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<document id="1A267EA741447D02B991ACEE4A8BE8C9" ID-DOI="10.11646/zootaxa.3635.1.9" ID-GBIF-Dataset="c2e1b77b-2a89-46f8-8b24-64ae71254792" ID-ISSN="1175-5326" ID-Zenodo-Dep="283539" ID-ZooBank="3B9C28A8-B64D-46AB-BE68-BF6CA85DBCAF" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460585776088" checkinUser="plazi" docAuthor="Shear, William A." docDate="2013" docId="03CE87AC9901FFD4F0C316D9FF5E6720" docLanguage="en" docName="zt03635p093.pdf" docOrigin="Zootaxa 3635 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dybasia Loomis" docType="treatment" docVersion="6" lastPageNumber="92" masterDocId="FFF7FFD49904FFD1F054164EFFBE657A" masterDocTitle="A new miniature species of the milliped genus Cleidogona from Costa Rica (Diplopoda, Chordeumatida, Cleidogonidae), with a review of Central American Cleidogonidae" masterLastPageNumber="93" masterPageNumber="87" pageNumber="92" updateTime="1698325116444" updateUser="plazi">
<mods:mods id="990D8029485938605BE6C52F4F7AC66E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A0F576909EB01DCFB9CE1DA6058846D4">
<mods:title id="DF3AD187A9B32D4E31DC4A639A3BA3A7">A new miniature species of the milliped genus Cleidogona from Costa Rica (Diplopoda, Chordeumatida, Cleidogonidae), with a review of Central American Cleidogonidae</mods:title>
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<mods:name id="AE0700E92C617950AF3E106645433E50" type="personal">
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<mods:namePart id="0A2A6CAD4DAA7EAA6EB429073410BDF1">Shear, William A.</mods:namePart>
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<mods:typeOfResource id="1FE72C538856E2E8040418463F929F65">text</mods:typeOfResource>
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<mods:title id="56A64BA6016F0475E1B560351DD43DD8">Zootaxa</mods:title>
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<mods:part id="BC634138555F0835E1FF282D6FF08B1B">
<mods:date id="B6502A19B99FEB3984E7E757216AD53A">2013</mods:date>
<mods:detail id="A568AAA4F830B31AD034BE472B1D30BB" type="volume">
<mods:number id="FA2D82E3B8B24FC4AE157789622E551E">3635</mods:number>
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<mods:classification id="124ED7C69E7C4BF16A868D3E145294FE">journal article</mods:classification>
<mods:identifier id="3D66F32594299078D82988FF4CC30A41" type="DOI">10.11646/zootaxa.3635.1.9</mods:identifier>
<mods:identifier id="50CBFD04B9AAA8EA38E174C13B8FDE2C" type="GBIF-Dataset">c2e1b77b-2a89-46f8-8b24-64ae71254792</mods:identifier>
<mods:identifier id="E4466F3949EC2EC6DD5DE606225699CB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E0106230786151F7796C8594BF457A4E" type="Zenodo-Dep">283539</mods:identifier>
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<treatment id="03CE87AC9901FFD4F0C316D9FF5E6720" ID-DOI="http://doi.org/10.5281/zenodo.6164072" ID-GBIF-Taxon="127663643" ID-Zenodo-Dep="6164072" LSID="urn:lsid:plazi:treatment:03CE87AC9901FFD4F0C316D9FF5E6720" httpUri="http://treatment.plazi.org/id/03CE87AC9901FFD4F0C316D9FF5E6720" lastPageNumber="92" pageId="5" pageNumber="92">
<subSubSection id="C37D65319901FFD4F0C316D9FF5E6720" pageId="5" pageNumber="92" type="nomenclature">
<paragraph id="8BD836BA9901FFD4F0C316D9FE0D65CB" blockId="5.[151,435,151,177]" box="[151,435,151,177]" pageId="5" pageNumber="92">
<heading id="D09081D69901FFD4F0C316D9FE0D65CB" bold="true" box="[151,435,151,177]" fontSize="11" level="1" pageId="5" pageNumber="92" reason="1">
<emphasis id="B913EAA89901FFD4F0C316D9FE0D65CB" bold="true" box="[151,435,151,177]" pageId="5" pageNumber="92">
Genus
