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<document id="1F3F9F68BA205FBA288264A9C320A010" ID-CLB-Dataset="23990" ID-DOI="10.1080/00222933.2019.1606355" ID-GBIF-Dataset="6ef4ad0e-2d1a-498f-b752-3b19c6068c4f" ID-ISSN="1464-5262" ID-Zenodo-Dep="3673146" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582027238037" checkinUser="valdenar" docAuthor="Penghai Qiao, Wen Qin, Huiqin Ma &amp; Tongzuo Zhang" docDate="2019" docId="03CBE07B4F02FFDFFE39FB01FCA4FBD2" docLanguage="en" docName="JNATHIST.53.15–16.897–921.pdf.imf" docOrigin="Journal of Natural History 53 (15)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Hessebius prospinosa Qiao, Qin, Ma &amp; Zhang, 2019, sp. nov." docType="treatment" docUuid="B1AD25A9-9C0B-47E6-8DB8-73BD56500320" docUuidSource="ZooBank" docVersion="11" lastPageNumber="904" masterDocId="FFF298034F06FFD7FFE3FF80FFBFFF9B" masterDocTitle="Two new species of lithobiid centipedes and the first record of Lamyctes africanus Porath (Chilopoda: Lithobiomorpha) in China" masterLastPageNumber="921" masterPageNumber="897" pageNumber="900" updateTime="1698784218181" updateUser="ExternalLinkService">
<mods:mods id="0D23A6F6780F5FF32FBE36630665A4F7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="714F6753A8E0820F3E9FF3CED41F40AA">
<mods:title id="EDE5E4C6E88956739679ACA8814143A8">Two new species of lithobiid centipedes and the first record of Lamyctes africanus Porath (Chilopoda: Lithobiomorpha) in China</mods:title>
</mods:titleInfo>
<mods:name id="B1997EB15267D10499B5F93ED6B8DFF5" type="personal">
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<mods:namePart id="BB1D8541E292C73C02E669DAB8F6C875">Penghai Qiao</mods:namePart>
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<mods:name id="B88B0AF15968F2C3C7907382D1F59B64" type="personal">
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<mods:roleTerm id="E29288FAA85BB98C31F326153C778AC8">Author</mods:roleTerm>
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<mods:namePart id="F3DDD22996E9A2D094104A799E3AC8B1">Wen Qin</mods:namePart>
</mods:name>
<mods:name id="A613A9B75E72A317D7891552EDC5B5C9" type="personal">
<mods:role id="9E94A208F106B5F846A23501F51C93B1">
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</mods:role>
<mods:namePart id="38B3C11AF70254769DCE3726CD5E6202">Huiqin Ma</mods:namePart>
</mods:name>
<mods:name id="7CE84A82F2B804DCEBA95F2B47377FBF" type="personal">
<mods:role id="D6A003D078A306F476DFB9180A196F0B">
<mods:roleTerm id="4C969329C5512D855C226C9BE2F28325">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C7C2800DCF36B168BF9B8772F8CFF605">Tongzuo Zhang</mods:namePart>
</mods:name>
<mods:typeOfResource id="FD92F6CA2CF14B14C7484F5DB3BE9693">text</mods:typeOfResource>
<mods:relatedItem id="803FA8D36F6EAA83C57E73206589BEBF" type="host">
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<mods:title id="BD99EE32E42FBAF51916F5CB3326AC92">Journal of Natural History</mods:title>
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<mods:part id="6881D3194A0DDD4758C7223A8284ACC4">
<mods:date id="43187A7A5750C59D326B03C3ECE8206B">2019</mods:date>
<mods:detail id="B1E32CE60F90FB2DAD1AD26DAF1A05AC" type="pubDate">
<mods:number id="CB0EE5BA0D5219A0437A747206293F01">2019-06-07</mods:number>
</mods:detail>
<mods:detail id="552F111BEAD454E2A848134AAAADBB89" type="volume">
<mods:number id="09E7F8777904AF5DE5CCAA53BD4C6C2D">53</mods:number>
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<mods:detail id="95CA64DC0E8E972D4D38F99DE2FD2287" type="issue">
<mods:number id="E018D6B0CC29D1489018636AD6DE8C66">15</mods:number>
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<mods:classification id="4BA632C8E0CBAE440BCFA76C168F76D6">journal article</mods:classification>
<mods:identifier id="6256FDF6EADD32D87E03C85C3C661137" type="CLB-Dataset">23990</mods:identifier>
<mods:identifier id="D2CEACBA4F5542D67480F78BAFA9C466" type="DOI">10.1080/00222933.2019.1606355</mods:identifier>
<mods:identifier id="BEF16AB331A73EBAFB8FA51D01EB0F5E" type="GBIF-Dataset">6ef4ad0e-2d1a-498f-b752-3b19c6068c4f</mods:identifier>
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<mods:identifier id="C2E734D19DAEF8425B04A52F3198415F" type="ZooBank">24364A2D-8D84-444D-9CFC-24336A211D2B</mods:identifier>
</mods:mods>
<treatment id="03CBE07B4F02FFDFFE39FB01FCA4FBD2" ID-DOI="http://doi.org/10.5281/zenodo.3679606" ID-GBIF-Taxon="162161534" ID-Zenodo-Dep="3679606" LSID="urn:lsid:zoobank.org:act:B1AD25A9-9C0B-47E6-8DB8-73BD56500320" httpUri="http://treatment.plazi.org/id/03CBE07B4F02FFDFFE39FB01FCA4FBD2" lastPageId="8" lastPageNumber="904" pageId="4" pageNumber="900">
<subSubSection id="C37802E64F02FFD3FE39FB01FCF6FB00" box="[474,841,1153,1179]" pageId="4" pageNumber="900" type="nomenclature">
<paragraph id="8BDD516D4F02FFD3FE39FB01FCF6FB00" blockId="4.[474,841,1153,1214]" box="[474,841,1153,1179]" pageId="4" pageNumber="900">
<heading id="D095E6014F02FFD3FE39FB01FCF6FB00" box="[474,841,1153,1179]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="900" reason="2">
<taxonomicName id="4C622AEE4F02FFD3FE39FB01FD5CFB00" authorityName="Qiao, Qin, Ma &amp; Zhang" authorityYear="2019" box="[474,739,1153,1179]" class="Chilopoda" family="Lithobiidae" genus="Hessebius" kingdom="Animalia" order="Lithobiomorpha" pageId="4" pageNumber="900" phylum="Arthropoda" rank="species" species="prospinosa" status="sp. nov.">
<emphasis id="B9168D7F4F02FFD3FE39FB01FD5CFB00" bold="true" box="[474,739,1153,1179]" italics="true" pageId="4" pageNumber="900">Hessebius prospinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22530044F02FFD3FD0EFB02FCF6FB00" box="[749,841,1154,1179]" pageId="4" pageNumber="900" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37802E64F02FFD3FDBAFB25FD74FB25" box="[601,715,1189,1214]" pageId="4" pageNumber="900" type="description">
<paragraph id="8BDD516D4F02FFD3FDBAFB25FD74FB25" blockId="4.[474,841,1153,1214]" box="[601,715,1189,1214]" pageId="4" pageNumber="900">
<heading id="D095E6014F02FFD3FDBAFB25FD74FB25" box="[601,715,1189,1214]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="900" reason="5">
(
