190 lines
34 KiB
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190 lines
34 KiB
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<document id="33E4A88D4C16E429FB964AC9E2779E7B" ID-DOI="10.5281/zenodo.191594" ID-GBIF-Dataset="6f65b06c-5600-4172-9ee2-854f09ef4455" ID-ISSN="1175-5326" ID-Zenodo-Dep="191594" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461129936904" checkinUser="plazi" docAuthor="Haddad, Charles R., Lyle, Robin, Bosselaers, Jan & Ramirez, Martin J." docDate="2009" docId="038087A2DA54450BFF4FFA9964D9F97A" docLanguage="en" docName="zt02296p038.pdf" docOrigin="Zootaxa 2296" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Austrachelas pondoensis Haddad, Lyle, Bosselaers & Ramirez, 2009, n. sp." docType="treatment" docVersion="10" lastPageNumber="24" masterDocId="FFB9FFDADA42451CFFD8FFC5641CFFEC" masterDocTitle="A revision of the endemic South African spider genus Austrachelas, with its transfer to the Gallieniellidae (Arachnida: Araneae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="23" updateTime="1698597546718" updateUser="plazi">
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<mods:titleInfo id="D09CAE1515CD2907033F32C9FA5792C1">
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<mods:title id="DD1EE4EE76697244A7C806A3A2DBD333">A revision of the endemic South African spider genus Austrachelas, with its transfer to the Gallieniellidae (Arachnida: Araneae)</mods:title>
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<mods:namePart id="986451EEAFEFCD2A1D273138F2E33B55">Haddad, Charles R.</mods:namePart>
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<mods:namePart id="36E75E6C8C3A2A81F72B276FC36D61AD">Lyle, Robin</mods:namePart>
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</mods:role>
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<mods:namePart id="8AC959E33A2B792A11E3734E8371E8B3">Bosselaers, Jan</mods:namePart>
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<mods:name id="00AC0B50264A47003FAF4DF4354A3DAE" type="personal">
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<mods:namePart id="38353673EA743C5996462611C1C55404">Ramirez, Martin J.</mods:namePart>
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<mods:typeOfResource id="9BB7B0A0A385E8F045A963C8BE494C0A">text</mods:typeOfResource>
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<mods:title id="9608ED3E994E8BC5FB7DE0A4987999BC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="85507E86F3CA40E9ACDE5046B63917C9">
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<mods:date id="0F40EA9969889ECE451EB9A8029EB3FE">2009</mods:date>
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<mods:detail id="C9F98A16264D20475E7A79DAC5A86DE9" type="volume">
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<mods:number id="2B7FBBD13D3514DC752D4529CD31C4C4">2296</mods:number>
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<mods:start id="84BBED93069AE4FFDEAB2A148AD671B7">1</mods:start>
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<mods:end id="17CFE88846E773204DA883CC92FC4E2E">38</mods:end>
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<mods:classification id="4ADBBA7EDAA09CF44EC2517A267E2826">journal article</mods:classification>
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<mods:identifier id="1E91F0C759A311FAD86C21A8C56E82AD" type="DOI">10.5281/zenodo.191594</mods:identifier>
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<mods:identifier id="9B43ECE44B756E6F1E9E5F0AA88D5BA2" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038087A2DA54450BFF4FFA9964D9F97A" ID-DOI="http://doi.org/10.5281/zenodo.6225461" ID-GBIF-Taxon="119626934" ID-Zenodo-Dep="6225461" LSID="urn:lsid:plazi:treatment:038087A2DA54450BFF4FFA9964D9F97A" httpUri="http://treatment.plazi.org/id/038087A2DA54450BFF4FFA9964D9F97A" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<subSubSection id="C333653FDA54450AFF4FFA99663AFA76" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8B9636B4DA54450AFF4FFA996611FA99" blockId="22.[151,550,1371,1434]" box="[151,525,1371,1398]" pageId="22" pageNumber="23">
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<heading id="D0DE81D8DA54450AFF4FFA996611FA99" bold="true" box="[151,525,1371,1398]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
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<emphasis id="B95DEAA6DA54450AFF4FFA996611FA99" bold="true" box="[151,525,1371,1398]" pageId="22" pageNumber="23">
