<documentid="A59214E1E495B8D1EEFEE19ED44793A2"ID-CLB-Dataset="305014"ID-DOI="10.11646/zootaxa.5527.1.1"ID-GBIF-Dataset="0d9d4ebd-5dd8-473c-80b9-19960053d8b4"ID-ISSN="1175-5334"ID-Zenodo-Dep="14021823"ID-ZooBank="771D2E7B-4025-45BF-B328-6EC8A8851ECD"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="guilherme"IM.materialsCitations_approvedBy="guilherme"IM.metadata_approvedBy="guilherme"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="guilherme"IM.treatmentCitations_approvedBy="guilherme"IM.treatments_approvedBy="guilherme"checkinTime="1730451569409"checkinUser="plazi"docAuthor="Ellenrieder, Natalia Von & Gill, Raymond J."docDate="2024"docId="039787AAFFB2FFD2FF4505EAFB79B4B5"docLanguage="en"docName="zootaxa.5527.1.1.pdf"docOrigin="Zootaxa 5527 (1)"docSource="http://dx.doi.org/10.11646/zootaxa.5527.1.1"docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph"docStyleId="5EBBA59367AD13919D70D935FA04F6A3"docStyleName="Zootaxa.2013-.monograph"docStyleVersion="19"docTitle="Tetralicia agrifoliae von Ellenrieder & Gill 2024, sp. nov."docType="treatment"docVersion="7"lastPageNumber="16"masterDocId="FFAEFFD2FFBFFFDDFFD2062DFFF1B368"masterDocTitle="The genus Tetralicia Harrison (Hemiptera: Sternorrhyncha: Aleyrodidae) in California, U. S. A., with the description of five new species and a redescription of Tetralicia granulata Sampson & Drews, 1941"masterLastPageNumber="129"masterPageNumber="1"pageNumber="14"updateTime="1730914976340"updateUser="ExternalLinkService">
<mods:titleid="700C0793723EB9AC1241A1BA38EABE47">The genus Tetralicia Harrison (Hemiptera: Sternorrhyncha: Aleyrodidae) in California, U. S. A., with the description of five new species and a redescription of Tetralicia granulata Sampson & Drews, 1941</mods:title>
<mods:affiliationid="AFD6585DA28BFC6754554A31F967DE06">Plant Pest Diagnostics Center, California Department of Food & Agriculture, 3294 Meadowview Road, Sacramento, CA 95832 - 1448, U. S. A. & natalia. von. ellenrieder @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 1159 - 2019</mods:affiliation>
<mods:affiliationid="0A9AC2A0C46FD26AD66FE83A52354E28">Plant Pest Diagnostics Center, California Department of Food & Agriculture, 3294 Meadowview Road, Sacramento, CA 95832 - 1448, U. S. A. & Plant Pest Diagnostics Center, California Department of Food & Agriculture, 3294 Meadowview Road, Sacramento, CA 95832 - 1448, U. S. A. & rayromagill @ gmail. com; https: // orcid. org / 0009 - 0005 - 9047 - 2742 * Corresponding author</mods:affiliation>
<taxonomicNameid="4C3E4D3FFFB2FFD0FF4505EAFD59B08A"authority="von Ellenrieder & Gill"authorityName="von Ellenrieder & Gill"authorityYear="2024"box="[151,680,967,994]"class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="13"pageNumber="14"phylum="Arthropoda"rank="species"species="agrifoliae"status="sp. nov.">
<figureCitationid="13052A39FFB2FFD0FF4505C1FF2FB76E"box="[151,222,1004,1030]"captionStart="FIGURES 1–4"captionStartId="12.[151,275,1849,1875]"captionTargetBox="[183,1405,391,1811]"captionTargetId="figure-135@12.[170,1418,378,1824]"captionText="FIGURES 1–4. Puparia of Tetralicia in situ: 1: T. abnormis, Santa Barbara Co., Los Olivos, on Quercus agrifolia; 2, 3: T. acaudata, 2: Riverside Co., Tin Canyon, on manzanita, 3: San Bernardino Co., Mountain Home Village, Mill Creek Canyon, on manzanita; 4: T. agrifoliae, Riverside Co., Blue Jay Road & Ortega highway, on Quercus agrifolia. Fig. 2, by N. von Ellenrieder & R.W. Garrison, 1, 3, 4 by R.J. Gill."figureDoi="http://doi.org/10.5281/zenodo.14046317"httpUri="https://zenodo.org/record/14046317/files/figure.png"pageId="13"pageNumber="14">Figs 4</figureCitation>
