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<document id="5C023B06A1722B76B89052DB5B43AE5D" ID-CLB-Dataset="27578" ID-DOI="10.11646/zootaxa.4554.1.5" ID-GBIF-Dataset="f0417544-ffc1-44fa-881d-7a46fc70b421" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623441" ID-ZooBank="682FE7E7-26D7-4A12-8424-942836D0585E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554271018513" checkinUser="plazi" docAuthor="Hsueh, Pan-Wen" docDate="2019" docId="03CE8794FFCB454BFF18FDC0FE9EFBA1" docLanguage="en" docName="zootaxa.4554.1.5.pdf" docOrigin="Zootaxa 4554 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neanthes donggungensis Hsueh 2019, n. sp." docType="treatment" docVersion="8" lastPageNumber="179" masterDocId="FFF7FFECFFCF454DFF8FFFE4FF86FFB2" masterDocTitle="Neanthes (Annelida: Nereididae) from Taiwanese waters, with description of seven new species and one new species record" masterLastPageNumber="198" masterPageNumber="173" pageNumber="177" updateTime="1698716235899" updateUser="ExternalLinkService">
<mods:mods id="5BA38118D7EB742C594A7C4C8BDB0D8B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D2ADA1E23F7F52BE0C4F26E9D3488498">
<mods:title id="2091BAAB8A3CEB30B216B88A0E198579">Neanthes (Annelida: Nereididae) from Taiwanese waters, with description of seven new species and one new species record</mods:title>
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<mods:namePart id="072D57D0585E6B1A5CF20969950B304A">Hsueh, Pan-Wen</mods:namePart>
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<mods:title id="FE851DDC4A3932152DF0AFCCC21F98E8">Zootaxa</mods:title>
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<mods:date id="984A7C0DE578DD500EE312DDF12839C3">2019</mods:date>
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<mods:number id="B86994F89457ABE85299D714B66F9771">2019-02-08</mods:number>
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<mods:number id="E9009FB3919FD82476B0993A8235EFA6">4554</mods:number>
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<mods:identifier id="79BE7028B2E57046BF1807DE6B277483" type="CLB-Dataset">27578</mods:identifier>
<mods:identifier id="36EA5E1CE5FA7AC05004C851FB640EBB" type="DOI">10.11646/zootaxa.4554.1.5</mods:identifier>
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<treatment id="03CE8794FFCB454BFF18FDC0FE9EFBA1" ID-DOI="http://doi.org/10.5281/zenodo.5924663" ID-GBIF-Taxon="154534677" ID-Zenodo-Dep="5924663" LSID="urn:lsid:plazi:treatment:03CE8794FFCB454BFF18FDC0FE9EFBA1" httpUri="http://treatment.plazi.org/id/03CE8794FFCB454BFF18FDC0FE9EFBA1" lastPageId="6" lastPageNumber="179" pageId="4" pageNumber="177">
<subSubSection id="C37D6509FFCB4549FF18FDC0FD96FD8C" box="[151,528,548,574]" pageId="4" pageNumber="177" type="nomenclature">
<paragraph id="8BD83682FFCB4549FF18FDC0FD96FD8C" blockId="4.[151,528,548,607]" box="[151,528,548,574]" pageId="4" pageNumber="177">
<heading id="D09081EEFFCB4549FF18FDC0FD96FD8C" bold="true" box="[151,528,548,574]" fontSize="11" level="1" pageId="4" pageNumber="177" reason="1">
<emphasis id="B913EA90FFCB4549FF18FDC0FD96FD8C" bold="true" box="[151,528,548,574]" pageId="4" pageNumber="177">
<taxonomicName id="4C674D01FFCB4549FF18FDC0FE40FD8C" ID-CoL="45Y47" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[151,454,548,574]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="177" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFCB4549FF18FDC0FE40FD8C" bold="true" box="[151,454,548,574]" italics="true" pageId="4" pageNumber="177">Neanthes donggungensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057EBFFCB4549FE41FDC0FD96FD8C" box="[462,528,548,574]" pageId="4" pageNumber="177" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37D6509FFCB4549FF18FDA2FE39FDED" box="[151,447,582,607]" pageId="4" pageNumber="177" type="description">
<paragraph id="8BD83682FFCB4549FF18FDA2FE39FDED" blockId="4.[151,528,548,607]" box="[151,447,582,607]" pageId="4" pageNumber="177">
<figureCitation id="135C2A07FFCB4549FF18FDA2FE94FDED" box="[151,274,582,607]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1694]" captionTargetBox="[150,1436,807,1664]" captionTargetId="figure@4.[150,1436,804,1664]" captionTargetPageId="4" captionText="FIGURE 3. Neanthes donggungensis n. sp.; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm." httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="4" pageNumber="177">
Figs 3A
<emphasis id="B913EA90FFCB4549FF7FFDA2FF78FDED" bold="true" box="[240,254,582,607]" pageId="4" pageNumber="177">–</emphasis>
G
</figureCitation>
,
<figureCitation id="135C2A07FFCB4549FE92FDA3FEE6FDEC" box="[285,352,582,607]" captionStart="FIGURE 4" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[167,1431,874,1903]" captionTargetId="figure@5.[167,1431,872,1903]" captionTargetPageId="5" captionText="FIGURE 4. Neanthes donggungensis n. sp.; paratype (NMNS 7946-015): A, notochaeta, homogomph spiniger, chaetiger 150; B, neurochaeta of dorsal fascicle, homogomph spiniger, chaetiger 74; C, neurochaeta of dorsal fascicle, medium heterogomph falciger, chaetiger 74; D, neurochaeta of ventral fascicle, heterogomph spiniger, chaetiger 74; E, neurochaeta of ventral fascicle, medium heterogomph falciger, chaetiger 74. Scale: A–E, 0.05 mm." httpUri="https://zenodo.org/record/2623449/files/figure.png" pageId="4" pageNumber="177">
4A
<emphasis id="B913EA90FFCB4549FEB0FDA2FECBFDED" bold="true" box="[319,333,582,607]" pageId="4" pageNumber="177">–</emphasis>
E
</figureCitation>
,
<tableCitation id="C6E50339FFCB4549FEE3FDA2FE39FDED" box="[364,447,582,607]" captionStart="TABLE 2" captionStartId="23.[147,231,531,553]" captionText="TABLE 2. Key morphological characters of Neanthes species in the present study. Abbreviations: DCAL = dorsal cirri on dorsal ligule; DFN = neurochaetae of dorsal fascicle; HoS = homogomph spinigers; HeF = heterogomph falcigers; LTJ = number of lateral teeth on jaws; NeL = number of neuropodial lobes; NPL = notopodial prechaetal lobe; NeL = neuropodial lobes; P/Ch1 =length ratio of peristomium/chaetiger 1; RCLPTC = range of chaetigers reached by the longest posterior tentacular cirri; SeS = sesquigomph spinigers; SeF = sesquigomph falcigers; THeF = types of heterogomph falcigers; VFN = neurochaetae of ventral fascicle." pageId="4" pageNumber="177">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37D6509FFCB4549FF18FD6AFB0BFD5D" pageId="4" pageNumber="177" type="materials_examined">
<paragraph id="8BD83682FFCB4549FF18FD6AFB0BFD5D" blockId="4.[151,1436,654,751]" pageId="4" pageNumber="177">
<emphasis id="B913EA90FFCB4549FF18FD6AFE00FD15" bold="true" box="[151,390,654,679]" pageId="4" pageNumber="177">Material examined.</emphasis>
<materialsCitation id="3B0F3CDFFFCB4549FE1CFD6AFBB6FD79" ID-GBIF-Occurrence="2234227399" collectingDate="2006-02-22" collectionCode="NMNS" country="Taiwan" elevation="220" latitude="22.4665" location="Donggung Port" longLatPrecision="12" longitude="120.44234" pageId="4" pageNumber="177" specimenCode="NMNS 7946-014" specimenCount="1" stateProvince="Pingtung" typeStatus="holotype">
<typeStatus id="54DC8820FFCB4549FE1CFD6AFE7DFD15" box="[403,507,654,679]" pageId="4" pageNumber="177" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DBC19EF9FFCB4549FD80FD6BFD58FD15" box="[527,734,654,679]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="4" pageNumber="177">NMNS 7946-014</specimenCode>
), offshore from
