plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/84/87/038487AF1F3DFFE112EAC33B7163FB5C.xml

479 lines
73 KiB
XML
Raw Normal View History

added 03
2024-06-21 12:22:17 +02:00
<document id="006CF40E80038D7B9C5C9488164C4C35" ID-DOI="10.5281/zenodo.5401510" ID-ISSN="1638-9387" ID-Zenodo-Dep="5401510" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630334724433" checkinUser="marcus" docAuthor="Cumberlidge, Neil &amp; Sternberg, Richard V." docDate="2002" docId="038487AF1F3DFFE112EAC33B7163FB5C" docLanguage="en" docName="Zoosystema.2002.24.1.41-79.pdf" docOrigin="Zoosystema 24 (1)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Skelosophusa gollhardi" docType="treatment" docVersion="3" lastPageNumber="71" masterDocId="FFBDFFD71F21FFFF120AC105702FFFF9" masterDocTitle="The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea)" masterLastPageNumber="79" masterPageNumber="41" pageNumber="69" updateTime="1699232587643" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="75BD4B5B1C93E3314A16DF42245BBC01" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A8B47BF8013A296F46CD1F9C9FF0724D">
<mods:title id="ABCE3673DBFA7B93BF4840C3D26C1DF5">The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea)</mods:title>
</mods:titleInfo>
<mods:name id="60A8E2DEBB85987E555782DB171F044C" type="personal">
<mods:role id="BA2332F1B38B88231B3307FA4E6B1CBB">
<mods:roleTerm id="2092A331CC287F6BBEF2F92E1131EFBC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5A69A875B7C43046AE923C8842F11BBC">Cumberlidge, Neil</mods:namePart>
<mods:affiliation id="6A43D86122FDC72EAF8D1A663D4D69A3">Department of Biology, Northern Michigan University, Marquette, MI, 49855 (USA) ncumberl @ nmu. edu</mods:affiliation>
<mods:nameIdentifier id="692FD4BD926C12621F18FD73B3F30A22" type="email">ncumberl@nmu.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="829722B1AA53B2AAD267AE1C9C02EBF6" type="personal">
<mods:role id="DD2B604C6C82C06A589F076ED67F1AC6">
<mods:roleTerm id="6AB07BED7119034E3256289C1969BD48">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E9AAA655EDFB5DD6CC093AA977F98D6">Sternberg, Richard V.</mods:namePart>
<mods:affiliation id="C9C6D9EFF04ED1B2F0F07EE632CDF7A4">NHB- 163, Department of Invertebrate Zoology, Smithsonian Institution, Washington D. C., 20560 (USA) Sternberg. Richard @ nmnh. si. edu.</mods:affiliation>
<mods:nameIdentifier id="5FCB4959A2AA0C3AC1CADC479A856E2D" type="email">ichard@nmnh.si.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="38A6BBF4C4ABFF22558F3EC690D243A1">text</mods:typeOfResource>
<mods:relatedItem id="97C97C2AFA8B72E6321DD352B990305C" type="host">
<mods:titleInfo id="9FC9FFCD759B6520F54817FABF24BCAA">
<mods:title id="802C8A820D9C2F8AFCF1D9594EE0C970">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="991E9BDD227727D9F4BA43766AA8B376">
<mods:date id="170D5001AD2B2BDDA9768093D1C91163">2002</mods:date>
<mods:detail id="066B1AF02EFAD08EA4C375AECF3343B7" type="volume">
<mods:number id="E61F6FD33E3155EF3D5EA99B8F8BED43">24</mods:number>
</mods:detail>
<mods:detail id="4D5AF6E1480B14F5FB2249043DC1F1FB" type="issue">
<mods:number id="4FE8EDAFBF9CD7D967CC86C18929C912">1</mods:number>
</mods:detail>
<mods:extent id="EA232F1A59BA7083A7ED4D3CE434C6C6" unit="page">
<mods:start id="E4242912264FEBEC81A252EBBF508153">41</mods:start>
<mods:end id="49E2B7E34F8EF4B8C8FD290A3F969BD9">79</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CC4A77E207F15251843AC9147536114F">journal article</mods:classification>
<mods:identifier id="1D8D9F86B80CF5557C2E7AB9E1DD90B2" type="DOI">10.5281/zenodo.5401510</mods:identifier>
<mods:identifier id="A12FDD5ACD7AAD1A577CA94D33809B11" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="968B064BA48F758CD18E64DB7437DB3B" type="Zenodo-Dep">5401510</mods:identifier>
</mods:mods>
<treatment id="038487AF1F3DFFE112EAC33B7163FB5C" LSID="urn:lsid:plazi:treatment:038487AF1F3DFFE112EAC33B7163FB5C" httpUri="http://treatment.plazi.org/id/038487AF1F3DFFE112EAC33B7163FB5C" lastPageId="30" lastPageNumber="71" pageId="28" pageNumber="69">
<subSubSection id="C33765321F3DFFE312EAC33B724CFDA1" box="[224,611,574,600]" pageId="28" pageNumber="69" type="nomenclature">
<paragraph id="8B9236B91F3DFFE312EAC33B724CFDA1" blockId="28.[151,684,574,1177]" box="[224,611,574,600]" pageId="28" pageNumber="69">
<heading id="D0DA81D51F3DFFE312EAC33B724CFDA1" box="[224,611,574,600]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="69" reason="2">
<taxonomicName id="4C2D4D3A1F3DFFE312EAC33B724CFDA1" authority="(Bott, 1965)" baseAuthorityName="Bott" baseAuthorityYear="1965" box="[224,611,574,600]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="gollhardi">
<emphasis id="B959EAAB1F3DFFE312EAC33B71FAFDA1" bold="true" box="[224,469,574,600]" italics="true" pageId="28" pageNumber="69">Skelosophusa gollhardi</emphasis>
(
<bibRefCitation id="EFBC4B481F3DFFE313E9C33B7274FDA1" author="BOTT R." box="[483,603,574,600]" pageId="28" pageNumber="69" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott, 1965</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE312CFC3587116FD40" pageId="28" pageNumber="69" type="description">
<paragraph id="8B9236B91F3DFFE312CFC3587116FD40" blockId="28.[151,684,574,1177]" pageId="28" pageNumber="69">
(
<figureCitation id="13162A3C1F3DFFE312C6C358710CFD8E" box="[204,291,605,631]" captionStart="FIG" captionStartId="11.[151,162,1354,1371]" captionTargetBox="[192,1245,168,1311]" captionTargetId="figure-26@11.[187,1212,162,1321]" captionTargetPageId="11" captionText="FIG. 2. — Dorsal view of the carapace of adult males of Madagascan freshwater crabs (A-C based on Ng &amp; Takeda 1994); A, Skeleosophusa gollhardi (Bott, 1965), holotype (MNHN B-5165); B, S. prolixa Ng &amp; Takeda, 1994, holotype, (MRSN); C, S. eumeces Ng &amp; Takeda, 1994, holotype, (MRSN); D, Boreas uglowi n. gen., n. sp., holotype (MNHN B- 11292); E, Madagapotamon humberti Bott, 1965; F, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002, holotype, (AMNH 17833). Scale bar: A, 7.2 mm; B, 7.1 mm; C, 6.5 mm; D, 9 mm; E, 9.4 mm; F, 6 mm." figureDoi="http://doi.org/10.5281/zenodo.5401514" httpUri="https://zenodo.org/record/5401514/files/figure.png" pageId="28" pageNumber="69">Figs 2A</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE31324C3587167FD8E" box="[302,328,605,631]" captionStart="FIG" captionStartId="16.[151,162,1071,1088]" captionTargetBox="[155,1241,171,1034]" captionTargetId="figure-282@16.[138,1257,160,1041]" captionTargetPageId="16" captionText="FIG. 4. — Left mandible of Madagascan freshwater crabs, frontal view showing the palp (J, K based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2000) n. comb.; G, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2000; H, Boreas uglowi n. gen., n. sp.; I, Skeleosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Madagapotamon humberti Bott, 1965. Scale bar: A-I, K, L, 1.0 mm; J, 1.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5401518" httpUri="https://zenodo.org/record/5401518/files/figure.png" pageId="28" pageNumber="69">4I</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE3135EC3587142FD8E" box="[340,365,605,631]" captionStart="FIG" captionStartId="21.[151,162,1262,1279]" captionTargetBox="[248,1159,165,1192]" captionTargetId="figure-109@21.