<materialsCitationid="3B563CE1FFB2FFD0FEA90275FAF1B7FE"collectedFrom="on Quercus agrifolia"collectingDate="1986-03-19"collectionCode="CSCA"collectorName="R. J. Gill"country="United States of America"county="Riverside County"location="Blue Jay Road & Ortega Highway"pageId="13"pageNumber="14"specimenCount="1"stateProvince="California"typeStatus="holotype">
26 slide mounted puparia:
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,
<collectingRegionid="49FAF85EFFB2FFD0FD2F0275FC82B71A"box="[765,883,1112,1138]"country="United States of America"name="California"pageId="13"pageNumber="14">California</collectingRegion>
<locationid="8EE16067FFB2FFD0FF370251FD9CB7FE"LSID="urn:lsid:plazi:treatment:039787AAFFB2FFD2FF4505EAFB79B4B5:8EE16067FFB2FFD0FF370251FD9CB7FE"box="[229,621,1148,1175]"country="United States of America"county="Riverside County"name="Blue Jay Road & Ortega Highway"pageId="13"pageNumber="14"stateProvince="California">Blue Jay Road & Ortega Highway</location>
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];
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<materialsCitationid="3B563CE1FFB2FFD0FADB0251FE08B7D2"collectedFrom="on Quercus agrifolia"collectingDate="1986-03-19"collectionCode="CSCA"collectorName="R. J. Gill"country="United States of America"county="Riverside County"location="Blue Jay Road & Ortega Highway"pageId="13"pageNumber="14"specimenCount="3"stateProvince="California"typeStatus="paratype">
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];
</materialsCitation>
<materialsCitationid="3B563CE1FFB2FFD0FDD3028DFCE8B7B6"collectedFrom="on Quercus agrifolia"collectingDate="1986-03-18"collectionCode="CSCA"collectorName="R. J. Gill"location="San Luis Rey River Day Use Area, Highway 76"municipality="San Diego County"pageId="13"pageNumber="14"specimenCount="9"typeStatus="paratype">
<collectingMunicipalityid="6BE5ACC6FFB2FFD0FDD3028DFD24B7D2"box="[513,725,1184,1210]"pageId="13"pageNumber="14">San Diego County</collectingMunicipality>
<locationid="8EE16067FFB2FFD0FCA8028DFA69B7D2"LSID="urn:lsid:plazi:treatment:039787AAFFB2FFD2FF4505EAFB79B4B5:8EE16067FFB2FFD0FCA8028DFA69B7D2"box="[890,1432,1184,1211]"municipality="San Diego County"name="San Luis Rey River Day Use Area, Highway 76"pageId="13"pageNumber="14">San Luis Rey River Day Use Area, Highway 76</location>
<collectionCodeid="ED2FAE79FFB2FFD0FD6F02E9FCFEB7B6"box="[701,783,1220,1246]"collectionName="USA, California, Sacramento, California State Collection of Arthropods"country="USA"lsid="urn:lsid:biocol.org:col:33156"name="California State Collection of Arthropods"pageId="13"pageNumber="14"type="Museum">CSCA</collectionCode>
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];
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<materialsCitationid="3B563CE1FFB2FFD0FADF02C4FB43B64E"collectedFrom="on Quercus sp."collectingDate="1930-06"collectionCode="CSCA"collectorName="Dan Gerling"county="Los Angeles County"location="Mt. Baldy"pageId="13"pageNumber="14"specimenCount="1"typeStatus="paratype">
<collectingCountyid="62E04E30FFB2FFD0FADF02C4FF1FB64E"pageId="13"pageNumber="14">Los Angeles County</collectingCounty>
<locationid="8EE16067FFB2FFD0FCF20320FC63B64E"LSID="urn:lsid:plazi:treatment:039787AAFFB2FFD2FF4505EAFB79B4B5:8EE16067FFB2FFD0FCF20320FC63B64E"box="[800,914,1292,1319]"country="United States of America"county="Los Angeles County"municipality="San Diego County"name="Mt. Baldy"pageId="13"pageNumber="14"stateProvince="California">Mt. Baldy</location>
