pageNumber="50" refId="ref37298" refString="Fei, L., C. - y. Ye, and Y. - z. Huang. 1990. Keys to Chinese Amphibians. Chongqing: Chongqing Branch, Science and Technology Materials Press, 364 pp. [in Chinese]" type="book" year="1990">Fei et al., 1990</bibRefCitation>
), is characterized by a combination of the following attributes: (1) body dorsoventrally compressed; (2) SVL means of males
<quantity id="4CBE5CCD007AFFDFFA7ADFC9FCCF1B9A" box="[776,859,1454,1477]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="49" pageNumber="50" unit="mm" value="52.0">52 mm</quantity>
(
<quantity id="4CBE5CCD007AFFDFFA1EDFC9FC651B9A" box="[876,1009,1454,1477]" metricMagnitude="-2" metricUnit="m" metricValue="5.15" metricValueMax="5.5" metricValueMin="4.8" pageId="49" pageNumber="50" unit="mm" value="51.5" valueMax="55.0" valueMin="48.0">4855 mm</quantity>
), females
<quantity id="4CBE5CCD007AFFDFFD18DFC9FB5F1B9A" box="[1130,1227,1454,1477]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="49" pageNumber="50" unit="mm" value="100.0">100 mm</quantity>
(
<quantity id="4CBE5CCD007AFFDFFBB1DFABFCC91BBC" box="[707,861,1484,1507]" metricMagnitude="-2" metricUnit="m" metricValue="9.9" metricValueMax="10.5" metricValueMin="9.3" pageId="49" pageNumber="50" unit="mm" value="99.0" valueMax="105.0" valueMin="93.0">93105 mm</quantity>
); (3) vomerine teeth in rows oblique to choanae; (4) yellow lip­stripe present in males, absent or indistinct in females; (5) head broad, bluntly rounded in profile; (6) tympanum round, distinct, TMP:EYE in males enormous (1.20), greater than in females (0.51); (7) supratympanic fold weak; (8) dorsal skin shagreened, dorsolateral folds present only in males; (9) dorsum olive to brown sometimes with black spots; flanks marbled yellow and brown­gray; forelimbs and hindlimbs barred; (10) ventromedial callous pad on fingers II, III, and IV to proximal tubercle, fringes on fingers II, III, and IV; (11) disks on fingers and toes greatly enlarged (&gt;2× base of phalanges); (12) feet fully webbed to toe disk, lateral fringes on I and V to terminal phalanges, webbing brown; (13) subarticular tubercles and an internal metatarsal tubercle distinct, conical; (14) terminal phalanges T­ shaped; (15) xiphisternum large, deeply notched posteriorly; (16) males with nuptial pads, paired gular pouches, pectoral spines absent; (17) eggs white.
</paragraph>
<paragraph id="8BF9F1280079FFDDF9EBD8F1FF401D8C" blockId="50.[124,653,193,1742]" lastBlockId="51.[123,652,193,1741]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
COMPARISONS:
<taxonomicName id="4C468AAB0079FFDCF822D8F0FDC31CF1" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[336,599,663,686]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCF822D8F0FDC31CF1" box="[336,599,663,686]" italics="true" pageId="50" pageNumber="51">Rana megatympanum</emphasis>
</taxonomicName>
superficially resembles other Asian cascade ranids, including
<taxonomicName id="4C468AAB0079FFDCF841D8B6FE561CB7" baseAuthorityName="Yang" baseAuthorityYear="1991" box="[307,450,721,744]" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="nasica">
<emphasis id="B9322D3A0079FFDCF841D8B6FE561CB7" box="[307,450,721,744]" italics="true" pageId="50" pageNumber="51">Huia nasica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF8A3D8B6FF181D59" authorityName="Boulenger" authorityYear="1882" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0079FFDCF8A3D8B6FF181D59" italics="true" pageId="50" pageNumber="51">Rana andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF9D0D888FECF1D59" authorityName="Inger &amp; Chan-ard" authorityYear="1997" box="[162,347,751,774]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="archotaphus">
<emphasis id="B9322D3A0079FFDCF9D0D888FECF1D59" box="[162,347,751,774]" italics="true" pageId="50" pageNumber="51">R. archotaphus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF803D888FD8E1D59" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[369,538,751,774]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0079FFDCF803D888FD8E1D59" box="[369,538,751,774]" italics="true" pageId="50" pageNumber="51">R. bacboensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB42D888FF5C1D7C" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="banaorum">
<emphasis id="B9322D3A0079FFDCFB42D888FF5C1D7C" italics="true" pageId="50" pageNumber="51">R. banaorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF9A5D96BFEED1D7C" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[215,377,780,803]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0079FFDCF9A5D96BFEED1D7C" box="[215,377,780,803]" italics="true" pageId="50" pageNumber="51">R. chalconota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF8FAD96BFDBC1D7C" box="[392,552,780,803]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A0079FFDCF8FAD96BFDBC1D7C" box="[392,552,780,803]" italics="true" pageId="50" pageNumber="51">R. chloronota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB45D96BFF391D1F" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="daorum">
<emphasis id="B9322D3A0079FFDCFB45D96BFF391D1F" italics="true" pageId="50" pageNumber="51">R. daorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF9A6D94EFEE21D1F" authorityName="Boulenger" authorityYear="1917" box="[212,374,809,832]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="grahami">
<emphasis id="B9322D3A0079FFDCF9A6D94EFEE21D1F" box="[212,374,809,832]" italics="true" pageId="50" pageNumber="51">R. grahami</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF8EED94EFDD81D1F" authorityName=": Boulenger" authorityYear="1899" box="[412,588,809,832]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B9322D3A0079FFDCF8EED94EFDD81D1F" box="[412,588,809,832]" italics="true" pageId="50" pageNumber="51">R. graminea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB00D94EFE931D01" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A0079FFDCFB00D94EFE931D01" italics="true" pageId="50" pageNumber="51">R. hainanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF865D920FE5D1D01" authorityName="Y.u.Deng" authorityYear="1992" box="[279,457,839,862]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hejiangensis">
<emphasis id="B9322D3A0079FFDCF865D920FE5D1D01" box="[279,457,839,862]" italics="true" pageId="50" pageNumber="51">R. hejiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF8AAD920FD111D01" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[472,645,839,862]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hmongorum">
<emphasis id="B9322D3A0079FFDCF8AAD920FD111D01" box="[472,645,839,862]" italics="true" pageId="50" pageNumber="51">R. hmongorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF90ED903FF4F1D24" authorityName="Boulenger" authorityYear="1891" box="[124,219,868,891]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hosii">
<emphasis id="B9322D3A0079FFDCF90ED903FF4F1D24" box="[124,219,868,891]" italics="true" pageId="50" pageNumber="51">R. hosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF983D903FE151D24" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[241,385,868,891]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="morafkai">
<emphasis id="B9322D3A0079FFDCF983D903FE151D24" box="[241,385,868,891]" italics="true" pageId="50" pageNumber="51">R. morafkai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF8E4D903FDCA1D24" box="[406,606,868,891]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0079FFDCF8E4D903FDCA1D24" box="[406,606,868,891]" italics="true" pageId="50" pageNumber="51">R. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB01D903FE9C1DC7" baseAuthorityName="Huang, Fei &amp; Ye" baseAuthorityYear="2001" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="junlianensis">
<emphasis id="B9322D3A0079FFDCFB01D903FE9C1DC7" italics="true" pageId="50" pageNumber="51">R. junlianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF868D9E6FE711DC7" box="[282,485,897,920]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="kwangwuensis">
<emphasis id="B9322D3A0079FFDCF868D9E6FE711DC7" box="[282,485,897,920]" italics="true" pageId="50" pageNumber="51">R. kwangwuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF885D9E6FD121DC7" authorityName=": Werner" authorityYear="1930" box="[503,646,897,920]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="leporipes">
<emphasis id="B9322D3A0079FFDCF885D9E6FD121DC7" box="[503,646,897,920]" italics="true" pageId="50" pageNumber="51">R. leporipes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF90ED9F8FF701DE9" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[124,228,927,950]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="livida">