[228,1169,158,1197]" captionTargetPageId="21" captionText="FIG. 5. — Left third maxilliped of Madagascan freshwater crabs, frontal view (J, K based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; G, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp. Scale bar: A-D, 5.0 mm; E-L, 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5401520" httpUri="https://zenodo.org/record/5401520/files/figure.png" pageId="28" pageNumber="69">5I</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE31373C35871BDFD8E" box="[377,402,605,631]" captionStart="FIG" captionStartId="23.[151,162,768,785]" captionTargetBox="[258,1160,169,728]" captionTargetId="figure-337@23.[233,1163,157,745]" captionTargetPageId="23" captionText="FIG. 6. — Superior view of the carpus of the cheliped (pereiopod 1) of adult males of Madagascan freshwater crabs (J, K based on Ng &amp; Takeda 1994); A Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; G, Madagapotamon humberti Bott, 1965; H, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; I, Skeleosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp. Scale bar: A-D, 5.5 mm; E-L, 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5401522" httpUri="https://zenodo.org/record/5401522/files/figure.png" pageId="28" pageNumber="69">6I</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE31395C35971C3FD8F" box="[415,492,604,630]" captionStart="FIG" captionStartId="25.[151,162,1567,1584]" captionTargetBox="[244,1126,230,1497]" captionTargetId="figure-22@25.[232,1187,219,1521]" captionTargetPageId="25" captionText="FIG. 7. — Superior and inferior views of the merus of the cheliped (pereiopod 1) of adult males of Madagascan freshwater crabs; A, B, Hydrothelphusa agilis A. Milne-Edwards, 1872; C, D, H. madagascariensis (A. Milne-Edwards, 1872); E, F, H. bombetokensis (Rathbun, 1904); G, H, H. goudoti (H. Milne Edwards, 1853) n. comb.; I, J, Malagasya antongilensis (Rathbun, 1905) n. comb.; K, L, Madagapotamon humberti Bott, 1965; M, N, Skelosophusa gollhardi (Bott, 1965); O, P, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; Q, R, Boreas uglowi n. gen., n. sp. Scale bar: A-H, 5.0 mm; I-T, 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5401524" httpUri="https://zenodo.org/record/5401524/files/figure.png" pageId="28" pageNumber="69">7M, N</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE313F2C358726CFD8F" box="[504,579,604,631]" captionStart="FIG" captionStartId="32.[151,162,1363,1380]" captionTargetBox="[149,1247,169,1337]" captionTargetId="figure-94@32.[141,1252,160,1342]" captionTargetPageId="32" captionText="FIG. 10. — Right first gonopod (dorsal and ventral views) of adult males of Madagascan freshwater crabs; A, B, Boreas uglowi n. gen., n. sp.; C, D, Malagasya antongilensis (Rathbun, 1905) n. comb.; E, F, Skeleosophusa gollhardi (Bott, 1965); G, H, Madagapotamon humberti Bott, 1965; I, J, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002. Scale bar: 4.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5401530" httpUri="https://zenodo.org/record/5401530/files/figure.png" pageId="28" pageNumber="69">10E, F</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE31045C358725AFD8E" box="[591,629,605,631]" captionStart="FIG" captionStartId="35.[151,162,1408,1425]" captionTargetBox="[170,1219,175,1376]" captionTargetId="figure-28@35.[157,1238,162,1394]" captionTargetPageId="35" captionText="FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853); E, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; F, Malagasya antongilensis (Rathbun, 1905) n. comb.; G, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="28" pageNumber="69">11I</figureCitation>
)
<taxonomicName id="4C2D4D3A1F3DFFE3129DC3827270FD66" authority="Bott, 1965: 346" authorityName="Bott" authorityPageNumber="346" authorityYear="1965" box="[151,607,646,671]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="gollhardi">
<emphasis id="B959EAAB1F3DFFE3129DC3827189FD66" box="[151,422,647,671]" italics="true" pageId="28" pageNumber="69">Madagapotamon gollhardi</emphasis>
<bibRefCitation id="EFBC4B481F3DFFE313A5C3837270FD66" author="BOTT R." box="[431,607,646,671]" pageId="28" pageNumber="69" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott, 1965: 346</bibRefCitation>
</taxonomicName>
, fig. 8, pl. 5, figs 19-22.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE3129DC3CC7117FD03" pageId="28" pageNumber="69" type="reference_group">
<paragraph id="8B9236B91F3DFFE3129DC3CC7117FD03" blockId="28.[151,684,574,1177]" pageId="28" pageNumber="69">
<treatmentCitationGroup id="AB3D11971F3DFFE3129DC3CC7117FD03" pageId="28" pageNumber="69">
<taxonomicName id="4C2D4D3A1F3DFFE3129DC3CC7148FD18" baseAuthorityName="Bott" baseAuthorityYear="1965" box="[151,359,713,737]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="gollhardi">
<emphasis id="B959EAAB1F3DFFE3129DC3CC7148FD18" box="[151,359,713,737]" italics="true" pageId="28" pageNumber="69">Skelosophusa gollhardi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A8C10A81F3DFFE3138AC3CD725AFD18" author="NG P. K. L. &amp; TAKEDA M." box="[384,629,712,737]" page="164" pageId="28" pageNumber="69" year="1994">
<bibRefCitation id="EFBC4B481F3DFFE3138AC3CD725AFD18" author="NG P. K. L. &amp; TAKEDA M." box="[384,629,712,737]" pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda 1994: 164</bibRefCitation>
</treatmentCitation>
, 165, fig. 1a-f, table 3.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE3129DC20F72BBFC6E" pageId="28" pageNumber="69" type="materials_examined">
<paragraph id="8B9236B91F3DFFE3129DC20F7217FC96" blockId="28.[151,684,574,1177]" pageId="28" pageNumber="69">
MATERIAL EXAMINED. —
<emphasis id="B959EAAB1F3DFFE313A4C20F7217FCDB" bold="true" box="[430,568,778,802]" pageId="28" pageNumber="69">
<collectingCountry id="F33A76291F3DFFE313A4C20F721BFCDB" box="[430,564,778,802]" name="Madagascar" pageId="28" pageNumber="69">Madagascar</collectingCountry>
.
</emphasis>
Grotte de l’Ankara,
<date id="FF9310791F3DFFE312FCC221715BFCC5" box="[246,372,804,828]" pageId="28" pageNumber="69" value="1939-07-23">23.VII.1939</date>
, coll. R. Decary,
<specimenCount id="9D2BFD301F3DFFE31014C221727FFCC5" box="[542,592,804,828]" pageId="28" pageNumber="69" type="generic">1 ad.</specimenCount>
,
<typeStatus id="5496881B1F3DFFE31072C22170EEFCAC" pageId="28" pageNumber="69" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2D4D3A1F3DFFE312ECC2387273FCAC" authority="Bott, 1965" authorityName="Bott" authorityYear="1965" box="[230,604,829,853]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="gollhardi">
<emphasis id="B959EAAB1F3DFFE312ECC23871CBFCAC" box="[230,484,829,853]" italics="true" pageId="28" pageNumber="69">Madagapotamon gollhardi</emphasis>
<bibRefCitation id="EFBC4B481F3DFFE313E6C2387273FCAC" author="BOTT R." box="[492,604,829,853]" pageId="28" pageNumber="69" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott, 1965</bibRefCitation>
</taxonomicName>
, cw 23, cl 16, ch 7.5, fw
<quantity id="4CD59B5C1F3DFFE31336C2527158FC94" box="[316,375,854,879]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="69" unit="mm" value="6.0">6 mm</quantity>
(
<collectionCode id="ED3CAE7C1F3DFFE3138EC25271DFFC96" box="[388,496,855,879]" pageId="28" pageNumber="69">MNHN-B</collectionCode>
5165).