<collectionCodeid="ED2FAE79FFB2FFD0FB8B0321FB5AB64E"box="[1113,1195,1292,1318]"collectionName="USA, California, Sacramento, California State Collection of Arthropods"country="USA"lsid="urn:lsid:biocol.org:col:33156"name="California State Collection of Arthropods"pageId="13"pageNumber="14"type="Museum">CSCA</collectionCode>
<materialsCitationid="3B563CE1FFB2FFD0FF15031DFC07B606"collectedFrom="on Quercus agrifolia"collectingDate="1986-03-19"collectionCode="CSCA"collectorName="R. J. Gill"country="United States of America"county="Riverside County"location="Blue Jay Road & Ortega Highway"pageId="13"pageNumber="14"specimenCount="1"stateProvince="California">
Additional material examined. 1 dry mounted puparium:
<collectingCountryid="F329762CFFB2FFD0FC99031DFC6BB622"box="[843,922,1328,1354]"name="United States of America"pageId="13"pageNumber="14">U.S.A.</collectingCountry>
,
<collectingRegionid="49FAF85EFFB2FFD0FC74031DFBEBB622"box="[934,1050,1328,1354]"country="United States of America"name="California"pageId="13"pageNumber="14">California</collectingRegion>
<locationid="8EE16067FFB2FFD0FB2A031DFE9EB606"LSID="urn:lsid:plazi:treatment:039787AAFFB2FFD2FF4505EAFB79B4B5:8EE16067FFB2FFD0FB2A031DFE9EB606"country="United States of America"county="Riverside County"name="Blue Jay Road & Ortega Highway"pageId="13"pageNumber="14"stateProvince="California">Blue Jay Road & Ortega Highway</location>
<collectionCodeid="ED2FAE79FFB2FFD0FC4F0379FC1EB606"box="[925,1007,1364,1390]"collectionName="USA, California, Sacramento, California State Collection of Arthropods"country="USA"lsid="urn:lsid:biocol.org:col:33156"name="California State Collection of Arthropods"pageId="13"pageNumber="14"type="Museum">CSCA</collectionCode>
<collectingRegionid="49FAF85EFFB2FFD0FE5D0355FDF0B6FA"box="[399,513,1400,1426]"country="United States of America"name="California"pageId="13"pageNumber="14">California</collectingRegion>
. Puparium black, rounded oval, lacking a caudal protuberance, lacking dorsal wax and with a wide lateral fringe of filamentous wax curling beyond submargin (
<figureCitationid="13052A39FFB2FFD0FC1E03C9FBE0B696"box="[972,1041,1508,1534]"captionStart="FIGURES 1–4"captionStartId="12.[151,275,1849,1875]"captionTargetBox="[183,1405,391,1811]"captionTargetId="figure-135@12.[170,1418,378,1824]"captionText="FIGURES 1–4. Puparia of Tetralicia in situ: 1: T. abnormis, Santa Barbara Co., Los Olivos, on Quercus agrifolia; 2, 3: T. acaudata, 2: Riverside Co., Tin Canyon, on manzanita, 3: San Bernardino Co., Mountain Home Village, Mill Creek Canyon, on manzanita; 4: T. agrifoliae, Riverside Co., Blue Jay Road & Ortega highway, on Quercus agrifolia. Fig. 2, by N. von Ellenrieder & R.W. Garrison, 1, 3, 4 by R.J. Gill."figureDoi="http://doi.org/10.5281/zenodo.14046317"httpUri="https://zenodo.org/record/14046317/files/figure.png"pageId="13"pageNumber="14">Fig. 4</figureCitation>
. TMS ending at submargin, not extending over it, 'U' shaped, with transverse section almost straight and distal ends bent at almost 90 degrees, not lined with tubercles medially (
<figureCitationid="13052A39FFB2FFD0FB300001FAF3B52E"box="[1250,1282,1580,1606]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="13"pageNumber="14">44</figureCitation>
); longitudinal molting suture lined with tubercles from TMS to head submargin; eyespots present, oval; Ce, T2 and T3 setae present; T3 setae arising well behind anterior margin of metanotum (
<figureCitationid="13052A39FFB2FFD0FC2E0059FBEDB5E6"box="[1020,1052,1652,1678]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="13"pageNumber="14">44</figureCitation>
); medial area lacking depressions, with large rounded tubercles forming a reticulated pattern on head, T1, and anterior portion of T2, and with a row of large squarish tubercles across anteromedial margin of each A1–A7 (