<emphasis id="B9322D3A0079FFDCF90ED9F8FF701DE9" box="[124,228,927,950]" italics="true" pageId="50" pageNumber="51">R. livida</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF985D9F8FE301DE9" authorityName="Liu" authorityYear="1950" box="[247,420,927,950]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="margaretae">
<emphasis id="B9322D3A0079FFDCF985D9F8FE301DE9" box="[247,420,927,950]" italics="true" pageId="50" pageNumber="51">R. margaretae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF8C5D9F8FDF41DE9" authorityName="Boettger" authorityYear="1892" box="[439,608,927,950]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="schmackeri">
<emphasis id="B9322D3A0079FFDCF8C5D9F8FDF41DE9" box="[439,608,927,950]" italics="true" pageId="50" pageNumber="51">R. schmackeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB01D9F8FF551D8C" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A0079FFDCFB01D9F8FF551D8C" italics="true" pageId="50" pageNumber="51">R. sinica</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0079FFDCF877D9DBFE081D8C" box="[261,412,956,979]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="tiannensis">
<emphasis id="B9322D3A0079FFDCF877D9DBFE081D8C" box="[261,412,956,979]" italics="true" pageId="50" pageNumber="51">R. tiannensis</emphasis>
</taxonomicName>
(table 12). It can be differentiated from all other
<taxonomicName id="4C468AAB0079FFDCF8BBD9BEFDA91DAF" authorityName="Fei, Ye, and Huang" authorityYear="1990" box="[457,573,985,1008]" class="Amphibia" family="Ranidae" genus="Odorrana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="genus">
<emphasis id="B9322D3A0079FFDCF8BBD9BEFDA91DAF" box="[457,573,985,1008]" italics="true" pageId="50" pageNumber="51">Odorrana</emphasis>
</taxonomicName>
by the enormous sexual dimorphism in tympanum size (TMP:EYE
<quantity id="4CBE5CCD0079FFDCF839DE73FE361A74" box="[331,418,1044,1067]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="50" pageNumber="51" unit="in" value="1.2">1.20 in</quantity>
males,
<quantity id="4CBE5CCD0079FFDCFB7ADE73FDCB1A74" box="[520,607,1044,1067]" metricMagnitude="-2" metricUnit="m" metricValue="1.2954" pageId="50" pageNumber="51" unit="in" value="0.51">0.51 in</quantity>
females). Male
<taxonomicName id="4C468AAB0079FFDCF853DE56FE691A17" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[289,509,1073,1096]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCF853DE56FE691A17" box="[289,509,1073,1096]" italics="true" pageId="50" pageNumber="51">R. megatympanum</emphasis>
</taxonomicName>
have a yellow lip­stripe and females have an indistinct or absent yellow lip­stripe, differentiating it from
<taxonomicName id="4C468AAB0079FFDCF9B3DEEEFEA21AFF" baseAuthorityName="Yang" baseAuthorityYear="1991" box="[193,310,1161,1184]" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="nasica">
<emphasis id="B9322D3A0079FFDCF9B3DEEEFEA21AFF" box="[193,310,1161,1184]" italics="true" pageId="50" pageNumber="51">H. nasica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF838DEEEFD951AFF" authorityName="Inger &amp; Chan-ard" authorityYear="1997" box="[330,513,1161,1184]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="archotaphus">
<emphasis id="B9322D3A0079FFDCF838DEEEFD951AFF" box="[330,513,1161,1184]" italics="true" pageId="50" pageNumber="51">R. archotaphus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB67DEEEFF3F1AE1" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="banaorum">
<emphasis id="B9322D3A0079FFDCFB67DEEEFF3F1AE1" italics="true" pageId="50" pageNumber="51">R. banaorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF9CDDEC0FEF11AE1" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[191,357,1191,1214]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0079FFDCF9CDDEC0FEF11AE1" box="[191,357,1191,1214]" italics="true" pageId="50" pageNumber="51">R. chalconota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF80BDEC0FD8B1AE1" box="[377,543,1191,1214]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A0079FFDCF80BDEC0FD8B1AE1" box="[377,543,1191,1214]" italics="true" pageId="50" pageNumber="51">R. chloronota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB40DEC0FF3F1A84" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="daorum">
<emphasis id="B9322D3A0079FFDCFB40DEC0FF3F1A84" italics="true" pageId="50" pageNumber="51">R. daorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF9CFDEA3FEC41A84" authorityName=": Boulenger" authorityYear="1899" box="[189,336,1220,1243]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B9322D3A0079FFDCF9CFDEA3FEC41A84" box="[189,336,1220,1243]" italics="true" pageId="50" pageNumber="51">R. graminea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF810DEA3FD831A84" authorityName="Y.u.Deng" authorityYear="1992" box="[354,535,1220,1243]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hejiangensis">
<emphasis id="B9322D3A0079FFDCF810DEA3FD831A84" box="[354,535,1220,1243]" italics="true" pageId="50" pageNumber="51">R. hejiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB5BDEA3FD121A84" authorityName="Boulenger" authorityYear="1891" box="[553,646,1220,1243]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hosii">
<emphasis id="B9322D3A0079FFDCFB5BDEA3FD121A84" box="[553,646,1220,1243]" italics="true" pageId="50" pageNumber="51">R. hosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF90EDE86FE9E1AA7" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[124,266,1249,1272]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="morafkai">
<emphasis id="B9322D3A0079FFDCF90EDE86FE9E1AA7" box="[124,266,1249,1272]" italics="true" pageId="50" pageNumber="51">R. morafkai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF86EDE86FE381AA7" authorityName=": Werner" authorityYear="1930" box="[284,428,1249,1272]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="leporipes">
<emphasis id="B9322D3A0079FFDCF86EDE86FE381AA7" box="[284,428,1249,1272]" italics="true" pageId="50" pageNumber="51">R. leporipes</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0079FFDCF886DE86FDCF1AA7" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[500,603,1249,1272]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="livida">
<emphasis id="B9322D3A0079FFDCF886DE86FDCF1AA7" box="[500,603,1249,1272]" italics="true" pageId="50" pageNumber="51">R. livida</emphasis>
</taxonomicName>
(all with white lip­stripes);
<taxonomicName id="4C468AAB0079FFDCF8FFDE98FDBE1B49" authorityName="Boulenger" authorityYear="1882" box="[397,554,1279,1302]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0079FFDCF8FFDE98FDBE1B49" box="[397,554,1279,1302]" italics="true" pageId="50" pageNumber="51">R. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB4BDE98FF471B6C" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0079FFDCFB4BDE98FF471B6C" italics="true" pageId="50" pageNumber="51">R. bacboensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF996DF7BFE051B6C" box="[228,401,1308,1331]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A0079FFDCF996DF7BFE051B6C" box="[228,401,1308,1331]" italics="true" pageId="50" pageNumber="51">R. hainanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF8D3DF7BFDF71B6C" box="[417,611,1308,1331]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0079FFDCF8D3DF7BFDF71B6C" box="[417,611,1308,1331]" italics="true" pageId="50" pageNumber="51">R. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB01DF7BFE901B0F" authorityName="Liu" authorityYear="1950" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="margaretae">
<emphasis id="B9322D3A0079FFDCFB01DF7BFE901B0F" italics="true" pageId="50" pageNumber="51">R. margaretae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0079FFDCF83FDF5EFE7C1B0F" box="[333,488,1337,1360]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="tiannensis">
<emphasis id="B9322D3A0079FFDCF83FDF5EFE7C1B0F" box="[333,488,1337,1360]" italics="true" pageId="50" pageNumber="51">R. tiannensis</emphasis>
</taxonomicName>
have vertical lip­bars;
<taxonomicName id="4C468AAB0079FFDCF99FDF30FE371B31" baseAuthorityName="Huang, Fei &amp; Ye" baseAuthorityYear="2001" box="[237,419,1367,1390]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="junlianensis">
<emphasis id="B9322D3A0079FFDCF99FDF30FE371B31" box="[237,419,1367,1390]" italics="true" pageId="50" pageNumber="51">R. junlianensis</emphasis>
</taxonomicName>
has a yellow lipstripe with brown lip­bars;