<figureCitation id="13594DE84F02FFD3FDBCFB25FD7CFB25" box="[607,707,1189,1214]" captionStart="Figure 2" captionStartId="5.[160,225,1535,1558]" captionTargetBox="[166,1144,161,1504]" captionTargetId="figure@5.[166,1149,142,1504]" captionTargetPageId="5" captionText="Figure 2. Hessebius prospinosa sp. nov.ı (a–c): paratypeı male: (a) habitusı dorsal viewı scale 1 mm; (b) forcipular coxosterniteı ventral viewı scale 500 μm; (c) posterior segments and gonopodsı ventral viewı scale 500 μm; (d–f) holotypeı female: (d) posterior segments and gonopodsı ventral viewı scale 500 μm; (e) ocelli and Tömösváry’s organı lateral viewı scale 250 μm; (f) right gonopodı lateral viewı scale 250 μm." figureDoi="http://doi.org/10.5281/zenodo.3673150" httpUri="https://zenodo.org/record/3673150/files/figure.png" pageId="4" pageNumber="900">Figure 2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37802E64F02FFD3FF43FB7FFCB9FA52" pageId="4" pageNumber="900" type="materials_examined">
<paragraph id="8BDD516D4F02FFD3FF43FB7FFE88FA83" blockId="4.[160,1157,1279,1374]" box="[160,311,1279,1304]" pageId="4" pageNumber="900">
<emphasis id="B9168D7F4F02FFD3FF43FB7FFE88FA83" bold="true" box="[160,311,1279,1304]" italics="true" pageId="4" pageNumber="900">Type locality</emphasis>
</paragraph>
<paragraph id="8BDD516D4F02FFD3FF43FAA3FD57FAC5" blockId="4.[160,1157,1279,1374]" pageId="4" pageNumber="900">
Haiyan Countyı
<collectingRegion id="49A69F8F4F02FFD3FEBAFAA3FE0EFAA0" box="[345,433,1315,1339]" country="China" name="Qinghai" pageId="4" pageNumber="900">Qinghai</collectingRegion>
Provinceı Chinaı 36.92ı769°Nı 100.95ı264°Eı 7 December
<date id="FFDC77AD4F02FFD3FBA4FAA3FB3AFAA0" box="[1095,1157,1315,1339]" pageId="4" pageNumber="900" value="2011-01">2011ı</date>
<quantity id="4C9AFC884F02FFD3FF43FAC6FF49FAC5" box="[160,246,1350,1374]" metricMagnitude="3" metricUnit="m" metricValue="3.0439999999999996" pageId="4" pageNumber="900" unit="m" value="3044.0">3044 m</quantity>
above sea levelı collected by Gonghua Lin.
</paragraph>
<materialsCitation id="3B0A5B304F02FFD3FF43FA0DFCB9FA52" ID-GBIF-Occurrence="2571176316" location="Material" pageId="4" pageNumber="900" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<paragraph id="8BDD516D4F02FFD3FF43FA0DFE3FFA3D" blockId="4.[160,774,1421,1481]" box="[160,384,1421,1446]" pageId="4" pageNumber="900">
<emphasis id="B9168D7F4F02FFD3FF43FA0DFE3FFA3D" bold="true" box="[160,384,1421,1446]" italics="true" pageId="4" pageNumber="900">
<location id="8EBD07B64F02FFD3FF43FA0DFEBCFA3D" LSID="urn:lsid:plazi:treatment:03CBE07B4F02FFDFFE39FB01FCA4FBD2:8EBD07B64F02FFD3FF43FA0DFEBCFA3D" box="[160,259,1421,1446]" name="Material" pageId="4" pageNumber="900">Material</location>
examined
</emphasis>
</paragraph>
<paragraph id="8BDD516D4F02FFD3FF43FA31FE6AFA52" blockId="4.[160,774,1421,1481]" box="[160,469,1457,1481]" pageId="4" pageNumber="900">
<specimenCount id="9D649AE44F02FFD3FF43FA31FF0BFA52" box="[160,180,1457,1481]" pageId="4" pageNumber="900" type="female">♀</specimenCount>
<typeStatus id="54D9EFCF4F02FFD3FF5EFA31FE9EFA52" box="[189,289,1457,1481]" pageId="4" pageNumber="900" type="holotype">holotype</typeStatus>
(HY4)ı
<specimenCount id="9D649AE44F02FFD3FE96FA31FE36FA52" box="[373,393,1457,1481]" pageId="4" pageNumber="900" type="female">♀</specimenCount>
(HY6)ı
</paragraph>
<paragraph id="8BDD516D4F02FFD3FE3EFA31FCB9FA52" blockId="4.[160,774,1421,1481]" box="[477,774,1457,1481]" pageId="4" pageNumber="900">
2 Ƌ
<typeStatus id="54D9EFCF4F02FFD3FDF3FA31FDC1FA52" box="[528,638,1457,1481]" pageId="4" pageNumber="900" type="paratype">paratypes</typeStatus>
(HY3ı HY8).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C37802E64F02FFD1FF43FA79FD5AFEAA" lastPageId="6" lastPageNumber="902" pageId="4" pageNumber="900" type="diagnosis">
<paragraph id="8BDD516D4F02FFD3FF43FA79FEABF989" blockId="4.[160,1156,1529,1727]" box="[160,276,1529,1554]" pageId="4" pageNumber="900">
<emphasis id="B9168D7F4F02FFD3FF43FA79FEABF989" bold="true" box="[160,276,1529,1554]" italics="true" pageId="4" pageNumber="900">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BDD516D4F02FFD1FF43F99DFD5AFEAA" blockId="4.[160,1156,1529,1727]" lastBlockId="6.[160,1155,142,305]" lastPageId="6" lastPageNumber="902" pageId="4" pageNumber="900">
Body length
<quantity id="4C9AFC884F02FFD3FED6F99DFE67F9AE" box="[309,472,1564,1589]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.58" metricValueMin="1.42" pageId="4" pageNumber="900" unit="mm" value="15.0" valueMax="15.8" valueMin="14.2">
14.2
<emphasis id="B9168D7F4F02FFD3FE80F99CFECFF9AF" box="[355,368,1564,1588]" italics="true" pageId="4" pageNumber="900">–</emphasis>
15.8 mm
</quantity>
. Antennae with 20 antennomeres; 9
<emphasis id="B9168D7F4F02FFD3FC99F99CFC38F9AF" box="[890,903,1564,1588]" italics="true" pageId="4" pageNumber="900">–</emphasis>
11 ocelli on each side arranged in 3 rows. Tömösváry
<emphasis id="B9168D7F4F02FFD3FE1CF9BFFDBAF9CC" box="[511,517,1599,1623]" italics="true" pageId="4" pageNumber="900">’</emphasis>
s organ ovalı almost equal in size to neighbouring ocelli (
<figureCitation id="13594DE84F02FFD3FF44F9E2FEB7F9E1" box="[167,264,1634,1658]" captionStart="Figure 2" captionStartId="5.[160,225,1535,1558]" captionTargetBox="[166,1144,161,1504]" captionTargetId="figure@5.[166,1149,142,1504]" captionTargetPageId="5" captionText="Figure 2. Hessebius prospinosa sp. nov.ı (a–c): paratypeı male: (a) habitusı dorsal viewı scale 1 mm; (b) forcipular coxosterniteı ventral viewı scale 500 μm; (c) posterior segments and gonopodsı ventral viewı scale 500 μm; (d–f) holotypeı female: (d) posterior segments and gonopodsı ventral viewı scale 500 μm; (e) ocelli and Tömösváry’s organı lateral viewı scale 250 μm; (f) right gonopodı lateral viewı scale 250 μm." figureDoi="http://doi.org/10.5281/zenodo.3673150" httpUri="https://zenodo.org/record/3673150/files/figure.png" pageId="4" pageNumber="900">Figure 2</figureCitation>
(e)). Lateral margins of forcipular coxosternite slantingı anterior margin with 2 + 2 smallı blunt teeth and with thick porodonts. Tergites without triangular posterolateral processı a line of setae along posterior border of TT 8ı 10ı 12 and 14. Legs 14 and 15 thicker than anterior ones in both sexes. Coxal pores 4
<emphasis id="B9168D7F4F00FFD1FCC5FF0EFC8CFF3D" box="[806,819,142,166]" italics="true" pageId="6" pageNumber="902">–</emphasis>
6ı round to ovateı arranged in 1 row. Female gonopods with 2 elongated conical spursı the second article of the female gonopods having a massive process and a strong slightly curved spine. Terminal claw of the third article of female gonopods simpleı with a small triangular protuberance on basal ventral side. Male gonopods short and small.