<taxonomicName id="4C674D399901FFD4F0BA16D9FE0D65CB" ID-CoL="7NXRJ" authority="Loomis" authorityName="Loomis" box="[238,435,151,177]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAA89901FFD4F0BA16D9FEEE65CB" bold="true" box="[238,336,151,177]" italics="true" pageId="5" pageNumber="92">Dybasia</emphasis>
Loomis
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD836BA9901FFD4F0C31691FE74658E" blockId="5.[151,458,222,245]" box="[151,458,222,245]" pageId="5" pageNumber="92">
<taxonomicName id="4C674D399901FFD4F0C31691FE3F658E" authority="Loomis, 1964" authorityName="Loomis" authorityYear="1964" box="[151,385,222,245]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAA89901FFD4F0C31691FF55658F" box="[151,235,223,245]" italics="true" pageId="5" pageNumber="92">Dybasia</emphasis>
Loomis, 1964
</taxonomicName>
, p. 100
</paragraph>
<paragraph id="8BD836BA9901FFD4F0C3176FFF5E6720" blockId="5.[151,1436,289,602]" pageId="5" pageNumber="92">
Loomis (1964) described three species of this Panamanian genus in three separate genera, which I synonymized under
<taxonomicName id="4C674D399901FFD4F08B170BFE826427" box="[223,316,325,349]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAA89901FFD4F08B170BFE826427" box="[223,316,325,349]" italics="true" pageId="5" pageNumber="92">Dybasia</emphasis>
</taxonomicName>
in 1972. He also established a separate family Dybasiidae for these obvious cleidogonids. Loomis extended the range of the genus into
<collectingCountry id="F370762A9901FFD4F26F1727FD0864FB" box="[571,694,361,385]" name="Costa Rica" pageId="5" pageNumber="92">Costa Rica</collectingCountry>
when he described
<taxonomicName id="4C674D399901FFD4F3CD1727FB1564FB" box="[921,1195,361,385]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="interamericana">
<emphasis id="B913EAA89901FFD4F3CD1727FB1564FB" box="[921,1195,361,385]" italics="true" pageId="5" pageNumber="92">Dybasia interamericana</emphasis>
</taxonomicName>
in 1974. The species of
<taxonomicName id="4C674D399901FFD4F0EF17C3FEA664DF" box="[187,280,397,421]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAA89901FFD4F0EF17C3FEA664DF" box="[187,280,397,421]" italics="true" pageId="5" pageNumber="92">Dybasia</emphasis>
</taxonomicName>
are small for cleidogonids, about
<quantity id="4C9F9B5F9901FFD4F2E317C3FC8B64DF" box="[695,821,397,421]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="5" pageNumber="92" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
in length. The gonopods have long anterior sternal extensions which parallel the angiocoxites, and very small colpocoxites. The ninth legs have a long median extension from the coxoprefemur. The Panamanian species have prominent segmental paranota, but these were not described by Loomis (1974) for
<emphasis id="B913EAA89901FFD4F27E17B4FCB6676B" box="[554,776,506,529]" italics="true" pageId="5" pageNumber="92">
<taxonomicName id="4C674D399901FFD4F27E17B4FCBA676B" box="[554,772,506,529]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="interamericana">D. interamericana</taxonomicName>
.
</emphasis>
Three of the four species are illustrated in my 1972 monograph, and
<taxonomicName id="4C674D399901FFD4F10C1450FD97674F" box="[344,553,542,565]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="interamericana">
<emphasis id="B913EAA89901FFD4F10C1450FD97674F" box="[344,553,542,565]" italics="true" pageId="5" pageNumber="92">D. interamericana</emphasis>
</taxonomicName>
by Loomis (1974).
<taxonomicName id="4C674D399901FFD4F3451453FC44674F" box="[785,1018,541,565]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="austrina">
<emphasis id="B913EAA89901FFD4F3451453FC44674F" box="[785,1018,541,565]" italics="true" pageId="5" pageNumber="92">Cleidogona austrina</emphasis>
</taxonomicName>
(see above) may also belong in this genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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