</paragraph>
<paragraph id="8B9236B91F3DFFE3129DC27B72BBFC6E" blockId="28.[151,684,574,1177]" box="[151,660,894,919]" pageId="28" pageNumber="69">
TYPELOCALITY. — Grotte de l’Ankara,
<collectingCountry id="F33A76291F3DFFE31010C27A72BFFC6E" box="[538,656,895,919]" name="Madagascar" pageId="28" pageNumber="69">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE3129DC2A2722EFC20" pageId="28" pageNumber="69" type="distribution">
<paragraph id="8B9236B91F3DFFE3129DC2A2722EFC20" blockId="28.[151,684,574,1177]" pageId="28" pageNumber="69">
DISTRIBUTION. —
<collectingCountry id="F33A76291F3DFFE3135BC2AD71EBFC39" box="[337,452,936,960]" name="Madagascar" pageId="28" pageNumber="69">Madagascar</collectingCountry>
(
<figureCitation id="13162A3C1F3DFFE313D8C2AD720BFC39" box="[466,548,936,960]" captionStart="FIG" captionStartId="35.[151,162,1408,1425]" captionTargetBox="[170,1219,175,1376]" captionTargetId="figure-28@35.[157,1238,162,1394]" captionTargetPageId="35" captionText="FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853); E, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; F, Malagasya antongilensis (Rathbun, 1905) n. comb.; G, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="28" pageNumber="69">Fig. 11I</figureCitation>
). This species is known only from the
<typeStatus id="5496881B1F3DFFE31389C2C57182FC21" box="[387,429,960,984]" pageId="28" pageNumber="69">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE3129DC2EC7128FB60" pageId="28" pageNumber="69" type="diagnosis">
<paragraph id="8B9236B91F3DFFE3129DC2EC7128FB60" blockId="28.[151,684,574,1177]" pageId="28" pageNumber="69">DIAGNOSIS. — Epigastric, postorbital crests weak, rugose, not joined. Cervical groove very short, shallow. Field of dense, short hair on carapace sidewall in pterygostomial region. Ratio length p5 merus to cw 0.63, ratio length p5 merus cl 0.91, ratio length first gonopod terminal article to subterminal segment 0.35.</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE3129DC5CF7272FA7C" pageId="28" pageNumber="69" type="description">
<paragraph id="8B9236B91F3DFFE3129DC5CF7107FB1B" blockId="28.[151,433,1226,1285]" box="[151,296,1226,1252]" pageId="28" pageNumber="69">DESCRIPTION</paragraph>
<paragraph id="8B9236B91F3DFFE3129DC5EE719EFAFC" blockId="28.[151,433,1226,1285]" box="[151,433,1258,1285]" pageId="28" pageNumber="69">
See
<bibRefCitation id="EFBC4B481F3DFFE312C8C5EF7182FAFC" author="NG P. K. L. &amp; TAKEDA M." box="[194,429,1258,1285]" pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda (1994)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9236B91F3DFFE3129DC42F70EEFABD" blockId="28.[151,684,1322,1413]" box="[151,193,1322,1348]" pageId="28" pageNumber="69">
<emphasis id="B959EAAB1F3DFFE3129DC42F70EEFABD" box="[151,193,1322,1348]" italics="true" pageId="28" pageNumber="69">Size</emphasis>
</paragraph>
<paragraph id="8B9236B91F3DFFE3129DC44E7272FA7C" blockId="28.[151,684,1322,1413]" pageId="28" pageNumber="69">
The adult size range is from cw
<quantity id="4CD59B5C1F3DFFE3101DC44E725FFA9D" box="[535,624,1354,1381]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="28" pageNumber="69" unit="mm" value="23.0">23 mm</quantity>
. The carapace proportions are given in
<tableCitation id="C6AF03021F3DFFE31009C46E7272FA7C" box="[515,605,1387,1413]" captionStart="TABLE" captionStartId="9.[151,162,169,186]" captionText="TABLE 3. — Carapace and other proportions of Madagascan freshwater crabs. Abbreviations: cw, carapace width; cl, carapace length; ch, carapace height; fw, front width; p5m, length of merus of right pereiopod 5." pageId="28" pageNumber="69">Table 3.</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE3129DC4AF70D4FA3B" box="[151,251,1450,1476]" pageId="28" pageNumber="69" type="discussion">
<paragraph id="8B9236B91F3DFFE3129DC4AF70D4FA3B" blockId="28.[151,684,1450,1669]" box="[151,251,1450,1476]" pageId="28" pageNumber="69">REMARKS</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE3129DC4CE7396FD18" pageId="28" pageNumber="69" type="reference_group">
<paragraph id="8B9236B91F3DFFE3129DC4CE744DFDDC" blockId="28.[151,684,1450,1669]" lastBlockId="28.[714,1247,170,549]" pageId="28" pageNumber="69">
<taxonomicName id="4C2D4D3A1F3DFFE3129DC4CE71A6FA1C" baseAuthorityName="Bott" baseAuthorityYear="1965" box="[151,393,1483,1509]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="gollhardi">
<emphasis id="B959EAAB1F3DFFE3129DC4CE71A6FA1C" box="[151,393,1483,1509]" italics="true" pageId="28" pageNumber="69">Skelosophusa gollhardi</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="5496881B1F3DFFE3129DC4EE70D9F9FC" box="[151,246,1515,1541]" pageId="28" pageNumber="69" type="holotype">holotype</typeStatus>
collected from a cave near Ankara.
<bibRefCitation id="EFBC4B481F3DFFE31083C4EF7170F9DC" author="NG P. K. L. &amp; TAKEDA M." pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda (1994)</bibRefCitation>
considered
<taxonomicName id="4C2D4D3A1F3DFFE313E0C70F724AF9DC" baseAuthorityName="Bott" baseAuthorityYear="1965" box="[490,613,1546,1573]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="gollhardi">
<emphasis id="B959EAAB1F3DFFE313E0C70F724AF9DC" box="[490,613,1546,1573]" italics="true" pageId="28" pageNumber="69">S. gollhardi</emphasis>
</taxonomicName>
to be closest to
<taxonomicName id="4C2D4D3A1F3DFFE31306C72F7158F9BC" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[268,375,1578,1605]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="prolixa">
<emphasis id="B959EAAB1F3DFFE31306C72F7158F9BC" box="[268,375,1578,1605]" italics="true" pageId="28" pageNumber="69">S. prolixa</emphasis>
</taxonomicName>
based on carapace characters but cited “significant” differences in the form of the first gonopod and in the relative length of the merus of p5. It is difficult to discern significant differences between gonopod 1 of the two species as illustrated by
<bibRefCitation id="EFBC4B481F3DFFE3169CC1EF7344FEDC" author="NG P. K. L. &amp; TAKEDA M." pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda (1994</bibRefCitation>
: fig. 1a, c-f, 2a-f, h, i), and the proposed species boundaries defined by the relative length of the merus of p5 may not be robust (
<bibRefCitation id="EFBC4B481F3DFFE31127C06E7402FE7C" author="NG P. K. L. &amp; TAKEDA M." box="[813,1069,363,389]" pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda 1994</bibRefCitation>
: table 3). For example, although the ratio of the length of the merus of p5 to cl of
<taxonomicName id="4C2D4D3A1F3DFFE311BCC0AF7433FE3C" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[950,1052,426,453]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="prolixa">
<emphasis id="B959EAAB1F3DFFE311BCC0AF7433FE3C" box="[950,1052,426,453]" italics="true" pageId="28" pageNumber="69">S. prolixa</emphasis>
</taxonomicName>
is 0.75 (which is close to 0.73, 0.74 of
<bibRefCitation id="EFBC4B481F3DFFE311C2C0CF7485FE1C" author="NG P. K. L. &amp; TAKEDA M." box="[968,1194,458,485]" pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda 1994</bibRefCitation>
) the ratio of the length of the merus of p5 to cw of
<taxonomicName id="4C2D4D3A1F3DFFE310C0C30F736AFDDC" baseAuthorityName="Bott" baseAuthorityYear="1965" box="[714,837,522,549]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="gollhardi">
<emphasis id="B959EAAB1F3DFFE310C0C30F736AFDDC" box="[714,837,522,549]" italics="true" pageId="28" pageNumber="69">S. gollhardi</emphasis>
</taxonomicName>
is 0.51 (not 0.73, 0.74).