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<taxonomicName id="4C294D37DA54450AFF4FFA9965DFFA9A" ID-CoL="67XJW" box="[151,451,1372,1398]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pondoensis" status="sp. nov.">
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<emphasis id="B95DEAA6DA54450AFF4FFA9965DFFA9A" bold="true" box="[151,451,1372,1398]" italics="true" pageId="22" pageNumber="23">Austrachelas pondoensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26E57DDDA54450AFE13FA9E6611FA99" box="[459,525,1371,1397]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9636B4DA54450AFF4FFA45663AFA76" blockId="22.[151,550,1371,1434]" box="[151,550,1408,1434]" pageId="22" pageNumber="23">
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(
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<figureCitation id="13122A31DA54450AFF47FA45650BFA76" box="[159,279,1408,1434]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,269,1353,1377]" captionTargetBox="[151,1435,315,1326]" captionTargetId="figure@2.[151,1436,313,1328]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Digital microscope photographs of the habitus of Austrachelas species: (1) A. bergi n. sp., female; (2) A. incertus Lawrence, 1938, female; (3) same, male; (4) A. kalaharinus n. sp., male; (5) A. merwei n. sp., female; (6) same, male; (7) A. natalensis Lawrence, 1942, female; (8) same, male; (9) A. pondoensis n. sp., female; (10) same, male; (11) A. reavelli n. sp., female; (12) same, male; (13) A. sexoculata n. sp., male; (14) A. wassenaari n. sp., female; (15) same, male. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/191595/files/figure.png" pageId="22" pageNumber="23">Figs 9, 10</figureCitation>
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,
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<figureCitation id="13122A31DA54450AFEFCFA456573FA76" box="[292,367,1408,1434]" captionStart-0="FIGURES 16 – 26" captionStart-1="FIGURES 27 – 35" captionStartId-0="4.[151,271,1757,1781]" captionStartId-1="6.[151,269,1569,1593]" captionTargetBox-0="[205,1391,198,1729]" captionTargetBox-1="[151,1436,197,1542]" captionTargetId-0="figure@4.[205,1392,197,1731]" captionTargetId-1="figure@6.[151,1436,197,1544]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="FIGURES 16 – 26. Digital microscope photographs of the somatic morphology of female Austrachelas pondoensis n. sp.. (16) cephalothorax, dorsal view; (17) same, ventral view; (18) same, lateral view; (19) abdomen, dorsal view; (20) abdominal venter, epigastric area; (21) trochanter I, ventral view; (22) trochanter IV, ventral view; (23) patella I, retrolateral view; (24) leg I, prolateral view, arrow indicating metatarsal and tarsal scopulae; (25 – 26) distal end of leg IV, indicating paired retrolateral dorsal spines and metatarsal and tarsal scopulae. Abbreviations: AH – anterior epigynal hood; EG – endite groove; IS – intercoxal sclerite; LH – lateral epigynal hood; MHT – maxillar hair tuft; PB – pleural bars; PDS – paired dorsal metatarsal spines; PES – post-epigastric sclerite; PI – patellar indentation; PT – precoxal triangle; SC – scopulae; SI – sigillum; TR – trichobothria." captionText-1="FIGURES 27 – 35. Scanning electron micrographs of Austrachelas pondoensis n. sp. somatic and genitalic morphology. (27) female chelicera, posterior view; (28) same, anterior view (29) male palp, ventral view; (30) same, dorsal view; (31) female epigyne, ventral view; (32) female left palp, retrolateral view, arrows indicating erect ventral setae; (33) female left leg 1, arrows indicating metatarsal and tarsal scopulae; (34) female left metatarsus 4, arrows indicating paired dorsal prolateral spines; (35) female, tarsal claw of left leg 4. Abbreviations: AH – anterior epigynal hood; CO – copulatory openings; EM – embolus; EP – embolar protuberance; LH – lateral epigynal hood; MA – median apophysis; PDS – paired dorsal metatarsal spines; PMT – promarginal teeth; RMT – retromarginal teeth; SC – scopulae; SS – scrappy seta; ST – subtegulum. Scale bars: 27, 29, 30 = 0.5 mm; 28, 31 = 0.2 mm; 32 – 34 = 1.0 mm; 35 = 0.1 mm." httpUri-0="https://zenodo.org/record/191596/files/figure.png" httpUri-1="https://zenodo.org/record/191597/files/figure.png" pageId="22" pageNumber="23">16–35</figureCitation>
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,