<taxonomicName id="4C468AAB0079FFDCF8C8DF13FDCB1BD4" authorityName="Boettger" authorityYear="1892" box="[442,607,1396,1419]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="schmackeri">
<emphasis id="B9322D3A0079FFDCF8C8DF13FDCB1BD4" box="[442,607,1396,1419]" italics="true" pageId="50" pageNumber="51">R. schmackeri</emphasis>
</taxonomicName>
has no lip­stripe or vertical lip­bars. The broad, rounded snout differentiates
<taxonomicName id="4C468AAB0079FFDCF8BFDFC8FF341BBC" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCF8BFDFC8FF341BBC" italics="true" pageId="50" pageNumber="51">R. megatympanum</emphasis>
</taxonomicName>
from
<taxonomicName id="4C468AAB0079FFDCF982DFABFEF11BBC" baseAuthorityName="Yang" baseAuthorityYear="1991" box="[240,357,1484,1507]" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="nasica">
<emphasis id="B9322D3A0079FFDCF982DFABFEF11BBC" box="[240,357,1484,1507]" italics="true" pageId="50" pageNumber="51">H. nasica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF80ADFABFD8C1BBC" authorityName="Boulenger" authorityYear="1882" box="[376,536,1484,1507]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0079FFDCF80ADFABFD8C1BBC" box="[376,536,1484,1507]" italics="true" pageId="50" pageNumber="51">R. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB5EDFABFF5F185F" baseAuthorityName="Schlegel" baseAuthorityYear="1837" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0079FFDCFB5EDFABFF5F185F" italics="true" pageId="50" pageNumber="51">R. chalconota</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0079FFDCF87EDF8EFE25185F" authorityName="Boettger" authorityYear="1892" box="[268,433,1513,1536]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="schmackeri">
<emphasis id="B9322D3A0079FFDCF87EDF8EFE25185F" box="[268,433,1513,1536]" italics="true" pageId="50" pageNumber="51">R. schmackeri</emphasis>
</taxonomicName>
(obtusely pointed) and from
<taxonomicName id="4C468AAB0079FFDCF98FDC60FE001841" authorityName=": Boulenger" authorityYear="1899" box="[253,404,1543,1566]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B9322D3A0079FFDCF98FDC60FE001841" box="[253,404,1543,1566]" italics="true" pageId="50" pageNumber="51">R. graminea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C468AAB0079FFDCF8AFDC60FD191841" authorityName="Liu" authorityYear="1950" box="[477,653,1543,1566]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="margaretae">
<emphasis id="B9322D3A0079FFDCF8AFDC60FD191841" box="[477,653,1543,1566]" italics="true" pageId="50" pageNumber="51">R. margaretae</emphasis>
</taxonomicName>
(depressed). Its gular pouches distinguish
<taxonomicName id="4C468AAB0079FFDCFB01DC43FEA01807" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFB01DC43FEA01807" italics="true" pageId="50" pageNumber="51">R. megatympanum</emphasis>
</taxonomicName>
from
<taxonomicName id="4C468AAB0079FFDCF8F2DC26FD8A1807" authorityName="Boulenger" authorityYear="1882" box="[384,542,1601,1624]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0079FFDCF8F2DC26FD8A1807" box="[384,542,1601,1624]" italics="true" pageId="50" pageNumber="51">R. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB5DDC26FF441829" baseAuthorityName="Schlegel" baseAuthorityYear="1837" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0079FFDCFB5DDC26FF441829" italics="true" pageId="50" pageNumber="51">R. chalconota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF998DC38FEEB1829" authorityName="Boulenger" authorityYear="1917" box="[234,383,1631,1654]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="grahami">
<emphasis id="B9322D3A0079FFDCF998DC38FEEB1829" box="[234,383,1631,1654]" italics="true" pageId="50" pageNumber="51">R. grahami</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF8EADC38FDCE1829" box="[408,602,1631,1654]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A0079FFDCF8EADC38FDCE1829" box="[408,602,1631,1654]" italics="true" pageId="50" pageNumber="51">R. hainanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB00DC38FE9C18CC" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hmongorum">
<emphasis id="B9322D3A0079FFDCFB00DC38FE9C18CC" italics="true" pageId="50" pageNumber="51">R. hmongorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF86DDC1BFEEA18CC" authorityName="Boulenger" authorityYear="1891" box="[287,382,1660,1683]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hosii">
<emphasis id="B9322D3A0079FFDCF86DDC1BFEEA18CC" box="[287,382,1660,1683]" italics="true" pageId="50" pageNumber="51">R. hosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF8E7DC1BFDC918CC" box="[405,605,1660,1683]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0079FFDCF8E7DC1BFDC918CC" box="[405,605,1660,1683]" italics="true" pageId="50" pageNumber="51">R. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB01DC1BFE9C18EF" baseAuthorityName="Huang, Fei &amp; Ye" baseAuthorityYear="2001" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="junlianensis">
<emphasis id="B9322D3A0079FFDCFB01DC1BFE9C18EF" italics="true" pageId="50" pageNumber="51">R. junlianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCF869DCFEFE7C18EF" box="[283,488,1689,1712]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="kwangwuensis">
<emphasis id="B9322D3A0079FFDCF869DCFEFE7C18EF" box="[283,488,1689,1712]" italics="true" pageId="50" pageNumber="51">R. kwangwuensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0079FFDCFB42DCFEFF411891" authorityName="Liu" authorityYear="1950" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="margaretae">
<emphasis id="B9322D3A0079FFDCFB42DCFEFF411891" italics="true" pageId="50" pageNumber="51">R. margaretae</emphasis>
</taxonomicName>
. Dorsolateral folds immediately differentiate male
<taxonomicName id="4C468AAB0079FFDCFA0DDAA6FBCA1E87" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[895,1118,193,216]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFA0DDAA6FBCA1E87" box="[895,1118,193,216]" italics="true" pageId="50" pageNumber="51">R. megatympanum</emphasis>
</taxonomicName>
from
<taxonomicName id="4C468AAB0079FFDCFDC1DAA6FCAC1EA9" authorityName="Boulenger" authorityYear="1882" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0079FFDCFDC1DAA6FCAC1EA9" italics="true" pageId="50" pageNumber="51">R. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA38DAB8FC7A1EA9" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[842,1006,223,246]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0079FFDCFA38DAB8FC7A1EA9" box="[842,1006,223,246]" italics="true" pageId="50" pageNumber="51">R. bacboensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFD72DAB8FB361EA9" box="[1024,1186,223,246]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A0079FFDCFD72DAB8FB361EA9" box="[1024,1186,223,246]" italics="true" pageId="50" pageNumber="51">R. chloronota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFDC1DAB8FCD31F4C" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A0079FFDCFDC1DAB8FCD31F4C" italics="true" pageId="50" pageNumber="51">R. hainanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA25DA9BFB9E1F4C" authorityName="Y.u.Deng" authorityYear="1992" box="[855,1034,252,275]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hejiangensis">
<emphasis id="B9322D3A0079FFDCFA25DA9BFB9E1F4C" box="[855,1034,252,275]" italics="true" pageId="50" pageNumber="51">R. hejiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFD68DA9BFB301F4C" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[1050,1188,252,275]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="morafkai">
<emphasis id="B9322D3A0079FFDCFD68DA9BFB301F4C" box="[1050,1188,252,275]" italics="true" pageId="50" pageNumber="51">R. morafkai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFDC1DA9BFCC91F6F" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0079FFDCFDC1DA9BFCC91F6F" italics="true" pageId="50" pageNumber="51">R. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA0ADB7EFBA51F6F" baseAuthorityName="Huang, Fei &amp; Ye" baseAuthorityYear="2001" box="[888,1073,281,304]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="junlianensis">
<emphasis id="B9322D3A0079FFDCFA0ADB7EFBA51F6F" box="[888,1073,281,304]" italics="true" pageId="50" pageNumber="51">R. junlianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFD39DB7EFC8C1F11" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="kwangwuensis">
<emphasis id="B9322D3A0079FFDCFD39DB7EFC8C1F11" italics="true" pageId="50" pageNumber="51">R. kwangwuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA58DB50FC041F11" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[810,912,311,334]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="livida">