</paragraph>
</subSubSection>
<caption id="DF1D01E54F03FFD2FF43FA7FFE94F917" ID-DOI="http://doi.org/10.5281/zenodo.3673150" ID-Zenodo-Dep="3673150" httpUri="https://zenodo.org/record/3673150/files/figure.png" pageId="5" pageNumber="901" startId="5.[160,225,1535,1558]" targetBox="[166,1144,161,1504]" targetPageId="5">
<paragraph id="8BDD516D4F03FFD2FF43FA7FFE94F917" blockId="5.[160,1156,1535,1676]" pageId="5" pageNumber="901">
<emphasis id="B9168D7F4F03FFD2FF43FA7FFF43F98D" bold="true" box="[160,252,1535,1558]" pageId="5" pageNumber="901">Figure 2.</emphasis>
<taxonomicName id="4C622AEE4F03FFD2FEE7FA7FFE6EF98D" authorityName="Qiao, Qin, Ma &amp; Zhang" authorityYear="2019" box="[260,465,1535,1558]" class="Chilopoda" family="Lithobiidae" genus="Hessebius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="901" phylum="Arthropoda" rank="species" species="prospinosa" status="sp. nov.">
<emphasis id="B9168D7F4F03FFD2FEE7FA7FFE6EF98D" box="[260,465,1535,1558]" italics="true" pageId="5" pageNumber="901">Hessebius prospinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22530044F03FFD2FE3AFA7FFD95F98D" box="[473,554,1535,1558]" pageId="5" pageNumber="901" rank="species">sp. nov.</taxonomicNameLabel>
ı (a
<emphasis id="B9168D7F4F03FFD2FDA5FA7FFDECF98D" box="[582,595,1535,1558]" italics="true" pageId="5" pageNumber="901">–</emphasis>
c): paratypeı male: (a) habitusı dorsal viewı scale 1 mm; (b) forcipular coxosterniteı ventral viewı scale 500 μm; (c) posterior segments and gonopodsı ventral viewı scale 500 μm; (d
<emphasis id="B9168D7F4F03FFD2FE98F9BAFE37F9CA" box="[379,392,1594,1617]" italics="true" pageId="5" pageNumber="901">–</emphasis>
f) holotypeı female: (d) posterior segments and gonopodsı ventral viewı scale 500 μm; (e) ocelli and Tömösváry
<emphasis id="B9168D7F4F03FFD2FE08F9D7FE4FF9F5" box="[491,496,1623,1646]" italics="true" pageId="5" pageNumber="901">’</emphasis>
s organı lateral viewı scale 250 μm; (f) right gonopodı lateral viewı scale 250 μm.
</paragraph>
</caption>
<subSubSection id="C37802E64F00FFD1FF43FEE7FD98FDB5" pageId="6" pageNumber="902" type="discussion">
<paragraph id="8BDD516D4F00FFD1FF43FEE7FEB9FE1B" blockId="6.[160,1155,359,558]" box="[160,262,359,384]" pageId="6" pageNumber="902">
<emphasis id="B9168D7F4F00FFD1FF43FEE7FEB9FE1B" bold="true" box="[160,262,359,384]" italics="true" pageId="6" pageNumber="902">Remarks</emphasis>
</paragraph>
<paragraph id="8BDD516D4F00FFD1FF43FE0AFD98FDB5" blockId="6.[160,1155,359,558]" pageId="6" pageNumber="902">
<taxonomicName id="4C622AEE4F00FFD1FF43FE0AFE32FE38" authorityName="Qiao, Qin, Ma &amp; Zhang" authorityYear="2019" box="[160,397,394,419]" class="Chilopoda" family="Lithobiidae" genus="Hessebius" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="902" phylum="Arthropoda" rank="species" species="prospinosa" status="sp. nov.">
<emphasis id="B9168D7F4F00FFD1FF43FE0AFE32FE38" box="[160,397,394,419]" italics="true" pageId="6" pageNumber="902">Hessebius prospinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22530044F00FFD1FE7FFE0BFE45FE38" box="[412,506,395,419]" pageId="6" pageNumber="902" rank="species">sp. nov.</taxonomicNameLabel>
can be easily distinguished from the other
<taxonomicName id="4C622AEE4F00FFD1FBF9FE0AFB3CFE38" authorityName="Verhoeffi" authorityYear="1941" box="[1050,1155,394,419]" class="Chilopoda" family="Lithobiidae" genus="Hessebius" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="902" phylum="Arthropoda" rank="genus">
<emphasis id="B9168D7F4F00FFD1FBF9FE0AFB3CFE38" box="[1050,1155,394,419]" italics="true" pageId="6" pageNumber="902">Hessebius</emphasis>
</taxonomicName>
species that are known from
<collectingCountry id="F37511FD4F00FFD1FE19FE2EFD85FE5D" box="[506,570,430,454]" name="China" pageId="6" pageNumber="902">China</collectingCountry>
by the well-defined tarsal articulation on all legsı the strong spine on the massive expansion projecting from the second article of the female gonopodsı and the terminal claw of the female gonopods having a triangular protuberance on the inner aspect.