<figureCitationid="13052A39FFB2FFD0FE7A00CDFE36B592"box="[424,455,1760,1786]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="13"pageNumber="14">46</figureCitation>
), with pores and porettes on each side of submedial area of head (0–4), T2 (0–2), T3 (0–2), A1 (0–1), A2 (0–1), A3 (0–1), A4 (0–1), A5 (0–2), A6 (0–1), A7 (0–1), and A8 (0–1); lateral areas of dorsal disc with large rounded tubercles forming a reticulated pattern, with pairs of pores and porettes along sides of dorsal disc; dorsal submargin with large rounded tubercles decreasing in size into small granulations towards margins (
<figureCitationid="13052A39FFB2FFD0FAB90161FF10B4E2"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="13"pageNumber="14">Figs 44, 48</figureCitation>
); deflexed submargin with a row of pores and covered by uniformly distributed granulations across its entire width (
<figureCitationid="13052A39FFB2FFD0FF3B01B9FEB1B4C7"box="[233,320,1940,1967]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="13"pageNumber="14">Fig. 45</figureCitation>
); marginal glandular teeth subquadragular with tips smoothly rounded (
<figureCitationid="13052A39FFB2FFD0FBAE01B9FB22B4C7"box="[1148,1235,1940,1967]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="13"pageNumber="14">Fig. 45</figureCitation>
); VO subcordate, inset from posterior margin by more than its own length; operculum cordate, its dorsal surface with longitudinal and transverse ridges and wavy striations in between, with microspinulae across distal third to fourth; lingula concealed by operculum; VO ring very narrow around lateral and posterior sections of VO, with anterior portion wide, open antero-medially, and forming an ear-like promontory on each anterior corner (
<figureCitationid="13052A39FFB1FFD3FBCD0696FB87B3BD"box="[1055,1142,187,213]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="14"pageNumber="15">Fig. 46</figureCitation>
), with dorsal setae of A8 arising anterior to level of anterior margin of operculum (
<figureCitationid="13052A39FFB1FFD3FCC506F2FC9AB391"box="[791,875,223,249]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="14"pageNumber="15">Fig. 48</figureCitation>
); bases of caudal setae close together, within level of operculum lateral margins (
<figureCitationid="13052A39FFB1FFD3FE39072EFDB1B275"box="[491,576,259,285]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="14"pageNumber="15">Fig. 49</figureCitation>
); with two membranous ventral sacs medially to bases of mesothoracic legs; leg bases and entire venter extensively covered with spinulae (
<figureCitationid="13052A39FFB1FFD3FC81070AFC56B229"box="[851,935,295,321]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="14"pageNumber="15">Fig. 47</figureCitation>
<taxonomicNameid="4C3E4D3FFFB1FFD3FD17010EFF13B408"authority=", San Bernardino Co., Mountain Home Village, Mill Creek."authorityName="San Bernardino Co., Mountain Home Village, Mill Creek."class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="ceanothi">
<taxonomicNameid="4C3E4D3FFFB1FFD3FED9016AFCD5B408"authority=", San Joaquin Co., Lake Camanche"authorityName="San Joaquin Co., Lake Camanche"box="[267,804,1862,1888]"class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="ceanothi">
<taxonomicNameid="4C3E4D3FFFB0FFD2FDD907F6FC00B29D"authority="Carapia-Ruiz & Sanchez-Flores"authorityName="Carapia-Ruiz & Sanchez-Flores"box="[523,1009,475,501]"class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="15"pageNumber="16"phylum="Arthropoda"rank="species"species="anahuac">