<emphasis id="B9322D3A0079FFDCFA58DB50FC041F11" box="[810,912,311,334]" italics="true" pageId="50" pageNumber="51">R. livida</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFAD1DB50FBDE1F11" authorityName="Boettger" authorityYear="1892" box="[931,1098,311,334]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="schmackeri">
<emphasis id="B9322D3A0079FFDCFAD1DB50FBDE1F11" box="[931,1098,311,334]" italics="true" pageId="50" pageNumber="51">R. schmackeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFD2FDB50FB511F11" box="[1117,1221,311,334]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A0079FFDCFD2FDB50FB511F11" box="[1117,1221,311,334]" italics="true" pageId="50" pageNumber="51">R. sinica</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0079FFDCFB86DB33FC051F34" box="[756,913,340,363]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="tiannensis">
<emphasis id="B9322D3A0079FFDCFB86DB33FC051F34" box="[756,913,340,363]" italics="true" pageId="50" pageNumber="51">R. tiannensis</emphasis>
</taxonomicName>
. The dorsolateral fold of male
<taxonomicName id="4C468AAB0079FFDCFB8DDB16FC4F1FD7" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[767,987,369,392]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFB8DDB16FC4F1FD7" box="[767,987,369,392]" italics="true" pageId="50" pageNumber="51">R. megatympanum</emphasis>
</taxonomicName>
differs from that of
<taxonomicName id="4C468AAB0079FFDCFBCEDBE8FCAB1FF9" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[700,831,399,422]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="daorum">
<emphasis id="B9322D3A0079FFDCFBCEDBE8FCAB1FF9" box="[700,831,399,422]" italics="true" pageId="50" pageNumber="51">R. daorum</emphasis>
</taxonomicName>
, which is composed of minute white granules,
<taxonomicName id="4C468AAB0079FFDCFAF2DBCBFBA51F9C" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[896,1073,428,451]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hmongorum">
<emphasis id="B9322D3A0079FFDCFAF2DBCBFBA51F9C" box="[896,1073,428,451]" italics="true" pageId="50" pageNumber="51">R. hmongorum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C468AAB0079FFDCFD07DBCBFD621FBF" authorityName="Boulenger" authorityYear="1917" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="grahami">
<emphasis id="B9322D3A0079FFDCFD07DBCBFD621FBF" italics="true" pageId="50" pageNumber="51">R. grahami</emphasis>
</taxonomicName>
, whose dorsolateral pustules sometimes form a dorsolateral fold, and
<taxonomicName id="4C468AAB0079FFDCFD5DDB80FB581FA1" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[1071,1228,487,510]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="banaorum">
<emphasis id="B9322D3A0079FFDCFD5DDB80FB581FA1" box="[1071,1228,487,510]" italics="true" pageId="50" pageNumber="51">R. banaorum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C468AAB0079FFDCFB9DD863FC061C44" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[751,914,516,539]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0079FFDCFB9DD863FC061C44" box="[751,914,516,539]" italics="true" pageId="50" pageNumber="51">R. chalconota</emphasis>
</taxonomicName>
, which have folds that are distinct and continuous, extending to the groin. The olive brown coloration with black spots of
<taxonomicName id="4C468AAB0079FFDCFA59D83BFB9D1C2C" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[811,1033,604,627]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFA59D83BFB9D1C2C" box="[811,1033,604,627]" italics="true" pageId="50" pageNumber="51">R. megatympanum</emphasis>
</taxonomicName>
differentiates it from
<taxonomicName id="4C468AAB0079FFDCFB8CD81EFCE41CCF" baseAuthorityName="Yang" baseAuthorityYear="1991" box="[766,880,633,656]" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="nasica">
<emphasis id="B9322D3A0079FFDCFB8CD81EFCE41CCF" box="[766,880,633,656]" italics="true" pageId="50" pageNumber="51">H. nasica</emphasis>
</taxonomicName>
(olive­brown dorsum, lighter brown laterally) and from
<taxonomicName id="4C468AAB0079FFDCFA80D8F0FB301CF1" authorityName="Inger &amp; Chan-ard" authorityYear="1997" box="[1010,1188,663,686]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="archotaphus">
<emphasis id="B9322D3A0079FFDCFA80D8F0FB301CF1" box="[1010,1188,663,686]" italics="true" pageId="50" pageNumber="51">R. archotaphus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFDC1D8F0FCA91C94" baseAuthorityName="Schlegel" baseAuthorityYear="1837" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0079FFDCFDC1D8F0FCA91C94" italics="true" pageId="50" pageNumber="51">R. chalconota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA27D8D3FC6A1C94" box="[853,1022,692,715]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A0079FFDCFA27D8D3FC6A1C94" box="[853,1022,692,715]" italics="true" pageId="50" pageNumber="51">R. chloronota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFD65D8D3FB0F1C94" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[1047,1179,692,715]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="daorum">
<emphasis id="B9322D3A0079FFDCFD65D8D3FB0F1C94" box="[1047,1179,692,715]" italics="true" pageId="50" pageNumber="51">R. daorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFDC1D8D3FCBF1CB7" authorityName=": Boulenger" authorityYear="1899" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B9322D3A0079FFDCFDC1D8D3FCBF1CB7" italics="true" pageId="50" pageNumber="51">R. graminea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA30D8B6FC6F1CB7" authorityName="Y.u.Deng" authorityYear="1992" box="[834,1019,721,744]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hejiangensis">
<emphasis id="B9322D3A0079FFDCFA30D8B6FC6F1CB7" box="[834,1019,721,744]" italics="true" pageId="50" pageNumber="51">R. hejiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFD63D8B6FB511CB7" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[1041,1221,721,744]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hmongorum">
<emphasis id="B9322D3A0079FFDCFD63D8B6FB511CB7" box="[1041,1221,721,744]" italics="true" pageId="50" pageNumber="51">R. hmongorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFBCED888FC821D59" authorityName="Boulenger" authorityYear="1891" box="[700,790,751,774]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hosii">
<emphasis id="B9322D3A0079FFDCFBCED888FC821D59" box="[700,790,751,774]" italics="true" pageId="50" pageNumber="51">R. hosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA57D888FC7C1D59" box="[805,1000,751,774]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0079FFDCFA57D888FC7C1D59" box="[805,1000,751,774]" italics="true" pageId="50" pageNumber="51">R. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA85D888FB311D59" baseAuthorityName="Huang, Fei &amp; Ye" baseAuthorityYear="2001" box="[1015,1189,751,774]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="junlianensis">
<emphasis id="B9322D3A0079FFDCFA85D888FB311D59" box="[1015,1189,751,774]" italics="true" pageId="50" pageNumber="51">R. junlianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFDC1D888FCF01D7C" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="kwangwuensis">
<emphasis id="B9322D3A0079FFDCFDC1D888FCF01D7C" italics="true" pageId="50" pageNumber="51">R. kwangwuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA05D96BFB931D7C" authorityName=": Werner" authorityYear="1930" box="[887,1031,780,803]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="leporipes">
<emphasis id="B9322D3A0079FFDCFA05D96BFB931D7C" box="[887,1031,780,803]" italics="true" pageId="50" pageNumber="51">R. leporipes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFD68D96BFB521D7C" authorityName="Liu" authorityYear="1950" box="[1050,1222,780,803]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="margaretae">
<emphasis id="B9322D3A0079FFDCFD68D96BFB521D7C" box="[1050,1222,780,803]" italics="true" pageId="50" pageNumber="51">R. margaretae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFBCED94EFCFE1D1F" authorityName="Boettger" authorityYear="1892" box="[700,874,809,832]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="schmackeri">
<emphasis id="B9322D3A0079FFDCFBCED94EFCFE1D1F" box="[700,874,809,832]" italics="true" pageId="50" pageNumber="51">R. schmackeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFAF0D94EFC641D1F" box="[898,1008,809,832]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A0079FFDCFAF0D94EFC641D1F" box="[898,1008,809,832]" italics="true" pageId="50" pageNumber="51">R. sinica</emphasis>
</taxonomicName>
(all with green).