</paragraph>
<paragraph id="8B9236B91F3DFFE310F8C35B745DFD6E" blockId="28.[754,1207,606,663]" pageId="28" pageNumber="69">
<taxonomicName id="4C2D4D3A1F3DFFE310F8C35B7498FD81" authority="Ng &amp; Takeda, 1994" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[754,1207,606,632]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="prolixa">
<emphasis id="B959EAAB1F3DFFE310F8C35B73FCFD81" bold="true" box="[754,979,606,632]" italics="true" pageId="28" pageNumber="69">Skelosophusa prolixa</emphasis>
<bibRefCitation id="EFBC4B481F3DFFE311D3C35B7498FD81" author="NG P. K. L. &amp; TAKEDA M." box="[985,1207,606,632]" pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda, 1994</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13162A3C1F3DFFE31134C37873BBFD6E" box="[830,916,637,663]" captionStart="FIG" captionStartId="11.[151,162,1354,1371]" captionTargetBox="[192,1245,168,1311]" captionTargetId="figure-26@11.[187,1212,162,1321]" captionTargetPageId="11" captionText="FIG. 2. — Dorsal view of the carapace of adult males of Madagascan freshwater crabs (A-C based on Ng &amp; Takeda 1994); A, Skeleosophusa gollhardi (Bott, 1965), holotype (MNHN B-5165); B, S. prolixa Ng &amp; Takeda, 1994, holotype, (MRSN); C, S. eumeces Ng &amp; Takeda, 1994, holotype, (MRSN); D, Boreas uglowi n. gen., n. sp., holotype (MNHN B- 11292); E, Madagapotamon humberti Bott, 1965; F, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002, holotype, (AMNH 17833). Scale bar: A, 7.2 mm; B, 7.1 mm; C, 6.5 mm; D, 9 mm; E, 9.4 mm; F, 6 mm." figureDoi="http://doi.org/10.5281/zenodo.5401514" httpUri="https://zenodo.org/record/5401514/files/figure.png" pageId="28" pageNumber="69">Figs 2B</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE311AAC3787396FD6E" box="[928,953,637,663]" captionStart="FIG" captionStartId="16.[151,162,1071,1088]" captionTargetBox="[155,1241,171,1034]" captionTargetId="figure-282@16.[138,1257,160,1041]" captionTargetPageId="16" captionText="FIG. 4. — Left mandible of Madagascan freshwater crabs, frontal view showing the palp (J, K based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2000) n. comb.; G, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2000; H, Boreas uglowi n. gen., n. sp.; I, Skeleosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Madagapotamon humberti Bott, 1965. Scale bar: A-I, K, L, 1.0 mm; J, 1.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5401518" httpUri="https://zenodo.org/record/5401518/files/figure.png" pageId="28" pageNumber="69">4J</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE311CFC37873F0FD6E" box="[965,991,637,663]" captionStart="FIG" captionStartId="21.[151,162,1262,1279]" captionTargetBox="[248,1159,165,1192]" captionTargetId="figure-109@21.[228,1169,158,1197]" captionTargetPageId="21" captionText="FIG. 5. — Left third maxilliped of Madagascan freshwater crabs, frontal view (J, K based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; G, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp. Scale bar: A-D, 5.0 mm; E-L, 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5401520" httpUri="https://zenodo.org/record/5401520/files/figure.png" pageId="28" pageNumber="69">5J</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE311E1C378742AFD6E" box="[1003,1029,637,663]" captionStart="FIG" captionStartId="23.[151,162,768,785]" captionTargetBox="[258,1160,169,728]" captionTargetId="figure-337@23.[233,1163,157,745]" captionTargetPageId="23" captionText="FIG. 6. — Superior view of the carpus of the cheliped (pereiopod 1) of adult males of Madagascan freshwater crabs (J, K based on Ng &amp; Takeda 1994); A Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; G, Madagapotamon humberti Bott, 1965; H, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; I, Skeleosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp. Scale bar: A-D, 5.5 mm; E-L, 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5401522" httpUri="https://zenodo.org/record/5401522/files/figure.png" pageId="28" pageNumber="69">6J</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE3161BC3787417FD6E" box="[1041,1080,637,663]" captionStart="FIG" captionStartId="29.[151,162,1545,1562]" captionTargetBox="[151,1247,251,1506]" captionTargetId="figure-43@29.[153,1248,238,1515]" captionTargetPageId="29" captionText="FIG. 8. — Last walking leg (pereiopod 5) of adult males of Madagascan freshwater crabs (H based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Boreas uglowi n. gen., n. sp.; F, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; G, Madagapotamon humberti Bott, 1965; H, Skelosophusa prolixa Ng &amp; Takeda, 1994; I, Malagasya antongilensis (Rathbun, 1905) n. comb.; J, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb. Scale bar: A-D, 5.0 mm; E-L, 2.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5401526" httpUri="https://zenodo.org/record/5401526/files/figure.png" pageId="28" pageNumber="69">8H</figureCitation>
;
<figureCitation id="13162A3C1F3DFFE31649C3787446FD6E" box="[1091,1129,637,663]" captionStart="FIG" captionStartId="35.[151,162,1408,1425]" captionTargetBox="[170,1219,175,1376]" captionTargetId="figure-28@35.[157,1238,162,1394]" captionTargetPageId="35" captionText="FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853); E, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; F, Malagasya antongilensis (Rathbun, 1905) n. comb.; G, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="28" pageNumber="69">11J</figureCitation>
)
</paragraph>
<paragraph id="8B9236B91F3DFFE310C0C3B57396FD18" blockId="28.[714,1248,687,1152]" pageId="28" pageNumber="69">
<treatmentCitationGroup id="AB3D11971F3DFFE310C0C3B57396FD18" pageId="28" pageNumber="69">
<treatmentCitation id="0A8C10A81F3DFFE310C0C3B574A3FD31" author="NG P. K. L. &amp; TAKEDA M." box="[714,1164,687,712]" page="166" pageId="28" pageNumber="69" year="1994">
<taxonomicName id="4C2D4D3A1F3DFFE310C0C3B574A3FD31" authority="Ng &amp; Takeda, 1994: 166" authorityName="Ng &amp; Takeda" authorityPageNumber="166" authorityYear="1994" box="[714,1164,687,712]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="prolixa">
<emphasis id="B959EAAB1F3DFFE310C0C3B573A6FD31" box="[714,905,688,712]" italics="true" pageId="28" pageNumber="69">Skelosophusa prolixa</emphasis>
<bibRefCitation id="EFBC4B481F3DFFE31185C3AA74A3FD31" author="NG P. K. L. &amp; TAKEDA M." box="[911,1164,687,712]" pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda, 1994: 166</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, table 3, figs 2a-c, 3a, 5a-i, 6 k-n.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE310C0C3F573AAFC6E" pageId="28" pageNumber="69" type="materials_examined">
<paragraph id="8B9236B91F3DFFE310C0C3F57336FCAC" blockId="28.[714,1248,687,1152]" pageId="28" pageNumber="69">
MATERIAL EXAMINED. —
<emphasis id="B959EAAB1F3DFFE311CAC3F57412FCF1" bold="true" box="[960,1085,752,776]" pageId="28" pageNumber="69">
<collectingCountry id="F33A76291F3DFFE311CAC3F57416FCF1" box="[960,1081,752,776]" name="Madagascar" pageId="28" pageNumber="69">Madagascar</collectingCountry>
.