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<figureCitation id="13122A31DA54450AFEA3FA4565D4FA76" box="[379,456,1408,1434]" captionStart="FIGURES 42 – 49" captionStartId="9.[151,269,1715,1739]" captionTargetBox="[151,1436,183,1685]" captionTargetId="figure@9.[151,1436,183,1686]" captionTargetPageId="9" captionText="FIGURES 42 – 49. Scanning electron micrographs of spinnerets of Austrachelas pondoensis n. sp. male (42, 44 – 46) and female (43, 47 – 49). (42, 43) spinneret arrangement; (44, 47) anterior lateral spinneret; (45, 48) posterior median spinneret; (46, 49) posterior lateral spinneret. Abbreviations: Ac – aciniform gland spigot (s); ALS – anterior lateral spinneret; Cy – cylindrical gland spigot (s); maAmp – major ampullate gland spigot (s); miAmp – minor ampullate gland spigot (s); MS – PLS modified spigot (s); nu – nubbin (s); Pi – piriform gland spigot; PLS – posterior lateral spinneret; PMS – posterior median spinneret; ta – tartipore (s). Scale bars: 42 = 0.2 mm; 43 = 0.5 mm; 44 – 47 = 0.05 mm; 48, 49 = 0.1 mm." httpUri="https://zenodo.org/record/191599/files/figure.png" pageId="22" pageNumber="23">42–49</figureCitation>
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,
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<figureCitation id="13122A31DA54450AFE0BFA456601FA76" box="[467,541,1408,1434]" captionStart="FIGURES 68 – 75" captionStartId="24.[151,269,1392,1416]" captionTargetBox="[193,1403,477,1365]" captionTargetId="figure@24.[192,1413,470,1366]" captionTargetPageId="24" captionText="FIGURES 68 – 75. Genitalic morphology of Austrachelas species. (68 – 71) A. pondoensis n. sp.: female epigyne, ventral view (68), dorsal view (69), male palp, ventral view (70), retrolateral view (71); (72 – 75) A. reavelli n. sp.: female epigyne, ventral view (72), dorsal view (73), male palp, ventral view (74), retrolateral view (75). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/191604/files/figure.png" pageId="22" pageNumber="23">68–71</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C333653FDA54450AFF4FFA0265C1F9A2" pageId="22" pageNumber="23" type="diagnosis">
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<paragraph id="8B9636B4DA54450AFF4FFA0265C1F9A2" blockId="22.[151,1437,1479,2036]" pageId="22" pageNumber="23">
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<emphasis id="B95DEAA6DA54450AFF4FFA02650CFA0D" bold="true" box="[151,272,1479,1505]" pageId="22" pageNumber="23">Diagnosis</emphasis>
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. This is the largest species in the genus, and females can be easily distinguished by the shape of the anterior epigynal hood and the widely separated spermathecae (
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<figureCitation id="13122A31DA54450AFC50FA2967F8F9EA" box="[904,996,1516,1542]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,269,1569,1593]" captionTargetBox="[151,1436,197,1542]" captionTargetId="figure@6.[151,1436,197,1544]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Scanning electron micrographs of Austrachelas pondoensis n. sp. somatic and genitalic morphology. (27) female chelicera, posterior view; (28) same, anterior view (29) male palp, ventral view; (30) same, dorsal view; (31) female epigyne, ventral view; (32) female left palp, retrolateral view, arrows indicating erect ventral setae; (33) female left leg 1, arrows indicating metatarsal and tarsal scopulae; (34) female left metatarsus 4, arrows indicating paired dorsal prolateral spines; (35) female, tarsal claw of left leg 4. Abbreviations: AH – anterior epigynal hood; CO – copulatory openings; EM – embolus; EP – embolar protuberance; LH – lateral epigynal hood; MA – median apophysis; PDS – paired dorsal metatarsal spines; PMT – promarginal teeth; RMT – retromarginal teeth; SC – scopulae; SS – scrappy seta; ST – subtegulum. Scale bars: 27, 29, 30 = 0.5 mm; 28, 31 = 0.2 mm; 32 – 34 = 1.0 mm; 35 = 0.1 mm." httpUri="https://zenodo.org/record/191597/files/figure.png" pageId="22" pageNumber="23">Figs 31</figureCitation>
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,
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<figureCitation id="13122A31DA54450AFC2AFA29605CF9EA" box="[1010,1088,1516,1542]" captionStart="FIGURES 68 – 75" captionStartId="24.[151,269,1392,1416]" captionTargetBox="[193,1403,477,1365]" captionTargetId="figure@24.[192,1413,470,1366]" captionTargetPageId="24" captionText="FIGURES 68 – 75. Genitalic morphology of Austrachelas species. (68 – 71) A. pondoensis n. sp.: female epigyne, ventral view (68), dorsal view (69), male palp, ventral view (70), retrolateral view (71); (72 – 75) A. reavelli n. sp.: female epigyne, ventral view (72), dorsal view (73), male palp, ventral view (74), retrolateral view (75). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/191604/files/figure.png" pageId="22" pageNumber="23">68, 69</figureCitation>