<location id="8EB86059FFCB4549FC2BFD6BFBD0FD14" LSID="urn:lsid:plazi:treatment:03CE8794FFCB454BFF18FDC0FE9EFBA1:8EB86059FFCB4549FC2BFD6BFBD0FD14" box="[932,1110,655,679]" country="Taiwan" latitude="22.4665" longLatPrecision="12" longitude="120.44234" name="Donggung Port" pageId="4" pageNumber="177" stateProvince="Pingtung">Donggung Port</location>
(
<geoCoordinate id="EE535045FFCB4549FBE4FD6BFB76FD15" box="[1131,1264,655,679]" direction="north" orientation="latitude" pageId="4" pageNumber="177" precision="9" value="22.4665">22°27.99´N</geoCoordinate>
<geoCoordinate id="EE535045FFCB4549FB72FD6AFA09FD15" box="[1277,1423,654,679]" direction="east" orientation="longitude" pageId="4" pageNumber="177" precision="9" value="120.44234">120°26.54´E</geoCoordinate>
),
<collectingRegion id="49A3F860FFCB4549FF18FD56FEDDFD79" box="[151,347,690,715]" country="Taiwan" name="Pingtung" pageId="4" pageNumber="177">Pingtung County</collectingRegion>
,
<collectingCountry id="F3707612FFCB4549FEE5FD56FE39FD79" box="[362,447,690,715]" name="Taiwan" pageId="4" pageNumber="177">Taiwan</collectingCountry>
, muddy bottom,
<quantity id="4C9F9B67FFCB4549FD07FD57FC8DFD78" box="[648,779,690,715]" metricMagnitude="2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="4" pageNumber="177" unit="m" value="220.0" valueMax="240.0" valueMin="200.0">
<elevation id="004AD1B1FFCB4549FD07FD57FC8DFD78" box="[648,779,690,715]" metricMagnitude="2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="4" pageNumber="177" unit="m" value="220.0" valueMax="240.0" valueMin="200.0">
200
<emphasis id="B913EA90FFCB4549FD3DFD56FD46FD79" bold="true" box="[690,704,690,715]" pageId="4" pageNumber="177">–</emphasis>
240 m
</elevation>
</quantity>
deep,
<date id="FFD91042FFCB4549FCD2FD57FBB6FD79" box="[861,1072,690,715]" pageId="4" pageNumber="177" value="2006-02-22">
<collectingDate id="EF9DE9AAFFCB4549FCD2FD57FBB6FD79" box="[861,1072,690,715]" pageId="4" pageNumber="177" value="2006-02-22">22 February 2006</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B0F3CDFFFCB4549FBB0FD57FB0FFD5D" ID-GBIF-Occurrence="2234227577" collectingDate="2001-04-26" collectionCode="NMNS" country="Taiwan" elevation="250" pageId="4" pageNumber="177" specimenCode="NMNS 7946-015" specimenCount="1" stateProvince="Pingtung" typeStatus="paratype">
<typeStatus id="54DC8820FFCB4549FBB0FD57FB20FD79" box="[1087,1190,691,715]" pageId="4" pageNumber="177" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D61FD0BFFCB4549FB3BFD57FAB1FD79" box="[1204,1335,690,715]" pageId="4" pageNumber="177" type="generic">1 specimen</specimenCount>
(
<specimenCode id="DBC19EF9FFCB4549FAC6FD57FE81FD5D" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="4" pageNumber="177">NMNS 7946-015</specimenCode>
), location and habitat information same as
<typeStatus id="54DC8820FFCB4549FD67FD32FCCBFD5D" box="[744,845,726,751]" pageId="4" pageNumber="177" type="holotype">holotype</typeStatus>
,
<quantity id="4C9F9B67FFCB4549FCD7FD32FC19FD5C" box="[856,927,726,751]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="177" unit="m" value="250.0">
<elevation id="004AD1B1FFCB4549FCD7FD32FC19FD5C" box="[856,927,726,751]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="177" unit="m" value="250.0">250 m</elevation>
</quantity>
deep,
<date id="FFD91042FFCB4549FC67FD32FB0FFD5D" box="[1000,1161,726,751]" pageId="4" pageNumber="177" value="2001-04-26">
<collectingDate id="EF9DE9AAFFCB4549FC67FD32FB0FFD5D" box="[1000,1161,726,751]" pageId="4" pageNumber="177" value="2001-04-26">26 April 2001</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF18660AFFCB4549FF18F96CFE06F892" httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="4" pageNumber="177" startId="4.[151,250,1672,1694]" targetBox="[150,1436,807,1664]" targetPageId="4">
<paragraph id="8BD83682FFCB4549FF18F96CFE06F892" blockId="4.[151,1437,1672,1824]" pageId="4" pageNumber="177">
<emphasis id="B913EA90FFCB4549FF18F96CFE95F92F" bold="true" box="[151,275,1672,1694]" pageId="4" pageNumber="177">FIGURE 3.</emphasis>
<taxonomicName id="4C674D01FFCB4549FE94F96CFD9EF92C" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[283,536,1672,1694]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="177" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFCB4549FE94F96CFD9EF92C" box="[283,536,1672,1694]" italics="true" pageId="4" pageNumber="177">Neanthes donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFCB4549FD90F96CFDDFF92F" bold="true" box="[543,601,1672,1693]" pageId="4" pageNumber="177">
<taxonomicNameLabel id="A22057EBFFCB4549FD90F96CFDDFF92F" box="[543,601,1672,1693]" pageId="4" pageNumber="177" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C37D6509FFCB4548FF48F8A3FDD6FC8A" lastPageId="5" lastPageNumber="178" pageId="4" pageNumber="177" type="description">
<paragraph id="8BD83682FFCB4548FF48F8A3FBBEFEF2" blockId="4.[151,1437,1863,2032]" lastBlockId="5.[151,1437,151,824]" lastPageId="5" lastPageNumber="178" pageId="4" pageNumber="177">
<emphasis id="B913EA90FFCB4549FF48F8A3FEDFF8D2" bold="true" box="[199,345,1863,1888]" pageId="4" pageNumber="177">Description.</emphasis>
<typeStatus id="54DC8820FFCB4549FEEBF8A3FE49F8D2" box="[356,463,1863,1888]" pageId="4" pageNumber="177" type="holotype">Holotype</typeStatus>
, complete, ovigerous female, body length 72.0 mm (74.0) with 98 (96) chaetigers, maximum width 6.0 mm (8.0) at chaetiger 17, excluding parapodia; beige in alcohol (
<figureCitation id="135C2A07FFCB4549FBD2F88FFB3EF831" box="[1117,1208,1899,1924]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1694]" captionTargetBox="[150,1436,807,1664]" captionTargetId="figure@4.[150,1436,804,1664]" captionTargetPageId="4" captionText="FIGURE 3. Neanthes donggungensis n. sp.; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm." httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="4" pageNumber="177">Fig. 3A</figureCitation>
). Prostomium wider than long, one pair antennae arise antero-laterally, palpophores spherical and biarticulate with sub-conical palpostyles, four pairs of tentacular cirri, longest posterior tentacular cirri reaching chaetiger 6 (7) (
<figureCitation id="135C2A07FFCB4549FB73F857FAD3F879" box="[1276,1365,1971,1996]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1694]" captionTargetBox="[150,1436,807,1664]" captionTargetId="figure@4.[150,1436,804,1664]" captionTargetPageId="4" captionText="FIGURE 3. Neanthes donggungensis n. sp.; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm." httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="4" pageNumber="177">Fig. 3B</figureCitation>
). Two pairs of eyes, in trapezoidal arrangement, subequal in size. One apodous anterior segment present, about 1.4 times as long as chaetiger 1. Pharynx with brown jaws, each with 7–8 (8–9) blunt-tipped lateral teeth; brown conical paragnaths as: I=2 (2), in longitudinal line, anterior cone smaller; II=9 (8) (left), 11 (10) (right), in 2 oblique rows; III=12 (15), in 2–3 transverse rows; IV=15 (17) (left), 17 (right), in 2–3 oblique rows; V=0; VI=10 (7) (left), 13 (7) (right), in a cluster, VII/VIII=91 (76), with 8 (9) large cones in 1 row anteriorly, 83 (67) small and medium cones in 2–3 rows posteriorly (
<figureCitation id="135C2A07FFCA4548FE01FEC3FE60FEF2" box="[398,486,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1694]" captionTargetBox="[150,1436,807,1664]" captionTargetId="figure@4.[150,1436,804,1664]" captionTargetPageId="4" captionText="FIGURE 3. Neanthes donggungensis n. sp.; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm." httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="5" pageNumber="178">Fig. 3C</figureCitation>
). Paragnaths mostly delicate, not strongly chitinized.