</emphasis>
Nosy-Bé, rivière de la route des lacs,
<date id="FF9310791F3DFFE31182C20E73D3FCDA" box="[904,1020,779,803]" pageId="28" pageNumber="69" value="1972-11-23">23.XI.1972</date>
, coll. A. Crosnier,
<specimenCount id="9D2BFD301F3DFFE316A5C20E74CFFCDA" box="[1199,1248,779,803]" pageId="28" pageNumber="69" type="generic">1 ad.</specimenCount>
cw 25.6, cl 18.8, ch 9.3, fw
<quantity id="4CD59B5C1F3DFFE31607C2217470FCC3" box="[1037,1119,803,828]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="28" pageNumber="69" unit="mm" value="7.1">7.1 mm</quantity>
(
<collectionCode id="ED3CAE7C1F3DFFE31667C22174F0FCC5" box="[1133,1247,804,828]" pageId="28" pageNumber="69">MNHN-B</collectionCode>
25734).
</paragraph>
<paragraph id="8B9236B91F3DFFE310C0C26073AAFC6E" blockId="28.[714,1248,687,1152]" pageId="28" pageNumber="69">
TYPELOCALITY. — Grotte nord de l’Ankara, Berges Sable,
<collectingCountry id="F33A76291F3DFFE31100C27A73AFFC6E" box="[778,896,895,919]" name="Madagascar" pageId="28" pageNumber="69">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE310C0C2A2741BFC20" pageId="28" pageNumber="69" type="distribution">
<paragraph id="8B9236B91F3DFFE310C0C2A2741BFC20" blockId="28.[714,1248,687,1152]" pageId="28" pageNumber="69">
DISTRIBUTION. —
<collectingCountry id="F33A76291F3DFFE3118EC2AD73D8FC39" box="[900,1015,936,960]" name="Madagascar" pageId="28" pageNumber="69">Madagascar</collectingCountry>
(
<figureCitation id="13162A3C1F3DFFE3160FC2AD7477FC39" box="[1029,1112,936,960]" captionStart="FIG" captionStartId="35.[151,162,1408,1425]" captionTargetBox="[170,1219,175,1376]" captionTargetId="figure-28@35.[157,1238,162,1394]" captionTargetPageId="35" captionText="FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853); E, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; F, Malagasya antongilensis (Rathbun, 1905) n. comb.; G, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="28" pageNumber="69">Fig. 11J</figureCitation>
). This species is known only from the
<typeStatus id="5496881B1F3DFFE311BCC2C573CFFC21" box="[950,992,960,984]" pageId="28" pageNumber="69">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE310C0C2ED7445FB79" pageId="28" pageNumber="69" type="diagnosis">
<paragraph id="8B9236B91F3DFFE310C0C2ED7445FB79" blockId="28.[714,1248,687,1152]" pageId="28" pageNumber="69">DIAGNOSIS. — Epigastric crests, postorbital crests weak, low, smooth. Cervical groove long, shallow. Dense field of hairs on carapace sidewall in pterygostomial region. Ratio length merus p5 to cw 0.51, ratio length p5 merus cl 0.75, ratio length first gonopod terminal article to subterminal segment 0.38.</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE310C0C5AF74BFFA9C" pageId="28" pageNumber="69" type="description">
<paragraph id="8B9236B91F3DFFE310C0C5AF7374FB3B" blockId="28.[714,996,1194,1253]" box="[714,859,1194,1220]" pageId="28" pageNumber="69">DESCRIPTION</paragraph>
<paragraph id="8B9236B91F3DFFE310C0C5CE73CBFB1C" blockId="28.[714,996,1194,1253]" box="[714,996,1226,1253]" pageId="28" pageNumber="69">
See
<bibRefCitation id="EFBC4B481F3DFFE310FFC5CF73CFFB1C" author="NG P. K. L. &amp; TAKEDA M." box="[757,992,1226,1253]" pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda (1994)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9236B91F3DFFE310C0C40F72DBFADD" blockId="28.[714,1247,1290,1381]" box="[714,756,1290,1316]" pageId="28" pageNumber="69">
<emphasis id="B959EAAB1F3DFFE310C0C40F72DBFADD" box="[714,756,1290,1316]" italics="true" pageId="28" pageNumber="69">Size</emphasis>
</paragraph>
<paragraph id="8B9236B91F3DFFE310C0C42E74BFFA9C" blockId="28.[714,1247,1290,1381]" pageId="28" pageNumber="69">
The adult size range is from cw
<quantity id="4CD59B5C1F3DFFE31640C42E748BFABD" box="[1098,1188,1322,1349]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="28" pageNumber="69" unit="mm" value="24.0">24 mm</quantity>
. The carapace proportions are given in
<tableCitation id="C6AF03021F3DFFE3163CC44E74BFFA9C" box="[1078,1168,1355,1381]" captionStart="TABLE" captionStartId="9.[151,162,169,186]" captionText="TABLE 3. — Carapace and other proportions of Madagascan freshwater crabs. Abbreviations: cw, carapace width; cl, carapace length; ch, carapace height; fw, front width; p5m, length of merus of right pereiopod 5." pageId="28" pageNumber="69">Table 3.</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3DFFE210C0C48F7388FF09" lastPageId="29" lastPageNumber="70" pageId="28" pageNumber="69" type="discussion">
<paragraph id="8B9236B91F3DFFE310C0C48F7301FA5B" blockId="28.[714,1247,1418,1669]" box="[714,814,1418,1444]" pageId="28" pageNumber="69">REMARKS</paragraph>
<paragraph id="8B9236B91F3DFFE310C0C4AE730CF97C" blockId="28.[714,1247,1418,1669]" pageId="28" pageNumber="69">
Our recalculations of the length of the merus of p5 relative to the carapace dimensions based on diagrams and photographs of
<taxonomicName id="4C2D4D3A1F3DFFE3164FC4EF7495F9FC" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[1093,1210,1514,1541]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="prolixa">
<emphasis id="B959EAAB1F3DFFE3164FC4EF7495F9FC" box="[1093,1210,1514,1541]" italics="true" pageId="28" pageNumber="69">S. prolixa</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFBC4B481F3DFFE310C0C70F73EDF9DC" author="NG P. K. L. &amp; TAKEDA M." box="[714,962,1546,1573]" pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda (1994)</bibRefCitation>
(see above) indicate different values for these diagnostic dimensions from those reported by
<bibRefCitation id="EFBC4B481F3DFFE311EFC74F74F3F99C" author="NG P. K. L. &amp; TAKEDA M." box="[997,1244,1610,1637]" pageId="28" pageNumber="69" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda (1994</bibRefCitation>
: table 3).