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); males can be recognised by the large partially hidden triangular lobe distally on the embolus and the denticulate retrolateral tibial apophysis (
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<figureCitation id="13122A31DA54450AFEC3F9F1656BF9A2" box="[283,375,1588,1614]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,269,1569,1593]" captionTargetBox="[151,1436,197,1542]" captionTargetId="figure@6.[151,1436,197,1544]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Scanning electron micrographs of Austrachelas pondoensis n. sp. somatic and genitalic morphology. (27) female chelicera, posterior view; (28) same, anterior view (29) male palp, ventral view; (30) same, dorsal view; (31) female epigyne, ventral view; (32) female left palp, retrolateral view, arrows indicating erect ventral setae; (33) female left leg 1, arrows indicating metatarsal and tarsal scopulae; (34) female left metatarsus 4, arrows indicating paired dorsal prolateral spines; (35) female, tarsal claw of left leg 4. Abbreviations: AH – anterior epigynal hood; CO – copulatory openings; EM – embolus; EP – embolar protuberance; LH – lateral epigynal hood; MA – median apophysis; PDS – paired dorsal metatarsal spines; PMT – promarginal teeth; RMT – retromarginal teeth; SC – scopulae; SS – scrappy seta; ST – subtegulum. Scale bars: 27, 29, 30 = 0.5 mm; 28, 31 = 0.2 mm; 32 – 34 = 1.0 mm; 35 = 0.1 mm." httpUri="https://zenodo.org/record/191597/files/figure.png" pageId="22" pageNumber="23">Figs 29</figureCitation>
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,
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<figureCitation id="13122A31DA54450AFE5BF9F165CCF9A2" box="[387,464,1588,1614]" captionStart="FIGURES 68 – 75" captionStartId="24.[151,269,1392,1416]" captionTargetBox="[193,1403,477,1365]" captionTargetId="figure@24.[192,1413,470,1366]" captionTargetPageId="24" captionText="FIGURES 68 – 75. Genitalic morphology of Austrachelas species. (68 – 71) A. pondoensis n. sp.: female epigyne, ventral view (68), dorsal view (69), male palp, ventral view (70), retrolateral view (71); (72 – 75) A. reavelli n. sp.: female epigyne, ventral view (72), dorsal view (73), male palp, ventral view (74), retrolateral view (75). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/191604/files/figure.png" pageId="22" pageNumber="23">70, 71</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C333653FDA54450AFF1EF9926180F99E" box="[198,1436,1623,1650]" pageId="22" pageNumber="23" type="etymology">
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<paragraph id="8B9636B4DA54450AFF1EF9926180F99E" blockId="22.[151,1437,1479,2036]" box="[198,1436,1623,1650]" pageId="22" pageNumber="23">
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<emphasis id="B95DEAA6DA54450AFF1EF9926557F99D" bold="true" box="[198,331,1623,1649]" pageId="22" pageNumber="23">Etymology</emphasis>
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. This species is named after the Pondoland region in the Eastern Cape Province,
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<collectingCountry id="F33E7624DA54450AFB26F99D6184F99E" box="[1278,1432,1624,1650]" name="South Africa" pageId="22" pageNumber="23">South Africa</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C333653FDA54450BFF1EF9BB673AFA8C" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
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<paragraph id="8B9636B4DA54450AFF1EF9BB6646F90A" blockId="22.[151,1437,1479,2036]" pageId="22" pageNumber="23">
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<emphasis id="B95DEAA6DA54450AFF1EF9BB653DF974" bold="true" box="[198,289,1662,1688]" pageId="22" pageNumber="23">Female</emphasis>
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.
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<emphasis id="B95DEAA6DA54450AFEE8F9BB65EDF974" bold="true" box="[304,497,1662,1688]" pageId="22" pageNumber="23">Measurements:</emphasis>
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CL 5.18–6.05, CW 3.70–4.28, FL 0.40–0.48, AL 6.50–9.20, AW 3.10–5.13, SL 2.88–3.05, SW 2.00–2.27, TL 11.80–15.30, AME–AME 0.08, AME–ALE 0.04, ALE–ALE 0.60, PME–PME 0.10, PME–PLE 0.22, PLE–PLE 0.89.