</paragraph>
</subSubSection>
<subSubSection id="C37802E64F00FFDFFF43FDE4FD7CFC35" lastPageId="8" lastPageNumber="904" pageId="6" pageNumber="902" type="description">
<paragraph id="8BDD516D4F00FFD1FF43FDE4FE99FDE6" blockId="6.[160,1155,612,706]" box="[160,294,612,637]" pageId="6" pageNumber="902">
<emphasis id="B9168D7F4F00FFD1FF43FDE4FE99FDE6" bold="true" box="[160,294,612,637]" italics="true" pageId="6" pageNumber="902">Description</emphasis>
</paragraph>
<paragraph id="8BDD516D4F00FFD1FF43FD08FF43FD59" blockId="6.[160,1155,612,706]" pageId="6" pageNumber="902">
Based on
<typeStatus id="54D9EFCF4F00FFD1FEFAFD08FEC2FD3B" box="[281,381,648,672]" pageId="6" pageNumber="902" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D649AE44F00FFD1FE72FD07FE1AFD04" box="[401,421,647,671]" pageId="6" pageNumber="902" type="female">♀</specimenCount>
)ı body length
<quantity id="4C9AFC884F00FFD1FDB9FD08FD7BFD3B" box="[602,708,648,672]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="6" pageNumber="902" unit="mm" value="14.2">14.2 mm</quantity>
; cephalic plate width 1.8 mmı length
<quantity id="4C9AFC884F00FFD1FF43FD2AFF46FD59" box="[160,249,682,706]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="902" unit="mm" value="1.6">1.6 mm</quantity>
.
</paragraph>
<paragraph id="8BDD516D4F00FFD1FF43FD79FEFEFCE1" blockId="6.[160,1155,761,890]" pageId="6" pageNumber="902">
<emphasis id="B9168D7F4F00FFD1FF43FD79FF4AFC89" bold="true" box="[160,245,761,786]" italics="true" pageId="6" pageNumber="902">Colour.</emphasis>
Antennae and sternites brown-yellow. Coxosternite light yellow. Distal part of forcipule reddish brown. Legs pale yellowı except legs 14 and 15 reddish brownı claws yellowish brown. Tergites rufousı with headı first tergiteı fifth tergiteı and terminal tergite darker.
</paragraph>
<paragraph id="8BDD516D4F00FFD1FF43FC30FBE3FBCF" blockId="6.[160,1156,944,1108]" pageId="6" pageNumber="902">
<emphasis id="B9168D7F4F00FFD1FF43FC30FEA6FC52" bold="true" box="[160,281,944,969]" italics="true" pageId="6" pageNumber="902">Antennae.</emphasis>
Length 4.2 mmı 29.5% of body length with 20 + 20 articles. Basal article the same width as lengthı second one markedly longer than wideı following articles gradually shortening and taperingı distal article markedly longer than wideı up to 1.8 times as long as penultimate article. Abundant setae on the surface of antennaeı less so on the basal articlesı gradually increasing in density to about article 5ı then more or less constant.
</paragraph>
<paragraph id="8BDD516D4F00FFD1FF43FB0AFBF0FB73" blockId="6.[160,1154,1162,1256]" pageId="6" pageNumber="902">
<emphasis id="B9168D7F4F00FFD1FF43FB0AFEEEFB38" bold="true" box="[160,337,1162,1187]" italics="true" pageId="6" pageNumber="902">Cephalic plate.</emphasis>
Heart-shapedı smoothı posterior margin concave without median thickening; several symmetric setae scattered along the marginal ridge of the cephalic plate; a strong transverse suture dividing into 2 branchesı projection of lateral marginal discontinuous.
</paragraph>
<paragraph id="8BDD516D4F00FFD1FF43FA9FFC08FAE6" blockId="6.[160,1156,1311,1474]" pageId="6" pageNumber="902">
<emphasis id="B9168D7F4F00FFD1FF43FA9FFF56FAA3" bold="true" box="[160,233,1311,1336]" italics="true" pageId="6" pageNumber="902">Ocelli.</emphasis>
1 + 4ı 4ı 2 oval to rounded ocelli on each side in 3 irregular rows. Posterior ocellus ovalı larger than the seriate ocelli; seriate ocelli moderately smallı approximately equal in size. All ocelli domedı translucentı usually darkly pigmented.
</paragraph>
<paragraph id="8BDD516D4F00FFD1FF23FA08FBEEFA59" blockId="6.[160,1156,1311,1474]" pageId="6" pageNumber="902">
Tömösváry
<emphasis id="B9168D7F4F00FFD1FED9FA08FE80FA3B" box="[314,319,1416,1440]" italics="true" pageId="6" pageNumber="902">’</emphasis>
s organ: ovateı almost the same size as the adjoining ocelliı some distance from the adjoining ocelliı situated on ventral side of cephalic pleurite (
<figureCitation id="13594DE84F00FFD1FC29FA2AFB98FA59" box="[970,1063,1450,1474]" captionStart="Figure 2" captionStartId="5.[160,225,1535,1558]" captionTargetBox="[166,1144,161,1504]" captionTargetId="figure@5.[166,1149,142,1504]" captionTargetPageId="5" captionText="Figure 2. Hessebius prospinosa sp. nov.ı (a–c): paratypeı male: (a) habitusı dorsal viewı scale 1 mm; (b) forcipular coxosterniteı ventral viewı scale 500 μm; (c) posterior segments and gonopodsı ventral viewı scale 500 μm; (d–f) holotypeı female: (d) posterior segments and gonopodsı ventral viewı scale 500 μm; (e) ocelli and Tömösváry’s organı lateral viewı scale 250 μm; (f) right gonopodı lateral viewı scale 250 μm." figureDoi="http://doi.org/10.5281/zenodo.3673150" httpUri="https://zenodo.org/record/3673150/files/figure.png" pageId="6" pageNumber="902">Figure 2</figureCitation>
(e)).
</paragraph>
<paragraph id="8BDD516D4F00FFD1FF43FA79FC29F924" blockId="6.[160,1156,1529,1727]" pageId="6" pageNumber="902">
<emphasis id="B9168D7F4F00FFD1FF43FA79FEF9F989" bold="true" box="[160,326,1529,1554]" italics="true" pageId="6" pageNumber="902">Coxosternites.</emphasis>
Subtrapezoidal with narrowı very slightly recurved dental margin. Median diastema moderately wideı weakly V-shaped; anterior margin having the teeth developed as smallı blunt bulbs that are encircled by a narrow rimı sclerotisation is markedly greater on the bulbs than on the rims. Porodonts thick and longı lying posterolateral to the most lateral teeth on the ventral side of the coxosternite; some short and long setae scattered on the ventral side of coxosternite.
</paragraph>
<paragraph id="8BDD516D4F01FFD0FF43FF0EFCC7FE27" blockId="7.[160,1157,142,444]" pageId="7" pageNumber="903">
<emphasis id="B9168D7F4F01FFD0FF43FF0EFEBBFF3C" bold="true" box="[160,260,142,167]" italics="true" pageId="7" pageNumber="903">Tergites.</emphasis>
Smoothı without wrinkles. Angles of posterolateral corners of tergites all rounded without triangular projections. T1 narrower than the cephalic plate and T3ı the cephalic plate almost the same width as T3. Posterior margin of TT 1ı 8 straightı continuous. Posterior marginal ridge of TT 3 and 5 concaveı continuous. Posterior margin of TT 7ı 9ı 11ı 12 and 14 slightly convexı T10 slightly concaveı all without ridge. All posterior angles generally roundedı without triangular projections. Lateral marginal ridge of all tergites continuous. Tiny setae scattered very sparsely over the surface. Short to long setae sparsely dispersed along lateral borders and posterior cornersı a band of setae on posterior border of TT 8ı 10ı 12ı 14.
