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<document id="B697D547B2C1D35DD7F87CBFCBF395CB" ID-CLB-Dataset="8554" ID-DOI="10.11646/zootaxa.4845.2.1" ID-GBIF-Dataset="25efbb90-245c-4391-8d17-fd75b5c35e86" ID-ISSN="1175-5326" ID-Zenodo-Dep="4406460" ID-ZooBank="184EB7D9-A6F8-44D4-B7BD-FBCC0B3466E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609422269866" checkinUser="plazi" docAuthor="Park, Kyu-Tek, Koo, Jun-Mo &amp; Minet, Joël" docDate="2020" docId="0392796AFFBEFFD65CE1992DFD42FE7C" docLanguage="en" docName="zootaxa.4845.2.1.pdf" docOrigin="Zootaxa 4845 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torodora ankasokella Park &amp; Koo &amp; Minet 2020, comb. nov." docType="treatment" docVersion="8" lastPageNumber="161" masterDocId="FFAB0112FFB7FFDC5C769F6AFFA0FFF6" masterDocTitle="Review of the Malagasy lecithocerid species described by Pierre Viette and deposited in MNHN (Paris), with new generic combinations and descriptions of a new subfamily and genus of Momphidae (Lepidoptera: Gelechioidea)" masterLastPageNumber="190" masterPageNumber="151" pageNumber="160" updateTime="1698850450038" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E0DF96400D058DE96B9F778CBB4019DA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7DBE87228FE66CE64DA4625D549F245B">
<mods:title id="09F7883BD0336D062DB463F453BE2C03">Review of the Malagasy lecithocerid species described by Pierre Viette and deposited in MNHN (Paris), with new generic combinations and descriptions of a new subfamily and genus of Momphidae (Lepidoptera: Gelechioidea)</mods:title>
</mods:titleInfo>
<mods:name id="DD35B6055CD714C57A5F792C2C2D7712" type="personal">
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<mods:roleTerm id="1D750EFD38A3B7F6F3BECB3B1E213A02">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0511E13FA841968F073DB3BFC56C79D3">Park, Kyu-Tek</mods:namePart>
<mods:affiliation id="84A45609B8017756A5049D23009152BA">Bioresource and Environmental Center, Incheon National University, Incheon, 22012 Korea;</mods:affiliation>
</mods:name>
<mods:name id="B637E84EEBFFD89202A54169ABEFE6FA" type="personal">
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<mods:namePart id="38472570CA3BA72B4776F951061B8BAE">Koo, Jun-Mo</mods:namePart>
<mods:affiliation id="08526023EEB1807E40135931D8FFA1F2">Department of Plant Medicine, Chungbuk National University, Cheongju, 28644 Korea.</mods:affiliation>
</mods:name>
<mods:name id="AAEE020B635B6B6E374C90F9DBC5D23D" type="personal">
<mods:role id="E9F2EDC6769D87BF54FECEF6EDB849FE">
<mods:roleTerm id="EA616D57154FD04190C02CFE5DC08777">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FDFE77448882B095693F7F7550B38598">Minet, Joël</mods:namePart>
<mods:affiliation id="0D19C1E8C5D050F0CE60D56246A022D3">Muséum National d’Histoire Naturelle (ISYEB, UMR 7205), Entomologie (C. P. 50), 45, rue Buffon, F- 75005 Paris, France.</mods:affiliation>
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<mods:typeOfResource id="BA9C1F3CAC7650F836EB45E960D7BB2E">text</mods:typeOfResource>
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<mods:title id="6993C89CE30A36CC3A4EE7EA2EA3DDD6">Zootaxa</mods:title>
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<mods:part id="C726276B37069E773C95DD088B22A3F1">
<mods:date id="0FAD3B9EC2C119D3F36A014203642203">2020</mods:date>
<mods:detail id="ECDCC254ECE5DAF60F9188659C35A396" type="pubDate">
<mods:number id="7C3DA226DC023BD5943D1E08A9C1AEBE">2020-09-02</mods:number>
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<mods:detail id="C4AA3A026AF2A63A56907A67DF99ACCD" type="volume">
<mods:number id="CEA45E0A751847B440C7315ED535CF40">4845</mods:number>