by its U-shaped transverse molting suture ending before submargin and dorsal reticulation pattern. It shares the combination of a rounded-oval puparium lacking a caudal protuberance, presence of eyespots, Ce setae, large rounded tubercles on lateral areas of dorsal disc, TMS not reaching apparent margin, and hosts with
<taxonomicNameid="4C3E4D3FFFB0FFD2FCF404A2FB11B1C1"authority="Sanchez-Flores et al. 2018"authorityName="Sanchez-Flores"authorityYear="2018"box="[806,1248,655,681]"class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="15"pageNumber="16"phylum="Arthropoda"rank="species"species="xalapensis">
<bibRefCitationid="EFAF4B4DFFB0FFD2FC6804A2FB11B1C1"author="Sanchez-Flores, O. A. & Carapia-Ruiz, V. E. & Garcia-Martinez, O. & Castillo-Gutierrez, A."box="[954,1248,655,681]"pageId="15"pageNumber="16"pagination="1 - 6"refId="ref60591"refString="Sanchez-Flores, O. A., Carapia-Ruiz, V. E., Garcia-Martinez, O. & Castillo-Gutierrez, A. (2018) Descripcion de una especie nueva del genero Aleuropleurocelus Drews y Sampson (Hemiptera: Aleyrodidae) de Mexico. Acta Zoologica Mexicana, Nueva Serie, 34, 1 - 6. https: // doi. org / 10.21829 / azm. 2018.3412104"type="journal article"year="2018">
<bibRefCitationid="EFAF4B4DFFB0FFD2FDFA04FAFCA6B199"author="Sanchez-Flores, O. A. & Carapia-Ruiz, V. E. & Garcia-Martinez, O. & Castillo-Gutierrez, A."box="[552,855,727,753]"pageId="15"pageNumber="16"pagination="1 - 6"refId="ref60591"refString="Sanchez-Flores, O. A., Carapia-Ruiz, V. E., Garcia-Martinez, O. & Castillo-Gutierrez, A. (2018) Descripcion de una especie nueva del genero Aleuropleurocelus Drews y Sampson (Hemiptera: Aleyrodidae) de Mexico. Acta Zoologica Mexicana, Nueva Serie, 34, 1 - 6. https: // doi. org / 10.21829 / azm. 2018.3412104"type="journal article"year="2018">
<figureCitationid="13052A39FFB0FFD2FE3704D6FDDAB07D"box="[485,555,763,789]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="15"pageNumber="16">48, 49</figureCitation>
) [puparium elliptical oval with a caudal protuberance;
<figureCitationid="13052A39FFB0FFD2FB5A04D6FB1BB07D"box="[1160,1258,763,789]"captionStart="FIGURE 142"captionStartId="91.[151,250,1765,1789]"captionTargetBox="[244,1338,195,1728]"captionTargetId="figure-25@91.[229,1357,181,1741]"captionTargetPageId="91"captionText="FIGURE 142. Tetralicia mexicana, California, on oak: Habitus."figureDoi="http://doi.org/10.5281/zenodo.14021912"httpUri="https://zenodo.org/record/14021912/files/figure.png"pageId="15"pageNumber="16">Figs 142</figureCitation>
,
<figureCitationid="13052A39FFB0FFD2FB2704D6FAD0B07D"box="[1269,1313,763,789]"captionStart="FIGURE 143"captionStartId="92.[151,250,1755,1779]"captionTargetBox="[261,1326,194,1719]"captionTargetId="figure-30@92.[248,1339,181,1732]"captionTargetPageId="92"captionText="FIGURE 143. Tetralicia mexicana, Sierra Madre, on Quercus: Habitus."figureDoi="http://doi.org/10.5281/zenodo.14021916"httpUri="https://zenodo.org/record/14021916/files/figure.png"pageId="15"pageNumber="16">143</figureCitation>
,
<figureCitationid="13052A39FFB0FFD2FAF904D6FA61B07D"box="[1323,1424,762,789]"captionStart="FIGURES 144–150"captionStartId="93.[151,264,1695,1719]"captionTargetBox="[164,1425,194,1661]"captionTargetId="figure-25@93.[151,1435,181,1671]"captionTargetPageId="93"captionText="FIGURES 144–150. Tetralicia mexicana: 144, 146, 149, 150: Sierra Madre, on Quercus; 145, 148: San Bernardino County, New York Mountains of E San Bernadino Co., on Quercus turbinella; 147, 149: California, on oak. 144, 145: Dorsal disc and submargin of cephalothorax; 147: Dorsal submargin; 147: Venter of T1–T3; 148: Dorsal disc of A3–A8; 149: Caudal end of abdomen, dorsal view; 150: Caudal end of abdomen, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021918"httpUri="https://zenodo.org/record/14021918/files/figure.png"pageId="15"pageNumber="16">149, 150</figureCitation>