<taxonomicName id="4C468AAB0079FFDCFBCED920FC2D1D01" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[700,953,839,862]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFBCED920FC2D1D01" box="[700,953,839,862]" italics="true" pageId="50" pageNumber="51">Rana megatympanum</emphasis>
</taxonomicName>
lacks an external metatarsal tubercle present in
<taxonomicName id="4C468AAB0079FFDCFA95D903FB0E1D24" authorityName="Inger &amp; Chan-ard" authorityYear="1997" box="[999,1178,868,891]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="archotaphus">
<emphasis id="B9322D3A0079FFDCFA95D903FB0E1D24" box="[999,1178,868,891]" italics="true" pageId="50" pageNumber="51">R. archotaphus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C468AAB0079FFDCFBCED9E6FCF61DC7" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[700,866,897,920]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0079FFDCFBCED9E6FCF61DC7" box="[700,866,897,920]" italics="true" pageId="50" pageNumber="51">R. chalconota</emphasis>
</taxonomicName>
. Absence of ventral spines in male
<taxonomicName id="4C468AAB0079FFDCFB8FD9F8FC431DE9" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[765,983,927,950]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFB8FD9F8FC431DE9" box="[765,983,927,950]" italics="true" pageId="50" pageNumber="51">R. megatympanum</emphasis>
</taxonomicName>
separates them from
<taxonomicName id="4C468AAB0079FFDCFBCED9DBFCCF1D8C" authorityName="Boulenger" authorityYear="1882" box="[700,859,956,979]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0079FFDCFBCED9DBFCCF1D8C" box="[700,859,956,979]" italics="true" pageId="50" pageNumber="51">R. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA1ED9DBFC661D8C" authorityName="Boulenger" authorityYear="1917" box="[876,1010,956,979]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="grahami">
<emphasis id="B9322D3A0079FFDCFA1ED9DBFC661D8C" box="[876,1010,956,979]" italics="true" pageId="50" pageNumber="51">R. grahami</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFD70D9DBFB511D8C" box="[1026,1221,956,979]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0079FFDCFD70D9DBFB511D8C" box="[1026,1221,956,979]" italics="true" pageId="50" pageNumber="51">R. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFBCED9BEFC151DAF" baseAuthorityName="Huang, Fei &amp; Ye" baseAuthorityYear="2001" box="[700,897,985,1008]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="junlianensis">
<emphasis id="B9322D3A0079FFDCFBCED9BEFC151DAF" box="[700,897,985,1008]" italics="true" pageId="50" pageNumber="51">R. junlianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFAEED9BEFBCD1DAF" authorityName="Liu" authorityYear="1950" box="[924,1113,985,1008]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="margaretae">
<emphasis id="B9322D3A0079FFDCFAEED9BEFBCD1DAF" box="[924,1113,985,1008]" italics="true" pageId="50" pageNumber="51">R. margaretae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0079FFDCFDC0D9BEFCD41A51" authorityName="Boettger" authorityYear="1892" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="schmackeri">
<emphasis id="B9322D3A0079FFDCFDC0D9BEFCD41A51" italics="true" pageId="50" pageNumber="51">R. schmackeri</emphasis>
</taxonomicName>
. The presence of white eggs differentiates
<taxonomicName id="4C468AAB0079FFDCFA32DE73FB891A74" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[832,1053,1044,1067]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFA32DE73FB891A74" box="[832,1053,1044,1067]" italics="true" pageId="50" pageNumber="51">R. megatympanum</emphasis>
</taxonomicName>
from
<taxonomicName id="4C468AAB0079FFDCFD19DE73FD431A17" baseAuthorityName="Yang" baseAuthorityYear="1991" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="nasica">
<emphasis id="B9322D3A0079FFDCFD19DE73FD431A17" italics="true" pageId="50" pageNumber="51">H. nasica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFB95DE56FC171A17" authorityName="Boulenger" authorityYear="1882" box="[743,899,1073,1096]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0079FFDCFB95DE56FC171A17" box="[743,899,1073,1096]" italics="true" pageId="50" pageNumber="51">R. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFAE0DE56FBA01A17" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[914,1076,1073,1096]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0079FFDCFAE0DE56FBA01A17" box="[914,1076,1073,1096]" italics="true" pageId="50" pageNumber="51">R. chalconota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFD31DE56FB511A17" authorityName="Boulenger" authorityYear="1917" box="[1091,1221,1073,1096]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="grahami">
<emphasis id="B9322D3A0079FFDCFD31DE56FB511A17" box="[1091,1221,1073,1096]" italics="true" pageId="50" pageNumber="51">R. grahami</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFBCEDE28FCF81A39" baseAuthorityName="Huang, Fei &amp; Ye" baseAuthorityYear="2001" box="[700,876,1103,1126]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="junlianensis">
<emphasis id="B9322D3A0079FFDCFBCEDE28FCF81A39" box="[700,876,1103,1126]" italics="true" pageId="50" pageNumber="51">R. junlianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFA0DDE28FBBD1A39" authorityName="Liu" authorityYear="1950" box="[895,1065,1103,1126]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="margaretae">
<emphasis id="B9322D3A0079FFDCFA0DDE28FBBD1A39" box="[895,1065,1103,1126]" italics="true" pageId="50" pageNumber="51">R. margaretae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0079FFDCFD4EDE28FD481ADC" authorityName="Boettger" authorityYear="1892" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="schmackeri">
<emphasis id="B9322D3A0079FFDCFD4EDE28FD481ADC" italics="true" pageId="50" pageNumber="51">R. schmackeri</emphasis>
</taxonomicName>
(white eggs with melanic poles), and
<taxonomicName id="4C468AAB0079FFDCFDC1DE0BFCA91AFF" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0079FFDCFDC1DE0BFCA91AFF" italics="true" pageId="50" pageNumber="51">R. bacboensis</emphasis>
</taxonomicName>
(eggs completely melanic).
<taxonomicName id="4C468AAB0079FFDCFDFDDEEEFCE01AE1" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFDFDDEEEFCE01AE1" italics="true" pageId="50" pageNumber="51">Rana megatympanum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C468AAB0079FFDCFD65DEC0FBEB1AE1" box="[1047,1151,1191,1214]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A0079FFDCFD65DEC0FBEB1AE1" box="[1047,1151,1191,1214]" italics="true" pageId="50" pageNumber="51">R. sinica</emphasis>
</taxonomicName>
by its distinct, uncovered tympanum (indistinct and covered with a layer of skin in
<taxonomicName id="4C468AAB0079FFDCFD43DE86FB031AA7" box="[1073,1175,1249,1272]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A0079FFDCFD43DE86FB031AA7" box="[1073,1175,1249,1272]" italics="true" pageId="50" pageNumber="51">R. sinica</emphasis>
</taxonomicName>
), its disk size (small in
<taxonomicName id="4C468AAB0079FFDCFAEDDE98FB921B49" box="[927,1030,1279,1302]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A0079FFDCFAEDDE98FB921B49" box="[927,1030,1279,1302]" italics="true" pageId="50" pageNumber="51">R. sinica</emphasis>
</taxonomicName>
), its relative finger lengths (I &lt;II &lt;IV for
<taxonomicName id="4C468AAB0079FFDCFD6CDF7BFB3F1B6C" authority=", II" authorityName="II" box="[1054,1195,1308,1331]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A0079FFDCFD6CDF7BFB121B6C" box="[1054,1158,1308,1331]" italics="true" pageId="50" pageNumber="51">R. sinica</emphasis>
, II
</taxonomicName>
&lt;I &lt;IV for
<taxonomicName id="4C468AAB0079FFDCFA26DF5EFBA71B0F" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[852,1075,1337,1360]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFA26DF5EFBA71B0F" box="[852,1075,1337,1360]" italics="true" pageId="50" pageNumber="51">R. megatympanum</emphasis>
</taxonomicName>
), and its T­ shaped distal phalanges (rounded in
<taxonomicName id="4C468AAB0079FFDCFD07DF30FD431BD4" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A0079FFDCFD07DF30FD431BD4" italics="true" pageId="50" pageNumber="51">R. sinica</emphasis>
</taxonomicName>
).
<taxonomicName id="4C468AAB0079FFDCFB8EDF13FB911BD4" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[764,1029,1396,1419]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFB8EDF13FB911BD4" box="[764,1029,1396,1419]" italics="true" pageId="50" pageNumber="51">Rana megatympanum</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C468AAB0079FFDCFA1CDFF6FB961BF7" authorityName=": Werner" authorityYear="1930" box="[878,1026,1425,1448]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="leporipes">
<emphasis id="B9322D3A0079FFDCFA1CDFF6FB961BF7" box="[878,1026,1425,1448]" italics="true" pageId="50" pageNumber="51">R. leporipes</emphasis>
</taxonomicName>
by its webbing (only to the basal end of the distal phalanx in
<taxonomicName id="4C468AAB0079FFDCFBADDFABFCE41BBC" authorityName=": Werner" authorityYear="1930" box="[735,880,1484,1507]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="leporipes">
<emphasis id="B9322D3A0079FFDCFBADDFABFCE41BBC" box="[735,880,1484,1507]" italics="true" pageId="50" pageNumber="51">R. leporipes</emphasis>
</taxonomicName>
, to the disk in
<taxonomicName id="4C468AAB0079FFDCFD47DFABFC9F185F" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFD47DFABFC9F185F" italics="true" pageId="50" pageNumber="51">R. megatympanum</emphasis>
</taxonomicName>
), supratympanic fold (colored white in
<taxonomicName id="4C468AAB0079FFDCFBAEDC60FCFF1841" authorityName=": Werner" authorityYear="1930" box="[732,875,1543,1566]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="leporipes">
<emphasis id="B9322D3A0079FFDCFBAEDC60FCFF1841" box="[732,875,1543,1566]" italics="true" pageId="50" pageNumber="51">R. leporipes</emphasis>
</taxonomicName>
), and its T­ shaped distal phalanges (oblong, somewhat rounded in
<taxonomicName id="4C468AAB0079FFDCFD0CDC43FC911807" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="leopripes">
<emphasis id="B9322D3A0079FFDCFD0CDC43FC911807" italics="true" pageId="50" pageNumber="51">R. leopripes</emphasis>
</taxonomicName>
).