</paragraph>
<paragraph id="8B9236B91F3CFFE2129DC1D27388FF09" blockId="29.[151,533,215,240]" box="[151,935,215,240]" lastBlockId="29.[916,935,215,240]" pageId="29" pageNumber="70">A B C</paragraph>
</subSubSection>
<caption id="DF5266311F3CFFE2129DC70C71ADF97C" ID-DOI="http://doi.org/10.5281/zenodo.5401526" ID-Zenodo-Dep="5401526" httpUri="https://zenodo.org/record/5401526/files/figure.png" pageId="29" pageNumber="70" startId="29.[151,162,1545,1562]" targetBox="[151,1247,251,1506]" targetPageId="29">
<paragraph id="8B9236B91F3CFFE2129DC70C71ADF97C" blockId="29.[151,1248,1545,1669]" pageId="29" pageNumber="70">
FIG. 8. — Last walking leg (pereiopod 5) of adult males of Madagascan freshwater crabs (H based on
<bibRefCitation id="EFBC4B481F3CFFE21627C70C74F7F9E3" author="NG P. K. L. &amp; TAKEDA M." box="[1069,1240,1545,1562]" pageId="29" pageNumber="70" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda 1994</bibRefCitation>
);
<emphasis id="B959EAAB1F3CFFE2129DC71A708BF9C9" bold="true" box="[151,164,1567,1584]" pageId="29" pageNumber="70">A</emphasis>
,
<taxonomicName id="4C2D4D3A1F3CFFE212A5C71A7261F9C9" authority="A. Milne-Edwards, 1872" authorityName="A. Milne-Edwards" authorityYear="1872" box="[175,590,1567,1584]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="70" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="B959EAAB1F3CFFE212A5C71A7143F9C9" box="[175,364,1567,1584]" italics="true" pageId="29" pageNumber="70">Hydrothelphusa agilis</emphasis>
A. Milne-Edwards, 1872
</taxonomicName>
;
<emphasis id="B959EAAB1F3CFFE21053C71A7249F9C9" bold="true" box="[601,614,1567,1584]" pageId="29" pageNumber="70">B</emphasis>
,
<taxonomicName id="4C2D4D3A1F3CFFE2107FC71A7438F9C9" authority="(A. Milne-Edwards, 1872)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1872" box="[629,1047,1567,1584]" class="Malacostraca" family="Potamidae" genus="Thelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="70" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B959EAAB1F3CFFE2107FC71A7304F9C9" box="[629,811,1567,1584]" italics="true" pageId="29" pageNumber="70">H. madagascariensis</emphasis>
(A. Milne-Edwards, 1872)
</taxonomicName>
;
<emphasis id="B959EAAB1F3CFFE21629C71A741EF9C9" bold="true" box="[1059,1073,1567,1584]" pageId="29" pageNumber="70">C</emphasis>
,
<taxonomicName id="4C2D4D3A1F3CFFE2164AC71A710CF9BC" authority="(Rathbun, 1904)" baseAuthorityName="Rathbun" baseAuthorityYear="1904" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="70" phylum="Arthropoda" rank="species" species="bombetokensis">
<emphasis id="B959EAAB1F3CFFE2164AC71A74F0F9C9" box="[1088,1247,1567,1584]" italics="true" pageId="29" pageNumber="70">H. bombetokensis</emphasis>
(
<bibRefCitation id="EFBC4B481F3CFFE21291C7317131F9BC" author="RATHBUN M. J." box="[155,286,1588,1605]" pageId="29" pageNumber="70" pagination="255 - 312" refId="ref26930" refString="RATHBUN M. J. 1904. - Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle 6 (4): 255 - 312." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
)
</taxonomicName>
;
<emphasis id="B959EAAB1F3CFFE21324C7317113F9BC" bold="true" box="[302,316,1588,1605]" pageId="29" pageNumber="70">D</emphasis>
,
<taxonomicName id="4C2D4D3A1F3CFFE21340C73172A4F9BC" authority="(H. Milne Edwards, 1853)" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[330,651,1588,1605]" class="Malacostraca" family="Potamidae" genus="Thelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="70" phylum="Arthropoda" rank="species" species="goudoti" status="comb. nov.">
<emphasis id="B959EAAB1F3CFFE21340C7317189F9BC" box="[330,422,1588,1605]" italics="true" pageId="29" pageNumber="70">H. goudoti</emphasis>
(H.
<bibRefCitation id="EFBC4B481F3CFFE213C5C73172A8F9BC" author="MILNE EDWARDS H." box="[463,647,1588,1605]" pageId="29" pageNumber="70" pagination="20" refId="ref26442" refString="MILNE EDWARDS H. 1853. - Observations sur les affinites zoologiques et la classification naturelle des Crustaces. Annales des Sciences naturelles, Zoologie, Serie 3, 20: 163 - 28." type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A26A57D01F3CFFE2109FC73172C8F9BC" box="[661,743,1588,1605]" pageId="29" pageNumber="70" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="B959EAAB1F3CFFE210FBC73172D2F9BC" bold="true" box="[753,765,1588,1605]" pageId="29" pageNumber="70">E</emphasis>
,
<taxonomicName id="4C2D4D3A1F3CFFE21101C73173A9F9BC" authority="Cumberlidge &amp; Sternberg, 2002" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[779,902,1588,1605]" class="Malacostraca" family="Potamonautidae" genus="Boreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="70" phylum="Arthropoda" rank="species" species="uglowi" status="n. gen., n. sp.">
<emphasis id="B959EAAB1F3CFFE21101C73173A9F9BC" box="[779,902,1588,1605]" italics="true" pageId="29" pageNumber="70">Boreas uglowi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57D01F3CFFE21184C7317420F9BC" box="[910,1039,1588,1605]" pageId="29" pageNumber="70" rank="species">n. gen., n. sp.</taxonomicNameLabel>
;
<emphasis id="B959EAAB1F3CFFE21610C731740AF9BC" bold="true" box="[1050,1061,1588,1605]" pageId="29" pageNumber="70">F</emphasis>
,
<taxonomicName id="4C2D4D3A1F3CFFE21639C73171F6F9A3" authority="Cumberlidge, Boyko &amp; Harvey, 2002" authorityName="Cumberlidge, Boyko &amp; Harvey" authorityYear="2002" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="70" phylum="Arthropoda" rank="species" species="longimerus">
<emphasis id="B959EAAB1F3CFFE21639C73174F0F9BC" box="[1075,1247,1588,1605]" italics="true" pageId="29" pageNumber="70">Marojejy longimerus</emphasis>
Cumberlidge, Boyko &amp; Harvey, 2002
</taxonomicName>
;
<emphasis id="B959EAAB1F3CFFE213EEC74C71DDF9A3" bold="true" box="[484,498,1609,1626]" pageId="29" pageNumber="70">G</emphasis>
,
<taxonomicName id="4C2D4D3A1F3CFFE2100BC74C7366F9A3" authority="Bott, 1965" authorityName="Bott" authorityYear="1965" box="[513,841,1609,1626]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="70" phylum="Arthropoda" rank="species" species="humberti">
<emphasis id="B959EAAB1F3CFFE2100BC74C72CDF9A3" box="[513,738,1609,1626]" italics="true" pageId="29" pageNumber="70">Madagapotamon humberti</emphasis>
<bibRefCitation id="EFBC4B481F3CFFE210E1C74C7366F9A3" author="BOTT R." box="[747,841,1609,1626]" pageId="29" pageNumber="70" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott, 1965</bibRefCitation>
</taxonomicName>
;
<emphasis id="B959EAAB1F3CFFE2115EC74C734DF9A3" bold="true" box="[852,866,1609,1626]" pageId="29" pageNumber="70">H</emphasis>
,
<taxonomicName id="4C2D4D3A1F3CFFE2117AC74C74F2F9A3" authority="Ng &amp; Takeda, 1994" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[880,1245,1609,1626]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="70" phylum="Arthropoda" rank="species" species="prolixa">
<emphasis id="B959EAAB1F3CFFE2117AC74C740AF9A3" box="[880,1061,1609,1626]" italics="true" pageId="29" pageNumber="70">Skelosophusa prolixa</emphasis>
<bibRefCitation id="EFBC4B481F3CFFE21624C74C74F2F9A3" author="NG P. K. L. &amp; TAKEDA M." box="[1070,1245,1609,1626]" pageId="29" pageNumber="70" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda, 1994</bibRefCitation>
</taxonomicName>
;
<emphasis id="B959EAAB1F3CFFE2129DC75A70B2F989" bold="true" box="[151,157,1631,1648]" pageId="29" pageNumber="70">I</emphasis>
,
<taxonomicName id="4C2D4D3A1F3CFFE212ADC75A722FF989" authority="(Rathbun, 1905)" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" baseAuthorityName="Rathbun" baseAuthorityYear="1905" box="[167,512,1631,1648]" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="70" phylum="Arthropoda" rank="species" species="antongilensis" status="comb. nov.">
<emphasis id="B959EAAB1F3CFFE212ADC75A715CF989" box="[167,371,1631,1648]" italics="true" pageId="29" pageNumber="70">Malagasya antongilensis</emphasis>
(
<bibRefCitation id="EFBC4B481F3CFFE21374C75A71D3F989" author="RATHBUN M. J." box="[382,508,1631,1648]" pageId="29" pageNumber="70" pagination="159 - 322" refId="ref26961" refString="RATHBUN M. J. 1905. - Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle 7 (4): 159 - 322." type="journal article" year="1905">Rathbun, 1905</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A26A57D01F3CFFE2100DC75A7278F989" box="[519,599,1631,1648]" pageId="29" pageNumber="70" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="B959EAAB1F3CFFE21055C75A7246F989" bold="true" box="[607,617,1631,1648]" pageId="29" pageNumber="70">J</emphasis>
,
<taxonomicName id="4C2D4D3A1F3CFFE2107CC75A7401F989" authority="(Cumberlidge, Boyko &amp; Harvey, 2002)" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" baseAuthorityName="Cumberlidge, Boyko &amp; Harvey" baseAuthorityYear="2002" box="[630,1070,1631,1648]" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="70" phylum="Arthropoda" rank="species" species="goodmani" status="comb. nov.">
<emphasis id="B959EAAB1F3CFFE2107CC75A72C9F989" box="[630,742,1631,1648]" italics="true" pageId="29" pageNumber="70">M. goodmani</emphasis>
(Cumberlidge, Boyko &amp; Harvey, 2002)
</taxonomicName>
<taxonomicNameLabel id="A26A57D01F3CFFE2163CC75A74ACF989" box="[1078,1155,1631,1648]" pageId="29" pageNumber="70" rank="species">n. comb.</taxonomicNameLabel>
Scale bar: A-D, 5.0 mm; E-L, 2.6 mm.