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<mods:classification id="33688F90339061E70E19B2169F8342F8">journal article</mods:classification>
<mods:identifier id="0F2EC3A294C0CC110C4E4B429A446785" type="CLB-Dataset">8554</mods:identifier>
<mods:identifier id="C899EAA27390D5DEFCD4620105A3FC60" type="DOI">10.11646/zootaxa.4845.2.1</mods:identifier>
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<treatment id="0392796AFFBEFFD65CE1992DFD42FE7C" ID-DOI="http://doi.org/10.5281/zenodo.4406793" ID-GBIF-Taxon="177202407" ID-Zenodo-Dep="4406793" LSID="urn:lsid:plazi:treatment:0392796AFFBEFFD65CE1992DFD42FE7C" httpUri="http://treatment.plazi.org/id/0392796AFFBEFFD65CE1992DFD42FE7C" lastPageId="10" lastPageNumber="161" pageId="9" pageNumber="160">
<subSubSection id="C3219BF7FFBEFFD55CE1992DFD42F994" box="[151,738,1607,1634]" pageId="9" pageNumber="160" type="nomenclature">
<paragraph id="8B84C87CFFBEFFD55CE1992DFD42F994" blockId="9.[151,738,1607,1670]" box="[151,738,1607,1634]" pageId="9" pageNumber="160">
<heading id="D0CC7F10FFBEFFD55CE1992DFD42F994" bold="true" box="[151,738,1607,1634]" fontSize="11" level="1" pageId="9" pageNumber="160" reason="1">
<emphasis id="B94F146EFFBEFFD55CE1992DFD42F994" bold="true" box="[151,738,1607,1634]" pageId="9" pageNumber="160">
<taxonomicName id="4C3BB3FFFFBEFFD55CE1992DFDEEF997" ID-CoL="9FPW9" authority="(Viette, 1968)" authorityName="Park &amp; Koo &amp; Minet" authorityYear="2020" baseAuthorityName="Viette" baseAuthorityYear="1968" box="[151,590,1607,1634]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="ankasokella" status="comb. nov.">
<emphasis id="B94F146EFFBEFFD55CE1992DFE3FF994" bold="true" box="[151,415,1607,1634]" italics="true" pageId="9" pageNumber="160">Torodora ankasokella</emphasis>
(Viette, 1968)
</taxonomicName>
,
<taxonomicNameLabel id="A27CA915FFBEFFD55E2A9922FD42F994" box="[604,738,1608,1634]" pageId="9" pageNumber="160" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFBEFFD55CE19901FE8AF970" box="[151,298,1643,1670]" pageId="9" pageNumber="160" type="description">
<paragraph id="8B84C87CFFBEFFD55CE19901FE8AF970" blockId="9.[151,738,1607,1670]" box="[151,298,1643,1670]" pageId="9" pageNumber="160">
(
<figureCitation id="1300D4F9FFBEFFD55CE99901FF4BF970" box="[159,235,1643,1670]" captionStart="FIGURE 5" captionStartId="10.[151,250,1409,1433]" captionTargetBox="[166,1422,432,1343]" captionTargetId="figure-118@10.[151,1435,417,1386]" captionTargetPageId="10" captionText="FIGURE 5. Torodora ankasokella (Viette, 1968), comb. nov.: A, holotype; B, labels; C, male genitalia, prep. J. Minet no. 1662; D, aedeagus; E, abdomen. Scale bar for the genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406470" httpUri="https://zenodo.org/record/4406470/files/figure.png" pageId="9" pageNumber="160">Figs. 5</figureCitation>
A–E)
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFBEFFD55CE199DAFD1EF93E" box="[151,702,1712,1736]" pageId="9" pageNumber="160" type="reference_group">
<paragraph id="8B84C87CFFBEFFD55CE199DAFD1EF93E" blockId="9.[151,702,1712,1736]" box="[151,702,1712,1736]" pageId="9" pageNumber="160">
<taxonomicName id="4C3BB3FFFFBEFFD55CE199DAFD9BF93E" ID-CoL="3SVDK" authority="Viette, 1968: 89" authorityName="Viette" authorityPageNumber="89" authorityYear="1968" box="[151,571,1712,1736]" class="Insecta" family="Lecithoceridae" genus="Lecithocera" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="ankasokella">
<emphasis id="B94F146EFFBEFFD55CE199DAFE30F93E" box="[151,400,1712,1736]" italics="true" pageId="9" pageNumber="160">Lecithocera ankasokella</emphasis>
Viette, 1968: 89
</taxonomicName>
. TL: Lakato.
