treatments-xml/data/03/E8/30/03E8307A6651027D9992FC79C472FCDF.xml

<document id="551FFCA62027B8001D1AE1C2A87422C4" ID-CLB-Dataset="291528" ID-DOI="10.11646/zootaxa.5428.2.5" ID-GBIF-Dataset="b0a98796-768e-43e2-be6b-1f79c24b9227" ID-ISSN="1175-5326" ID-Zenodo-Dep="10847635" ID-ZooBank="135D8285-BEC0-467E-BCD5-4DA116C6DFBC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1711009984219" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Baňař, Petr" docDate="2024" docId="03E8307A6651027D9992FC79C472FCDF" docLanguage="en" docName="zootaxa.5428.2.5.pdf" docOrigin="Zootaxa 5428 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5428.2.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paulianidia tanala Janak &amp; Banar 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="258" masterDocId="FFD14802665202789905FFF3C74BFFCA" masterDocTitle="Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="268" masterPageNumber="253" pageNumber="256" updateTime="1711133149338" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C1F053904649A85B822C79BE02775AC0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C226394181A1F4D58F0370220370843F">
<mods:title id="648F6CDF6E12F91B824A4C36AC7BAFFD">Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae)</mods:title>
</mods:titleInfo>
<mods:name id="514E4F9D703F7C4E05DF6D4DBB7FE775" type="personal">
<mods:role id="F8100FB5626334D35FDBA919883C0954">
<mods:roleTerm id="ADF0CFBE70F12FDFA430A2E76DFC0805">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5AFF506735587AD65405787F3261701C">Janák, Jiří</mods:namePart>
<mods:affiliation id="A39A99D3E35A55DCA80B8C88C3DD5992">CZ- 417 62 Rtyně nad Bílinou, č. 4, Czech Republic</mods:affiliation>
</mods:name>
<mods:name id="0E11C130ABC408305B7389A59CC18C0D" type="personal">
<mods:role id="81382E4C4D8128232D4734A4120EABBD">
<mods:roleTerm id="9CF0B4342C4FC9D676414777D74DB967">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D9C167202A9B9109217259E20B350DBF">Baňař, Petr</mods:namePart>
<mods:affiliation id="110A5945F23713A10E3ECFFB32E33D2E">Moravian Museum, Department of Entomology, Hviezdoslavova 29 a, Brno, CZ- 627 00, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0B6B4F454E0453AE6294E1F2A3D5FE8B">text</mods:typeOfResource>
<mods:relatedItem id="F7650C57B4B759279C6213C4FF094481" type="host">
<mods:titleInfo id="365D0D632B531C9641A2DECB89D83690">
<mods:title id="C27469A8340E0564E68FE24963849BFC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8BEA16B0949B379CE1CAAB64D07C9803">
<mods:date id="90F7FC9A5D33CE346E9A083616DA4C33">2024</mods:date>
<mods:detail id="1F511F0BBFC5E83E2D75E4DFB1A212C5" type="pubDate">
<mods:number id="A2D69E48F52120A4EB8EE4B9F2B3883F">2024-03-21</mods:number>
</mods:detail>
<mods:detail id="A0FC7B48A418A1325A12452B858788DC" type="volume">
<mods:number id="0509D8C6FCD2831BE63F1AB1A7219B62">5428</mods:number>
</mods:detail>
<mods:detail id="AD43EAA600E2985533A5A41251D8007D" type="issue">
<mods:number id="3D7CE41C3ED50A4FD21342E20DCCC2B9">2</mods:number>
</mods:detail>
<mods:extent id="97BDFC8A9E4073F5A919A3C59137B0AE" unit="page">
<mods:start id="FFFB11F0B78E3E113B1CC415DDFD2009">253</mods:start>
<mods:end id="DC06876199120E5E525ACDD61411B577">268</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3524BCE8C91A0AF1E26F362BC297329D">
<mods:url id="9D14E5BCCB0D8CCCE92925A61BED78AC">http://dx.doi.org/10.11646/zootaxa.5428.2.5</mods:url>
</mods:location>
<mods:classification id="DC5910BF6BD09DB2E1C0F52B12EB8335">journal article</mods:classification>
<mods:identifier id="DC0FEE9C9FF346A5EDC1754912101C48" type="CLB-Dataset">291528</mods:identifier>
<mods:identifier id="91B6400ACC6F45912F43C939AF70700F" type="DOI">10.11646/zootaxa.5428.2.5</mods:identifier>
<mods:identifier id="4FA4DDF8E8EF3A11F035D9DF7A8B735E" type="GBIF-Dataset">b0a98796-768e-43e2-be6b-1f79c24b9227</mods:identifier>
<mods:identifier id="C110FCB908ECBA621AD3C88DC36A45FD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="04E9A26F7F180ADB379B545C04AD5699" type="Zenodo-Dep">10847635</mods:identifier>
<mods:identifier id="CEF14844FCE363859D53D8566C641398" type="ZooBank">135D8285-BEC0-467E-BCD5-4DA116C6DFBC</mods:identifier>
</mods:mods>
<treatment id="03E8307A6651027D9992FC79C472FCDF" ID-DOI="http://doi.org/10.5281/zenodo.10847573" ID-Zenodo-Dep="10847573" LSID="urn:lsid:plazi:treatment:03E8307A6651027D9992FC79C472FCDF" httpUri="http://treatment.plazi.org/id/03E8307A6651027D9992FC79C472FCDF" lastPageId="5" lastPageNumber="258" pageId="3" pageNumber="256">
<subSubSection id="C35BD2E76651027B9992FC79C6F0FC03" pageId="3" pageNumber="256" type="nomenclature">
<paragraph id="8BFE816C6651027B9992FC79C5E5FC6F" blockId="3.[151,686,906,969]" box="[151,686,906,933]" pageId="3" pageNumber="256">
<heading id="D0B636006651027B9992FC79C5E5FC6F" bold="true" box="[151,686,906,933]" fontSize="11" level="1" pageId="3" pageNumber="256" reason="1">
<emphasis id="B9355D7E6651027B9992FC79C5E5FC6F" bold="true" box="[151,686,906,933]" pageId="3" pageNumber="256">
<taxonomicName id="4C41FAEF6651027B9992FC79C50FFC6E" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2024" box="[151,580,906,933]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="tanala" status="sp. nov.">