], dorsal A8 setae longer than caudal setae (
<figureCitationid="13052A39FFB0FFD2FD8C0532FD2DB051"box="[606,732,799,825]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="15"pageNumber="16">Figs 48, 49</figureCitation>
<figureCitationid="13052A39FFB0FFD2FDF6056EFD87B035"box="[548,630,835,861]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="15"pageNumber="16">Fig. 47</figureCitation>
) [smooth except for wide irregular band of spinules medial to leg bases and around mouthparts;
<figureCitationid="13052A39FFB0FFD2FE77054AFDF7B0E9"box="[421,518,871,897]"captionStart="FIGURES 144–150"captionStartId="93.[151,264,1695,1719]"captionTargetBox="[164,1425,194,1661]"captionTargetId="figure-25@93.[151,1435,181,1671]"captionTargetPageId="93"captionText="FIGURES 144–150. Tetralicia mexicana: 144, 146, 149, 150: Sierra Madre, on Quercus; 145, 148: San Bernardino County, New York Mountains of E San Bernadino Co., on Quercus turbinella; 147, 149: California, on oak. 144, 145: Dorsal disc and submargin of cephalothorax; 147: Dorsal submargin; 147: Venter of T1–T3; 148: Dorsal disc of A3–A8; 149: Caudal end of abdomen, dorsal view; 150: Caudal end of abdomen, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021918"httpUri="https://zenodo.org/record/14021918/files/figure.png"pageId="15"pageNumber="16">Fig. 147</figureCitation>
<figureCitationid="13052A39FFB0FFD2FE9205A6FEACB0CD"box="[320,349,907,933]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="15"pageNumber="16">44</figureCitation>
) [absent] and VO ring subtriangular with each anterior corner partially separated from remainder of ring, not forming entirely independent protuberances (
<figureCitationid="13052A39FFB0FFD2FE2B0236FDCFB75D"box="[505,574,1051,1077]"captionStart="FIGURES 1–4"captionStartId="12.[151,275,1849,1875]"captionTargetBox="[183,1405,391,1811]"captionTargetId="figure-135@12.[170,1418,378,1824]"captionText="FIGURES 1–4. Puparia of Tetralicia in situ: 1: T. abnormis, Santa Barbara Co., Los Olivos, on Quercus agrifolia; 2, 3: T. acaudata, 2: Riverside Co., Tin Canyon, on manzanita, 3: San Bernardino Co., Mountain Home Village, Mill Creek Canyon, on manzanita; 4: T. agrifoliae, Riverside Co., Blue Jay Road & Ortega highway, on Quercus agrifolia. Fig. 2, by N. von Ellenrieder & R.W. Garrison, 1, 3, 4 by R.J. Gill."figureDoi="http://doi.org/10.5281/zenodo.14046317"httpUri="https://zenodo.org/record/14046317/files/figure.png"pageId="15"pageNumber="16">Fig. 4</figureCitation>
<figureCitationid="13052A39FFB0FFD2FCCB024EFCC7B715"box="[793,822,1123,1149]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="15"pageNumber="16">44</figureCitation>
) transverse molting suture [W-shaped;
<figureCitationid="13052A39FFB0FFD2FB26024EFAA4B715"box="[1268,1365,1123,1149]"captionStart="FIGURE 143"captionStartId="92.[151,250,1755,1779]"captionTargetBox="[261,1326,194,1719]"captionTargetId="figure-30@92.[248,1339,181,1732]"captionTargetPageId="92"captionText="FIGURE 143. Tetralicia mexicana, Sierra Madre, on Quercus: Habitus."figureDoi="http://doi.org/10.5281/zenodo.14021916"httpUri="https://zenodo.org/record/14021916/files/figure.png"pageId="15"pageNumber="16">Fig. 143</figureCitation>
,