</paragraph>
<paragraph id="8BDD516D4F01FFD0FF43FE6AFEB7FD16" blockId="7.[160,1156,490,653]" pageId="7" pageNumber="903">
<emphasis id="B9168D7F4F01FFD0FF43FE6AFEB2FD98" bold="true" box="[160,269,490,515]" italics="true" pageId="7" pageNumber="903">Sternites.</emphasis>
Trapeziformı posterior side narrower. SS 1
<emphasis id="B9168D7F4F01FFD0FD10FE6AFCBFFD99" box="[755,768,490,514]" italics="true" pageId="7" pageNumber="903">–</emphasis>
11 glabrousı about 8
<emphasis id="B9168D7F4F01FFD0FC0AFE6AFC49FD99" box="[1001,1014,490,514]" italics="true" pageId="7" pageNumber="903">–</emphasis>
10 comparatively short to long setae scattered sparsely on anterior and posterior lateral margin of each sternite. Three irregular lines of setae on posterior part of SS 10ı 11. Numerous minute seta surround the surface of SS 12
<emphasis id="B9168D7F4F01FFD0FCFCFDD2FC93FDF1" box="[799,812,594,618]" italics="true" pageId="7" pageNumber="903">–</emphasis>
15 except that the middle is glabrous.
</paragraph>
<paragraph id="8BDD516D4F01FFD0FF43FD3AFCCBFBB5" blockId="7.[160,1155,698,1070]" pageId="7" pageNumber="903">
<emphasis id="B9168D7F4F01FFD0FF43FD3AFF64FD48" bold="true" box="[160,219,698,723]" italics="true" pageId="7" pageNumber="903">Legs.</emphasis>
Clear tarsal articulation on all legs. Anterior and posterior accessory spines on legs 1
<emphasis id="B9168D7F4F01FFD0FF4DFD5EFF04FD6D" box="[174,187,734,758]" italics="true" pageId="7" pageNumber="903">–</emphasis>
14ı no accessory spurs on leg 15. Anterior accessory spines long and slenderı about 0.5 times the length of principal clawı posterior one slightly strongerı about 0.3 times the length of principal claw. The anterior accessory spines form relatively large angles with the pretarsus. The posterior accessory spines form relatively small angles with the pretarsus. Leg pairs 14 and 15 longer and thicker than 1
<emphasis id="B9168D7F4F01FFD0FCA9FCE9FCE8FC1A" box="[842,855,873,897]" italics="true" pageId="7" pageNumber="903">–</emphasis>
13. Leg 15 longest of allı with tarsus 1 about 4.0 times longer than wide. Tarsus 2 about 60% length of tarsus 1. Abundant glandular pores on ventral surfaces of femurı tibia and tarsus of legs 14 and 15. Short to moderately long setae scattered over the surface of legs 1
<emphasis id="B9168D7F4F01FFD0FBF0FC51FB9FFC72" box="[1043,1056,977,1001]" italics="true" pageId="7" pageNumber="903">–</emphasis>
13. Tarsi generally more setoseı few setae on legs 14
<emphasis id="B9168D7F4F01FFD0FD5EFC73FD75FB90" box="[701,714,1011,1035]" italics="true" pageId="7" pageNumber="903">–</emphasis>
15. Tarsi 1 and 2 of legs 1
<emphasis id="B9168D7F4F01FFD0FBF8FC73FB97FB90" box="[1051,1064,1011,1035]" italics="true" pageId="7" pageNumber="903">–</emphasis>
13 with combs of setae on ventral side. Legs
<emphasis id="B9168D7F4F01FFD0FDA1FB96FDF7FBB5" box="[578,584,1046,1070]" italics="true" pageId="7" pageNumber="903">’</emphasis>
plectrotaxy: as in
<tableCitation id="C6E064D64F01FFD0FCF9FB96FCD1FBB5" box="[794,878,1046,1070]" captionStart="Table A" captionStartId="23.[159,215,212,235]" captionTargetBox="[169,1143,336,1762]" captionText="Table A1. Species used for COI sequence analysisı sequence referencesı GenBank accession numbers Voucher and Locality. ZSM = Zoologische Staatssammlung Münchenı Germany; Vouchers in Australian Museum prefix with AM KS; MCZ = Australian Museumı Sydney; SMNG = Senckenberg Museum of Natural History; MQ = Maqin; HY = Haiyan; RKZ = Rikaze." httpUri="http://table.plazi.org/id/DF1D01E54F11FFC0FF7CFF54FCE1FED8" pageId="7" pageNumber="903" tableUuid="DF1D01E54F11FFC0FF7CFF54FCE1FED8">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BDD516D4F01FFD0FF43FBDBFE8DFB21" blockId="7.[160,1156,1115,1210]" pageId="7" pageNumber="903">
<emphasis id="B9168D7F4F01FFD0FF43FBDBFE8DFBEF" bold="true" box="[160,306,1115,1140]" italics="true" pageId="7" pageNumber="903">Coxal pores.</emphasis>
On the left 5664 and on the right 5665ı arranged in a singleı regular rowı circular or ovate. Coxal pore field set in a shallow grooveı with the fringe of coxal pore field distinct.
</paragraph>
<paragraph id="8BDD516D4F01FFDFFF43FB67FD80FE45" blockId="7.[160,1156,1255,1419]" lastBlockId="8.[160,1155,142,478]" lastPageId="8" lastPageNumber="904" pageId="7" pageNumber="903">
<emphasis id="B9168D7F4F01FFD0FF43FB67FE60FA9B" bold="true" box="[160,479,1255,1280]" italics="true" pageId="7" pageNumber="903">Female posterior segment.</emphasis>
Sternite of female genital segment usually well chitinisedı wider than long; nearly glabrousı fringed with longer setae along the posterior lateral margin. Posterior margin of genital sternite deeply concave between the condyles of gonopodsı except for a smallı medianı approximately tongue-shaped bulge. Female gonopods divided into 3 articles as usual in Lithobiidaeı the first article moderately broadı bearing 28
<emphasis id="B9168D7F4F0EFFDFFE84FF0EFECBFF3D" box="[359,372,142,166]" italics="true" pageId="8" pageNumber="904">–</emphasis>
30 short to moderately long setaeı arranged in 4 irregular rows. Seven short to long blunt spines along the dorsolateral ridge of the first article. The first article bearing 2 + 2 moderately longı bullet-shaped spursı inner spur slightly smaller and more anterior than the outer (