<emphasis id="B9355D7E6651027B9992FC79C632FC6F" bold="true" box="[151,377,906,933]" italics="true" pageId="3" pageNumber="256">Paulianidia tanala</emphasis>
Janák &amp; Baňař
</taxonomicName>
, 
<taxonomicNameLabel id="A206E0056651027B9B54FC78C5E5FC6F" box="[593,686,907,933]" pageId="3" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE816C6651027B9992FC5CC6F0FC03" blockId="3.[151,686,906,969]" box="[151,443,943,969]" pageId="3" pageNumber="256">
(
<figureCitation id="137A9DE96651027B999AFC5CC637FC03" box="[159,380,943,969]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="3" pageNumber="256">Figs. 19–27, 29–30</figureCitation>
, 
<figureCitation id="137A9DE96651027B9882FC5CC6F9FC03" box="[391,434,943,969]" captionStart="FIGURES 116–122" captionStartId="12.[151,264,1780,1804]" captionTargetBox="[180,1408,194,1743]" captionTargetId="figure-34@12.[167,1421,181,1756]" captionText="FIGURES 116–122. 116–120: Paulianidia steineri Lecoq, Andasibe: 116–118, aedeagus (116, ventral; 117, 1/3 lateral; 118, lateral); 119, male sternite VII; 120, male sternite VIII. Scale 0.5 mm: 116–120. 121: Distribution of P. densepunctata sp. nov., (rhombus), P. valohoaka sp. nov. (star), P. tanala sp. nov. and P. vohimana sp. nov. (rectangle). 122: Distribution of Paulianidia species in Madagascar." figureDoi="http://doi.org/10.5281/zenodo.10854991" httpUri="https://zenodo.org/record/10854991/files/figure.png" pageId="3" pageNumber="256">121</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76651027B9992FC04C658FA0B" pageId="3" pageNumber="256" type="materials_examined">
<paragraph id="8BFE816C6651027B9992FC04C5B4FBDB" blockId="3.[151,1437,1014,1977]" box="[151,767,1014,1041]" pageId="3" pageNumber="256">
<materialsCitation id="3B298B316651027B9992FC04C5B4FBDB" box="[151,767,1014,1041]" country="Madagascar" location="Ranomafana National Park" pageId="3" pageNumber="256" specimenCount="1" typeStatus="holotype">
<emphasis id="B9355D7E6651027B9992FC04C67DFBDA" bold="true" box="[151,310,1014,1041]" pageId="3" pageNumber="256">
<typeStatus id="54FA3FCE6651027B9992FC04C799FBDB" box="[151,210,1015,1041]" pageId="3" pageNumber="256">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F356C1FC6651027B9839FC04C68EFBDB" box="[316,453,1015,1041]" name="Madagascar" pageId="3" pageNumber="256">Madagascar</collectingCountry>
, 
<location id="8E9ED7B76651027B98D5FC04C5B4FBDB" LSID="urn:lsid:plazi:treatment:03E8307A6651027D9992FC79C472FCDF:8E9ED7B76651027B98D5FC04C5B4FBDB" box="[464,767,1015,1041]" country="Madagascar" name="Ranomafana National Park" pageId="3" pageNumber="256">Ranomafana National Park</location>
</materialsCitation>
</paragraph>
<paragraph id="8BFE816C6651027B99C2FBE8C658FA0B" blockId="3.[151,1437,1014,1977]" pageId="3" pageNumber="256">
<emphasis id="B9355D7E6651027B99C2FBE8C63DFBFE" bold="true" box="[199,374,1050,1077]" pageId="3" pageNumber="256">Type material.</emphasis>
<materialsCitation id="3B298B316651027B987AFBE8C3DAFBB7" collectingDate="2012-09" collectingDateMax="2012-10-08" collectingDateMin="2012-09" collectionCode="MMBC" collectorName="L. S. Rahanitriniaina" country="Madagascar" determinerName="Banar" elevation="1003" latitude="-21.265112" location="MADAGASCAR" longLatPrecision="1" longitude="47.417168" pageId="3" pageNumber="256" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B9355D7E6651027B987AFBE8C6A7FBFF" bold="true" box="[383,492,1051,1077]" pageId="3" pageNumber="256">
<typeStatus id="54FA3FCE6651027B987AFBE8C6A7FBFF" box="[383,492,1051,1077]" pageId="3" pageNumber="256" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D474AE56651027B98F1FBE8C547FBFF" box="[500,524,1051,1077]" count="1" pageId="3" pageNumber="256" type="male">♂</specimenCount>
: 
<emphasis id="B9355D7E6651027B9B1CFBE9C5ADFBFE" bold="true" box="[537,742,1050,1076]" pageId="3" pageNumber="256">
<collectingCountry id="F356C1FC6651027B9B1CFBE9C5ADFBFE" box="[537,742,1050,1076]" name="Madagascar" pageId="3" pageNumber="256">MADAGASCAR</collectingCountry>
</emphasis>
: “RNF/ 
<date id="FFFFA7AC6651027B9A46FBE8C4F6FBFF" box="[835,957,1051,1077]" pageId="3" pageNumber="256" value="2012-09">
<collectingDate id="EFBB5E446651027B9A46FBE8C4F6FBFF" box="[835,957,1051,1077]" pageId="3" pageNumber="256" value="2012-09">Sept. 2012</collectingDate>
</date>
/11 MADAGASCAR, RANOMAFANA N. P.; 
<date id="FFFFA7AC6651027B9992FBCCC7B2FB93" box="[151,249,1087,1113]" pageId="3" pageNumber="256" value="2012-10-08">
<collectingDate id="EFBB5E446651027B9992FBCCC7B2FB93" box="[151,249,1087,1113]" pageId="3" pageNumber="256" value="2012-10-08">8.x.2012</collectingDate>
</date>
<geoCoordinate id="EE75E7AB6651027B9801FBCDC6D5FB93" box="[260,414,1086,1113]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="3" pageNumber="256" precision="1" seconds="54.4" value="-21.265112">S21°15’54.4”</geoCoordinate>
<geoCoordinate id="EE75E7AB6651027B98ACFBCCC50CFB93" box="[425,583,1086,1113]" degrees="47" direction="east" minutes="25" orientation="longitude" pageId="3" pageNumber="256" precision="1" seconds="01.8" value="47.417168">E47°25’01.8”</geoCoordinate>
; 
<quantity id="4CB92C896651027B9B52FBCCC5EEFB93" box="[599,677,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="1.003" pageId="3" pageNumber="256" unit="m" value="1003.0">1003m</quantity>
TALATAKELY, sifting litter, Winkler app. extr. 