</paragraph>
</caption>
<subSubSection id="C33765321F3FFFE112B1C1AF71F3FEDA" pageId="30" pageNumber="71" type="reference_group">
<paragraph id="8B9236B91F3FFFE112B1C1AF71F3FEDA" blockId="30.[151,684,169,632]" pageId="30" pageNumber="71">
<taxonomicName id="4C2D4D3A1F3FFFE112B1C1AF72A7FF3D" authority="Ng &amp; Takeda, 1994" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[187,648,169,196]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="71" phylum="Arthropoda" rank="species" species="eumeces">
<emphasis id="B959EAAB1F3FFFE112B1C1AF718CFF3A" bold="true" box="[187,419,169,196]" italics="true" pageId="30" pageNumber="71">Skelosophusa eumeces</emphasis>
<bibRefCitation id="EFBC4B481F3FFFE113A0C1AC72A7FF3D" author="NG P. K. L. &amp; TAKEDA M." box="[426,648,169,196]" pageId="30" pageNumber="71" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda, 1994</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13162A3C1F3FFFE1131BC1CC7146FF1A" box="[273,361,201,227]" captionStart="FIG" captionStartId="11.[151,162,1354,1371]" captionTargetBox="[192,1245,168,1311]" captionTargetId="figure-26@11.[187,1212,162,1321]" captionTargetPageId="11" captionText="FIG. 2. — Dorsal view of the carapace of adult males of Madagascan freshwater crabs (A-C based on Ng &amp; Takeda 1994); A, Skeleosophusa gollhardi (Bott, 1965), holotype (MNHN B-5165); B, S. prolixa Ng &amp; Takeda, 1994, holotype, (MRSN); C, S. eumeces Ng &amp; Takeda, 1994, holotype, (MRSN); D, Boreas uglowi n. gen., n. sp., holotype (MNHN B- 11292); E, Madagapotamon humberti Bott, 1965; F, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002, holotype, (AMNH 17833). Scale bar: A, 7.2 mm; B, 7.1 mm; C, 6.5 mm; D, 9 mm; E, 9.4 mm; F, 6 mm." figureDoi="http://doi.org/10.5281/zenodo.5401514" httpUri="https://zenodo.org/record/5401514/files/figure.png" pageId="30" pageNumber="71">Figs 2C</figureCitation>
;
<figureCitation id="13162A3C1F3FFFE1137FC1CC71B7FF1A" box="[373,408,201,227]" captionStart="FIG" captionStartId="16.[151,162,1071,1088]" captionTargetBox="[155,1241,171,1034]" captionTargetId="figure-282@16.[138,1257,160,1041]" captionTargetPageId="16" captionText="FIG. 4. — Left mandible of Madagascan freshwater crabs, frontal view showing the palp (J, K based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2000) n. comb.; G, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2000; H, Boreas uglowi n. gen., n. sp.; I, Skeleosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Madagapotamon humberti Bott, 1965. Scale bar: A-I, K, L, 1.0 mm; J, 1.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5401518" httpUri="https://zenodo.org/record/5401518/files/figure.png" pageId="30" pageNumber="71">4K</figureCitation>
;
<figureCitation id="13162A3C1F3FFFE113AEC1CC71E8FF1A" box="[420,455,201,227]" captionStart="FIG" captionStartId="21.[151,162,1262,1279]" captionTargetBox="[248,1159,165,1192]" captionTargetId="figure-109@21.[228,1169,158,1197]" captionTargetPageId="21" captionText="FIG. 5. — Left third maxilliped of Madagascan freshwater crabs, frontal view (J, K based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; G, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp. Scale bar: A-D, 5.0 mm; E-L, 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5401520" httpUri="https://zenodo.org/record/5401520/files/figure.png" pageId="30" pageNumber="71">5K</figureCitation>
;
<figureCitation id="13162A3C1F3FFFE113D8C1CC71D9FF1A" box="[466,502,201,227]" captionStart="FIG" captionStartId="23.[151,162,768,785]" captionTargetBox="[258,1160,169,728]" captionTargetId="figure-337@23.[233,1163,157,745]" captionTargetPageId="23" captionText="FIG. 6. — Superior view of the carpus of the cheliped (pereiopod 1) of adult males of Madagascan freshwater crabs (J, K based on Ng &amp; Takeda 1994); A Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; G, Madagapotamon humberti Bott, 1965; H, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; I, Skeleosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp. Scale bar: A-D, 5.5 mm; E-L, 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5401522" httpUri="https://zenodo.org/record/5401522/files/figure.png" pageId="30" pageNumber="71">6K</figureCitation>
;
<figureCitation id="13162A3C1F3FFFE1100BC1CC721EFF1A" box="[513,561,201,227]" captionStart="FIG" captionStartId="35.[151,162,1408,1425]" captionTargetBox="[170,1219,175,1376]" captionTargetId="figure-28@35.[157,1238,162,1394]" captionTargetPageId="35" captionText="FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853); E, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; F, Malagasya antongilensis (Rathbun, 1905) n. comb.; G, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="30" pageNumber="71">11K</figureCitation>
)
<taxonomicName id="4C2D4D3A1F3FFFE1129DC1F6725CFEF2" authority="Ng &amp; Takeda, 1994: 166" authorityName="Ng &amp; Takeda" authorityPageNumber="166" authorityYear="1994" box="[151,627,242,267]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="71" phylum="Arthropoda" rank="species" species="eumeces">
<emphasis id="B959EAAB1F3FFFE1129DC1F6714CFEF3" box="[151,355,243,267]" italics="true" pageId="30" pageNumber="71">Skelosophusa eumeces</emphasis>
<bibRefCitation id="EFBC4B481F3FFFE11361C1F7725CFEF2" author="NG P. K. L. &amp; TAKEDA M." box="[363,627,242,267]" pageId="30" pageNumber="71" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda, 1994: 166</bibRefCitation>
</taxonomicName>
, 169, 171, table 3, figs 3b, 4, 6a-g, j, o.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3FFFE1129DC0317267FE51" pageId="30" pageNumber="71" type="materials_examined">
<paragraph id="8B9236B91F3FFFE1129DC0317279FE79" blockId="30.[151,684,169,632]" pageId="30" pageNumber="71">
MATERIAL EXAMINED. —
<emphasis id="B959EAAB1F3FFFE113AFC0317206FEB5" bold="true" box="[421,553,308,332]" pageId="30" pageNumber="71">
<collectingCountry id="F33A76291F3FFFE113AFC0317209FEB5" box="[421,550,308,332]" name="Madagascar" pageId="30" pageNumber="71">Madagascar</collectingCountry>
.