<figureCitation id="13594DE84F0EFFDFFE58FF76FDA4FE95" box="[443,539,246,270]" captionStart="Figure 2" captionStartId="5.[160,225,1535,1558]" captionTargetBox="[166,1144,161,1504]" captionTargetId="figure@5.[166,1149,142,1504]" captionTargetPageId="5" captionText="Figure 2. Hessebius prospinosa sp. nov.ı (a–c): paratypeı male: (a) habitusı dorsal viewı scale 1 mm; (b) forcipular coxosterniteı ventral viewı scale 500 μm; (c) posterior segments and gonopodsı ventral viewı scale 500 μm; (d–f) holotypeı female: (d) posterior segments and gonopodsı ventral viewı scale 500 μm; (e) ocelli and Tömösváry’s organı lateral viewı scale 250 μm; (f) right gonopodı lateral viewı scale 250 μm." figureDoi="http://doi.org/10.5281/zenodo.3673150" httpUri="https://zenodo.org/record/3673150/files/figure.png" pageId="8" pageNumber="904">Figure 2</figureCitation>
(d)). The second article with 5 moderately long setaeı arranged in 2 irregular rows. Nine short to long blunt spines along the dorsolateral ridge of the second article; the dorsal terminal part of the second article of the female gonopods strongly extending backwards and a robust slightly curved spine at the end of the massive projection (
<figureCitation id="13594DE84F0EFFDFFE28FE01FD97FE02" box="[459,552,385,409]" captionStart="Figure 2" captionStartId="5.[160,225,1535,1558]" captionTargetBox="[166,1144,161,1504]" captionTargetId="figure@5.[166,1149,142,1504]" captionTargetPageId="5" captionText="Figure 2. Hessebius prospinosa sp. nov.ı (a–c): paratypeı male: (a) habitusı dorsal viewı scale 1 mm; (b) forcipular coxosterniteı ventral viewı scale 500 μm; (c) posterior segments and gonopodsı ventral viewı scale 500 μm; (d–f) holotypeı female: (d) posterior segments and gonopodsı ventral viewı scale 500 μm; (e) ocelli and Tömösváry’s organı lateral viewı scale 250 μm; (f) right gonopodı lateral viewı scale 250 μm." figureDoi="http://doi.org/10.5281/zenodo.3673150" httpUri="https://zenodo.org/record/3673150/files/figure.png" pageId="8" pageNumber="904">Figure 2</figureCitation>
(f)). Five moderately long setae on the surface of third article. Terminal claw simpleı slender and sharp (
<figureCitation id="13594DE84F0EFFDFFD39FE24FC85FE27" box="[730,826,420,444]" captionStart="Figure 2" captionStartId="5.[160,225,1535,1558]" captionTargetBox="[166,1144,161,1504]" captionTargetId="figure@5.[166,1149,142,1504]" captionTargetPageId="5" captionText="Figure 2. Hessebius prospinosa sp. nov.ı (a–c): paratypeı male: (a) habitusı dorsal viewı scale 1 mm; (b) forcipular coxosterniteı ventral viewı scale 500 μm; (c) posterior segments and gonopodsı ventral viewı scale 500 μm; (d–f) holotypeı female: (d) posterior segments and gonopodsı ventral viewı scale 500 μm; (e) ocelli and Tömösváry’s organı lateral viewı scale 250 μm; (f) right gonopodı lateral viewı scale 250 μm." figureDoi="http://doi.org/10.5281/zenodo.3673150" httpUri="https://zenodo.org/record/3673150/files/figure.png" pageId="8" pageNumber="904">Figure 2</figureCitation>
(f))ı having moderately small protuberance on ventral-lateral side.
</paragraph>
<caption id="DF1D01E54F01FFD0FF43FA45FB3AFA47" ID-Table-UUID="DF1D01E54F01FFD0FF43FA45FB3AFA47" box="[160,1157,1476,1500]" httpUri="http://table.plazi.org/id/DF1D01E54F01FFD0FF43FA45FB3AFA47" pageId="7" pageNumber="903" targetBox="[171,1145,1512,1717]" targetIsTable="true" targetPageId="7">
<paragraph id="8BDD516D4F01FFD0FF43FA45FB3AFA47" blockId="7.[160,1157,1476,1500]" box="[160,1157,1476,1500]" pageId="7" pageNumber="903">
<emphasis id="B9168D7F4F01FFD0FF43FA45FF4CFA47" bold="true" box="[160,243,1477,1500]" pageId="7" pageNumber="903">Table 1.</emphasis>
<taxonomicName id="4C622AEE4F01FFD0FF18FA45FE78FA47" authorityName="Qiao, Qin, Ma &amp; Zhang" authorityYear="2019" box="[251,455,1477,1500]" class="Chilopoda" family="Lithobiidae" genus="Hessebius" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="903" phylum="Arthropoda" rank="species" species="prospinosa" status="sp. nov.">
<emphasis id="B9168D7F4F01FFD0FF18FA45FE78FA47" box="[251,455,1477,1500]" italics="true" pageId="7" pageNumber="903">Hessebius prospinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22530044F01FFD0FE2CFA45FD9FFA47" box="[463,544,1477,1500]" pageId="7" pageNumber="903" rank="species">sp. nov.</taxonomicNameLabel>
: legs
<emphasis id="B9168D7F4F01FFD0FDB1FA44FDE8FA40" box="[594,599,1476,1499]" italics="true" pageId="7" pageNumber="903">’</emphasis>
plectrotaxy; letters in brackets indicate variable spines.
</paragraph>
</caption>
<paragraph id="8BDD516D4F01FFD0FE24FA68FBBFF92E" pageId="7" pageNumber="903">
<table id="F962A3CD4F010028FF48FA68FBC6F92E" box="[171,1145,1512,1717]" colsContinueIn="12.[171,1144,207,436]" gridcols="11" gridrows="8" pageId="7" pageNumber="903">
<tr id="3552532F4F010028FF48FA68FBC6FA60" box="[171,1145,1512,1531]" gridrow="0" pageId="7" pageNumber="903" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="76833A534F010028FE20FA68FDBDFA60" box="[451,514,1512,1531]" gridcol="3" gridrow="0" pageId="7" pageNumber="903">Ventral</th>
<th id="76833A534F010028FC6EFA68FC73FA60" box="[909,972,1512,1531]" gridcol="8" gridrow="0" pageId="7" pageNumber="903">Dorsal</th>
</tr>
<tr id="3552532F4F010028FF48F98AFBC6F986" box="[171,1145,1546,1565]" gridrow="1" pageId="7" pageNumber="903">
<th id="76833A534F010028FF48F98AFF61F986" box="[171,222,1546,1565]" gridcol="0" gridrow="1" pageId="7" pageNumber="903">Legs</th>
<td id="76833A534F010028FEC2F98AFE93F986" box="[289,300,1546,1565]" gridcol="1" gridrow="1" pageId="7" pageNumber="903">C</td>
<td id="76833A534F010028FE8CF98AFEC0F986" box="[367,383,1546,1565]" gridcol="2" gridrow="1" pageId="7" pageNumber="903">Tr</td>
<td id="76833A534F010028FE20F98AFDBDF986" box="[451,514,1546,1565]" gridcol="3" gridrow="1" pageId="7" pageNumber="903">P</td>
<td id="76833A534F010028FDD5F98AFDE4F986" box="[566,603,1546,1565]" gridcol="4" gridrow="1" pageId="7" pageNumber="903">F</td>
<td id="76833A534F010028FD6DF98AFD17F986" box="[654,680,1546,1565]" gridcol="5" gridrow="1" pageId="7" pageNumber="903">Ti</td>