<collectorName id="26B4E4BA6651027B9DB3FBCCC2D6FB93" box="[1206,1437,1087,1113]" pageId="3" pageNumber="256">L.S. Rahanitriniaina</collectorName>
lgt.” // “HOLOTYPUS 
<taxonomicName id="4C41FAEF6651027B98AFFB90C53EFBB7" authority="Janák &amp; Baňař, 2024" authorityName="Janák &amp; Baňař" authorityYear="2024" box="[426,629,1123,1149]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="tanala" status="sp. nov.">Paulianidia tanala</taxonomicName>
<taxonomicNameLabel id="A206E0056651027B9B7AFB90C590FBB7" box="[639,731,1123,1149]" pageId="3" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
, Janák &amp; 
<determinerName id="7E226DA86651027B9A51FB90C4DCFBB7" box="[852,919,1123,1149]" pageId="3" pageNumber="256">Baňař</determinerName>
det. 2023” (
<collectionCode id="ED5019A96651027B9D2EFB90C3C2FBB7" box="[1067,1161,1123,1149]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="256">MMBC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B298B316651027B9D9AFB90C380FB0F" collectingDate="2012-09" collectingDateMax="2012-09-27" collectingDateMin="2012-09" collectionCode="MMBC, JJRC" country="Madagascar" elevation="1017" latitude="-21.261223" longLatPrecision="1" longitude="47.42125" pageId="3" pageNumber="256" specimenCount="8" specimenCount-female="3" specimenCount-male="5" typeStatus="paratype">
<emphasis id="B9355D7E6651027B9D9AFB90C25CFBB7" bold="true" box="[1183,1303,1123,1149]" pageId="3" pageNumber="256">
<typeStatus id="54FA3FCE6651027B9D9AFB90C25CFBB7" box="[1183,1303,1123,1149]" pageId="3" pageNumber="256" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="9D474AE56651027B9C2DFB91C213FBB7" box="[1320,1368,1122,1149]" count="5" pageId="3" pageNumber="256" type="male">5 ♂</specimenCount>
, 
<specimenCount id="9D474AE56651027B9C62FB90C2DDFBB7" box="[1383,1430,1123,1149]" count="3" pageId="3" pageNumber="256" type="female">3 ♀</specimenCount>
: “RNF/ 
<date id="FFFFA7AC6651027B99E1FB74C628FB6B" box="[228,355,1159,1185]" pageId="3" pageNumber="256" value="2012-09">
<collectingDate id="EFBB5E446651027B99E1FB74C628FB6B" box="[228,355,1159,1185]" pageId="3" pageNumber="256" value="2012-09">Sept. 2012</collectingDate>
</date>
/01 MADAGASCAR, RANOMAFANA N. P.; 
<date id="FFFFA7AC6651027B9A68FB74C4AEFB6B" box="[877,997,1159,1185]" pageId="3" pageNumber="256" value="2012-09-27">
<collectingDate id="EFBB5E446651027B9A68FB74C4AEFB6B" box="[877,997,1159,1185]" pageId="3" pageNumber="256" value="2012-09-27">27.ix.2012</collectingDate>
</date>
<geoCoordinate id="EE75E7AB6651027B9AF1FB75C3C6FB6B" box="[1012,1165,1158,1185]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="3" pageNumber="256" precision="1" seconds="40.4" value="-21.261223">S21°15’40.4”</geoCoordinate>
<geoCoordinate id="EE75E7AB6651027B9D98FB74C271FB6B" box="[1181,1338,1158,1185]" degrees="47" direction="east" minutes="25" orientation="longitude" pageId="3" pageNumber="256" precision="1" seconds="16.5" value="47.42125">E47°25’16.5”</geoCoordinate>
; 
<quantity id="4CB92C896651027B9C4BFB74C2D7FB6B" box="[1358,1436,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.017" pageId="3" pageNumber="256" unit="m" value="1017.0">1017m</quantity>
AMBATOLAHIDIMY; sifting litter, Winkler app. extr.; L.S. Rahanitriniaina lgt.” (
<collectionCode id="ED5019A96651027B9D1CFB58C33FFB0F" box="[1049,1140,1195,1221]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="256">MMBC</collectionCode>
, 
<collectionCode id="ED5019A96651027B9D86FB58C388FB0F" box="[1155,1219,1195,1221]" pageId="3" pageNumber="256">JJRC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B298B316651027B9DDFFB58C467FAC7" collectingDate="2012-09" collectingDateMax="2012-10-06" collectingDateMin="2012-09" collectionCode="JJRC" country="Madagascar" elevation="1008" latitude="-21.297722" longLatPrecision="1" longitude="47.443085" pageId="3" pageNumber="256" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D474AE56651027B9DDFFB58C241FB0F" box="[1242,1290,1195,1221]" count="3" pageId="3" pageNumber="256" type="male">3 ♂</specimenCount>
: “RNF/ 
<date id="FFFFA7AC6651027B9C63FB58C79BFB23" pageId="3" pageNumber="256" value="2012-09">
<collectingDate id="EFBB5E446651027B9C63FB58C79BFB23" pageId="3" pageNumber="256" value="2012-09">Sept. 2012</collectingDate>
</date>
/10 MADAGASCAR, RANOMAFANA N. P.; 
<date id="FFFFA7AC6651027B9BB0FB3CC45DFB23" box="[693,790,1231,1257]" pageId="3" pageNumber="256" value="2012-10-06">
<collectingDate id="EFBB5E446651027B9BB0FB3CC45DFB23" box="[693,790,1231,1257]" pageId="3" pageNumber="256" value="2012-10-06">6.x.2012</collectingDate>
</date>