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</paragraph>
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<paragraph id="8B9636B4DA54450AFF1DF93765ECF8B6" blockId="22.[151,1437,1479,2036]" pageId="22" pageNumber="23">Length of leg segments (sequence from femur to tarsus, and total): I 3.95 + 2.38 + 2.95 + 2.12 + 1.38 = 12.78; II 3.50 + 2.10 + 2.58 + 2.05 + 1.28 = 11.51; III 2.78 + 1.65 + 1.70 + 2.32 + 1.05 = 9.50; IV 4.07 + 2.10 + 3.08 + 3.72 + 1.20 = 14.17.</paragraph>
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<paragraph id="8B9636B4DA54450BFF1DF8A36507FDB7" blockId="22.[151,1437,1479,2036]" lastBlockId="23.[151,1437,152,1686]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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Carapace deep red-brown, slightly paler medially (
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<figureCitation id="13122A31DA54450AFCFEF8A36773F86C" box="[806,879,1894,1920]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,269,1353,1377]" captionTargetBox="[151,1435,315,1326]" captionTargetId="figure@2.[151,1436,313,1328]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Digital microscope photographs of the habitus of Austrachelas species: (1) A. bergi n. sp., female; (2) A. incertus Lawrence, 1938, female; (3) same, male; (4) A. kalaharinus n. sp., male; (5) A. merwei n. sp., female; (6) same, male; (7) A. natalensis Lawrence, 1942, female; (8) same, male; (9) A. pondoensis n. sp., female; (10) same, male; (11) A. reavelli n. sp., female; (12) same, male; (13) A. sexoculata n. sp., male; (14) A. wassenaari n. sp., female; (15) same, male. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/191595/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
|
|||
|
); AER strongly procurved, laterals larger than medians; AME separated by distance equal to ½ their diameter, AME separated from ALE by distance equal to ¼ AME diameter; clypeus height equal to 1½ AME diameter at AME, equal to 1¼ ALE diameter at ALE; PER straight, laterals larger than medians; PME separated by distance equal to ½ their diameter, PME separated from PLE by distance equal to 1¼ PME diameter. Chelicerae deep orange; promargin with three teeth, proximal tooth smallest, median tooth largest; proximal and median teeth closer to each other than to distal tooth; retromargin with two small denticles, situated either side of distal promarginal tooth. Legs all deep orange in colour, slightly paler distally. Leg spination: femora: I pl 1 do 1, II pl 1 do 1, III pl 1 do 2 rl 1, IV do 2; patellae: spineless; tibiae: I & II spineless, III pl 2 rl 2 plv 2 rlv 1 vt 2, IV pl 2 rl 2 plv 3 rlv 2 vt 2; metatarsi: I & II spineless, strongly scopulate, III & IV scopulate in distal half, III pl 6 rl 5 plv 1 rlv 1 vt 2, IV pl 6 rl 6 plv 2 rlv 1 vt 2; palp: femora do 2, patellae spineless, with prominent pl 1 seta, tibiae pl 3 do 2, tarsus pl 3 do 2 rl 3 plv 1 rlv 1. Abdomen dark grey dorsally, with mottled pale grey chevron (
|
|||
|
<figureCitation id="13122A31DA55450BFB18FE636116FE2C" box="[1216,1290,422,448]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,269,1353,1377]" captionTargetBox="[151,1435,315,1326]" captionTargetId="figure@2.[151,1436,313,1328]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Digital microscope photographs of the habitus of Austrachelas species: (1) A. bergi n. sp., female; (2) A. incertus Lawrence, 1938, female; (3) same, male; (4) A. kalaharinus n. sp., male; (5) A. merwei n. sp., female; (6) same, male; (7) A. natalensis Lawrence, 1942, female; (8) same, male; (9) A. pondoensis n. sp., female; (10) same, male; (11) A. reavelli n. sp., female; (12) same, male; (13) A. sexoculata n. sp., male; (14) A. wassenaari n. sp., female; (15) same, male. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/191595/files/figure.png" pageId="23" pageNumber="24">Fig. 9</figureCitation>
|
|||
|
), paler grey laterally and ventrally; dorsum and venter lacking scutum or other sclerites. Female epigyne with short narrow anterior hood and small lateral hoods (
|
|||
|
