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145 lines
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<document id="FF09B88DFAA1E6520A4484CEFDF7B173" ID-DOI="10.11646/zootaxa.4027.3.6" ID-GBIF-Dataset="aeea1e53-a725-44d7-a0c7-0c5811c12d6c" ID-ISSN="1175-5326" ID-Zenodo-Dep="241194" ID-ZooBank="A3F9E54F-A59B-4DFC-8F88-AFD5FEDE5B2C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461346717439" checkinUser="plazi" docAuthor="Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo & Bragagnolo, Cibele" docDate="2015" docId="03926F62FFEDFFE6D2EDFF1CFA763827" docLanguage="en" docName="zt04027p436.pdf" docOrigin="Zootaxa 4027 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holcobunus" docType="key" docVersion="8" lastPageNumber="427" masterDocId="FFAB171AFFEFFFE4D27AFF84FFED3B5E" masterDocTitle="Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae)" masterLastPageNumber="436" masterPageNumber="425" pageNumber="427" updateTime="1698620615736" updateUser="plazi">
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<mods:title id="10D5442A25935AFACE36CAE10D687F76">Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae)</mods:title>
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<mods:namePart id="43BCD5AEB1CF6041013DA4098C5F94E4">Tourinho, Ana Lúcia</mods:namePart>
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<mods:namePart id="FC9209E275DF72B5354FE6A3C5DA3ED9">Pinto-Da-Rocha, Ricardo</mods:namePart>
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<mods:name id="BA3409217EFDB65AC745F0506DEA4FD5" type="personal">
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<mods:namePart id="FBBC0A5FBAE33990276D43BC9CFC5B38">Bragagnolo, Cibele</mods:namePart>
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<mods:typeOfResource id="1357E8046BB978A6194BF2382D948703">text</mods:typeOfResource>
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<mods:title id="00A525C2E850395BB673CA88C6039462">Zootaxa</mods:title>
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<mods:part id="49617A754B8C53B7DB38544C36890B56">
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<mods:date id="B93306B435BFF5CBFF02AD5054576D45">2015</mods:date>
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<mods:number id="FC1F82CE5566A0E0921E6BAAF562CB49">4027</mods:number>
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<mods:number id="1223510D3A74713685DC4708D909DDA4">3</mods:number>
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<mods:start id="BFD30C915FAB771C37164575A3DBF285">425</mods:start>
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<mods:end id="BDEE0BE3ABF40BB9F6D0951412FC54DF">436</mods:end>
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<mods:identifier id="16B9567CC0240DB9F5CEDBCAA91EDC46" type="DOI">10.11646/zootaxa.4027.3.6</mods:identifier>
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<treatment id="03926F62FFEDFFE6D2EDFF1CFA763827" ID-DOI="http://doi.org/10.5281/zenodo.6119978" ID-GBIF-Taxon="119645381" ID-Zenodo-Dep="6119978" LSID="urn:lsid:plazi:treatment:03926F62FFEDFFE6D2EDFF1CFA763827" httpUri="http://treatment.plazi.org/id/03926F62FFEDFFE6D2EDFF1CFA763827" lastPageNumber="427" pageId="2" pageNumber="427">
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<subSubSection id="C3218DFFFFEDFFE6D2EDFF1CFDD03BEF" box="[151,573,151,178]" pageId="2" pageNumber="427" type="nomenclature">
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<paragraph id="8B84DE74FFEDFFE6D2EDFF1CFDD03BEF" blockId="2.[151,573,151,178]" box="[151,573,151,178]" pageId="2" pageNumber="427">
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<heading id="D0CC6918FFEDFFE6D2EDFF1CFDD03BEF" bold="true" box="[151,573,151,178]" fontSize="11" level="1" pageId="2" pageNumber="427" reason="1">
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<emphasis id="B94F0266FFEDFFE6D2EDFF1CFDD03BEF" bold="true" box="[151,573,151,178]" pageId="2" pageNumber="427">
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Key for the species of
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<taxonomicName id="4C3BA5F7FFEDFFE6D3D3FF13FDD03BEF" ID-CoL="7PKJ8" box="[425,573,151,177]" class="Arachnida" family="Sclerosomatidae" genus="Holcobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="427" phylum="Arthropoda" rank="genus">