<geoCoordinate id="EE75E7AB6651027B9A1BFB3DC4FCFB23" box="[798,951,1230,1257]" degrees="21" direction="south" minutes="17" orientation="latitude" pageId="3" pageNumber="256" precision="1" seconds="51.8" value="-21.297722">S21°17’51.8”</geoCoordinate>
<geoCoordinate id="EE75E7AB6651027B9ABAFB3CC316FB23" box="[959,1117,1230,1257]" degrees="47" direction="east" minutes="26" orientation="longitude" pageId="3" pageNumber="256" precision="1" seconds="35.1" value="47.443085">E47°26’35.1”</geoCoordinate>
; 
<quantity id="4CB92C896651027B9D6CFB3CC3FCFB23" box="[1129,1207,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.008" pageId="3" pageNumber="256" unit="m" value="1008.0">1008m</quantity>
VALOHOAKA, sifting litter, Winkler app. extr. L.S. Rahanitriniaina lgt.” (
<collectionCode id="ED5019A96651027B9BE6FB00C46FFAC7" box="[739,804,1267,1293]" pageId="3" pageNumber="256">JJRC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B298B316651027B9A3FFB00C36DFAFB" collectingDate="2010-11-16" collectingDateMax="2010-11-18" collectingDateMin="2010-11-16" collectionCode="MMBC" country="Madagascar" elevation="1079" latitude="-21.2475" longLatPrecision="21" longitude="47.403614" pageId="3" pageNumber="256" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D474AE56651027B9A3FFB00C421FAC7" box="[826,874,1267,1293]" count="2" pageId="3" pageNumber="256" type="male">2 ♂</specimenCount>
: “MADAGASCAR—CE, Ranomafana N.P., 
<date id="FFFFA7AC6651027B9C77FB00C659FAFB" pageId="3" pageNumber="256" value="2010-11-16" valueMax="2010-11-18" valueMin="2010-11-16">
<collectingDate id="EFBB5E446651027B9C77FB00C659FAFB" pageId="3" pageNumber="256" value="2010-11-16" valueMax="2010-11-18" valueMin="2010-11-16">16– 18.xi.2010</collectingDate>
</date>
; 
<quantity id="4CB92C896651027B9825FAE4C63CFAFB" box="[288,375,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.079" pageId="3" pageNumber="256" unit="m" value="1079.0">1079 m</quantity>
// 
<geoCoordinate id="EE75E7AB6651027B989CFAE5C555FAFB" box="[409,542,1302,1329]" degrees="21" direction="south" minutes="14" orientation="latitude" pageId="3" pageNumber="256" precision="15" seconds="51" value="-21.2475">S21°14’51”</geoCoordinate>
<geoCoordinate id="EE75E7AB6651027B9B22FAE4C5E4FAFB" box="[551,687,1303,1329]" degrees="47" direction="east" minutes="24" orientation="longitude" pageId="3" pageNumber="256" precision="15" seconds="13" value="47.403614">E47°24’13”</geoCoordinate>
, sifting; P. Baňař lgt.” (
<collectionCode id="ED5019A96651027B9AC5FAE4C356FAFB" box="[960,1053,1303,1329]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="256">MMBC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B298B316651027B9D36FAE4C211FA9F" collectionCode="MMBC" country="Madagascar" elevation="1079" latitude="-21.2475" longLatPrecision="21" longitude="47.403614" pageId="3" pageNumber="256" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D474AE56651027B9D36FAE4C32AFAFB" box="[1075,1121,1303,1329]" count="1" pageId="3" pageNumber="256" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D474AE56651027B9D75FAE4C3D5FAFB" box="[1136,1182,1303,1329]" count="1" pageId="3" pageNumber="256" type="female">1 ♀</specimenCount>
: “MADAGASCAR— CE 2010, RANOMAFANA N.P., 16–18.xi., 
<geoCoordinate id="EE75E7AB6651027B9B6CFAC9C5A5FA9F" box="[617,750,1338,1365]" degrees="21" direction="south" minutes="14" orientation="latitude" pageId="3" pageNumber="256" precision="15" seconds="51" value="-21.2475">S21°14’51”</geoCoordinate>
<geoCoordinate id="EE75E7AB6651027B9BF2FAC8C4CBFA9F" box="[759,896,1339,1365]" degrees="47" direction="east" minutes="24" orientation="longitude" pageId="3" pageNumber="256" precision="15" seconds="13" value="47.403614">E47°24’13”</geoCoordinate>
, 
<quantity id="4CB92C896651027B9A8BFAC8C4ADFA9F" box="[910,998,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.079" pageId="3" pageNumber="256" unit="m" value="1079.0">1079 m</quantity>
sifting; P. Baňař lgt.” (
<collectionCode id="ED5019A96651027B9DF1FAC8C21AFA9F" box="[1268,1361,1339,1365]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="256">MMBC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B298B316651027B9C6DFAC8C674FA57" collectingDate="2010-11-17" collectionCode="MMBC" country="Madagascar" elevation="960" pageId="3" pageNumber="256" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D474AE56651027B9C6DFAC8C2DCFA9F" box="[1384,1431,1339,1365]" count="1" pageId="3" pageNumber="256" type="male">1 ♂</specimenCount>
: “MADAGASCAR—CE, RANOMAFANA N.P., 
<date id="FFFFA7AC6651027B9B96FAACC445FAB3" box="[659,782,1375,1401]" pageId="3" pageNumber="256" value="2010-11-17">
<collectingDate id="EFBB5E446651027B9B96FAACC445FAB3" box="[659,782,1375,1401]" pageId="3" pageNumber="256" value="2010-11-17">17.xi.2010</collectingDate>