</paragraph>
<paragraph id="8BD83682FFCA4548FF48FEA8FA0EFD36" blockId="5.[151,1437,151,824]" pageId="5" pageNumber="178">
Notopodial dorsal ligule triangular with a pointed tip on anterior chaetigers, wide triangular with a pointed tip on posterior chaetigers; dorsal ligule with base dorsal edge greatly expanded on posterior chaetigers (
<figureCitation id="135C2A07FFCA4548FB48FE8BFA83FE3A" box="[1223,1285,367,392]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1694]" captionTargetBox="[150,1436,807,1664]" captionTargetId="figure@4.[150,1436,804,1664]" captionTargetPageId="4" captionText="FIGURE 3. Neanthes donggungensis n. sp.; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm." httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="5" pageNumber="178">Fig. 3</figureCitation>
D–G). Dorsal cirri mid-dorsally attached to dorsal notopodial ligule, longer than dorsal ligule throughout (
<figureCitation id="135C2A07FFCA4548FB1DFE70FB54FE1E" box="[1170,1234,404,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1694]" captionTargetBox="[150,1436,807,1664]" captionTargetId="figure@4.[150,1436,804,1664]" captionTargetPageId="4" captionText="FIGURE 3. Neanthes donggungensis n. sp.; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm." httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="5" pageNumber="178">Fig. 3</figureCitation>
D–G). Notopodial prechaetal lobe present on chaetiger 3 to mid-body chaetigers, small triangular flap on anterior chaetigers, less developed on latter chaetigers (
<figureCitation id="135C2A07FFCA4548FE61FE38FDC0FE46" box="[494,582,476,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1694]" captionTargetBox="[150,1436,807,1664]" captionTargetId="figure@4.[150,1436,804,1664]" captionTargetPageId="4" captionText="FIGURE 3. Neanthes donggungensis n. sp.; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm." httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="5" pageNumber="178">Fig. 3D</figureCitation>
). Notopodial ventral ligule narrowed triangular with a pointed tip throughout. Neuropodium with prominent inferior lobe throughout, blunt-tipped. Neuropodial superior lobe present, inconspicuous on anterior chaetigers, slightly shorter than inferior lobe on posterior chaetigers. Neuropodial postchaetal lobe present throughout, triangular with a pointed tip. Neuropodial ventral ligule digitiform throughout; ventral cirri mid-ventrally attached to ventral edge of parapodia, shorter than ventral ligule throughout (
<figureCitation id="135C2A07FFCA4548FA8EFD88FAC6FD36" box="[1281,1344,620,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1694]" captionTargetBox="[150,1436,807,1664]" captionTargetId="figure@4.[150,1436,804,1664]" captionTargetPageId="4" captionText="FIGURE 3. Neanthes donggungensis n. sp.; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm." httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="5" pageNumber="178">Fig. 3</figureCitation>
D–G).
</paragraph>
<paragraph id="8BD83682FFCA4548FF48FD6BFDD6FC8A" blockId="5.[151,1437,151,824]" pageId="5" pageNumber="178">
Notochaetae present from chaetiger 3 to posterior end, homogomph spinigers (
<figureCitation id="135C2A07FFCA4548FBCAFD6BFB27FD1A" box="[1093,1185,655,680]" captionStart="FIGURE 4" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[167,1431,874,1903]" captionTargetId="figure@5.[167,1431,872,1903]" captionTargetPageId="5" captionText="FIGURE 4. Neanthes donggungensis n. sp.; paratype (NMNS 7946-015): A, notochaeta, homogomph spiniger, chaetiger 150; B, neurochaeta of dorsal fascicle, homogomph spiniger, chaetiger 74; C, neurochaeta of dorsal fascicle, medium heterogomph falciger, chaetiger 74; D, neurochaeta of ventral fascicle, heterogomph spiniger, chaetiger 74; E, neurochaeta of ventral fascicle, medium heterogomph falciger, chaetiger 74. Scale: A–E, 0.05 mm." httpUri="https://zenodo.org/record/2623449/files/figure.png" pageId="5" pageNumber="178">Fig. 4A</figureCitation>
). Neurochaetae dorsal fascicle: homogomph spinigers (
<figureCitation id="135C2A07FFCA4548FD89FD50FDD9FD7E" box="[518,607,692,717]" captionStart="FIGURE 4" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[167,1431,874,1903]" captionTargetId="figure@5.[167,1431,872,1903]" captionTargetPageId="5" captionText="FIGURE 4. Neanthes donggungensis n. sp.; paratype (NMNS 7946-015): A, notochaeta, homogomph spiniger, chaetiger 150; B, neurochaeta of dorsal fascicle, homogomph spiniger, chaetiger 74; C, neurochaeta of dorsal fascicle, medium heterogomph falciger, chaetiger 74; D, neurochaeta of ventral fascicle, heterogomph spiniger, chaetiger 74; E, neurochaeta of ventral fascicle, medium heterogomph falciger, chaetiger 74. Scale: A–E, 0.05 mm." httpUri="https://zenodo.org/record/2623449/files/figure.png" pageId="5" pageNumber="178">Fig. 4B</figureCitation>
), and heterogomph falcigers with medium-length, serrated terminal blade (
<figureCitation id="135C2A07FFCA4548FF10FD33FF79FD42" box="[159,255,727,752]" captionStart="FIGURE 4" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[167,1431,874,1903]" captionTargetId="figure@5.[167,1431,872,1903]" captionTargetPageId="5" captionText="FIGURE 4. Neanthes donggungensis n. sp.; paratype (NMNS 7946-015): A, notochaeta, homogomph spiniger, chaetiger 150; B, neurochaeta of dorsal fascicle, homogomph spiniger, chaetiger 74; C, neurochaeta of dorsal fascicle, medium heterogomph falciger, chaetiger 74; D, neurochaeta of ventral fascicle, heterogomph spiniger, chaetiger 74; E, neurochaeta of ventral fascicle, medium heterogomph falciger, chaetiger 74. Scale: A–E, 0.05 mm." httpUri="https://zenodo.org/record/2623449/files/figure.png" pageId="5" pageNumber="178">Fig. 4C</figureCitation>
). Neurochaetae ventral fascicle: heterogomph spinigers (
<figureCitation id="135C2A07FFCA4548FC28FD33FB8EFD42" box="[935,1032,727,752]" captionStart="FIGURE 4" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[167,1431,874,1903]" captionTargetId="figure@5.[167,1431,872,1903]" captionTargetPageId="5" captionText="FIGURE 4. Neanthes donggungensis n. sp.; paratype (NMNS 7946-015): A, notochaeta, homogomph spiniger, chaetiger 150; B, neurochaeta of dorsal fascicle, homogomph spiniger, chaetiger 74; C, neurochaeta of dorsal fascicle, medium heterogomph falciger, chaetiger 74; D, neurochaeta of ventral fascicle, heterogomph spiniger, chaetiger 74; E, neurochaeta of ventral fascicle, medium heterogomph falciger, chaetiger 74. Scale: A–E, 0.05 mm." httpUri="https://zenodo.org/record/2623449/files/figure.png" pageId="5" pageNumber="178">Fig. 4D</figureCitation>
), and heterogomph falcigers with medium-length, serrated terminal blade (
<figureCitation id="135C2A07FFCA4548FDFFFD18FD4AFCA6" box="[624,716,764,789]" captionStart="FIGURE 4" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[167,1431,874,1903]" captionTargetId="figure@5.[167,1431,872,1903]" captionTargetPageId="5" captionText="FIGURE 4. Neanthes donggungensis n. sp.; paratype (NMNS 7946-015): A, notochaeta, homogomph spiniger, chaetiger 150; B, neurochaeta of dorsal fascicle, homogomph spiniger, chaetiger 74; C, neurochaeta of dorsal fascicle, medium heterogomph falciger, chaetiger 74; D, neurochaeta of ventral fascicle, heterogomph spiniger, chaetiger 74; E, neurochaeta of ventral fascicle, medium heterogomph falciger, chaetiger 74. Scale: A–E, 0.05 mm." httpUri="https://zenodo.org/record/2623449/files/figure.png" pageId="5" pageNumber="178">Fig. 4E</figureCitation>
). Pygidium round, one pair of filamentous anal cirri attached ventrally, extending about 9 chaetigers.