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<emphasis id="B94F0266FFEDFFE6D3D3FF13FDD03BEF" bold="true" box="[425,573,151,177]" italics="true" pageId="2" pageNumber="427">Holcobunus</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3218DFFFFEDFFE6D2EDFF5AFA763827" pageId="2" pageNumber="427" type="key">
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<keyStep id="30CFC6D1FFEDFFE6D2EDFF5AFA763A15" pageId="2" pageNumber="427">
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<paragraph id="8B84DE74FFEDFFE6D2EDFF5AFA763A4F" blockId="2.[151,1436,222,889]" pageId="2" pageNumber="427">
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<keyLead id="30CA7D41FFEDFFE6D2EDFF5AFA763A4F" pageId="2" pageNumber="427">
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1. Eye mound armed with two anterior blunt, single-blunt pointed spines (Se
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<bibRefCitation id="EFAAA385FFEDFFE6D1D1FF5AFB623BAD" author="Tourinho" box="[939,1167,222,243]" pageId="2" pageNumber="427" refString="Tourinho, A. L. & Kury, A. B. (2001) Notes on Holcobunus Roewer, 1910 (Arachnida, Opiliones, Sclerosomatidae). Boletim do Museu Nacional, Zoologia, 461, 1 - 22." type="journal article" year="2001">Tourinho & Kury, 2001</bibRefCitation>
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), femur of leg I shorter than body, femur of leg II with six pseudoarticular nodules, femur of leg IV with three pseudoarticular nodules......
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<taxonomicName id="4C3BA5F7FFEDFFE6D762FF78FA763A4F" ID-CoL="7XXJ7" box="[1304,1435,251,273]" class="Arachnida" family="Sclerosomatidae" genus="Paratamboicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="427" phylum="Arthropoda" rank="species" species="bicornutus">
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<emphasis id="B94F0266FFEDFFE6D762FF78FA763A4F" box="[1304,1435,251,273]" italics="true" pageId="2" pageNumber="427">H. bicornutus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8B84DE74FFEDFFE6D2EDFE9DFA763A15" blockId="2.[151,1436,222,889]" pageId="2" pageNumber="427">
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<keyLead id="30CA7D41FFEDFFE6D2EDFE9DFA763A15" pageId="2" pageNumber="427">
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- Eye mound with two rows of more than three short granules (
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<figureCitation id="1300C2F1FFEDFFE6D151FE9DFC893A70" box="[811,868,281,302]" captionStart="FIGURE 3" captionStartId="5.[151,250,1618,1640]" captionTargetBox="[160,1427,380,1586]" captionTargetId="figure@5.[151,1436,375,1595]" captionTargetPageId="5" captionText="FIGURE 3. Holcobunus ibitirama sp. nov. Male holotype: A, dorsal view; B, ventral view; C, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/241197/files/figure.png" pageId="2" pageNumber="427">Fig. 3</figureCitation>
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C), femur of leg I longer than the body, femur of leg II with four, five or six pseudoarticular nodules.................................................................... 2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30CFC6D1FFEDFFE6D2EDFED7FA763AA5" pageId="2" pageNumber="427">
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<paragraph id="8B84DE74FFEDFFE6D2EDFED7FA763AFD" blockId="2.[151,1436,222,889]" pageId="2" pageNumber="427">
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<keyLead id="30CA7D41FFEDFFE6D2EDFED7FA763AFD" pageId="2" pageNumber="427">
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2. Body color black eye mound black or mound with a pair of yellowish stripes, females with light dorsal stripes and spots (
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<figureCitation id="1300C2F1FFEDFFE6D70EFED7FF033AD8" captionStart="FIGURE 5" captionStartId="7.[151,250,1980,2002]" captionTargetBox="[163,1419,312,1946]" captionTargetId="figure@7.[151,1436,303,1957]" captionTargetPageId="7" captionText="FIGURE 5. Holcobunus uaisoh sp. nov. Male holotype: A, dorsal view; B, ventral view; C, lateral view. Female paratype: D, dorsal view; E, ventral view; F, lateral view; Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/241199/files/figure.png" pageId="2" pageNumber="427">Fig. 5</figureCitation>
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), eye, femur of leg II with five or six pseudoarticular nodules, femur of leg IV with two or three pseudoarticular nodules................................................................................................