</date>
, ~ 
<quantity id="4CB92C896651027B9A22FAACC424FAB3" box="[807,879,1375,1401]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="256" unit="m" value="960.0">960 m</quantity>
sifting by stream close to main entrance; P. Baňař lgt.” (
<collectionCode id="ED5019A96651027B99DCFA70C67CFA57" box="[217,311,1411,1437]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="256">MMBC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B298B316651027B984FFA70C65BFA0B" country="Madagascar" determinerName="Janak &amp; Banar" location="All" pageId="3" pageNumber="256" specimenCount="2" typeStatus="paratype">
<location id="8E9ED7B76651027B984FFA70C625FA57" LSID="urn:lsid:plazi:treatment:03E8307A6651027D9992FC79C472FCDF:8E9ED7B76651027B984FFA70C625FA57" box="[330,366,1411,1437]" country="Madagascar" name="All" pageId="3" pageNumber="256">All</location>
paratypes with an additional label “PARATYPUS 
<taxonomicName id="4C41FAEF6651027B9AB3FA70C3CBFA57" authority="Janák &amp; Baňař, 2024" authorityName="Janák &amp; Baňař" authorityYear="2024" box="[950,1152,1411,1437]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="tanala" status="sp. nov.">Paulianidia tanala</taxonomicName>
<taxonomicNameLabel id="A206E0056651027B9D8CFA70C3AFFA57" box="[1161,1252,1411,1437]" pageId="3" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
, 
<determinerName id="7E226DA86651027B9DF4FA70C27AFA57" box="[1265,1329,1411,1437]" pageId="3" pageNumber="256">Janák</determinerName>
&amp; 
<determinerName id="7E226DA86651027B9C5CFA70C2D7FA57" box="[1369,1436,1411,1437]" pageId="3" pageNumber="256">Baňař</determinerName>
det. 2023”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76651027D99C2FA39C5B0FE43" lastPageId="5" lastPageNumber="258" pageId="3" pageNumber="256" type="description">
<paragraph id="8BFE816C6651027B99C2FA39C6E6F9E7" blockId="3.[151,1437,1014,1977]" pageId="3" pageNumber="256">
<emphasis id="B9355D7E6651027B99C2FA39C61AFA2E" bold="true" box="[199,337,1482,1508]" pageId="3" pageNumber="256">Description</emphasis>
: Body length 5.2–6.0 mm (M = 
<quantity id="4CB92C896651027B9BCEFA39C46EFA2F" box="[715,805,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="256" unit="mm" value="5.5">5.5 mm</quantity>
, HT = 
<quantity id="4CB92C896651027B9A7DFA39C49FFA2F" box="[888,980,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="256" unit="mm" value="5.9">5.9 mm</quantity>
), forebody length 
<quantity id="4CB92C896651027B9DA0FA38C266FA2F" box="[1189,1325,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="3" pageNumber="256" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 mm</quantity>
(M = 3.0 mm, HT = 3.0 mm). Reddish brown to brown, abdomen, mouthparts and antennomeres reddish brown, legs light reddish yellow (
<figureCitation id="137A9DE96651027B9849F9E0C6EBF9E7" box="[332,416,1555,1581]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="3" pageNumber="256">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8BFE816C6651027B99C2F9C4C3DCF8CF" blockId="3.[151,1437,1014,1977]" pageId="3" pageNumber="256">
Head (
<figureCitation id="137A9DE96651027B9816F9C4C62DF99B" box="[275,358,1591,1617]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="3" pageNumber="256">Fig. 20</figureCitation>
) moderately convex, only slightly longer than wide (L/W = 1.03–1.10, M = 1.06, HT = 1.06). Surface between eyes with coarse and dense umbilicate punctures, in centre of disc with punctation markedly sparser and finer and here and there with interstices larger than diameter of punctures. Eyes moderately large, temples on average about 1.5 times as long as eyes (R = 1.46–1.68, M = 1.56, HT = 1.46), markedly narrowed posteriorly. Antennae long and slender, fifth antennomere about one and half as long as wide (L/W = 1.34–1.65, M = 1.55, HT = 1.55), tenth antennomere about as long as wide (L/W = 0.88–1.05, M = 0.96, HT = 1.05).
</paragraph>
<paragraph id="8BFE816C6651027B99C2F8FCC4D1F873" blockId="3.[151,1437,1014,1977]" pageId="3" pageNumber="256">
Pronotum (
<figureCitation id="137A9DE96651027B9843F8FCC6D3F8E3" box="[326,408,1807,1833]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="3" pageNumber="256">Fig. 20</figureCitation>
) egg-shaped, moderately convex, with shallow lateral impressions, about 1.1 times as long as wide (L/W = 1.10–1.16, M = 1.13, HT = 1.14) and almost as wide as head (PW/HW = 0.94–0.99, M = 0.96, HT = 0.97). Sides slightly widened posteriorly. Surface with moderately coarse and sparse punctures and with distinct interstices, punctation coarse and less dense than that on head and less dense along moderately narrow, in anterior part of pronotum more than one puncture wide unpunctured midline.