</paragraph>
</subSubSection>
<caption id="DF18660AFFCA4548FF18F89FFC1EF85F" httpUri="https://zenodo.org/record/2623449/files/figure.png" pageId="5" pageNumber="178" startId="5.[151,250,1915,1937]" targetBox="[167,1431,874,1903]" targetPageId="5">
<paragraph id="8BD83682FFCA4548FF18F89FFC1EF85F" blockId="5.[151,1436,1915,2029]" pageId="5" pageNumber="178">
<emphasis id="B913EA90FFCA4548FF18F89FFE94F822" bold="true" box="[151,274,1915,1937]" pageId="5" pageNumber="178">FIGURE 4.</emphasis>
<taxonomicName id="4C674D01FFCA4548FE96F89FFD93F823" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[281,533,1915,1937]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="178" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFCA4548FE96F89FFD93F823" box="[281,533,1915,1937]" italics="true" pageId="5" pageNumber="178">Neanthes donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFCA4548FD93F898FDD3F823" bold="true" box="[540,597,1916,1937]" pageId="5" pageNumber="178">
<taxonomicNameLabel id="A22057EBFFCA4548FD93F898FDD3F823" box="[540,597,1916,1937]" pageId="5" pageNumber="178" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; paratype (NMNS 7946-015): A, notochaeta, homogomph spiniger, chaetiger 150; B, neurochaeta of dorsal fascicle, homogomph spiniger, chaetiger 74; C, neurochaeta of dorsal fascicle, medium heterogomph falciger, chaetiger 74; D, neurochaeta of ventral fascicle, heterogomph spiniger, chaetiger 74; E, neurochaeta of ventral fascicle, medium heterogomph falciger, chaetiger 74. Scale: A–E, 0.05 mm.
</paragraph>
</caption>
<subSubSection id="C37D6509FFC9454BFF48FF73FE82FF67" pageId="6" pageNumber="179" type="etymology">
<paragraph id="8BD83682FFC9454BFF48FF73FE82FF67" blockId="6.[151,1437,151,1043]" pageId="6" pageNumber="179">
<emphasis id="B913EA90FFC9454BFF48FF73FEC8FF02" bold="true" box="[199,334,151,176]" pageId="6" pageNumber="179">Etymology.</emphasis>
The name is derived from the name of nearby township, Donggung, where the worm was collected.
</paragraph>
</subSubSection>
<subSubSection id="C37D6509FFC9454BFF48FF3BFC74FF4A" box="[199,1010,223,248]" pageId="6" pageNumber="179" type="materials_examined">
<paragraph id="8BD83682FFC9454BFF48FF3BFC74FF4A" blockId="6.[151,1437,151,1043]" box="[199,1010,223,248]" pageId="6" pageNumber="179">
<materialsCitation id="3B0F3CDFFFC9454BFF48FF3BFC68FF4A" ID-GBIF-Occurrence="2234227531" box="[199,1006,223,248]" country="Taiwan" location="Offshore" municipality="Donggung Port" pageId="6" pageNumber="179" specimenCount="1" stateProvince="Pingtung" typeStatus="holotype">
<emphasis id="B913EA90FFC9454BFF48FF3BFEE3FF4A" bold="true" box="[199,357,223,248]" pageId="6" pageNumber="179">
<typeStatus id="54DC8820FFC9454BFF48FF3BFE84FF4A" box="[199,258,223,248]" pageId="6" pageNumber="179">Type</typeStatus>
locality.
</emphasis>
<location id="8EB86059FFC9454BFEE3FF3BFE49FF4A" LSID="urn:lsid:plazi:treatment:03CE8794FFCB454BFF18FDC0FE9EFBA1:8EB86059FFC9454BFEE3FF3BFE49FF4A" box="[364,463,223,248]" country="Taiwan" municipality="Donggung Port" name="Offshore" pageId="6" pageNumber="179" stateProvince="Pingtung">Offshore</location>
from
<collectingMunicipality id="6BBCACF8FFC9454BFD9CFF04FD44FF45" box="[531,706,224,248]" pageId="6" pageNumber="179">Donggung Port</collectingMunicipality>
,
<collectingRegion id="49A3F860FFC9454BFD42FF3BFC0BFF4A" box="[717,909,223,248]" country="Taiwan" name="Pingtung" pageId="6" pageNumber="179">Pingtung County</collectingRegion>
,
<collectingCountry id="F3707612FFC9454BFC17FF3BFC68FF4A" box="[920,1006,223,248]" name="Taiwan" pageId="6" pageNumber="179">Taiwan</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D6509FFC9454BFF48FEE0FD77FEAF" box="[199,753,260,285]" pageId="6" pageNumber="179" type="distribution">
<paragraph id="8BD83682FFC9454BFF48FEE0FD77FEAF" blockId="6.[151,1437,151,1043]" box="[199,753,260,285]" pageId="6" pageNumber="179">
<emphasis id="B913EA90FFC9454BFF48FEE0FEE6FEAF" bold="true" box="[199,352,260,285]" pageId="6" pageNumber="179">Distribution.</emphasis>
Only known from the
<typeStatus id="54DC8820FFC9454BFDEFFEE1FD16FEAF" box="[608,656,261,285]" pageId="6" pageNumber="179">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C37D6509FFC9454BFF48FEC3FE9EFBA1" pageId="6" pageNumber="179" type="discussion">
<paragraph id="8BD83682FFC9454BFF48FEC3FE9EFBA1" blockId="6.[151,1437,151,1043]" pageId="6" pageNumber="179">
<emphasis id="B913EA90FFC9454BFF48FEC3FEBDFEF2" bold="true" box="[199,315,295,320]" pageId="6" pageNumber="179">Remarks.</emphasis>
The most noticeable morphological characters of
<taxonomicName id="4C674D01FFC9454BFCE2FECDFBB0FEF2" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[877,1078,296,320]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFCE2FECDFC06FEF2" box="[877,896,297,320]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFC01FECCFBB0FEF2" box="[910,1078,296,320]" italics="true" pageId="6" pageNumber="179">donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFBB2FEC3FBFAFEF2" bold="true" box="[1085,1148,295,320]" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFBB2FEC3FBFAFEF2" box="[1085,1148,295,320]" pageId="6" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are the numerous conical paragnaths (76–91) on Area VII/VIII of the pharynx (
<tableCitation id="C6E50339FFC9454BFD79FEA8FCCBFED6" box="[758,845,332,357]" captionStart="TABLE 2" captionStartId="23.[147,231,531,553]" captionText="TABLE 2. Key morphological characters of Neanthes species in the present study. Abbreviations: DCAL = dorsal cirri on dorsal ligule; DFN = neurochaetae of dorsal fascicle; HoS = homogomph spinigers; HeF = heterogomph falcigers; LTJ = number of lateral teeth on jaws; NeL = number of neuropodial lobes; NPL = notopodial prechaetal lobe; NeL = neuropodial lobes; P/Ch1 =length ratio of peristomium/chaetiger 1; RCLPTC = range of chaetigers reached by the longest posterior tentacular cirri; SeS = sesquigomph spinigers; SeF = sesquigomph falcigers; THeF = types of heterogomph falcigers; VFN = neurochaetae of ventral fascicle." pageId="6" pageNumber="179">Table 2</tableCitation>
) and the greatly expanded dorsal ligule on posterior chaetigers (
<figureCitation id="135C2A07FFC9454BFE94FE8BFE11FE3A" box="[283,407,367,392]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1694]" captionTargetBox="[150,1436,807,1664]" captionTargetId="figure@4.[150,1436,804,1664]" captionTargetPageId="4" captionText="FIGURE 3. Neanthes donggungensis n. sp.; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm." httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="6" pageNumber="179">Fig. 3E, F</figureCitation>
). Of the 22
<taxonomicName id="4C674D01FFC9454BFDA6FE94FD17FE3A" authorityName="Kinberg" authorityYear="1865" box="[553,657,368,392]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="genus">