<figureCitation id="13122A31DA55450BFDB8FE3166A1FDE2" box="[608,701,500,526]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,269,1569,1593]" captionTargetBox="[151,1436,197,1542]" captionTargetId="figure@6.[151,1436,197,1544]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Scanning electron micrographs of Austrachelas pondoensis n. sp. somatic and genitalic morphology. (27) female chelicera, posterior view; (28) same, anterior view (29) male palp, ventral view; (30) same, dorsal view; (31) female epigyne, ventral view; (32) female left palp, retrolateral view, arrows indicating erect ventral setae; (33) female left leg 1, arrows indicating metatarsal and tarsal scopulae; (34) female left metatarsus 4, arrows indicating paired dorsal prolateral spines; (35) female, tarsal claw of left leg 4. Abbreviations: AH – anterior epigynal hood; CO – copulatory openings; EM – embolus; EP – embolar protuberance; LH – lateral epigynal hood; MA – median apophysis; PDS – paired dorsal metatarsal spines; PMT – promarginal teeth; RMT – retromarginal teeth; SC – scopulae; SS – scrappy seta; ST – subtegulum. Scale bars: 27, 29, 30 = 0.5 mm; 28, 31 = 0.2 mm; 32 – 34 = 1.0 mm; 35 = 0.1 mm." httpUri="https://zenodo.org/record/191597/files/figure.png" pageId="23" pageNumber="24">Figs 31</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="13122A31DA55450BFD12FE3166F7FDE2" box="[714,747,500,526]" captionStart="FIGURES 68 – 75" captionStartId="24.[151,269,1392,1416]" captionTargetBox="[193,1403,477,1365]" captionTargetId="figure@24.[192,1413,470,1366]" captionTargetPageId="24" captionText="FIGURES 68 – 75. Genitalic morphology of Austrachelas species. (68 – 71) A. pondoensis n. sp.: female epigyne, ventral view (68), dorsal view (69), male palp, ventral view (70), retrolateral view (71); (72 – 75) A. reavelli n. sp.: female epigyne, ventral view (72), dorsal view (73), male palp, ventral view (74), retrolateral view (75). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/191604/files/figure.png" pageId="23" pageNumber="24">68</figureCitation>
|
|||
|
); copulatory openings laterally situated, entering ST 2 via narrow entrance ducts; ST 2 small, globose, connected broadly to widely separated elongate oval posterior ST 1 (
|
|||
|
<figureCitation id="13122A31DA55450BFF6DFD846511FDB7" box="[181,269,577,603]" captionStart="FIGURES 68 – 75" captionStartId="24.[151,269,1392,1416]" captionTargetBox="[193,1403,477,1365]" captionTargetId="figure@24.[192,1413,470,1366]" captionTargetPageId="24" captionText="FIGURES 68 – 75. Genitalic morphology of Austrachelas species. (68 – 71) A. pondoensis n. sp.: female epigyne, ventral view (68), dorsal view (69), male palp, ventral view (70), retrolateral view (71); (72 – 75) A. reavelli n. sp.: female epigyne, ventral view (72), dorsal view (73), male palp, ventral view (74), retrolateral view (75). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/191604/files/figure.png" pageId="23" pageNumber="24">Fig. 69</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B9636B4DA55450BFF1EFDA26142FD44" blockId="23.[151,1437,152,1686]" pageId="23" pageNumber="24">
|
|||
|
<emphasis id="B95DEAA6DA55450BFF1EFDA265C4FD6D" bold="true" box="[198,472,615,641]" pageId="23" pageNumber="24">Male. Measurements:</emphasis>
|
|||
|
CL 4.78, CW 3.58, FL 0.32, AL 5.30, AW 2.55, SL 2.60, SW 1.92, TL 10.32, AME–AME 0.10, AME–ALE 0.03, ALE–ALE 0.43, PME–PME 0.07, PME–PLE 0.14, PLE–PLE 0.63.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B9636B4DA55450BFF1DFD7065ECFCF0" blockId="23.[151,1437,152,1686]" pageId="23" pageNumber="24">Length of leg segments (sequence from femur to tarsus, and total): I 3.30 + 1.88 + 2.65 + 2.15 + 1.25 = 11.23; II 2.95 + 1.88 + 2.28 + 1.90 + 1.28 = 10.29; III 2.48 + 1.38 + 1.50 + 2.05 + 1.08 = 8.49; IV 3.40 + 1.70 + 2.55 + 3.30 + 1.25 = 12.20.</paragraph>
|
|||
|
<paragraph id="8B9636B4DA55450BFF1DFCEC673AFA8C" blockId="23.[151,1437,152,1686]" pageId="23" pageNumber="24">
|
|||
|
Carapace deep wine-red (
|
|||
|
<figureCitation id="13122A31DA55450BFE2FFCEC664CFCAF" box="[503,592,809,835]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,269,1353,1377]" captionTargetBox="[151,1435,315,1326]" captionTargetId="figure@2.[151,1436,313,1328]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Digital microscope photographs of the habitus of Austrachelas species: (1) A. bergi n. sp., female; (2) A. incertus Lawrence, 1938, female; (3) same, male; (4) A. kalaharinus n. sp., male; (5) A. merwei n. sp., female; (6) same, male; (7) A. natalensis Lawrence, 1942, female; (8) same, male; (9) A. pondoensis n. sp., female; (10) same, male; (11) A. reavelli n. sp., female; (12) same, male; (13) A. sexoculata n. sp., male; (14) A. wassenaari n. sp., female; (15) same, male. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/191595/files/figure.png" pageId="23" pageNumber="24">Fig. 10</figureCitation>