<figureCitationid="13052A39FFB0FFD2FAB0024EFA61B715"box="[1378,1424,1123,1149]"captionStart="FIGURES 144–150"captionStartId="93.[151,264,1695,1719]"captionTargetBox="[164,1425,194,1661]"captionTargetId="figure-25@93.[151,1435,181,1671]"captionTargetPageId="93"captionText="FIGURES 144–150. Tetralicia mexicana: 144, 146, 149, 150: Sierra Madre, on Quercus; 145, 148: San Bernardino County, New York Mountains of E San Bernadino Co., on Quercus turbinella; 147, 149: California, on oak. 144, 145: Dorsal disc and submargin of cephalothorax; 147: Dorsal submargin; 147: Venter of T1–T3; 148: Dorsal disc of A3–A8; 149: Caudal end of abdomen, dorsal view; 150: Caudal end of abdomen, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021918"httpUri="https://zenodo.org/record/14021918/files/figure.png"pageId="15"pageNumber="16">144</figureCitation>
], dorsal setae of A8 (
<figureCitationid="13052A39FFB0FFD2FEA402AAFE38B7C9"box="[374,457,1159,1185]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="15"pageNumber="16">Fig. 48</figureCitation>
) arising anterior to level of anterior margin of operculum [at that level or posterior to it;
<figureCitationid="13052A39FFB0FFD2FF6A0286FEEAB7AD"box="[184,283,1195,1221]"captionStart="FIGURES 144–150"captionStartId="93.[151,264,1695,1719]"captionTargetBox="[164,1425,194,1661]"captionTargetId="figure-25@93.[151,1435,181,1671]"captionTargetPageId="93"captionText="FIGURES 144–150. Tetralicia mexicana: 144, 146, 149, 150: Sierra Madre, on Quercus; 145, 148: San Bernardino County, New York Mountains of E San Bernadino Co., on Quercus turbinella; 147, 149: California, on oak. 144, 145: Dorsal disc and submargin of cephalothorax; 147: Dorsal submargin; 147: Venter of T1–T3; 148: Dorsal disc of A3–A8; 149: Caudal end of abdomen, dorsal view; 150: Caudal end of abdomen, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021918"httpUri="https://zenodo.org/record/14021918/files/figure.png"pageId="15"pageNumber="16">Fig. 149</figureCitation>
], and bases of caudal setae close together, within level of operculum lateral margins (
<figureCitationid="13052A39FFB0FFD2FB260286FAB9B7AD"box="[1268,1352,1195,1221]"captionStart="FIGURES 44–49"captionStartId="63.[151,264,1705,1729]"captionTargetBox="[161,1430,194,1666]"captionTargetId="figure-25@63.[151,1435,181,1681]"captionTargetPageId="63"captionText="FIGURES 44–49. Tetralicia agrifoliae sp. nov.: 44–46: Holotype; 47–49: Paratype: San Diego Co., San Luis Rey River Day Use Area, on Quercus agrifolia. 44: Anterodorsal section; 45: Detail deflexed submargin; 46: Detail dorsal view of vasiform orifice; 47: Detail ventral spinulae; 48: Detail caudal end, dorsal view; 49: Detail caudal end, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021836"httpUri="https://zenodo.org/record/14021836/files/figure.png"pageId="15"pageNumber="16">Fig. 49</figureCitation>
) [more widely separated, outside level of lateral margins of VO ring;
<figureCitationid="13052A39FFB0FFD2FC9502E2FC5BB781"box="[839,938,1231,1257]"captionStart="FIGURE 143"captionStartId="92.[151,250,1755,1779]"captionTargetBox="[261,1326,194,1719]"captionTargetId="figure-30@92.[248,1339,181,1732]"captionTargetPageId="92"captionText="FIGURE 143. Tetralicia mexicana, Sierra Madre, on Quercus: Habitus."figureDoi="http://doi.org/10.5281/zenodo.14021916"httpUri="https://zenodo.org/record/14021916/files/figure.png"pageId="15"pageNumber="16">Figs 143</figureCitation>
,
<figureCitationid="13052A39FFB0FFD2FC6402E3FC14B781"box="[950,997,1230,1257]"captionStart="FIGURES 144–150"captionStartId="93.[151,264,1695,1719]"captionTargetBox="[164,1425,194,1661]"captionTargetId="figure-25@93.[151,1435,181,1671]"captionTargetPageId="93"captionText="FIGURES 144–150. Tetralicia mexicana: 144, 146, 149, 150: Sierra Madre, on Quercus; 145, 148: San Bernardino County, New York Mountains of E San Bernadino Co., on Quercus turbinella; 147, 149: California, on oak. 144, 145: Dorsal disc and submargin of cephalothorax; 147: Dorsal submargin; 147: Venter of T1–T3; 148: Dorsal disc of A3–A8; 149: Caudal end of abdomen, dorsal view; 150: Caudal end of abdomen, ventral view."figureDoi="http://doi.org/10.5281/zenodo.14021918"httpUri="https://zenodo.org/record/14021918/files/figure.png"pageId="15"pageNumber="16">150</figureCitation>