<emphasis id="B913EA90FFC9454BFDA6FE94FD17FE3A" box="[553,657,368,392]" italics="true" pageId="6" pageNumber="179">Neanthes</emphasis>
</taxonomicName>
species reported from East and Southeastern Asia, three also have numerous conical paragnaths on Area VII/VIII of the pharynx:
<taxonomicName id="4C674D01FFC9454BFCEDFE71FB24FE1F" authority="(Ehlers, 1868)" authorityName="Ehlers" authorityYear="1868" baseAuthorityName="Ehlers" baseAuthorityYear="1868" box="[866,1186,404,429]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="acuminata">
<emphasis id="B913EA90FFC9454BFCEDFE71FCF3FE1E" box="[866,885,405,428]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFC0CFE71FC7FFE1E" box="[899,1017,405,428]" italics="true" pageId="6" pageNumber="179">acuminata</emphasis>
(
<bibRefCitation id="EFF64B73FFC9454BFB86FE70FB1FFE1F" author="Ehlers, E." box="[1033,1177,404,429]" pageId="6" pageNumber="179" refId="ref12363" refString="Ehlers, E. (1868) Die Borstenwurmer (Annelida Chaetopoda) nach systematischen und anatomischen Untersuchungen dargestellt. Vol. 2. Wilhelm Engelmann, Leipzig, pp. 269 - 748, pls. XII-XXIV." type="book" year="1868">Ehlers, 1868</bibRefCitation>
)
</taxonomicName>
(
<typeStatus id="54DC8820FFC9454BFB3EFE70FB65FE1E" box="[1201,1251,404,428]" pageId="6" pageNumber="179">type</typeStatus>
locality in
<collectingCountry id="F3707612FFC9454BFAEBFE70FA1EFE1F" box="[1380,1432,404,429]" name="Italy" pageId="6" pageNumber="179">Italy</collectingCountry>
, reported from
<collectingCountry id="F3707612FFC9454BFEB4FE53FE07FE62" box="[315,385,439,464]" name="China" pageId="6" pageNumber="179">China</collectingCountry>
,
<collectingCountry id="F3707612FFC9454BFE01FE5CFE48FE7D" box="[398,462,440,463]" name="Japan" pageId="6" pageNumber="179">Japan</collectingCountry>
&amp;
<collectingCountry id="F3707612FFC9454BFE7BFE53FDF2FE62" box="[500,628,439,464]" name="Philippines" pageId="6" pageNumber="179">Philippines</collectingCountry>
) (
<emphasis id="B913EA90FFC9454BFD02FE5DFD1AFE62" box="[653,668,441,464]" italics="true" pageId="6" pageNumber="179">=</emphasis>
<taxonomicName id="4C674D01FFC9454BFD12FE5DFB96FE62" authority="(Delle Chiaje, 1828)" authorityName="Delle Chiaje" authorityYear="1828" baseAuthorityName="Delle Chiaje" baseAuthorityYear="1828" box="[669,1040,439,464]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="caudata">
<emphasis id="B913EA90FFC9454BFD12FE5DFD35FE62" box="[669,691,441,464]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFD4DFE5CFC9AFE62" box="[706,796,440,464]" italics="true" pageId="6" pageNumber="179">caudata</emphasis>
(Delle Chiaje, 1828)
</taxonomicName>
, from
<collectingCountry id="F3707612FFC9454BFBD3FE5CFB26FE62" box="[1116,1184,440,464]" name="Japan" pageId="6" pageNumber="179">Japan</collectingCountry>
),
<taxonomicName id="4C674D01FFC9454BFB3BFE5DFF5EFE46" authority="(Grube, 1878)" authorityName="Grube" authorityYear="1878" baseAuthorityName="Grube" baseAuthorityYear="1878" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="crucifera">
<emphasis id="B913EA90FFC9454BFB3BFE5DFB41FE62" box="[1204,1223,441,464]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFB59FE5CFABBFE62" box="[1238,1341,440,464]" italics="true" pageId="6" pageNumber="179">crucifera</emphasis>
(
<bibRefCitation id="EFF64B73FFC9454BFAC2FE53FF56FE46" author="Grube, A. E." pageId="6" pageNumber="179" pagination="1 - 300" refId="ref12574" refString="Grube, A. E. (1878) Annulata Semperiana. Beitrage zur Kenntniss der Annelidenfauna der Philippinen. Memoires de L'Academie Imperiale des Sciences de St. Petersbourg, Serie 7, 25 (8), 1 - 300." type="journal article" year="1878">Grube, 1878</bibRefCitation>
)
</taxonomicName>
(from
<collectingCountry id="F3707612FFC9454BFEA3FE38FE2AFE47" box="[300,428,476,501]" name="Philippines" pageId="6" pageNumber="179">Philippines</collectingCountry>
), and
<taxonomicName id="4C674D01FFC9454BFE78FE39FC9BFE46" authority="(Grube, 1878)" authorityName="Grube" authorityYear="1878" baseAuthorityName="Grube" baseAuthorityYear="1878" box="[503,797,476,501]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="verrillii">
<emphasis id="B913EA90FFC9454BFE78FE39FD8CFE46" box="[503,522,477,500]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFD94FE38FDF6FE46" box="[539,624,476,500]" italics="true" pageId="6" pageNumber="179">verrillii</emphasis>
(
<bibRefCitation id="EFF64B73FFC9454BFD0CFE38FC93FE46" author="Grube, A. E." box="[643,789,476,501]" pageId="6" pageNumber="179" pagination="1 - 300" refId="ref12574" refString="Grube, A. E. (1878) Annulata Semperiana. Beitrage zur Kenntniss der Annelidenfauna der Philippinen. Memoires de L'Academie Imperiale des Sciences de St. Petersbourg, Serie 7, 25 (8), 1 - 300." type="journal article" year="1878">Grube, 1878</bibRefCitation>
)
</taxonomicName>
(from
<collectingCountry id="F3707612FFC9454BFCFEFE38FC77FE47" box="[881,1009,476,501]" name="Philippines" pageId="6" pageNumber="179">Philippines</collectingCountry>
) (
<bibRefCitation id="EFF64B73FFC9454BFB85FE38FB1FFE47" author="Ehlers, E." box="[1034,1177,476,501]" pageId="6" pageNumber="179" refId="ref12363" refString="Ehlers, E. (1868) Die Borstenwurmer (Annelida Chaetopoda) nach systematischen und anatomischen Untersuchungen dargestellt. Vol. 2. Wilhelm Engelmann, Leipzig, pp. 269 - 748, pls. XII-XXIV." type="book" year="1868">Ehlers 1868</bibRefCitation>
: 552, pl. XXII fig 24;
<bibRefCitation id="EFF64B73FFC9454BFF18FDE6FEA4FDA9" author="Grube, A. E." box="[151,290,514,539]" pageId="6" pageNumber="179" pagination="1 - 300" refId="ref12574" refString="Grube, A. E. (1878) Annulata Semperiana. Beitrage zur Kenntniss der Annelidenfauna der Philippinen. Memoires de L'Academie Imperiale des Sciences de St. Petersbourg, Serie 7, 25 (8), 1 - 300." type="journal article" year="1878">Grube 1878</bibRefCitation>
: 79, 81, pl. V, figs 2, 2
<superScript id="7C129BCAFFC9454BFDBDFE1BFDBFFDBF" attach="none" box="[562,569,511,525]" fontSize="6" pageId="6" pageNumber="179">a</superScript>
, 6;
<bibRefCitation id="EFF64B73FFC9454BFDE6FDE6FC8AFDA9" author="Imajima, M." box="[617,780,514,539]" pageId="6" pageNumber="179" pagination="37 - 153" refId="ref12668" refString="Imajima, M. (1972) Review of the annelid worms of the family Nereidae of Japan, with descriptions of five new species or subspecies. Bulletin of the National Science Museum, Tokyo, 15 (1), 37 - 153." type="journal article" year="1972">Imajima 1972</bibRefCitation>
: 105–107, fig. 31b–e;
<bibRefCitation id="EFF64B73FFC9454BFB9DFDE7FB71FDA9" author="Sun, R. &amp; Yang, D." box="[1042,1271,515,539]" pageId="6" pageNumber="179" pagination="1 - 520" refId="ref13285" refString="Sun, R. &amp; Yang, D. (2004) Annelida. Polychaeta II. Nereidida (= Nereimorpha). Nereididae, Syllidae, Hesionidae, Pilargidae, Nephtyidae. In: Huo, C. &amp; Zhao, G. (Eds.), Fauna Sinica, Invertebrata. Science Press, Beijing, pp. 1 - 520 pp. [in Chinese with English abstract]" type="book chapter" year="2004">Sun &amp; Young 2004</bibRefCitation>