|
|||
|
); AER strongly procurved, laterals slightly larger than medians; AME separated by distance equal to ½ their diameter, separated from ALE by distance slightly smaller than ¼ AME diameter; clypeus height equal to 1½ AME diameter at AME, equal to ALE diameter at ALE; PER straight, laterals larger than medians; PME separated by distance equal to ½ their diameter, PME separated from PLE by distance equal to PME diameter. Chelicerae deep red-brown, anterior margin with coarse ridges; promargin with three teeth, median tooth largest and proximal tooth smallest; median and proximal teeth closest together; retromargin with two small teeth, situated either side of distal promarginal tooth. Legs all deep red, slightly paler orange-red distally. Leg spination: femora: I pl 1 do 1, II pl 1 do 1, III pl 1 do 2 rl 1, IV do 2; patellae: spineless; tibiae: I & II spineless, III pl 2 rl 2 plv 2 rlv 1 vt 2, IV pl 1 rl 2 plv 2 rlv 2 vt 2; metatarsi: I & II spineless, strongly scopulate, III & IV scopulate in distal half, III pl 6 rl 6 plv 1 rlv 1 vt 2, IV pl 6 rl 6 plv 2 rlv 1 vt 2. Abdomen dark grey with pale grey chevron dorsally (
|
|||
|
<figureCitation id="13122A31DA55450BFC1AFB696001FB2A" box="[962,1053,1196,1222]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,269,1353,1377]" captionTargetBox="[151,1435,315,1326]" captionTargetId="figure@2.[151,1436,313,1328]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Digital microscope photographs of the habitus of Austrachelas species: (1) A. bergi n. sp., female; (2) A. incertus Lawrence, 1938, female; (3) same, male; (4) A. kalaharinus n. sp., male; (5) A. merwei n. sp., female; (6) same, male; (7) A. natalensis Lawrence, 1942, female; (8) same, male; (9) A. pondoensis n. sp., female; (10) same, male; (11) A. reavelli n. sp., female; (12) same, male; (13) A. sexoculata n. sp., male; (14) A. wassenaari n. sp., female; (15) same, male. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/191595/files/figure.png" pageId="23" pageNumber="24">Fig. 10</figureCitation>
|
|||
|
), cream laterally and ventrally; dorsum with small anterior scutum, venter without sclerites. Male palpal tibia with curved retrolateral apophysis with three distinctive tooth-like apophyses along inner margin; median tegular apophysis broad, hook-like with blunt tip; embolus arc-shaped, with large triangular lobe partially hidden behind distal margin of embolus (
|
|||
|
<figureCitation id="13122A31DA55450BFEF1FA8365AEFA8C" box="[297,434,1350,1376]" captionStart="FIGURES 68 – 75" captionStartId="24.[151,269,1392,1416]" captionTargetBox="[193,1403,477,1365]" captionTargetId="figure@24.[192,1413,470,1366]" captionTargetPageId="24" captionText="FIGURES 68 – 75. Genitalic morphology of Austrachelas species. (68 – 71) A. pondoensis n. sp.: female epigyne, ventral view (68), dorsal view (69), male palp, ventral view (70), retrolateral view (71); (72 – 75) A. reavelli n. sp.: female epigyne, ventral view (72), dorsal view (73), male palp, ventral view (74), retrolateral view (75). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/191604/files/figure.png" pageId="23" pageNumber="24">Figs 70, 71</figureCitation>
|
|||
|
); palpal spination: femora do 2.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="C333653FDA55450BFF1DFAA867C4F9A4" pageId="23" pageNumber="24" type="materials_examined">
|
|||
|
<paragraph id="8B9636B4DA55450BFF1DFAA86684FA38" blockId="23.[151,1437,152,1686]" pageId="23" pageNumber="24">
|
|||
|
<emphasis id="B95DEAA6DA55450BFF1DFAA8656AFA6B" bold="true" box="[197,374,1389,1415]" pageId="23" pageNumber="24">
|
|||
|
<typeStatus id="54928816DA55450BFF1DFAA8651FFA6B" box="[197,259,1389,1415]" pageId="23" pageNumber="24">Type</typeStatus>
|
|||
|
material
|
|||
|
</emphasis>
|
|||
|
:
|
|||
|
<typeStatus id="54928816DA55450BFE5FFAA865EAFA6B" box="[391,502,1389,1415]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