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<taxonomicName id="4C3BA5F7FFEDFFE6D744FE0AFA763AFD" ID-CoL="7WMVN" box="[1342,1435,397,419]" class="Arachnida" family="Sclerosomatidae" genus="Holcobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="427" phylum="Arthropoda" rank="species" species="uaisoh">
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<emphasis id="B94F0266FFEDFFE6D744FE0AFA763AFD" box="[1342,1435,397,419]" italics="true" pageId="2" pageNumber="427">H. uaisoh</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8B84DE74FFEDFFE6D2EDFE2FFA763AA5" blockId="2.[151,1436,222,889]" pageId="2" pageNumber="427">
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<keyLead id="30CA7D41FFEDFFE6D2EDFE2FFA763AA5" pageId="2" pageNumber="427">
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- Body uniformly yellow (
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<figureCitation id="1300C2F1FFEDFFE6D3B7FE2FFDE83A9E" box="[461,517,427,448]" captionStart="FIGURE 3" captionStartId="5.[151,250,1618,1640]" captionTargetBox="[160,1427,380,1586]" captionTargetId="figure@5.[151,1436,375,1595]" captionTargetPageId="5" captionText="FIGURE 3. Holcobunus ibitirama sp. nov. Male holotype: A, dorsal view; B, ventral view; C, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/241197/files/figure.png" pageId="2" pageNumber="427">Fig. 3</figureCitation>
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) or background yellow or brown with white or dark stripes or stains (
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<figureCitation id="1300C2F1FFEDFFE6D6E3FE2FFB3D3A9E" box="[1177,1232,427,448]" captionStart="FIGURE 1" captionStartId="3.[151,250,1958,1980]" captionTargetBox="[219,1370,203,1923]" captionTargetId="figure@3.[211,1376,195,1934]" captionTargetPageId="3" captionText="FIGURE 1. Holcobunus dissimilis sp. nov. Male holotype: A, dorsal view; B, ventral view; C, lateral view. Female paratype: D, dorsal view; E, ventral view; F lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/241195/files/figure.png" pageId="2" pageNumber="427">Fig. 1</figureCitation>
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), eye mound with 8– 9 short granules, femur of leg II with four or five pseudoarticular nodules, femur of leg IV with two, three or four pseudoarticular nodules.......................................................................................... 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30CFC6D1FFEDFFE6D2EDFD87FA7639D0" pageId="2" pageNumber="427">
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<paragraph id="8B84DE74FFEDFFE6D2EDFD87FA763968" blockId="2.[151,1436,222,889]" pageId="2" pageNumber="427">
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<keyLead id="30CA7D41FFEDFFE6D2EDFD87FA763968" pageId="2" pageNumber="427">
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3. Body color uniformly yellow without stripes or stains, femur of leg II with four pseudoarticular nodules, femur of leg IV with three or four pseudoarticular nodules.............................................................