<taxonomicName id="4C468AAB0079FFDCFA5ADC26FBBB1807" box="[808,1071,1601,1624]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatypmanum">
<emphasis id="B9322D3A0079FFDCFA5ADC26FBBB1807" box="[808,1071,1601,1624]" italics="true" pageId="50" pageNumber="51">Rana megatypmanum</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="4C468AAB0079FFDCFB8FDC38FC3B1829" authorityName="Y.u.Deng" authorityYear="1992" box="[765,943,1631,1654]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hejiangensis">
<emphasis id="B9322D3A0079FFDCFB8FDC38FC3B1829" box="[765,943,1631,1654]" italics="true" pageId="50" pageNumber="51">R. hejiangensis</emphasis>
</taxonomicName>
by its finger formula (II &lt;I &lt;III &lt;IV for
<taxonomicName id="4C468AAB0079FFDCFADEDC1BFBF418CC" authorityName="Y.u.Deng" authorityYear="1992" box="[940,1120,1660,1683]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hejiangensis">
<emphasis id="B9322D3A0079FFDCFADEDC1BFBF418CC" box="[940,1120,1660,1683]" italics="true" pageId="50" pageNumber="51">R. hejiangensis</emphasis>
</taxonomicName>
), and by its large disks (small for
<taxonomicName id="4C468AAB0079FFDCFD71DCFEFB2818EF" authorityName="Y.u.Deng" authorityYear="1992" box="[1027,1212,1689,1712]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="hejiangensis">
<emphasis id="B9322D3A0079FFDCFD71DCFEFB2818EF" box="[1027,1212,1689,1712]" italics="true" pageId="50" pageNumber="51">R. hejiangensis</emphasis>
</taxonomicName>
).
<taxonomicName id="4C468AAB0079FFDCFBCEDCD0FC551891" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[700,961,1719,1742]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0079FFDCFBCEDCD0FC551891" box="[700,961,1719,1742]" italics="true" pageId="50" pageNumber="51">Rana megatympanum</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="4C468AAB0079FFDDFDC1DCD0FF631E87" authorityName="Boulenger" authorityYear="1882" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" lastPageId="51" lastPageNumber="52" order="Anura" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0079FFDDFDC1DCD0FF631E87" italics="true" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">R. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0078FFDDF874DAA6FE271E87" box="[262,435,193,216]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A0078FFDDF874DAA6FE271E87" box="[262,435,193,216]" italics="true" pageId="51" pageNumber="52">R. hainanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0078FFDDF8B0DAA6FD171E87" box="[450,643,193,216]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0078FFDDF8B0DAA6FD171E87" box="[450,643,193,216]" italics="true" pageId="51" pageNumber="52">R. jingdongensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0078FFDDF9C2DAB8FEDF1EA9" box="[176,331,223,246]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="tiannensis">
<emphasis id="B9322D3A0078FFDDF9C2DAB8FEDF1EA9" box="[176,331,223,246]" italics="true" pageId="51" pageNumber="52">R. tiannensis</emphasis>
</taxonomicName>
. In addition,
<taxonomicName id="4C468AAB0078FFDDF886DAB8FF5E1F4C" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0078FFDDF886DAB8FF5E1F4C" italics="true" pageId="51" pageNumber="52">R. megatympanum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C468AAB0078FFDDF81CDA9BFD9A1F4C" authorityName="Boulenger" authorityYear="1882" box="[366,526,252,275]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0078FFDDF81CDA9BFD9A1F4C" box="[366,526,252,275]" italics="true" pageId="51" pageNumber="52">R. andersonii</emphasis>
</taxonomicName>
in that its males are smaller (
<quantity id="4CBE5CCD0078FFDDF817DB7EFE7C1F6F" box="[357,488,281,304]" metricMagnitude="-2" metricUnit="m" metricValue="6.55" metricValueMax="7.8" metricValueMin="5.3" pageId="51" pageNumber="52" unit="mm" value="65.5" valueMax="78.0" valueMin="53.0">5378 mm</quantity>
for males of
<taxonomicName id="4C468AAB0078FFDDF909DB50FE891F11" authorityName="Boulenger" authorityYear="1882" box="[123,285,311,334]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0078FFDDF909DB50FE891F11" box="[123,285,311,334]" italics="true" pageId="51" pageNumber="52">R. andersonii</emphasis>
</taxonomicName>
), and its finger and toe­disks are relatively larger.
<taxonomicName id="4C468AAB0078FFDDF8F7DB33FD1F1F34" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[389,651,340,363]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0078FFDDF8F7DB33FD1F1F34" box="[389,651,340,363]" italics="true" pageId="51" pageNumber="52">Rana megatympanum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C468AAB0078FFDDF86DDB16FE401FD7" box="[287,468,369,392]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A0078FFDDF86DDB16FE401FD7" box="[287,468,369,392]" italics="true" pageId="51" pageNumber="52">R. hainanensis</emphasis>
</taxonomicName>
in its smaller SVL (
<taxonomicName id="4C468AAB0078FFDDF9B5DBE8FEE31FF9" box="[199,375,399,422]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A0078FFDDF9B5DBE8FEE31FF9" box="[199,375,399,422]" italics="true" pageId="51" pageNumber="52">R. hainanensis</emphasis>
</taxonomicName>
males
<quantity id="4CBE5CCD0078FFDDF8A1DBE9FDCE1FFA" box="[467,602,398,421]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" metricValueMax="6.2" metricValueMin="4.9" pageId="51" pageNumber="52" unit="mm" value="55.5" valueMax="62.0" valueMin="49.0">4962 mm</quantity>
, females
<quantity id="4CBE5CCD0078FFDDF9B5DBCBFECC1F9C" box="[199,344,428,451]" metricMagnitude="-2" metricUnit="m" metricValue="9.85" metricValueMax="12.2" metricValueMin="7.5" pageId="51" pageNumber="52" unit="mm" value="98.5" valueMax="122.0" valueMin="75.0">75122 mm</quantity>
,
<taxonomicName id="4C468AAB0078FFDDF817DBCBFDAB1F9C" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[357,575,428,451]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0078FFDDF817DBCBFDAB1F9C" box="[357,575,428,451]" italics="true" pageId="51" pageNumber="52">R. megatympanum</emphasis>
</taxonomicName>
males
<quantity id="4CBE5CCD0078FFDDF909DBAEFF681FBF" box="[123,252,457,480]" metricMagnitude="-2" metricUnit="m" metricValue="5.15" metricValueMax="5.5" metricValueMin="4.8" pageId="51" pageNumber="52" unit="mm" value="51.5" valueMax="55.0" valueMin="48.0">4855 mm</quantity>
, females
<quantity id="4CBE5CCD0078FFDDF81FDBAEFE681FBF" box="[365,508,457,480]" metricMagnitude="-2" metricUnit="m" metricValue="9.9" metricValueMax="10.5" metricValueMin="9.3" pageId="51" pageNumber="52" unit="mm" value="99.0" valueMax="105.0" valueMin="93.0">93105 mm</quantity>
) and its relative finger lengths (II &lt;IV &lt;I &lt;III for
<taxonomicName id="4C468AAB0078FFDDFB00DB80FEB81C44" authority=", II" authorityName="II" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A0078FFDDFB00DB80FE921C44" italics="true" pageId="51" pageNumber="52">R. hainanensis</emphasis>
, II
</taxonomicName>
&lt;I &lt;IV &lt;III for
<taxonomicName id="4C468AAB0078FFDDFB48D863FE851C67" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0078FFDDFB48D863FE851C67" italics="true" pageId="51" pageNumber="52">R. megatympanum</emphasis>
</taxonomicName>
).