</emphasis>
Andrafiabe,
<specimenCount id="9D2BFD301F3FFFE1129DC04B70E2FE9F" box="[151,205,334,358]" pageId="30" pageNumber="71" type="generic">1 ad.</specimenCount>
,
<typeStatus id="5496881B1F3FFFE112F4C04B7172FE9F" box="[254,349,334,358]" pageId="30" pageNumber="71" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2D4D3A1F3FFFE1138EC04B7130FE79" authority="Ng &amp; Takeda, 1994" authorityName="Ng &amp; Takeda" authorityYear="1994" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="71" phylum="Arthropoda" rank="species" species="eumeces">
<emphasis id="B959EAAB1F3FFFE1138EC04B724CFE9C" box="[388,611,334,358]" italics="true" pageId="30" pageNumber="71">Skelosophusa eumeces</emphasis>
<bibRefCitation id="EFBC4B481F3FFFE11067C0487130FE79" author="NG P. K. L. &amp; TAKEDA M." pageId="30" pageNumber="71" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda, 1994</bibRefCitation>
</taxonomicName>
cw 22.7, cl
<quantity id="4CD59B5C1F3FFFE1139CC06D71DFFE87" box="[406,496,359,384]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="30" pageNumber="71" unit="mm" value="15.8">15.8 mm</quantity>
(
<collectionCode id="ED3CAE7C1F3FFFE113F4C06D7264FE79" box="[510,587,360,384]" pageId="30" pageNumber="71">MRSN</collectionCode>
).
</paragraph>
<paragraph id="8B9236B91F3FFFE1129DC08A7267FE51" blockId="30.[151,684,169,632]" box="[151,584,399,424]" pageId="30" pageNumber="71">
TYPELOCALITY. — Andrafiabe,
<collectingCountry id="F33A76291F3FFFE113C4C095726BFE51" box="[462,580,400,424]" name="Madagascar" pageId="30" pageNumber="71">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3FFFE1129DC0BD7266FE13" pageId="30" pageNumber="71" type="distribution">
<paragraph id="8B9236B91F3FFFE1129DC0BD7266FE13" blockId="30.[151,684,169,632]" pageId="30" pageNumber="71">
DISTRIBUTION. —
<collectingCountry id="F33A76291F3FFFE11360C0BC71C4FE28" box="[362,491,441,465]" name="Madagascar" pageId="30" pageNumber="71">Madagascar</collectingCountry>
(
<figureCitation id="13162A3C1F3FFFE113F6C0BC724AFE28" box="[508,613,441,465]" captionStart="FIG" captionStartId="35.[151,162,1408,1425]" captionTargetBox="[170,1219,175,1376]" captionTargetId="figure-28@35.[157,1238,162,1394]" captionTargetPageId="35" captionText="FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853); E, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; F, Malagasya antongilensis (Rathbun, 1905) n. comb.; G, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="30" pageNumber="71">Fig. 11K</figureCitation>
). This species is known only from the
<typeStatus id="5496881B1F3FFFE113C1C0D471DAFE10" box="[459,501,465,489]" pageId="30" pageNumber="71">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3FFFE1129DC0FC7272FCFC" pageId="30" pageNumber="71" type="diagnosis">
<paragraph id="8B9236B91F3FFFE1129DC0FC71B3FD81" blockId="30.[151,684,169,632]" pageId="30" pageNumber="71">DIAGNOSIS. — Epigastric, postorbital crests distinct, rugose. Cervical groove broad, distinct, short. Ratio length p5 merus to cw 0.56, ratio length p5 merus to cl 0.82, ratio length first gonopod terminal article to subterminal segment 0.35.</paragraph>
<paragraph id="8B9236B91F3FFFE1129DC3AF70EEFD3D" blockId="30.[151,684,682,773]" box="[151,193,682,708]" pageId="30" pageNumber="71">
<emphasis id="B959EAAB1F3FFFE1129DC3AF70EEFD3D" box="[151,193,682,708]" italics="true" pageId="30" pageNumber="71">Size</emphasis>
</paragraph>
<paragraph id="8B9236B91F3FFFE1129DC3CE7272FCFC" blockId="30.[151,684,682,773]" pageId="30" pageNumber="71">
The adult size range is from cw
<quantity id="4CD59B5C1F3FFFE1101DC3CE725FFD1D" box="[535,624,714,741]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="30" pageNumber="71" unit="mm" value="24.0">24 mm</quantity>
. The carapace proportions are given in
<tableCitation id="C6AF03021F3FFFE11009C3EE7272FCFC" box="[515,605,747,773]" captionStart="TABLE" captionStartId="9.[151,162,169,186]" captionText="TABLE 3. — Carapace and other proportions of Madagascan freshwater crabs. Abbreviations: cw, carapace width; cl, carapace length; ch, carapace height; fw, front width; p5m, length of merus of right pereiopod 5." pageId="30" pageNumber="71">Table 3.</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33765321F3FFFE1129DC22F7163FB5C" pageId="30" pageNumber="71" type="discussion">
<paragraph id="8B9236B91F3FFFE1129DC22F70D4FCBB" blockId="30.[151,684,810,1189]" box="[151,251,810,836]" pageId="30" pageNumber="71">REMARKS</paragraph>
<paragraph id="8B9236B91F3FFFE1129DC24E7163FB5C" blockId="30.[151,684,810,1189]" pageId="30" pageNumber="71">
Our recalculations of the length of the merus of p5 relative to the carapace dimensions based on the diagrams and photographs of
<taxonomicName id="4C2D4D3A1F3FFFE11010C28F72A5FC5A" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[538,650,906,932]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="71" phylum="Arthropoda" rank="species" species="eumeces">
<emphasis id="B959EAAB1F3FFFE11010C28F72A5FC5A" box="[538,650,906,932]" italics="true" pageId="30" pageNumber="71">S. eumeces</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFBC4B481F3FFFE1129DC2AF71BEFC3C" author="NG P. K. L. &amp; TAKEDA M." box="[151,401,938,965]" pageId="30" pageNumber="71" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda (1994)</bibRefCitation>
indicate different values for these diagnostic dimensions from those reported by
<bibRefCitation id="EFBC4B481F3FFFE11328C2EF723FFBFC" author="NG P. K. L. &amp; TAKEDA M." box="[290,528,1002,1029]" pageId="30" pageNumber="71" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda (1994</bibRefCitation>
: table 3). For example, the ratio of the length of the merus of p5 to cw in
<taxonomicName id="4C2D4D3A1F3FFFE11327C52F71B1FBBA" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[301,414,1066,1092]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="71" phylum="Arthropoda" rank="species" species="eumeces">
<emphasis id="B959EAAB1F3FFFE11327C52F71B1FBBA" box="[301,414,1066,1092]" italics="true" pageId="30" pageNumber="71">S. eumeces</emphasis>
</taxonomicName>
is 0.56 (not 0.82) and the ratio of the terminal article to subterminal segment length of gonopod
<quantity id="4CD59B5C1F3FFFE113EAC56E723FFB7C" box="[480,528,1131,1157]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="30" pageNumber="71" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4C2D4D3A1F3FFFE11017C56F72A0FB7A" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[541,655,1130,1156]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="71" phylum="Arthropoda" rank="species" species="eumeces">
<emphasis id="B959EAAB1F3FFFE11017C56F72A0FB7A" box="[541,655,1130,1156]" italics="true" pageId="30" pageNumber="71">S. eumeces</emphasis>
</taxonomicName>
is 0.35 (not 0.43).
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue Copy permalink