|
|||
|
Ƥ, deposited together with 13 and 7Ƥ
|
|||
|
<typeStatus id="54928816DA55450BFC0CFAA86057FA6B" box="[980,1099,1389,1415]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
|
|||
|
:
|
|||
|
<collectingCountry id="F33E7624DA55450BFB8FFAA86128FA6B" box="[1111,1332,1389,1415]" name="South Africa" pageId="23" pageNumber="24">SOUTH AFRICA</collectingCountry>
|
|||
|
:
|
|||
|
<emphasis id="B95DEAA6DA55450BFAE7FAA86553FA41" italics="true" pageId="23" pageNumber="24">Eastern Cape Province</emphasis>
|
|||
|
, Lusikisiki district, Mzimhlava River mouth,
|
|||
|
<geoCoordinate id="EE1D5073DA55450BFC48FA5167E8FA42" box="[912,1012,1428,1454]" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="925" value="-31.333334">31°20'S</geoCoordinate>
|
|||
|
,
|
|||
|
<geoCoordinate id="EE1D5073DA55450BFBDAFA516074FA42" box="[1026,1128,1428,1454]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="925" value="29.666666">29°40'E</geoCoordinate>
|
|||
|
,
|
|||
|
<date id="FF971074DA55450BFBAEFA5160CFFA42" box="[1142,1235,1428,1454]" pageId="23" pageNumber="24" value="1980-02">II.1980</date>
|
|||
|
, M.E. Baddeley (coastal evergreen forest) (
|
|||
|
<collectionCode id="ED38AE71DA55450BFE17FA7F663BFA38" box="[463,551,1466,1492]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="23" pageNumber="24">MRAC</collectionCode>
|
|||
|
163974).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B9636B4DA55450BFF1EFA2567C4F9A4" blockId="23.[151,1437,152,1686]" pageId="23" pageNumber="24">
|
|||
|
<emphasis id="B95DEAA6DA55450BFF1EFA256620FA16" bold="true" box="[198,572,1504,1530]" pageId="23" pageNumber="24">Additional material examined</emphasis>
|
|||
|
: Same data as
|
|||
|
<typeStatus id="54928816DA55450BFD34FA246731FA17" box="[748,813,1505,1531]" pageId="23" pageNumber="24">types</typeStatus>
|
|||
|
,
|
|||
|
<date id="FF971074DA55450BFCE0FA2467ACFA17" box="[824,944,1505,1531]" pageId="23" pageNumber="24" value="1980-01-12">I.1980, 12</date>
|
|||
|
Ƥ 13 (
|
|||
|
<collectionCode id="ED38AE71DA55450BFBD9FA246045FA17" box="[1025,1113,1505,1531]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="23" pageNumber="24">MRAC</collectionCode>
|
|||
|
159047, SEM preparations MJR-
|
|||
|
<date id="FF971074DA55450BFF06F9CD6540F9CE" box="[222,348,1544,1570]" pageId="23" pageNumber="24" value="1031" valueMax="1038">1031-1038</date>
|
|||
|
,
|
|||
|
<date id="FF971074DA55450BFEB1F9CD65F7F9CE" box="[361,491,1544,1570]" pageId="23" pageNumber="24" value="1063" valueMax="1065">1063-1065</date>
|
|||
|
, temporary mount CJG-00154);
|
|||
|
<date id="FF971074DA55450BFCB5F9CD67F1F9CC" box="[877,1005,1542,1570]" pageId="23" pageNumber="24" value="1980-01-13">I.1980, 13</date>
|
|||
|
(
|
|||
|
<collectionCode id="ED38AE71DA55450BFC26F9CD604BF9CE" box="[1022,1111,1544,1570]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="23" pageNumber="24">MRAC</collectionCode>
|
|||
|
166821, SEM preparations MJR-01039, MJR-01040, temporary mounts CJG-00155, CJG-00156).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="C333653FDA55450BFF1EF99064D9F97A" pageId="23" pageNumber="24" type="distribution">
|
|||
|
<paragraph id="8B9636B4DA55450BFF1EF99064D9F97A" blockId="23.[151,1437,152,1686]" pageId="23" pageNumber="24">
|
|||
|
<emphasis id="B95DEAA6DA55450BFF1EF9906543F983" bold="true" box="[198,351,1621,1647]" pageId="23" pageNumber="24">Distribution</emphasis>
|
|||
|
. Known only from the
|
|||
|
<typeStatus id="54928816DA55450BFDB7F99066BDF983" box="[623,673,1621,1647]" pageId="23" pageNumber="24">type</typeStatus>
|
|||
|
locality in Pondoland, Eastern Cape Province,
|
|||
|
<collectingCountry id="F33E7624DA55450BFB1FF9906141F983" box="[1223,1373,1621,1647]" name="South Africa" pageId="23" pageNumber="24">South Africa</collectingCountry>
|
|||
|
(
|
|||
|
<figureCitation id="13122A31DA55450BFAB4F99064A4F97A" captionStart="FIGURE 82" captionStartId="28.[151,255,1122,1146]" captionTargetBox="[274,1329,188,1095]" captionTargetId="figure@28.[268,1335,181,1109]" captionTargetPageId="28" captionText="FIGURE 82. Distribution of the genus Austrachelas, endemic to South Africa." httpUri="https://zenodo.org/record/191606/files/figure.png" pageId="23" pageNumber="24">Fig. 82</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|