). However,
<taxonomicName id="4C674D01FFC9454BFA0DFDE7FE8BFD8D" baseAuthorityName="Ehlers" baseAuthorityYear="1868" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="acuminata">
<emphasis id="B913EA90FFC9454BFA0DFDE7FA13FDA8" box="[1410,1429,515,538]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFF18FDCCFE8BFD8D" box="[151,269,552,575]" italics="true" pageId="6" pageNumber="179">acuminata</emphasis>
</taxonomicName>
has a greater number of conical paragnaths on Area I of the pharynx than that of
<taxonomicName id="4C674D01FFC9454BFB3CFDCCFAFBFD8D" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[1203,1405,551,575]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFB3CFDCCFB40FD8D" box="[1203,1222,552,575]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFB5AFDC3FAFBFD8D" box="[1237,1405,551,575]" italics="true" pageId="6" pageNumber="179">donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFA0AFDC2FF3EFDD1" bold="true" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFA0AFDC2FF3EFDD1" pageId="6" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(10 versus 3) (
<bibRefCitation id="EFF64B73FFC9454BFEFFFDAEFD22FDD1" author="Ehlers, E." box="[368,676,586,611]" pageId="6" pageNumber="179" refId="ref12363" refString="Ehlers, E. (1868) Die Borstenwurmer (Annelida Chaetopoda) nach systematischen und anatomischen Untersuchungen dargestellt. Vol. 2. Wilhelm Engelmann, Leipzig, pp. 269 - 748, pls. XII-XXIV." type="book" year="1868">
Ehlers 1868: 552;
<tableCitation id="C6E50339FFC9454BFDC4FDAEFD22FDD1" box="[587,676,586,611]" captionStart="TABLE 2" captionStartId="23.[147,231,531,553]" captionText="TABLE 2. Key morphological characters of Neanthes species in the present study. Abbreviations: DCAL = dorsal cirri on dorsal ligule; DFN = neurochaetae of dorsal fascicle; HoS = homogomph spinigers; HeF = heterogomph falcigers; LTJ = number of lateral teeth on jaws; NeL = number of neuropodial lobes; NPL = notopodial prechaetal lobe; NeL = neuropodial lobes; P/Ch1 =length ratio of peristomium/chaetiger 1; RCLPTC = range of chaetigers reached by the longest posterior tentacular cirri; SeS = sesquigomph spinigers; SeF = sesquigomph falcigers; THeF = types of heterogomph falcigers; VFN = neurochaetae of ventral fascicle." pageId="6" pageNumber="179">Table 2</tableCitation>
</bibRefCitation>
). Conical paragnaths on Areas V-VIII of
<taxonomicName id="4C674D01FFC9454BFB01FDAFFAAFFDD0" baseAuthorityName="Ehlers" baseAuthorityYear="1868" box="[1166,1321,587,610]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="acuminata">
<emphasis id="B913EA90FFC9454BFB01FDAFFB27FDD0" box="[1166,1185,587,610]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFB3CFDAFFAAFFDD0" box="[1203,1321,587,610]" italics="true" pageId="6" pageNumber="179">acuminata</emphasis>
</taxonomicName>
formed a continuous broad band, which is not observed in
<taxonomicName id="4C674D01FFC9454BFD41FD94FC1FFD35" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[718,921,623,647]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFD41FD94FD67FD35" box="[718,737,624,647]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFD7EFD8BFC1FFD35" box="[753,921,623,647]" italics="true" pageId="6" pageNumber="179">donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFC2DFD8AFC65FD35" bold="true" box="[930,995,622,647]" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFC2DFD8AFC65FD35" box="[930,995,622,647]" pageId="6" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<bibRefCitation id="EFF64B73FFC9454BFC7BFD8AFAA7FD35" author="Ehlers, E." box="[1012,1313,622,647]" pageId="6" pageNumber="179" refId="ref12363" refString="Ehlers, E. (1868) Die Borstenwurmer (Annelida Chaetopoda) nach systematischen und anatomischen Untersuchungen dargestellt. Vol. 2. Wilhelm Engelmann, Leipzig, pp. 269 - 748, pls. XII-XXIV." type="book" year="1868">
Ehlers 1868: 552;
<figureCitation id="135C2A07FFC9454BFB44FD8AFAA7FD35" box="[1227,1313,622,647]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1694]" captionTargetBox="[150,1436,807,1664]" captionTargetId="figure@4.[150,1436,804,1664]" captionTargetPageId="4" captionText="FIGURE 3. Neanthes donggungensis n. sp.; A–C, holotype (NMNS 7946-014), D–G paratype (NMNS 7946-015): A, whole animal; B, anterior body; C, paragnaths of the pharynx; D, right parapodium, anterior view, chaetiger 10; E, right parapodium, posterior view, chaetiger 10; F, right parapodium, anterior view, chaetiger 74; G, right parapodium, posterior view, chaetiger 74. The Roman numeral represents the pharyngeal area VI. Abbreviation: NoPL, notopodial prechaetal lobe. Scale: A, 10.0 mm; B, 1.0 mm; C–G, 0.5 mm." httpUri="https://zenodo.org/record/2623447/files/figure.png" pageId="6" pageNumber="179">Fig. 3C</figureCitation>
</bibRefCitation>
)
<taxonomicName id="4C674D01FFC9454BFABBFD8BFF78FD19" baseAuthorityName="Grube" baseAuthorityYear="1878" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="crucifera">
<emphasis id="B913EA90FFC9454BFABBFD8BFF78FD19" italics="true" pageId="6" pageNumber="179">Neanthes crucifera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D01FFC9454BFEBAFD77FE2AFD19" baseAuthorityName="Grube" baseAuthorityYear="1878" box="[309,428,659,683]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="verrillii">
<emphasis id="B913EA90FFC9454BFEBAFD77FECEFD18" box="[309,328,659,682]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFED8FD77FE2AFD19" box="[343,428,659,683]" italics="true" pageId="6" pageNumber="179">verrillii</emphasis>
</taxonomicName>
have similar numbers of conical paragnaths on Area I as in
<taxonomicName id="4C674D01FFC9454BFBD7FD77FAA4FD19" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[1112,1314,659,683]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFBD7FD77FBEDFD18" box="[1112,1131,659,682]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFBF5FD77FAA4FD19" box="[1146,1314,659,683]" italics="true" pageId="6" pageNumber="179">donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFAA6FD76FAEFFD19" bold="true" box="[1321,1385,658,683]" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFAA6FD76FAEFFD19" box="[1321,1385,658,683]" pageId="6" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but both have several conical paragnaths on Area V (6 &amp; 8–9, respectively) in comparison to none in