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFBEFFD65CE1999CFDD7FEEB" lastPageId="10" lastPageNumber="161" pageId="9" pageNumber="160" type="materials_examined">
<paragraph id="8B84C87CFFBEFFD55CE1999CFBFBF8C2" blockId="9.[151,1437,1782,2025]" pageId="9" pageNumber="160">
<emphasis id="B94F146EFFBEFFD55CE1999CFF72F8E6" bold="true" box="[151,210,1782,1808]" pageId="9" pageNumber="160">Type</emphasis>
.
<materialsCitation id="3B53C221FFBEFFD55CA9999CFBFBF8C2" ID-GBIF-Occurrence="3027957331" collectingDate="1959-01-02" collectingDateMax="1959-01-10" collectingDateMin="1959-01-02" collectorName="P. Viette &amp; J. Minet" country="Madagascar" county="Ankasoka" elevation="1100" location="East" municipality="Lakato" pageId="9" pageNumber="160" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D3D03F5FFBEFFD55CA9999CFEA2F8E6" box="[223,258,1782,1808]" pageId="9" pageNumber="160" type="male">1♂</specimenCount>
(
<typeStatus id="548076DEFFBEFFD55D79999CFED7F8E7" box="[271,375,1782,1809]" pageId="9" pageNumber="160" type="holotype">holotype</typeStatus>
),
<location id="8EE49EA7FFBEFFD55DFF999DFE19F8E7" LSID="urn:lsid:plazi:treatment:0392796AFFBEFFD65CE1992DFD42FE7C:8EE49EA7FFBEFFD55DFF999DFE19F8E7" box="[393,441,1783,1809]" country="Madagascar" county="Ankasoka" municipality="Lakato" name="East" pageId="9" pageNumber="160">East</location>
<collectingCountry id="F32C88ECFFBEFFD55DC9999CFDE8F8E6" box="[447,584,1782,1808]" name="Madagascar" pageId="9" pageNumber="160">Madagascar</collectingCountry>
, road to
<collectingMunicipality id="6BE05206FFBEFFD55ED0999CFD57F8E6" box="[678,759,1782,1808]" pageId="9" pageNumber="160">Lakato</collectingMunicipality>
, at km 15,
<collectingCounty id="62E5B0F0FFBEFFD55F1A999CFC7FF8E6" box="[876,991,1782,1808]" pageId="9" pageNumber="160">Ankasoka</collectingCounty>
, alt.
<quantity id="4CC36599FFBEFFD55867999CFBCEF8E7" box="[1041,1134,1782,1809]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="160" unit="m" value="1100.0">
<elevation id="00162F4FFFBEFFD55867999CFBCEF8E7" box="[1041,1134,1782,1809]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="160" unit="m" value="1100.0">1,100 m</elevation>
</quantity>
,
<date id="FF85EEBCFFBEFFD5580F999CFAA0F8E7" box="[1145,1280,1782,1809]" pageId="9" pageNumber="160" value="1959-01-02" valueMax="1959-01-10" valueMin="1959-01-02">
<collectingDate id="EFC11754FFBEFFD5580F999CFAA0F8E7" box="[1145,1280,1782,1809]" pageId="9" pageNumber="160" value="1959-01-02" valueMax="1959-01-10" valueMin="1959-01-02">2–10 i 1959</collectingDate>
</date>
,
<collectorName id="26CEADAAFFBEFFD5597C999DFAC6F8E6" box="[1290,1382,1782,1809]" pageId="9" pageNumber="160">P. Viette</collectorName>
leg.;
<taxonomicName id="4C3BB3FFFFBEFFD55CE19870FE0AF8C2" authority="Park &amp; Koo &amp; Minet, 2020" authorityName="Park &amp; Koo &amp; Minet" authorityYear="2020" box="[151,426,1818,1844]" class="Insecta" family="Lecithoceridae" genus="Lecithocera" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="ankasokella" status="sp. nov.">
<emphasis id="B94F146EFFBEFFD55CE19870FE0AF8C2" box="[151,426,1818,1844]" italics="true" pageId="9" pageNumber="160">Lecithocera ankasokella</emphasis>
</taxonomicName>
<emphasis id="B94F146EFFBEFFD55DC79870FE4FF8C2" bold="true" box="[433,495,1818,1844]" pageId="9" pageNumber="160">
<taxonomicNameLabel id="A27CA915FFBEFFD55DC79870FE4FF8C2" box="[433,495,1818,1844]" pageId="9" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="548076DEFFBEFFD55D889870FDC9F8C2" box="[510,617,1818,1844]" pageId="9" pageNumber="160" type="holotype">Holotype</typeStatus>
,
<collectorName id="26CEADAAFFBEFFD55E029871FD74F8C2" box="[628,724,1818,1845]" pageId="9" pageNumber="160">P. Viette</collectorName>
; genitalia: prep.