<taxonomicName id="4C468AAB0078FFDDF85CD846FDA41C67" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[302,560,545,568]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0078FFDDF85CD846FDA41C67" box="[302,560,545,568]" italics="true" pageId="51" pageNumber="52">Rana megatympanum</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C468AAB0078FFDDF864D858FE4F1C09" box="[278,475,575,598]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0078FFDDF864D858FE4F1C09" box="[278,475,575,598]" italics="true" pageId="51" pageNumber="52">R. jingdongensis</emphasis>
</taxonomicName>
by its smaller males (
<taxonomicName id="4C468AAB0078FFDDF9ABD83BFE7F1C2C" authority="SVL" authorityName="SVL" box="[217,491,604,627]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0078FFDDF9ABD83BFE371C2C" box="[217,419,604,627]" italics="true" pageId="51" pageNumber="52">R. jingdongensis</emphasis>
SVL
</taxonomicName>
<quantity id="4CBE5CCD0078FFDDF889D83BFD171C2C" box="[507,643,604,627]" metricMagnitude="-2" metricUnit="m" metricValue="7.15" metricValueMax="8.1" metricValueMin="6.2" pageId="51" pageNumber="52" unit="mm" value="71.5" valueMax="81.0" valueMin="62.0">6281 mm</quantity>
) and skin (
<taxonomicName id="4C468AAB0078FFDDF986D81EFE2C1CCF" box="[244,440,633,656]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0078FFDDF986D81EFE2C1CCF" box="[244,440,633,656]" italics="true" pageId="51" pageNumber="52">R. jingdongensis</emphasis>
</taxonomicName>
dorsum scattered with tubercles and large warts, lips and sides of heads with white spines, all absent in
<taxonomicName id="4C468AAB0078FFDDFB00D8D3FEA71CB7" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0078FFDDFB00D8D3FEA71CB7" italics="true" pageId="51" pageNumber="52">R. megatympanum</emphasis>
</taxonomicName>
).
<taxonomicName id="4C468AAB0078FFDDF83ED8B6FDDE1CB7" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[332,586,721,744]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A0078FFDDF83ED8B6FDDE1CB7" box="[332,586,721,744]" italics="true" pageId="51" pageNumber="52">Rana megatympanum</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C468AAB0078FFDDF83DD888FE711D59" box="[335,485,751,774]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="tiannensis">
<emphasis id="B9322D3A0078FFDDF83DD888FE711D59" box="[335,485,751,774]" italics="true" pageId="51" pageNumber="52">R. tiannensis</emphasis>
</taxonomicName>
, another large brown cascade ranid, but differs from it by having shagreened dorsal skin with small lateral granulations (dorsum of
<taxonomicName id="4C468AAB0078FFDDF8A4D920FDFB1D01" box="[470,623,839,862]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="tiannensis">
<emphasis id="B9322D3A0078FFDDF8A4D920FDFB1D01" box="[470,623,839,862]" italics="true" pageId="51" pageNumber="52">R. tiannensis</emphasis>
</taxonomicName>
is rough with large, prominent lateral granulations), and toe disks are smaller than those on fingers (the opposite condition of
<taxonomicName id="4C468AAB0078FFDDFB40D9F8FF501D8C" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="tiannensis">
<emphasis id="B9322D3A0078FFDDFB40D9F8FF501D8C" italics="true" pageId="51" pageNumber="52">R. tiannensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35CA2A30078FFDAF9EAD9BEFDEA1A74" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="description">
<paragraph id="8BF9F1280078FFDDF9EAD9BEFDDB1864" blockId="51.[123,652,193,1741]" pageId="51" pageNumber="52">
<materialsCitation id="3B2EFB750078FFDDF9EAD9BEFDDB1864" ID-GBIF-Occurrence="3408947302" accessionNumber="ROM 39684" pageId="51" pageNumber="52" specimenCount="1" typeStatus="holotype">
DESCRIPTION OF
<typeStatus id="54FD4F8A0078FFDDF827D9BEFE5E1DB0" box="[341,458,985,1008]" pageId="51" pageNumber="52" type="holotype">HOLOTYPE</typeStatus>
: ROM 39684, a gravid female, head width 77% of head length, length 50% of SVL; snout short, acutely rounded in dorsal view, bluntly rounded in profile, protruding beyond margin of lower jaw; eye very large, prominent, 73% of snout length; eyelid broader than interorbital distance. Top of head flat; canthus rostralis rounded; loreal region concave; lip flared just anterior to orbit; nostril about three­fourths distance from eye to tip of snout; supratympanic fold curving posteroventrally from posterior corner of eye to a level above the insertion of arm; tympanum round, distinctly visible, separated from eye by distance equal to TMP, 52% of EYE. Choanae ovoid; vomerine dentigerous processes prominent, oblique, posteromedial to choanae, each bearing numerous teeth. Tongue cordiform, distinctly notched posteriorly, free for approximately two­thirds its length.
</materialsCitation>
</paragraph>
<paragraph id="8BF9F1280078FFDDF9EADC26FCE01C94" blockId="51.[123,652,193,1741]" lastBlockId="51.[699,1228,192,1741]" pageId="51" pageNumber="52">
Forearms moderately robust; fingers moderately short, slender, hands 27% of SVL, relative lengths of fingers II &lt;I &lt;IV &lt;III, lateral fringes on finger II, III, and IV, with median callous pads to proximal tubercle; disks greatly expanded (&gt;2× base of phalanges), relative pad size II &lt;I &lt;IV &lt;III, pad width (III) 87% of pad length, ventral circummarginal grooves present; terminal phalanges T­ shaped; subarticular tubercles conical. Hindlimbs moderately robust; tibia length 69% of SVL; foot length 53% of SVL; relative toe lengths I &lt;II &lt;III &lt;V &lt;IV; inner tarsal fold absent; feet fully webbed to base of toe pads, lateral fringes on I and V to terminal phalanges; toes long, slender, with large, rounded triangular disks, relative pad size I = II = III&gt; IV
<emphasis id="B9322D3A0078FFDDFD2FD845FBEC1C66" bold="true" box="[1117,1144,546,569]" italics="true" pageId="51" pageNumber="52">k</emphasis>
V, pad width (IV) 75% of pad length, each with ventral circummarginal grooves; subarticular tubercles prominent and conical; inner metatarsal tubercle ovoid, long; outer metatarsal tubercle absent.
</paragraph>
<paragraph id="8BF9F1280078FFDDFBAAD8B6FC831D5A" blockId="51.[699,1228,192,1741]" pageId="51" pageNumber="52">Xiphisternum large, deeply notched posteriorly.</paragraph>
<paragraph id="8BF9F1280078FFDDFBAAD96BFC9D1D8C" blockId="51.[699,1228,192,1741]" pageId="51" pageNumber="52">Skin on dorsum shagreened, becoming increasingly granular laterally; dorsolateral folds absent; small tubercles posteroventral to tympanum; prominent granules on flanks and around cloaca; cloacal opening unmodified, directed posteriorly, at upper level of thighs.</paragraph>
<paragraph id="8BF9F1280078FFDDFBAAD9BEFC211B6C" blockId="51.[699,1228,192,1741]" pageId="51" pageNumber="52">COLOR IN LIFE (in preservative): Dorsum olive­brown, flanks yellow and brown­gray (gray to olive); lip­stripe absent (tympanum beige with dark brown center ring); loreal brown (black); iris gold; top one­third red, dorsal limbs brown with black banding (brown); posterior surface of thighs brown with black marbling (cloacal region black, thighs gray with white mottling); webbing marbled white on dark brown (brown on white); venter creamy white (creamy yellow with black mottling).</paragraph>
<paragraph id="8BF9F1280078FFDDFBAADF5EFC0D18CC" blockId="51.[699,1228,192,1741]" pageId="51" pageNumber="52">
SECONDARY SEXUAL CHARACTERS: The eggs of the
<typeStatus id="54FD4F8A0078FFDDFA35DF31FC391B32" box="[839,941,1366,1389]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
are creamy white and
<quantity id="4CBE5CCD0078FFDDFDCEDF31FD7C1BD4" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="51" pageNumber="52" unit="mm" value="2.0">2 mm</quantity>
in diameter. Adult females have SVL approximately twice that of males. Males have a yellow lip­stripe, and females either lack one or have an indistinctly yellow lip. Males also possess a weak dorsolateral fold, whereas females do not. Males have a larger tympanum than females, velvety nuptial pads extending across the thumb, paired gular pouches located at the angle of the jaw, and no pectoral spines.
</paragraph>
<paragraph id="8BF9F1280078FFDAFBAADCFEFDF01F4C" blockId="51.[699,1228,192,1741]" lastBlockId="52.[124,653,193,1742]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
MEASUREMENTS OF
<typeStatus id="54FD4F8A0078FFDDFABDDCFEFBD118F0" box="[975,1093,1689,1712]" pageId="51" pageNumber="52" type="holotype">HOLOTYPE</typeStatus>
(in mm): SVL 93.6; SNT 15.0; HDL 46.5; HDW 35.7; EYE 10.8; IOD 6.8; TMP 5.6; TEY 4.1; HND 25.7; FGR 19.6; FPL 3.0; FPW 2.6; TIB 65.0; FTL 49.7; TPL 3.2; TPW 2.4.