</paragraph>
<caption id="DF3ED1E46656027C9992F934C3BFF8ED" ID-DOI="http://doi.org/10.5281/zenodo.10854989" ID-Zenodo-Dep="10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="4" pageNumber="257" startId="4.[151,264,1735,1759]" targetBox="[173,1414,195,1698]" targetPageId="4">
<paragraph id="8BFE816C6656027C9992F934C3BFF8ED" blockId="4.[151,1436,1735,1832]" pageId="4" pageNumber="257">
<emphasis id="B9355D7E6656027C9992F934C604F915" bold="true" box="[151,335,1735,1759]" pageId="4" pageNumber="257">FIGURES 19–30</emphasis>
. 19–27, 29–30: 
<taxonomicName id="4C41FAEF6656027C98F3F934C5FDF915" authority="Janák &amp; Baňař, 2024" authorityName="Janák &amp; Baňař" authorityYear="2024" box="[502,694,1735,1759]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="species" species="tanala" status="sp. nov.">
<emphasis id="B9355D7E6656027C98F3F934C5FDF915" box="[502,694,1735,1759]" italics="true" pageId="4" pageNumber="257">Paulianidia tanala</emphasis>
</taxonomicName>
<emphasis id="B9355D7E6656027C9BB8F934C445F915" bold="true" box="[701,782,1735,1759]" pageId="4" pageNumber="257">
<taxonomicNameLabel id="A206E0056656027C9BB8F934C445F915" box="[701,782,1735,1759]" pageId="4" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 19–23, 26–27, 29–30, 
<typeStatus id="54FA3FCE6656027C9D06F934C315F915" box="[1027,1118,1735,1759]" pageId="4" pageNumber="257" type="holotype">holotype</typeStatus>
, 24–25, 
<typeStatus id="54FA3FCE6656027C9DB3F93BC251F92A" box="[1206,1306,1736,1760]" pageId="4" pageNumber="257" type="paratype">paratypes</typeStatus>
; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: 
<taxonomicName id="4C41FAEF6656027C989BF8E3C57BF8ED" authority="Lecoq" authorityName="Lecoq" authorityYear="2008" box="[414,560,1807,1831]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B9355D7E6656027C989BF8E3C6ACF8ED" box="[414,487,1807,1831]" italics="true" pageId="4" pageNumber="257">P. hova</emphasis>
Lecoq
</taxonomicName>
, 
<typeStatus id="54FA3FCE6656027C9B3FF8E3C5DFF8E2" box="[570,660,1808,1832]" pageId="4" pageNumber="257" type="paratype">paratype</typeStatus>
, head. Scales 
<quantity id="4CB92C896656027C9A24F8FCC414F8ED" box="[801,863,1807,1831]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="257" unit="mm" value="1.0">1 mm</quantity>
: 19, 20; 
<quantity id="4CB92C896656027C9AB3F8FCC34CF8ED" box="[950,1031,1807,1831]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="257" unit="mm" value="0.5">0.5 mm</quantity>
: 21–25, 26–27, 28–30.
</paragraph>
</caption>
<paragraph id="8BFE816C6656027C99C2F880C53CF833" blockId="4.[151,1436,1907,2041]" pageId="4" pageNumber="257">
Elytra (
<figureCitation id="137A9DE96656027C9818F880C63BF847" box="[285,368,1907,1933]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="4" pageNumber="257">Fig. 20</figureCitation>
) subrectangular, with very slightly rounded sides and distinct depression along suture in first quarter behind scutellum, slightly flattened along suture, about 1.3 times as long as wide (R = 1.29–1.42, M = 1.36, HT = 1.33). Surface moderately coarsely and densely punctured. Interstices mostly as large as diameter of punctures, shining, without microscupture.
</paragraph>
<paragraph id="8BFE816C6657027D99C2FF64C4B5FF1F" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="258">
Abdomen (
<figureCitation id="137A9DE96657027D984CFF64C6D5FF7B" box="[329,414,151,177]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="5" pageNumber="258">Fig. 19</figureCitation>
) slightly widened to segment V, very finely and densely punctured. Interstices shining, without microsculpture. Tergite VII with distinct membranous fringe apically.
</paragraph>
<paragraph id="8BFE816C6657027D99C2FF2CC62BFEAF" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="258">
Male. Posterior margin of sternite VII broadly rounded (
<figureCitation id="137A9DE96657027D9A4DFF2CC4D7FF33" box="[840,924,223,249]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="5" pageNumber="258">Fig. 26</figureCitation>
), sternite VIII with relatively wide triangular emargination in posterior third (
<figureCitation id="137A9DE96657027D98FCFEF0C507FED7" box="[505,588,259,285]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="5" pageNumber="258">Fig. 27</figureCitation>
). Aedeagus (
<figureCitation id="137A9DE96657027D9BDCFEF0C414FED7" box="[729,863,258,285]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="5" pageNumber="258">Figs. 21–25</figureCitation>
) long and slender, with relatively small roof shaped apical structures, about 
<quantity id="4CB92C896657027D989AFED5C549FE8B" box="[415,514,294,321]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="258" unit="mm" value="0.85">0.85 mm</quantity>
long (n = 6, AL = 
<quantity id="4CB92C896657027D9BC8FED4C43AFE8B" box="[717,881,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" metricValueMax="9.0" metricValueMin="8.4" pageId="5" pageNumber="258" unit="mm" value="0.87" valueMax="0.9" valueMin="0.84">0.84–0.90 mm</quantity>
, M = 
<quantity id="4CB92C896657027D9AB4FED4C35DFE8B" box="[945,1046,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="5" pageNumber="258" unit="mm" value="0.87">0.87 mm</quantity>
, HT = 
<quantity id="4CB92C896657027D9D67FED4C382FE8B" box="[1122,1225,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="5" pageNumber="258" unit="mm" value="0.88">0.88 mm</quantity>
), parameres longer than median lobe.