<collectorName id="26CEADAAFFBEFFD55FF89871FC4AF8C2" box="[910,1002,1818,1845]" pageId="9" pageNumber="160">J. Minet</collectorName>
no. 1662.
</materialsCitation>
</paragraph>
<paragraph id="8B84C87CFFBEFFD55CB19854FD64F81F" blockId="9.[151,1437,1782,2025]" pageId="9" pageNumber="160">
<emphasis id="B94F146EFFBEFFD55CB19854FEABF8AE" bold="true" box="[199,267,1854,1880]" pageId="9" pageNumber="160">Adult</emphasis>
(
<figureCitation id="1300D4F9FFBEFFD55D6A9854FE02F8AF" box="[284,418,1854,1881]" captionStart="FIGURE 5" captionStartId="10.[151,250,1409,1433]" captionTargetBox="[166,1422,432,1343]" captionTargetId="figure-118@10.[151,1435,417,1386]" captionTargetPageId="10" captionText="FIGURE 5. Torodora ankasokella (Viette, 1968), comb. nov.: A, holotype; B, labels; C, male genitalia, prep. J. Minet no. 1662; D, aedeagus; E, abdomen. Scale bar for the genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406470" httpUri="https://zenodo.org/record/4406470/files/figure.png" pageId="9" pageNumber="160">Figs. 5A, B</figureCitation>
). Wingspan,
<quantity id="4CC36599FFBEFFD55E419854FD3FF8AF" box="[567,671,1854,1881]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="9" pageNumber="160" unit="mm" value="22.5">22.5 mm</quantity>
. This species is superficially similar to
<taxonomicName id="4C3BB3FFFFBEFFD558299854FF7BF88A" authority="(Viette, 1988)" authorityName="Park &amp; Koo &amp; Minet" authorityYear="2020" baseAuthorityName="Viette" baseAuthorityYear="1988" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="hiarakella">
<emphasis id="B94F146EFFBEFFD558299854FAE1F8AE" box="[1119,1345,1854,1880]" italics="true" pageId="9" pageNumber="160">Torodora hiarakella</emphasis>
(
<bibRefCitation id="EFAAB58DFFBEFFD559279854FF72F88A" author="Viette, P. E. L." pageId="9" pageNumber="160" pagination="99 - 109" refId="ref24718" refString="Viette, P. E. L. (1988) Nouveaux Microlepidopteres de Madagascar (Xyloryctidae, Stenomidae, Lecithoceridae, Gelechiidae) [Lep.]. Bulletin de la Societe entomologique de France, 93 (3 - 4), 99 - 109. https: // doi. org / 10.3406 / linly. 1986.10753" type="journal article" year="1988">Viette, 1988</bibRefCitation>
)
</taxonomicName>
, especially in the forewing colour pattern with orange-white costal band along margin, however, it can be distinguished from the latter by having two small, triangular costal patches beyond half. Hind wing has M
<subScript id="17BFCA39FFBEFFD5594E98F3FAE0F851" attach="left" box="[1336,1344,1945,1959]" fontSize="6" pageId="9" pageNumber="160">2</subScript>
present, M
<subScript id="17BFCA39FFBEFFD55CC698D7FF18F83D" attach="left" box="[176,184,1981,1995]" fontSize="6" pageId="9" pageNumber="160">3</subScript>
and CuA
<subScript id="17BFCA39FFBEFFD55D5298D7FE8CF83D" attach="left" box="[292,300,1981,1995]" fontSize="6" pageId="9" pageNumber="160">1</subScript>
stalked. Abdominal segments (
<figureCitation id="1300D4F9FFBEFFD55EFD98C0FD40F833" box="[651,736,1962,1989]" captionStart="FIGURE 5" captionStartId="10.[151,250,1409,1433]" captionTargetBox="[166,1422,432,1343]" captionTargetId="figure-118@10.[151,1435,417,1386]" captionTargetPageId="10" captionText="FIGURE 5. Torodora ankasokella (Viette, 1968), comb. nov.: A, holotype; B, labels; C, male genitalia, prep. J. Minet no. 1662; D, aedeagus; E, abdomen. Scale bar for the genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406470" httpUri="https://zenodo.org/record/4406470/files/figure.png" pageId="9" pageNumber="160">Fig. 5E</figureCitation>
) have strong spines triangularly distributed on segment II–VII dorsally, but rare in the segment I and not in
<collectionCode id="ED2A50B9FFBEFFD55EFA98A5FD60F81F" box="[652,704,1999,2025]" pageId="9" pageNumber="160">VIII</collectionCode>
.