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<taxonomicName id="4C3BA5F7FFEDFFE6D75EFDA5FA763968" ID-CoL="7WMV3" box="[1316,1435,544,566]" class="Arachnida" family="Sclerosomatidae" genus="Holcobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="427" phylum="Arthropoda" rank="species" species="ibitirama">
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<emphasis id="B94F0266FFEDFFE6D75EFDA5FA763968" box="[1316,1435,544,566]" italics="true" pageId="2" pageNumber="427">H. ibitirama</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8B84DE74FFEDFFE6D2EDFDBAFA7639D0" blockId="2.[151,1436,222,889]" pageId="2" pageNumber="427">
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<keyLead id="30CA7D41FFEDFFE6D2EDFDBAFA7639D0" pageId="2" pageNumber="427">
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- Body color yellow with dark stripes or stains on carapace and/or dorsal scute or background brown with a white arch and several white spots and stripes distributed on scute (
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<figureCitation id="1300C2F1FFEDFFE6D0C9FDD8FD06392F" box="[691,747,604,625]" captionStart="FIGURE 1" captionStartId="3.[151,250,1958,1980]" captionTargetBox="[219,1370,203,1923]" captionTargetId="figure@3.[211,1376,195,1934]" captionTargetPageId="3" captionText="FIGURE 1. Holcobunus dissimilis sp. nov. Male holotype: A, dorsal view; B, ventral view; C, lateral view. Female paratype: D, dorsal view; E, ventral view; F lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/241195/files/figure.png" pageId="2" pageNumber="427">Fig. 1</figureCitation>
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), femur of leg II with four or five pseudoarticular nodules, femur of leg IV with two, three or four pseudoarticular nodules........................................................... 4
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30CFC6D1FFEDFFE6D2EDFD12FA763827" pageId="2" pageNumber="427">
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<paragraph id="8B84DE74FFEDFFE6D2EDFD12FA76385D" blockId="2.[151,1436,222,889]" pageId="2" pageNumber="427">
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<keyLead id="30CA7D41FFEDFFE6D2EDFD12FA76385D" pageId="2" pageNumber="427">
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4. Body color yellow (when recently collected it tends to be orange), with median posterior region of dorsal scute and anterior border of coxae covered by a faint black spot in males, females with a lateral rounded spot in the dorsal scute, dorsal black faint spot lacking (See
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<bibRefCitation id="EFAAA385FFEDFFE6D3C4FD55FD4F39B8" author="Tourinho" box="[446,674,721,742]" pageId="2" pageNumber="427" refString="Tourinho, A. L. & Kury, A. B. (2001) Notes on Holcobunus Roewer, 1910 (Arachnida, Opiliones, Sclerosomatidae). Boletim do Museu Nacional, Zoologia, 461, 1 - 22." type="journal article" year="2001">Tourinho & Kury, 2001</bibRefCitation>
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), femur of leg II with five pseudoarticular nodules, femur of leg IV with two or three pseudoarticular nodules..................................................................
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<taxonomicName id="4C3BA5F7FFEDFFE6D76FFD6AFA76385D" ID-CoL="7WMVF" box="[1301,1435,749,771]" class="Arachnida" family="Sclerosomatidae" genus="Holcobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="427" phylum="Arthropoda" rank="species" species="nigripalpis">
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<emphasis id="B94F0266FFEDFFE6D76FFD6AFA76385D" box="[1301,1435,749,771]" italics="true" pageId="2" pageNumber="427">H. nigripalpis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8B84DE74FFEDFFE6D2EDFC88FA763827" blockId="2.[151,1436,222,889]" pageId="2" pageNumber="427">
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<keyLead id="30CA7D41FFEDFFE6D2EDFC88FA763827" pageId="2" pageNumber="427">
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- Body color yellow, with abdomen with large dark median stripe along its full extent, or background brown with a white arch surrounding anterior prosoma and several white spots and stripes distributed on scute, eye mound darkish brown (
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<figureCitation id="1300C2F1FFEDFFE6D728FCADFA603860" box="[1362,1421,809,830]" captionStart="FIGURE 1" captionStartId="3.[151,250,1958,1980]" captionTargetBox="[219,1370,203,1923]" captionTargetId="figure@3.[211,1376,195,1934]" captionTargetPageId="3" captionText="FIGURE 1. Holcobunus dissimilis sp. nov. Male holotype: A, dorsal view; B, ventral view; C, lateral view. Female paratype: D, dorsal view; E, ventral view; F lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/241195/files/figure.png" pageId="2" pageNumber="427">Fig. 1</figureCitation>
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), femur of leg II with four or five pseudoarticular nodules, femur of leg IV with two, three or four pseudoarticular nodules femoral.......................................................................................
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<taxonomicName id="4C3BA5F7FFEDFFE6D759FCE0FA763827" ID-CoL="7WMTW" box="[1315,1435,867,889]" class="Arachnida" family="Sclerosomatidae" genus="Holcobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="427" phylum="Arthropoda" rank="species" species="dissimilis">
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<emphasis id="B94F0266FFEDFFE6D759FCE0FA763827" box="[1315,1435,867,889]" italics="true" pageId="2" pageNumber="427">H. dissimilis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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