<taxonomicName id="4C674D01FFC9454BFB5BFD5CFA1AFD7D" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[1236,1436,695,719]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFB5BFD5CFB61FD7D" box="[1236,1255,696,719]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFB7BFD53FA1AFD7D" box="[1268,1436,695,719]" italics="true" pageId="6" pageNumber="179">donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFF18FD3EFF74FD41" bold="true" box="[151,242,730,755]" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFF18FD3EFF74FD41" box="[151,242,730,755]" pageId="6" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<bibRefCitation id="EFF64B73FFC9454BFE8DFD3EFD7BFD41" author="Grube, A. E." box="[258,765,730,755]" pageId="6" pageNumber="179" pagination="1 - 300" refId="ref12574" refString="Grube, A. E. (1878) Annulata Semperiana. Beitrage zur Kenntniss der Annelidenfauna der Philippinen. Memoires de L'Academie Imperiale des Sciences de St. Petersbourg, Serie 7, 25 (8), 1 - 300." type="journal article" year="1878">
Grube 1878: 78, 81, pl. V, figs 2, 5;
<tableCitation id="C6E50339FFC9454BFD2AFD3EFD7BFD41" box="[677,765,730,755]" captionStart="TABLE 2" captionStartId="23.[147,231,531,553]" captionText="TABLE 2. Key morphological characters of Neanthes species in the present study. Abbreviations: DCAL = dorsal cirri on dorsal ligule; DFN = neurochaetae of dorsal fascicle; HoS = homogomph spinigers; HeF = heterogomph falcigers; LTJ = number of lateral teeth on jaws; NeL = number of neuropodial lobes; NPL = notopodial prechaetal lobe; NeL = neuropodial lobes; P/Ch1 =length ratio of peristomium/chaetiger 1; RCLPTC = range of chaetigers reached by the longest posterior tentacular cirri; SeS = sesquigomph spinigers; SeF = sesquigomph falcigers; THeF = types of heterogomph falcigers; VFN = neurochaetae of ventral fascicle." pageId="6" pageNumber="179">Table 2</tableCitation>
</bibRefCitation>
). Of the nine species reported in the present study, only
<taxonomicName id="4C674D01FFC9454BFA0DFD3FFEB9FCA5" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFA0DFD3FFA13FD40" box="[1410,1429,731,754]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFF18FD1BFEB9FCA5" box="[151,319,767,791]" italics="true" pageId="6" pageNumber="179">donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFEC8FD1AFE00FCA5" bold="true" box="[327,390,766,791]" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFEC8FD1AFE00FCA5" box="[327,390,766,791]" pageId="6" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C674D01FFC9454BFE19FCE4FDC7FCA5" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[406,577,767,791]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="kaomeiensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFE19FCE4FE2FFCA5" box="[406,425,768,791]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFE37FD1BFDC7FCA5" box="[440,577,767,791]" italics="true" pageId="6" pageNumber="179">kaomeiensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFDC6FD1AFD0EFCA5" bold="true" box="[585,648,766,791]" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFDC6FD1AFD0EFCA5" box="[585,648,766,791]" pageId="6" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C674D01FFC9454BFD4EFCE4FC08FCA5" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[705,910,766,791]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="nanhaiensis">
<emphasis id="B913EA90FFC9454BFD4EFCE4FD52FCA5" box="[705,724,768,791]" italics="true" pageId="6" pageNumber="179">N</emphasis>
. cf.
<emphasis id="B913EA90FFC9454BFC87FD1BFC08FCA5" box="[776,910,767,791]" italics="true" pageId="6" pageNumber="179">nanhaiensis</emphasis>
</taxonomicName>
have no paragnaths on Area V of the pharynx (
<tableCitation id="C6E50339FFC9454BFF11FCC6FF73FC89" box="[158,245,802,827]" captionStart="TABLE 2" captionStartId="23.[147,231,531,553]" captionText="TABLE 2. Key morphological characters of Neanthes species in the present study. Abbreviations: DCAL = dorsal cirri on dorsal ligule; DFN = neurochaetae of dorsal fascicle; HoS = homogomph spinigers; HeF = heterogomph falcigers; LTJ = number of lateral teeth on jaws; NeL = number of neuropodial lobes; NPL = notopodial prechaetal lobe; NeL = neuropodial lobes; P/Ch1 =length ratio of peristomium/chaetiger 1; RCLPTC = range of chaetigers reached by the longest posterior tentacular cirri; SeS = sesquigomph spinigers; SeF = sesquigomph falcigers; THeF = types of heterogomph falcigers; VFN = neurochaetae of ventral fascicle." pageId="6" pageNumber="179">Table 2</tableCitation>
).
<taxonomicName id="4C674D01FFC9454BFE86FCC7FD99FC89" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[265,543,803,827]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFE86FCC7FD99FC89" box="[265,543,803,827]" italics="true" pageId="6" pageNumber="179">Neanthes donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFDA9FCC6FDE2FC89" bold="true" box="[550,612,802,827]" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFDA9FCC6FDE2FC89" box="[550,612,802,827]" pageId="6" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, however, can be distinguished from
<taxonomicName id="4C674D01FFC9454BFB8BFCC7FB2BFC89" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[1028,1197,803,827]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="kaomeiensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFB8BFCC7FB91FC88" box="[1028,1047,803,826]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFBABFCC7FB2BFC89" box="[1060,1197,803,827]" italics="true" pageId="6" pageNumber="179">kaomeiensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFB3BFCC6FB74FC89" bold="true" box="[1204,1266,802,827]" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFB3BFCC6FB74FC89" box="[1204,1266,802,827]" pageId="6" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the absence of neuropodial sesquigomph spinigers and falcigers and by the absence of short heterogomph falcigers in the neuropodia (
<tableCitation id="C6E50339FFC9454BFEABFC8EFEFBFC31" box="[292,381,874,899]" captionStart="TABLE 2" captionStartId="23.[147,231,531,553]" captionText="TABLE 2. Key morphological characters of Neanthes species in the present study. Abbreviations: DCAL = dorsal cirri on dorsal ligule; DFN = neurochaetae of dorsal fascicle; HoS = homogomph spinigers; HeF = heterogomph falcigers; LTJ = number of lateral teeth on jaws; NeL = number of neuropodial lobes; NPL = notopodial prechaetal lobe; NeL = neuropodial lobes; P/Ch1 =length ratio of peristomium/chaetiger 1; RCLPTC = range of chaetigers reached by the longest posterior tentacular cirri; SeS = sesquigomph spinigers; SeF = sesquigomph falcigers; THeF = types of heterogomph falcigers; VFN = neurochaetae of ventral fascicle." pageId="6" pageNumber="179">Table 2</tableCitation>
). Unlike
<taxonomicName id="4C674D01FFC9454BFE68FC8FFD32FC31" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[487,692,874,899]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="nanhaiensis">
<emphasis id="B913EA90FFC9454BFE68FC8FFE7CFC30" box="[487,506,875,898]" italics="true" pageId="6" pageNumber="179">N</emphasis>
. cf.