</paragraph>
<paragraph id="8BFE816C6657027D99C2FE9CC5B0FE43" blockId="5.[151,1437,151,789]" box="[199,763,367,393]" pageId="5" pageNumber="258">Female. Posterior margin of sternite VIII rounded.</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76657027D99C2FE61C6E9FD4F" pageId="5" pageNumber="258" type="diagnosis">
<paragraph id="8BFE816C6657027D99C2FE61C6E9FD4F" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="258">
<emphasis id="B9355D7E6657027D99C2FE61C69AFE66" bold="true" box="[199,465,402,428]" pageId="5" pageNumber="258">Differential diagnosis.</emphasis>
The new species is arranged in a small group of species with normally (depressed) punctured elytra, impunctate or sparsely punctate disc of the head and integument of tergites smooth between punctures together with 
<taxonomicName id="4C41FAEF6657027D98A2FE28C5C4FE3F" authority="Lecoq, 2008" authorityName="Lecoq" authorityYear="2008" box="[423,655,475,501]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B9355D7E6657027D98A2FE28C6B2FE3F" box="[423,505,475,501]" italics="true" pageId="5" pageNumber="258">P. hova</emphasis>
<bibRefCitation id="EFD0FC9D6657027D9B05FE28C5C4FE3F" author="Lecoq, J. C." box="[512,655,475,501]" pageId="5" pageNumber="258" pagination="579 - 603" refId="ref12275" refString="Lecoq, J. C. (2008) Revision du genre Paulianidia Jarrige, 1978 de Madagascar (Insecta, Coleoptera, Staphylinidae, Paederinae). Zoosystema, 30 (3), 579 - 603." type="journal article" year="2008">Lecoq, 2008</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C41FAEF6657027D9B9FFE28C4D6FE3F" authority="Lecoq, 2008" authorityName="Lecoq" authorityYear="2008" box="[666,925,475,501]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="B9355D7E6657027D9B9FFE28C441FE3E" box="[666,778,475,500]" italics="true" pageId="5" pageNumber="258">P. minima</emphasis>
<bibRefCitation id="EFD0FC9D6657027D9A14FE28C4D6FE3F" author="Lecoq, J. C." box="[785,925,475,501]" pageId="5" pageNumber="258" pagination="579 - 603" refId="ref12275" refString="Lecoq, J. C. (2008) Revision du genre Paulianidia Jarrige, 1978 de Madagascar (Insecta, Coleoptera, Staphylinidae, Paederinae). Zoosystema, 30 (3), 579 - 603." type="journal article" year="2008">Lecoq, 2008</bibRefCitation>
</taxonomicName>
and a species described below— 
<taxonomicName id="4C41FAEF6657027D9C17FE28C608FDD3" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="vohimana" status="sp. nov.">
<emphasis id="B9355D7E6657027D9C17FE28C2D7FE3F" box="[1298,1436,475,501]" italics="true" pageId="5" pageNumber="258">P. vohimana</emphasis>
Janák &amp; Baňař
</taxonomicName>
, 
<emphasis id="B9355D7E6657027D984AFE0CC6E1FDD3" bold="true" box="[335,426,511,537]" pageId="5" pageNumber="258">
<taxonomicNameLabel id="A206E0056657027D984AFE0CC6E1FDD3" box="[335,426,511,537]" pageId="5" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The aedeagus is similar to that of 
<taxonomicName id="4C41FAEF6657027D9A30FE0CC4C3FDD3" authorityName="Lecoq" authorityYear="2008" box="[821,904,511,537]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B9355D7E6657027D9A30FE0CC4C3FDD3" box="[821,904,511,537]" italics="true" pageId="5" pageNumber="258">P. hova</emphasis>
</taxonomicName>
, but the apical roof-shaped part of aedeagus is smaller and the species differs also externally by the head markedly narrowed behind eyes (cf. 
<figureCitation id="137A9DE96657027D9DC1FDD0C25FFDF7" box="[1220,1300,547,573]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="5" pageNumber="258">
Fig. 
<quantity id="4CB92C896657027D9DFDFDD0C279FDF7" box="[1272,1330,547,573]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" pageId="5" pageNumber="258" unit="in" value="29.0">29 in</quantity>
</figureCitation>
<taxonomicName id="4C41FAEF6657027D9C3CFDD0C2D6FDF7" authorityName="Janák &amp; Baňař" authorityYear="2024" box="[1337,1437,547,573]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B9355D7E6657027D9C3CFDD0C2D6FDF7" box="[1337,1437,547,573]" italics="true" pageId="5" pageNumber="258">P. tanala</emphasis>
</taxonomicName>
and 
<figureCitation id="137A9DE96657027D99C2FDB4C65CFDAB" box="[199,279,583,609]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="5" pageNumber="258">
Fig. 
<quantity id="4CB92C896657027D99FEFDB4C67EFDAB" box="[251,309,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" pageId="5" pageNumber="258" unit="in" value="28.0">28 in</quantity>
</figureCitation>
<taxonomicName id="4C41FAEF6657027D9839FDB4C6C5FDAB" authorityName="Lecoq" authorityYear="2008" box="[316,398,583,609]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B9355D7E6657027D9839FDB4C6C5FDAB" box="[316,398,583,609]" italics="true" pageId="5" pageNumber="258">P. hova</emphasis>
</taxonomicName>
) and by different shape of posterior margin of male sternite VII (cf. 