</paragraph>
<paragraph id="8B84C87CFFBDFFD65CB19FFDFDD7FEEB" blockId="10.[151,1437,150,394]" pageId="10" pageNumber="161">
<emphasis id="B94F146EFFBDFFD65CB19FFDFECAFF47" box="[199,362,151,177]" italics="true" pageId="10" pageNumber="161">Male genitalia</emphasis>
(
<figureCitation id="1300D4F9FFBDFFD65D0E9FFDFE5BFF47" box="[376,507,150,177]" captionStart="FIGURE 5" captionStartId="10.[151,250,1409,1433]" captionTargetBox="[166,1422,432,1343]" captionTargetId="figure-118@10.[151,1435,417,1386]" captionTargetPageId="10" captionText="FIGURE 5. Torodora ankasokella (Viette, 1968), comb. nov.: A, holotype; B, labels; C, male genitalia, prep. J. Minet no. 1662; D, aedeagus; E, abdomen. Scale bar for the genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406470" httpUri="https://zenodo.org/record/4406470/files/figure.png" pageId="10" pageNumber="161">Figs. 5C, D</figureCitation>
): Uncus narrowly elongated, with an expansion medially. Valva broad basally; costa strongly concaved; ventral margin emarginated before cucullus; cucullus narrow basally, then slightly broadened distally with rounded outer margin. Juxta large, without caudal lobes. Aedeagus slender, as long as valva; cornuti consist of numerous spinules in a long sac.
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFBDFFD65CB19E4DFB76FE93" pageId="10" pageNumber="161" type="discussion">
<paragraph id="8B84C87CFFBDFFD65CB19E4DFB76FE93" blockId="10.[151,1437,150,394]" pageId="10" pageNumber="161">
<emphasis id="B94F146EFFBDFFD65CB19E4DFE9BFEB7" bold="true" box="[199,315,295,321]" pageId="10" pageNumber="161">Remarks.</emphasis>
The species was originally described, based on a single specimen, but the genitalia of the
<typeStatus id="548076DEFFBDFFD6594C9E4DFA3CFEB7" box="[1338,1436,295,321]" pageId="10" pageNumber="161" type="holotype">holotype</typeStatus>
was not dissected by Viette (1968) and the third author dissected and illustrated for the first time.
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFBDFFD65CB19E04FD42FE7C" box="[199,738,366,394]" pageId="10" pageNumber="161" type="distribution">
<paragraph id="8B84C87CFFBDFFD65CB19E04FD42FE7C" blockId="10.[151,1437,150,394]" box="[199,738,366,394]" pageId="10" pageNumber="161">
<emphasis id="B94F146EFFBDFFD65CB19E04FEC1FE7E" bold="true" box="[199,353,366,392]" pageId="10" pageNumber="161">Distribution.</emphasis>
<collectingCountry id="F32C88ECFFBDFFD65D119E05FE4EFE7F" box="[359,494,367,393]" name="Madagascar" pageId="10" pageNumber="161">Madagascar</collectingCountry>
(East) (Viette, 1968).
</paragraph>
</subSubSection>
</treatment>
</document>
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