<emphasis id="B913EA90FFC9454BFDA1FC8FFD32FC31" box="[558,692,875,899]" italics="true" pageId="6" pageNumber="179">nanhaiensis</emphasis>
</taxonomicName>
, which have zero to one conical paragnath on Areas I, VI &amp; VII/ VIII, the number of conical paragnaths on that of
<taxonomicName id="4C674D01FFC9454BFD6FFC74FC2AFC15" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[736,940,911,935]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFD6FFC74FD75FC15" box="[736,755,912,935]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFC8BFC6BFC2AFC15" box="[772,940,911,935]" italics="true" pageId="6" pageNumber="179">donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFC39FC6AFC71FC15" bold="true" box="[950,1015,910,935]" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFC39FC6AFC71FC15" box="[950,1015,910,935]" pageId="6" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are 3–7, 2–4 &amp; 51–66, respectively (
<tableCitation id="C6E50339FFC9454BFF11FC56FF7EFC79" box="[158,248,946,971]" captionStart="TABLE 2" captionStartId="23.[147,231,531,553]" captionText="TABLE 2. Key morphological characters of Neanthes species in the present study. Abbreviations: DCAL = dorsal cirri on dorsal ligule; DFN = neurochaetae of dorsal fascicle; HoS = homogomph spinigers; HeF = heterogomph falcigers; LTJ = number of lateral teeth on jaws; NeL = number of neuropodial lobes; NPL = notopodial prechaetal lobe; NeL = neuropodial lobes; P/Ch1 =length ratio of peristomium/chaetiger 1; RCLPTC = range of chaetigers reached by the longest posterior tentacular cirri; SeS = sesquigomph spinigers; SeF = sesquigomph falcigers; THeF = types of heterogomph falcigers; VFN = neurochaetae of ventral fascicle." pageId="6" pageNumber="179">Table 2</tableCitation>
). Moreover,
<taxonomicName id="4C674D01FFC9454BFE02FC57FDDAFC79" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[397,604,946,971]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="nanhaiensis">
<emphasis id="B913EA90FFC9454BFE02FC57FE26FC78" box="[397,416,947,970]" italics="true" pageId="6" pageNumber="179">N</emphasis>
. cf.
<emphasis id="B913EA90FFC9454BFE59FC57FDDAFC79" box="[470,604,947,971]" italics="true" pageId="6" pageNumber="179">nanhaiensis</emphasis>
</taxonomicName>
has both medium-length and long heterogomph falcigers in neuropodia, whereas
<taxonomicName id="4C674D01FFC9454BFF75FC3CFE45FC5D" authority="Hsueh, 2019" authorityName="Hsueh" authorityYear="2019" box="[250,451,983,1007]" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="179" phylum="Annelida" rank="species" species="donggungensis" status="sp. nov.">
<emphasis id="B913EA90FFC9454BFF75FC3CFE8BFC5D" box="[250,269,984,1007]" italics="true" pageId="6" pageNumber="179">N</emphasis>
.
<emphasis id="B913EA90FFC9454BFE94FC33FE45FC5D" box="[283,451,983,1007]" italics="true" pageId="6" pageNumber="179">donggungensis</emphasis>
</taxonomicName>
<emphasis id="B913EA90FFC9454BFE45FC32FD8CFC5D" bold="true" box="[458,522,982,1007]" pageId="6" pageNumber="179">
<taxonomicNameLabel id="A22057EBFFC9454BFE45FC32FD8CFC5D" box="[458,522,982,1007]" pageId="6" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has only medium-length heterogomph falcigers in neuropodia (
<figureCitation id="135C2A07FFC9454BFB58FC32FAD5FC5D" box="[1239,1363,982,1007]" captionStart="FIGURE 4" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[167,1431,874,1903]" captionTargetId="figure@5.[167,1431,872,1903]" captionTargetPageId="5" captionText="FIGURE 4. Neanthes donggungensis n. sp.; paratype (NMNS 7946-015): A, notochaeta, homogomph spiniger, chaetiger 150; B, neurochaeta of dorsal fascicle, homogomph spiniger, chaetiger 74; C, neurochaeta of dorsal fascicle, medium heterogomph falciger, chaetiger 74; D, neurochaeta of ventral fascicle, heterogomph spiniger, chaetiger 74; E, neurochaeta of ventral fascicle, medium heterogomph falciger, chaetiger 74. Scale: A–E, 0.05 mm." httpUri="https://zenodo.org/record/2623449/files/figure.png" pageId="6" pageNumber="179">Figs 4C, E</figureCitation>
;
<figureCitation id="135C2A07FFC9454BFAEFFC33FF2CFBA0" captionStart="FIGURE 12" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[304,1288,193,1864]" captionTargetId="figure@14.[304,1288,193,1864]" captionTargetPageId="14" captionText="FIGURE 12. Neanthes cf. nanhaiensis Wu, Sun &amp; Yang, 1981 (NMNS 7946-029): A, notochaeta, homogomph spiniger, chaetiger 32; B, neurochaeta of dorsal fascicle, homogomph spiniger, chaetiger 32; C, neurochaeta of dorsal fascicle, medium heterogomph falciger, chaetiger 32; D, neurochaeta of ventral fascicle, homogomph spiniger, chaetiger 32; E, neurochaeta of ventral fascicle, medium heterogomph falciger, chaetiger 32; F, neurochaeta of ventral fascicle, long heterogomph falciger, chaetiger 32. Scale: A–B, D, 0.05 mm; C, E–F, 0.01 mm." httpUri="https://zenodo.org/record/2623468/files/figure.png" pageId="6" pageNumber="179">
12C
<emphasis id="B913EA90FFC9454BFA01FC32FA1AFC5D" bold="true" box="[1422,1436,982,1007]" pageId="6" pageNumber="179">–</emphasis>
F
</figureCitation>
;
<tableCitation id="C6E50339FFC9454BFF3AFC1EFE8DFBA1" box="[181,267,1018,1043]" captionStart="TABLE 2" captionStartId="23.[147,231,531,553]" captionText="TABLE 2. Key morphological characters of Neanthes species in the present study. Abbreviations: DCAL = dorsal cirri on dorsal ligule; DFN = neurochaetae of dorsal fascicle; HoS = homogomph spinigers; HeF = heterogomph falcigers; LTJ = number of lateral teeth on jaws; NeL = number of neuropodial lobes; NPL = notopodial prechaetal lobe; NeL = neuropodial lobes; P/Ch1 =length ratio of peristomium/chaetiger 1; RCLPTC = range of chaetigers reached by the longest posterior tentacular cirri; SeS = sesquigomph spinigers; SeF = sesquigomph falcigers; THeF = types of heterogomph falcigers; VFN = neurochaetae of ventral fascicle." pageId="6" pageNumber="179">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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