<figureCitation id="137A9DE96657027D9D91FDB4C3AFFDAB" box="[1172,1252,583,609]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1735,1759]" captionTargetBox="[173,1414,195,1698]" captionTargetId="figure-34@4.[160,1427,181,1711]" captionText="FIGURES 19–30. 19–27, 29–30: Paulianidia tanala sp. nov.: 19–23, 26–27, 29–30, holotype, 24–25, paratypes; 19, habitus; 20, forebody; 21–25, aedeagus (21, 24–25, ventral; 22, 1/3 lateral; 23, lateral); 26, male sternite VII; 27, male sternite VIII; 29, head; 30, left elytron. 28: P. hova Lecoq, paratype, head. Scales 1 mm: 19, 20; 0.5 mm: 21–25, 26–27, 28–30." figureDoi="http://doi.org/10.5281/zenodo.10854989" httpUri="https://zenodo.org/record/10854989/files/figure.png" pageId="5" pageNumber="258">
Fig. 
<quantity id="4CB92C896657027D9DCDFDB4C24AFDAB" box="[1224,1281,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="5" pageNumber="258" unit="in" value="26.0">26 in</quantity>
</figureCitation>
<taxonomicName id="4C41FAEF6657027D9C0CFDB4C226FDAB" authorityName="Janák &amp; Baňař" authorityYear="2024" box="[1289,1389,583,609]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B9355D7E6657027D9C0CFDB4C226FDAB" box="[1289,1389,583,609]" italics="true" pageId="5" pageNumber="258">P. tanala</emphasis>
</taxonomicName>
and 
<figureCitation id="137A9DE96657027D9992FD98C657FD4F" box="[151,284,619,645]" captionStart="FIGURES 1–18" captionStartId="2.[151,264,1696,1720]" captionTargetBox="[176,1412,191,1653]" captionTargetId="figure-34@2.[163,1424,181,1672]" captionTargetPageId="2" captionText="FIGURES 1–18. 1–8: Paulianidia densepunctata sp. nov., holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18: P. hova Lecoq, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3." figureDoi="http://doi.org/10.5281/zenodo.10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="5" pageNumber="258">
Figs. 
<quantity id="4CB92C896657027D99D0FD98C657FD4F" box="[213,284,619,645]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="5" pageNumber="258" unit="in" value="16.5" valueMax="17.0" valueMin="16.0">16–17</quantity>
</figureCitation>
in 
<taxonomicName id="4C41FAEF6657027D983AFD98C6DAFD4F" authorityName="Lecoq" authorityYear="2008" box="[319,401,619,645]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B9355D7E6657027D983AFD98C6DAFD4F" box="[319,401,619,645]" italics="true" pageId="5" pageNumber="258">P. hova</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76657027D99CBFD7CC35FFD63" box="[206,1044,655,681]" pageId="5" pageNumber="258" type="etymology">
<paragraph id="8BFE816C6657027D99CBFD7CC35FFD63" blockId="5.[151,1437,151,789]" box="[206,1044,655,681]" pageId="5" pageNumber="258">
<emphasis id="B9355D7E6657027D99CBFD7CC61EFD63" bold="true" box="[206,341,655,681]" pageId="5" pageNumber="258">Etymology.</emphasis>
This species is named after the Malagasy ethnic group Tanala.
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76657027D99C2FD41C472FCDF" pageId="5" pageNumber="258" type="distribution">
<paragraph id="8BFE816C6657027D99C2FD41C434FD3B" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="258">
<emphasis id="B9355D7E6657027D99C2FD41C62AFD06" bold="true" box="[199,353,690,716]" pageId="5" pageNumber="258">Distribution.</emphasis>
<taxonomicName id="4C41FAEF6657027D986FFD40C5B1FD07" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2024" box="[362,762,691,717]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="tanala" status="sp. nov.">
<emphasis id="B9355D7E6657027D986FFD40C50AFD07" box="[362,577,691,717]" italics="true" pageId="5" pageNumber="258">Paulianidia tanala</emphasis>
Janák &amp; Baňař
</taxonomicName>
, 
<emphasis id="B9355D7E6657027D9A0DFD40C42FFD07" bold="true" box="[776,868,691,717]" pageId="5" pageNumber="258">
<taxonomicNameLabel id="A206E0056657027D9A0DFD40C42FFD07" box="[776,868,691,717]" pageId="5" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently recorded only from the Ranomafana National Park in the central eastern part of 
<collectingCountry id="F356C1FC6657027D9B7CFD24C44BFD3B" box="[633,768,727,753]" name="Madagascar" pageId="5" pageNumber="258">Madagascar</collectingCountry>
(
<figureCitation id="137A9DE96657027D9A15FD24C43AFD3B" box="[784,881,727,753]" captionStart="FIGURES 116–122" captionStartId="12.[151,264,1780,1804]" captionTargetBox="[180,1408,194,1743]" captionTargetId="figure-34@12.[167,1421,181,1756]" captionText="FIGURES 116–122. 116–120: Paulianidia steineri Lecoq, Andasibe: 116–118, aedeagus (116, ventral; 117, 1/3 lateral; 118, lateral); 119, male sternite VII; 120, male sternite VIII. Scale 0.5 mm: 116–120. 121: Distribution of P. densepunctata sp. nov., (rhombus), P. valohoaka sp. nov. (star), P. tanala sp. nov. and P. vohimana sp. nov. (rectangle). 122: Distribution of Paulianidia species in Madagascar." figureDoi="http://doi.org/10.5281/zenodo.10854991" httpUri="https://zenodo.org/record/10854991/files/figure.png" pageId="5" pageNumber="258">Fig. 121</figureCitation>
).
</paragraph>
<paragraph id="8BFE816C6657027D99C2FD09C472FCDF" blockId="5.[151,1437,151,789]" box="[199,825,762,789]" pageId="5" pageNumber="258">
<emphasis id="B9355D7E6657027D99C2FD09C600FCDE" bold="true" box="[199,331,762,788]" pageId="5" pageNumber="258">Bionomics.</emphasis>
The 
<typeStatus id="54FA3FCE6657027D9881FD08C6FFFCDF" box="[388,436,763,789]" pageId="5" pageNumber="258">type</typeStatus>
series was sifted from forest litter.
</paragraph>
</subSubSection>
</treatment>
</document>