<td id="76833A534F010028FD08F98AFD49F986" box="[747,758,1546,1565]" gridcol="6" gridrow="1" pageId="7" pageNumber="903">C</td>
<td id="76833A534F010028FCDAF98AFCF6F986" box="[825,841,1546,1565]" gridcol="7" gridrow="1" pageId="7" pageNumber="903">Tr</td>
<td id="76833A534F010028FC6EF98AFC73F986" box="[909,972,1546,1565]" gridcol="8" gridrow="1" pageId="7" pageNumber="903">P</td>
<td id="76833A534F010028FC16F98AFBAAF986" box="[1013,1045,1546,1565]" gridcol="9" gridrow="1" pageId="7" pageNumber="903">F</td>
<td id="76833A534F010028FBBBF98AFBC6F986" box="[1112,1145,1546,1565]" gridcol="10" gridrow="1" pageId="7" pageNumber="903">Ti</td>
</tr>
<tr id="3552532F4F010028FF48F9AAFBC6F9A6" box="[171,1145,1578,1597]" gridrow="2" pageId="7" pageNumber="903" rowspan-1="1" rowspan-2="1" rowspan-6="1" rowspan-7="1">
<th id="76833A534F010028FF48F9AAFF61F9A6" box="[171,222,1578,1597]" gridcol="0" gridrow="2" pageId="7" pageNumber="903">1</th>
<td id="76833A534F010028FE20F9AAFDBDF9A6" box="[451,514,1578,1597]" gridcol="3" gridrow="2" pageId="7" pageNumber="903">mp</td>
<td id="76833A534F010028FDD5F9AAFDE4F9A6" box="[566,603,1578,1597]" gridcol="4" gridrow="2" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FD6DF9AAFD17F9A6" box="[654,680,1578,1597]" gridcol="5" gridrow="2" pageId="7" pageNumber="903">am</td>
<td id="76833A534F010028FC6EF9AAFC73F9A6" box="[909,972,1578,1597]" gridcol="8" gridrow="2" pageId="7" pageNumber="903">ap</td>
<td id="76833A534F010028FC16F9AAFBAAF9A6" box="[1013,1045,1578,1597]" gridcol="9" gridrow="2" pageId="7" pageNumber="903">ap</td>
<td id="76833A534F010028FBBBF9AAFBC6F9A6" box="[1112,1145,1578,1597]" gridcol="10" gridrow="2" pageId="7" pageNumber="903">a(p)</td>
</tr>
<tr id="3552532F4F010028FF48F9C2FBC6F9CE" box="[171,1145,1602,1621]" gridrow="3" pageId="7" pageNumber="903" rowspan-1="1" rowspan-2="1" rowspan-6="1" rowspan-7="1">
<th id="76833A534F010028FF48F9C2FF61F9CE" box="[171,222,1602,1621]" gridcol="0" gridrow="3" pageId="7" pageNumber="903">
2
<emphasis id="B9168D7F4F01FFD0FF56F9C2FF7FF9CE" box="[181,192,1602,1621]" italics="true" pageId="7" pageNumber="903">–</emphasis>
10
</th>
<td id="76833A534F010028FE20F9C2FDBDF9CE" box="[451,514,1602,1621]" gridcol="3" gridrow="3" pageId="7" pageNumber="903">mp</td>
<td id="76833A534F010028FDD5F9C2FDE4F9CE" box="[566,603,1602,1621]" gridcol="4" gridrow="3" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FD6DF9C2FD17F9CE" box="[654,680,1602,1621]" gridcol="5" gridrow="3" pageId="7" pageNumber="903">am</td>
<td id="76833A534F010028FC6EF9C2FC73F9CE" box="[909,972,1602,1621]" gridcol="8" gridrow="3" pageId="7" pageNumber="903">ap</td>
<td id="76833A534F010028FC16F9C2FBAAF9CE" box="[1013,1045,1602,1621]" gridcol="9" gridrow="3" pageId="7" pageNumber="903">ap</td>
<td id="76833A534F010028FBBBF9C2FBC6F9CE" box="[1112,1145,1602,1621]" gridcol="10" gridrow="3" pageId="7" pageNumber="903">ap</td>
</tr>
<tr id="3552532F4F010028FF48F9DAFBC6F9F6" box="[171,1145,1626,1645]" gridrow="4" pageId="7" pageNumber="903" rowspan-1="1" rowspan-2="1" rowspan-6="1" rowspan-7="1">
<th id="76833A534F010028FF48F9DAFF61F9F6" box="[171,222,1626,1645]" gridcol="0" gridrow="4" pageId="7" pageNumber="903">11</th>
<td id="76833A534F010028FE20F9DAFDBDF9F6" box="[451,514,1626,1645]" gridcol="3" gridrow="4" pageId="7" pageNumber="903">(a)mp</td>
<td id="76833A534F010028FDD5F9DAFDE4F9F6" box="[566,603,1626,1645]" gridcol="4" gridrow="4" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FD6DF9DAFD17F9F6" box="[654,680,1626,1645]" gridcol="5" gridrow="4" pageId="7" pageNumber="903">am</td>
<td id="76833A534F010028FC6EF9DAFC73F9F6" box="[909,972,1626,1645]" gridcol="8" gridrow="4" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FC16F9DAFBAAF9F6" box="[1013,1045,1626,1645]" gridcol="9" gridrow="4" pageId="7" pageNumber="903">ap</td>
<td id="76833A534F010028FBBBF9DAFBC6F9F6" box="[1112,1145,1626,1645]" gridcol="10" gridrow="4" pageId="7" pageNumber="903">ap</td>
</tr>
<tr id="3552532F4F010028FF48F9F2FBC6F91E" box="[171,1145,1650,1669]" gridrow="5" pageId="7" pageNumber="903" rowspan-1="1" rowspan-2="1" rowspan-7="1">
<th id="76833A534F010028FF48F9F2FF61F91E" box="[171,222,1650,1669]" gridcol="0" gridrow="5" pageId="7" pageNumber="903">
12
<emphasis id="B9168D7F4F01FFD0FF5CF9F2FF75F91E" box="[191,202,1650,1669]" italics="true" pageId="7" pageNumber="903">–</emphasis>
13
</th>
<td id="76833A534F010028FE20F9F2FDBDF91E" box="[451,514,1650,1669]" gridcol="3" gridrow="5" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FDD5F9F2FDE4F91E" box="[566,603,1650,1669]" gridcol="4" gridrow="5" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FD6DF9F2FD17F91E" box="[654,680,1650,1669]" gridcol="5" gridrow="5" pageId="7" pageNumber="903">am</td>
<td id="76833A534F010028FD08F9F2FD49F91E" box="[747,758,1650,1669]" gridcol="6" gridrow="5" pageId="7" pageNumber="903">a</td>
<td id="76833A534F010028FC6EF9F2FC73F91E" box="[909,972,1650,1669]" gridcol="8" gridrow="5" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FC16F9F2FBAAF91E" box="[1013,1045,1650,1669]" gridcol="9" gridrow="5" pageId="7" pageNumber="903">(a)p</td>
<td id="76833A534F010028FBBBF9F2FBC6F91E" box="[1112,1145,1650,1669]" gridcol="10" gridrow="5" pageId="7" pageNumber="903">(a)p</td>
</tr>
<tr id="3552532F4F010028FF48F90AFBC6F906" box="[171,1145,1674,1693]" gridrow="6" pageId="7" pageNumber="903" rowspan-1="1" rowspan-7="1">
<th id="76833A534F010028FF48F90AFF61F906" box="[171,222,1674,1693]" gridcol="0" gridrow="6" pageId="7" pageNumber="903">14</th>
<td id="76833A534F010028FE8CF90AFEC0F906" box="[367,383,1674,1693]" gridcol="2" gridrow="6" pageId="7" pageNumber="903">m</td>