</paragraph>
<paragraph id="8BF9F128007FFFDAF9EBDB7EFD841F12" blockId="52.[124,653,193,1742]" pageId="52" pageNumber="53">
VARIATION OF
<typeStatus id="54FD4F8A007FFFDAF835DB7EFE511F70" box="[327,453,281,304]" pageId="52" pageNumber="53" type="paratype">PARATYPES</typeStatus>
: Variation in all type material is given in
<tableCitation id="C6C4C493007FFFDAF8DBDB51FD841F12" box="[425,528,310,333]" captionStart="TABLE 15" captionStartId="46.[620,694,194,213]" captionTargetId="figure-50@46.[121,1231,304,1673]" captionTargetPageId="46" captionText="TABLE 15 Variation in Body Proportions for Rana chloronota sensu stricto and Vietnamese Species of the Rana chloronota Complex Given as Means (in mm) ± 1 SD and ranges See Materials and Methods for abbreviations." pageId="52" pageNumber="53">table 15.</tableCitation>
</paragraph>
<paragraph id="8BF9F128007FFFDAF9EBDB33FE271C94" blockId="52.[124,653,193,1742]" pageId="52" pageNumber="53">
MEASUREMENTS OF FEMALE
<typeStatus id="54FD4F8A007FFFDAF896DB33FDF61F35" box="[484,610,340,363]" pageId="52" pageNumber="53" type="paratype">PARATYPES</typeStatus>
(in mm, n = 10, ROM 2639826400, 39685 39691): SVL 100.3 ± 4.2 (93.6105.3); SNT 14.8 ± 0.8 (13.816.5); HDL 45.1 ± 3.3 (41.347.6); HDW 35.2 ± 0.7 (34.135.7); EYE 10.2 ± 0.7 (9.311.6); IOD 8.8 ± 1.0 (6.810.0); TMP 5.3 ± 0.4 (4.65.9); TEY 4.7 ± 0.3 (4.15.0); HND 25.2 ± 2.5 (20.6 29.7); FGR 21.0 ± 1.0 (19.622.6); FPL 3.5 ± 0.6 (2.84.4); FPW 3.0 ± 0.5 (2.53.8); TIB 63.0 ± 3.3 (55.867.7); FTL 72.0 ± 12.1 (49.788.0) TPL 3.4 ± 1.0 (1.95.2); TPW 2.8 ± 0.4 (2.23.4).
</paragraph>
<paragraph id="8BF9F128007FFFDAF9EBD8B6FDEA1A74" blockId="52.[124,653,193,1742]" pageId="52" pageNumber="53">
MEASUREMENTS OF MALE
<typeStatus id="54FD4F8A007FFFDAF8ACD8B6FDC81CB8" box="[478,604,721,744]" pageId="52" pageNumber="53" type="paratype">PARATYPES</typeStatus>
(in mm, n = 4, ROM 3923739240): SVL 52.3 ± 3.4 (48.655.2); SNT 8.7 ± 0.8 (8.39.6); HDL 28.0 ± 1.4 (24.627.1); HDW 18.5 ± 0.8 (18.019.1); EYE 4.1 ± 0.9 (3.174.6); IOD 5.0 ± 0.5 (4.75.6); TMP 4.7 ± 0.3 (4.35.1); TEY 1.7 ± 0.3 (1.52.0); HND 15.2 ± 0.4 (14.815.5); FGR 12.5 ± 0.4 (12.213.0); FPL 2.0 ± 0.5 (1.62.5); FPW 1.9 ± 0.4 (1.42.3); TIB 32.7 ± 0.4 (32.3 33.1); FTL 38.3 ± 7.3 (29.942.8) TPL 1.8 ± 0.6 (1.32.5); TPW 1.6 ± 0.1 (1.51.6).
</paragraph>
</subSubSection>
<subSubSection id="C35CA2A3007FFFDAF9EBDE56FE201AE2" pageId="52" pageNumber="53" type="etymology">
<paragraph id="8BF9F128007FFFDAF9EBDE56FE201AE2" blockId="52.[124,653,193,1742]" pageId="52" pageNumber="53">ETYMOLOGY: The specific name is a noun in opposition, derived from the Latin prefix mega (meaning very large) and tympanum, in reference to the relatively large tympanum of this species.</paragraph>
</subSubSection>
<subSubSection id="C35CA2A3007FFFDAF9EBDEA3FF47185F" pageId="52" pageNumber="53" type="distribution">
<paragraph id="8BF9F128007FFFDAF9EBDEA3FF47185F" blockId="52.[124,653,193,1742]" pageId="52" pageNumber="53">
DISTRIBUTION AND ECOLOGY:
<taxonomicName id="4C468AAB007FFFDAF888DEA3FF691AA7" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A007FFFDAF888DEA3FF691AA7" italics="true" pageId="52" pageNumber="53">Rana megatympanum</emphasis>
</taxonomicName>
is known from northern and north­central
<collectingCountry id="F351B1B8007FFFDAF86CDE99FE111B4A" box="[286,389,1278,1301]" name="Vietnam" pageId="52" pageNumber="53">Vietnam</collectingCountry>
. It occurs in montane rivers that vary from shallow and slow moving to torrential and deep. It may be found on boulders and logs, both in and around the water as well as in the adjacent forest. Radiographs revealed that large invertebrates (both insects and crustaceans) form part of the diet. Vocalizations and tadpoles are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35CA2A3007FFFDAF9EBDC61FDA21892" pageId="52" pageNumber="53" type="discussion">
<paragraph id="8BF9F128007FFFDAF9EBDC61FDA21892" blockId="52.[124,653,193,1742]" pageId="52" pageNumber="53">
REMARKS: The dorsolateral fold and very large tympanum of male
<taxonomicName id="4C468AAB007FFFDAF8C2DC43FD191864" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[432,653,1572,1595]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A007FFFDAF8C2DC43FD191864" box="[432,653,1572,1595]" italics="true" pageId="52" pageNumber="53">R. megatympanum</emphasis>
</taxonomicName>
potentially make it
<bibRefCitation id="EFD78CD9007FFFDAF815DC26FDBE1807" author="Bourret, R." box="[359,554,1601,1624]" pageId="52" pageNumber="53" refId="ref36841" refString="Bourret, R. 1942. Les batrachians de l'Indochine. Hanoi, Vietnam: Gouvernment General de L'Indochine, Hanoi, Vietnam, 547 pp." type="book" year="1942">Bourrets (1942)</bibRefCitation>
northern form of
<taxonomicName id="4C468AAB007FFFDAF86EDC38FE291829" box="[284,445,1631,1654]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A007FFFDAF86EDC38FE291829" box="[284,445,1631,1654]" italics="true" pageId="52" pageNumber="53">R. chloronota</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C468AAB007FFFDAF89EDC38FDEF1829" authorityName=": Boulenger" authorityYear="1899" box="[492,635,1631,1654]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B9322D3A007FFFDAF89EDC38FDEF1829" box="[492,635,1631,1654]" italics="true" pageId="52" pageNumber="53">R. graminea</emphasis>
</taxonomicName>
). However,
<taxonomicName id="4C468AAB007FFFDAF987DC1BFE1318CC" authorityName=": Boulenger" authorityYear="1899" box="[245,391,1660,1683]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B9322D3A007FFFDAF987DC1BFE1318CC" box="[245,391,1660,1683]" italics="true" pageId="52" pageNumber="53">R. graminea</emphasis>
</taxonomicName>
is bright green above (
<bibRefCitation id="EFD78CD9007FFFDAF9F6DCFEFEDA18EF" author="Boulenger, G. A." box="[132,334,1689,1712]" pageId="52" pageNumber="53" pagination="956 - 962" refId="ref36680" refString="Boulenger, G. A. 1899. On the reptiles, batrachians, and fishes collected by the late Mr. John Whitehead in the interior of Hainan. Proceedings of the Zoological Society of London 1899: 956 - 962, 4 pls." type="book chapter" year="1899">Boulenger, 1899</bibRefCitation>
) in contrast to the olivebrown dorsum of
<taxonomicName id="4C468AAB007FFFDAF826DCD0FDBB1891" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[340,559,1719,1742]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B9322D3A007FFFDAF826DCD0FDBB1891" box="[340,559,1719,1742]" italics="true" pageId="52" pageNumber="53">R. megatympanum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>