<taxonomicNameid="4C3E4D3FFFB0FFD2FF1503AEFE5AB6F5"authorityName="von Ellenrieder & Gill"authorityYear="2024"box="[199,427,1411,1437]"class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="15"pageNumber="16"phylum="Arthropoda"rank="species"species="agrifoliae">
<figureCitationid="13052A39FFB0FFD2FCA0038AFC49B6A9"box="[882,952,1447,1473]"captionStart="FIGURES 1–4"captionStartId="12.[151,275,1849,1875]"captionTargetBox="[183,1405,391,1811]"captionTargetId="figure-135@12.[170,1418,378,1824]"captionText="FIGURES 1–4. Puparia of Tetralicia in situ: 1: T. abnormis, Santa Barbara Co., Los Olivos, on Quercus agrifolia; 2, 3: T. acaudata, 2: Riverside Co., Tin Canyon, on manzanita, 3: San Bernardino Co., Mountain Home Village, Mill Creek Canyon, on manzanita; 4: T. agrifoliae, Riverside Co., Blue Jay Road & Ortega highway, on Quercus agrifolia. Fig. 2, by N. von Ellenrieder & R.W. Garrison, 1, 3, 4 by R.J. Gill."figureDoi="http://doi.org/10.5281/zenodo.14046317"httpUri="https://zenodo.org/record/14046317/files/figure.png"pageId="15"pageNumber="16">Fig. 4</figureCitation>
<figureCitationid="13052A39FFB0FFD2FB02038AFAE4B6A9"box="[1232,1301,1447,1473]"captionStart="FIGURES 1–4"captionStartId="12.[151,275,1849,1875]"captionTargetBox="[183,1405,391,1811]"captionTargetId="figure-135@12.[170,1418,378,1824]"captionText="FIGURES 1–4. Puparia of Tetralicia in situ: 1: T. abnormis, Santa Barbara Co., Los Olivos, on Quercus agrifolia; 2, 3: T. acaudata, 2: Riverside Co., Tin Canyon, on manzanita, 3: San Bernardino Co., Mountain Home Village, Mill Creek Canyon, on manzanita; 4: T. agrifoliae, Riverside Co., Blue Jay Road & Ortega highway, on Quercus agrifolia. Fig. 2, by N. von Ellenrieder & R.W. Garrison, 1, 3, 4 by R.J. Gill."figureDoi="http://doi.org/10.5281/zenodo.14046317"httpUri="https://zenodo.org/record/14046317/files/figure.png"pageId="15"pageNumber="16">Fig. 1</figureCitation>
, or without noticeable wax formation respectively], TMS approximately 'U' shaped, with transverse section almost straight and distal ends bent at almost 90 degrees (
<bibRefCitationid="EFAF4B4DFFB0FFD2FE91011EFDBDB425"author="Dooley, J. W. & Lambrecht, S. & Honda, J."box="[323,588,1843,1869]"pageId="15"pageNumber="16"pagination="1 - 36"refId="ref59301"refString="Dooley, J. W., Lambrecht, S. & Honda, J. (2010) Eight new state records of aleyrodine whiteflies found in Clark County, Nevada and three newly described taxa (Hemiptera: Aleyrodidae, Aleyrodinae). Insecta Mundi, 140, 1 - 36. [https: // digitalcommons. unl. edu / insectamundi / 660 /]"type="journal article"year="2010">
<collectingRegionid="49FAF85EFFB0FFD2FB98011EFB51B425"box="[1098,1184,1843,1869]"country="United States of America"name="Nevada"pageId="15"pageNumber="16">Nevada</collectingRegion>
with an oval puparium lacking a caudal protuberance and with oval eyespots. This short characterization could apply to
<taxonomicNameid="4C3E4D3FFFB0FFD2FDE00156FD4BB4FD"authorityName="von Ellenrieder & Gill"authorityYear="2024"box="[562,698,1915,1941]"class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="15"pageNumber="16"phylum="Arthropoda"rank="species"species="agrifoliae">
<collectingRegionid="49FAF85EFFB0FFD2FE0601EEFDB7B4B5"box="[468,582,1987,2013]"country="United States of America"name="California"pageId="15"pageNumber="16">California</collectingRegion>