<td id="76833A534F010028FE20F90AFDBDF906" box="[451,514,1674,1693]" gridcol="3" gridrow="6" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FDD5F90AFDE4F906" box="[566,603,1674,1693]" gridcol="4" gridrow="6" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FD6DF90AFD17F906" box="[654,680,1674,1693]" gridcol="5" gridrow="6" pageId="7" pageNumber="903">am</td>
<td id="76833A534F010028FD08F90AFD49F906" box="[747,758,1674,1693]" gridcol="6" gridrow="6" pageId="7" pageNumber="903">a</td>
<td id="76833A534F010028FC6EF90AFC73F906" box="[909,972,1674,1693]" gridcol="8" gridrow="6" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FC16F90AFBAAF906" box="[1013,1045,1674,1693]" gridcol="9" gridrow="6" pageId="7" pageNumber="903">p</td>
<td id="76833A534F010028FBBBF90AFBC6F906" box="[1112,1145,1674,1693]" gridcol="10" gridrow="6" pageId="7" pageNumber="903">(p)</td>
</tr>
<tr id="3552532F4F010028FF48F922FBC6F92E" box="[171,1145,1698,1717]" gridrow="7" pageId="7" pageNumber="903" rowspan-1="1" rowspan-10="1" rowspan-7="1">
<th id="76833A534F010028FF48F922FF61F92E" box="[171,222,1698,1717]" gridcol="0" gridrow="7" pageId="7" pageNumber="903">15</th>
<td id="76833A534F010028FE8CF922FEC0F92E" box="[367,383,1698,1717]" gridcol="2" gridrow="7" pageId="7" pageNumber="903">m</td>
<td id="76833A534F010028FE20F922FDBDF92E" box="[451,514,1698,1717]" gridcol="3" gridrow="7" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FDD5F922FDE4F92E" box="[566,603,1698,1717]" gridcol="4" gridrow="7" pageId="7" pageNumber="903">am</td>
<td id="76833A534F010028FD6DF922FD17F92E" box="[654,680,1698,1717]" gridcol="5" gridrow="7" pageId="7" pageNumber="903">a</td>
<td id="76833A534F010028FD08F922FD49F92E" box="[747,758,1698,1717]" gridcol="6" gridrow="7" pageId="7" pageNumber="903">a</td>
<td id="76833A534F010028FC6EF922FC73F92E" box="[909,972,1698,1717]" gridcol="8" gridrow="7" pageId="7" pageNumber="903">amp</td>
<td id="76833A534F010028FC16F922FBAAF92E" box="[1013,1045,1698,1717]" gridcol="9" gridrow="7" pageId="7" pageNumber="903">p</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDD516D4F0EFFDFFF43FD9BFE4CFD00" blockId="8.[160,1155,539,667]" pageId="8" pageNumber="904">
<emphasis id="B9168D7F4F0EFFDFFF43FD9BFE7EFDAF" bold="true" box="[160,449,539,564]" italics="true" pageId="8" pageNumber="904">Male posterior segment.</emphasis>
Posterior margin of the sternite of the genital segment quite deeply concave between gonopodsı no bulge medially. Gonopods short and smallı only a small hemisphere bulgeı with 4 long setae on surfaceı terminal surface of male gonopods slightly sclerotised.
</paragraph>
<paragraph id="8BDD516D4F0EFFDFFF43FD58FEB1FD6A" blockId="8.[160,1157,728,822]" box="[160,270,728,753]" pageId="8" pageNumber="904">
<emphasis id="B9168D7F4F0EFFDFFF43FD58FEB1FD6A" bold="true" box="[160,270,728,753]" italics="true" pageId="8" pageNumber="904">Variation</emphasis>
</paragraph>
<paragraph id="8BDD516D4F0EFFDFFF43FD7BFDC3FCAD" blockId="8.[160,1157,728,822]" pageId="8" pageNumber="904">
Ocelli 1 + 4ı 4ı 2 or 1 + 4ı 3ı 1 or 1 + 3ı 3ı 2 on the cephalic plate. Coxal pores 5664ı 5555ı
<quantity id="4C9AFC884F0EFFDFFF43FC9EFF4AFCAD" box="[160,245,798,822]" metricMagnitude="2" metricUnit="m" metricValue="1.43891" pageId="8" pageNumber="904" unit="in" value="5665.0">5665 in</quantity>
female; 5565ı 5554ı
<quantity id="4C9AFC884F0EFFDFFE01FC9EFD89FCAD" box="[482,566,798,822]" metricMagnitude="2" metricUnit="m" metricValue="1.41097" pageId="8" pageNumber="904" unit="in" value="5555.0">5555 in</quantity>
male.
</paragraph>
<paragraph id="8BDD516D4F0EFFDFFF43FCF3FF46FC17" blockId="8.[160,707,883,942]" box="[160,249,883,908]" pageId="8" pageNumber="904">
<emphasis id="B9168D7F4F0EFFDFFF43FCF3FF46FC17" bold="true" box="[160,249,883,908]" italics="true" pageId="8" pageNumber="904">Habitat</emphasis>
</paragraph>
<paragraph id="8BDD516D4F0EFFDFFF43FC16FD7CFC35" blockId="8.[160,707,883,942]" box="[160,707,918,942]" pageId="8" pageNumber="904">Specimens were collected under railway stones.</paragraph>
</subSubSection>
<subSubSection id="C37802E64F0EFFDFFF43FC6BFCA4FBD2" pageId="8" pageNumber="904" type="etymology">
<paragraph id="8BDD516D4F0EFFDFFF43FC6BFEA2FB9F" blockId="8.[160,1155,1003,1097]" box="[160,285,1003,1028]" pageId="8" pageNumber="904">
<emphasis id="B9168D7F4F0EFFDFFF43FC6BFEA2FB9F" bold="true" box="[160,285,1003,1028]" italics="true" pageId="8" pageNumber="904">Etymology</emphasis>
</paragraph>
<paragraph id="8BDD516D4F0EFFDFFF43FB8EFCA4FBD2" blockId="8.[160,1155,1003,1097]" pageId="8" pageNumber="904">
The specific name refers to the strong spine on the massive expansion projecting from the second article of the female gonopods (
<figureCitation id="13594DE84F0EFFDFFD7BFBB1FD48FBD2" box="[664,759,1073,1097]" captionStart="Figure 2" captionStartId="5.[160,225,1535,1558]" captionTargetBox="[166,1144,161,1504]" captionTargetId="figure@5.[166,1149,142,1504]" captionTargetPageId="5" captionText="Figure 2. Hessebius prospinosa sp. nov.ı (a–c): paratypeı male: (a) habitusı dorsal viewı scale 1 mm; (b) forcipular coxosterniteı ventral viewı scale 500 μm; (c) posterior segments and gonopodsı ventral viewı scale 500 μm; (d–f) holotypeı female: (d) posterior segments and gonopodsı ventral viewı scale 500 μm; (e) ocelli and Tömösváry’s organı lateral viewı scale 250 μm; (f) right gonopodı lateral viewı scale 250 μm." figureDoi="http://doi.org/10.5281/zenodo.3673150" httpUri="https://zenodo.org/record/3673150/files/figure.png" pageId="8" pageNumber="904">Figure 2</figureCitation>
(f)).
</paragraph>
</subSubSection>
</treatment>
</document>
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