treatments-xml/data/0C/5B/5E/0C5B5E9EEFEA57C5B22CE5BF562C853C.xml

<document ID-DOI="http://dx.doi.org/10.3897/vz.71.e73146" ID-GBIF-Dataset="974e70ec-c0c2-4498-9112-117d40c09f29" ID-GBIF-Taxon="191979921" ID-Pensoft-Pub="2625-8498-71-711" ID-Pensoft-UUID="0F850A993C0A576CB6FCDBBD57F0E018" ID-ZooBank="BF0FB6C4FC3647128D70394030988F8B" LSID="urn:lsid:zoobank.org:act:625C3200-0AA6-4611-868A-3C21A628C71F" ModsDocID="2625-8498-71-711" checkinTime="1641857257033" checkinUser="pensoft" docAuthor="Novaes, Roberto Leonan M., Wilson, Don E. &amp; Moratelli, Ricardo" docDate="2021" docId="0C5B5E9EEFEA57C5B22CE5BF562C853C" docLanguage="en" docName="VertZool 71: 711-722" docOrigin="Vertebrate Zoology 71" docPubDate="2021-11-25" docSource="http://dx.doi.org/10.3897/vz.71.e73146" docTitle="Myotis pampa Novaes &amp; Wilson &amp; Moratelli 2021, sp. nov." docType="treatment" docUuid="625C3200-0AA6-4611-868A-3C21A628C71F" docUuidSource="ZooBank" docVersion="4" id="0F850A993C0A576CB6FCDBBD57F0E018" lastPageNumber="711" masterDocId="0F850A993C0A576CB6FCDBBD57F0E018" masterDocTitle="A new species of Myotis (Chiroptera, Vespertilionidae) from Uruguay" masterLastPageNumber="722" masterPageNumber="711" pageId="0" pageNumber="711" updateTime="1641863261047" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A new species of Myotis (Chiroptera, Vespertilionidae) from Uruguay</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Novaes, Roberto Leonan M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1657-2807</mods:nameIdentifier>
<mods:affiliation>Fiocruz Mata Atlantica, Fundacao Oswaldo Cruz, 22713 - 375, Rio de Janeiro, RJ, Brazil; https: // orcid. org / 0000 - 0003 - 1657 - 2807</mods:affiliation>
<mods:nameIdentifier type="email">robertoleonan@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wilson, Don E.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4228-5334</mods:nameIdentifier>
<mods:affiliation>Division of Mammals, National Museum of Natural History, Smithsonian Institution, 20560, Washington, DC, USA; https: // orcid. org / 0000 - 0003 - 4228 - 5334</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moratelli, Ricardo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0942-6633</mods:nameIdentifier>
<mods:affiliation>Fiocruz Mata Atlantica, Fundacao Oswaldo Cruz, 22713 - 375, Rio de Janeiro, RJ, Brazil; https: // orcid. org / 0000 - 0003 - 0942 - 6633</mods:affiliation>
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<mods:titleInfo>
<mods:title>Vertebrate Zoology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-11-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>71</mods:number>
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<mods:extent unit="page">
<mods:start>711</mods:start>
<mods:end>722</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/vz.71.e73146</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/vz.71.e73146</mods:identifier>
<mods:identifier type="Pensoft-Pub">2625-8498-71-711</mods:identifier>
<mods:identifier type="ZooBank">BF0FB6C4FC3647128D70394030988F8B</mods:identifier>
<mods:identifier type="Pensoft-UUID">0F850A993C0A576CB6FCDBBD57F0E018</mods:identifier>
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<treatment ID-GBIF-Taxon="191979921" LSID="urn:lsid:zoobank.org:act:625C3200-0AA6-4611-868A-3C21A628C71F" httpUri="http://treatment.plazi.org/id/0C5B5E9EEFEA57C5B22CE5BF562C853C" lastPageNumber="711" pageId="0" pageNumber="711">
<subSubSection pageId="0" pageNumber="711" type="nomenclature">
<paragraph pageId="0" pageNumber="711">
<taxonomicName LSID="http://zoobank.org/625C3200-0AA6-4611-868A-3C21A628C71F" authority="Novaes &amp; Wilson &amp; Moratelli, 2021" authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa" status="sp. nov.">Myotis pampa</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="711">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="711" type="holotype">
<paragraph pageId="0" pageNumber="711">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="711">
The holotype (AMNH 205471) comprises the skin and skull of an adult female (Figs 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal (upper left), ventral (upper right), and lateral (lower) views of the skull and mandible of Myotis pampa holotype (AMNH 205471) from the Uruguayan Pampas." figureDoi="10.3897/vz.71.e73146.figure4" httpUri="https://binary.pensoft.net/fig/615060" pageId="0" pageNumber="711">4</figureCitation>
- 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skin in dorsal (left) and ventral (right) views of the holotype of Myotis pampa (AMNH 205471) from the Uruguayan Pampas." figureDoi="10.3897/vz.71.e73146.figure5" httpUri="https://binary.pensoft.net/fig/615061" pageId="0" pageNumber="711">5</figureCitation>
), collected by 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M." order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="genus">M.</taxonomicName>
D. Tuttle in January, 1963. The specimen is deposited in the mammal collection at the American Museum of Natural History (New York, USA).
</paragraph>
<caption doi="10.3897/vz.71.e73146.figure4" httpUri="https://binary.pensoft.net/fig/615060" pageId="0" pageNumber="711" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="711">
<emphasis bold="true" pageId="0" pageNumber="711">Figure 4.</emphasis>
Dorsal (upper left), ventral (upper right), and lateral (lower) views of the skull and mandible of 
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
holotype (AMNH 205471) from the Uruguayan Pampas.
</paragraph>
</caption>
<caption doi="10.3897/vz.71.e73146.figure5" httpUri="https://binary.pensoft.net/fig/615061" pageId="0" pageNumber="711" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="711">
<emphasis bold="true" pageId="0" pageNumber="711">Figure 5.</emphasis>
Skin in dorsal (left) and ventral (right) views of the holotype of 
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
(AMNH 205471) from the Uruguayan Pampas.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="711" type="type locality">
<paragraph pageId="0" pageNumber="711">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="711">
Ca. 6 km NW from 
<normalizedToken originalValue="Belén">Belen</normalizedToken>
, Artigas, Uruguay (
<geoCoordinate degrees="30" direction="south" minutes="37" orientation="latitude" precision="925" value="-30.616667">30°37'S</geoCoordinate>
; 
<geoCoordinate degrees="57" direction="west" minutes="50" orientation="longitude" precision="925" value="-57.833332">57°50'W</geoCoordinate>
; 32 m elevation).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="711" type="paratypes">
<paragraph pageId="0" pageNumber="711">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="711">
Five individuals from ca. 6 km NW from 
<normalizedToken originalValue="Belén">Belen</normalizedToken>
, Artigas, Uruguay (
<geoCoordinate degrees="30" direction="south" minutes="37" orientation="latitude" precision="925" value="-30.616667">30°37'S</geoCoordinate>
; 
<geoCoordinate degrees="57" direction="west" minutes="50" orientation="longitude" precision="925" value="-57.833332">57°50'W</geoCoordinate>
, 32 m of elevation; AMNH 205461 ♀, AMNH 205464 ♀, AMNH 205467 ♀, AMNH 205472 ♀, AMNH 205476 ♀); and nine individuals from 40 km NW to Tacuarembo city, 
<normalizedToken originalValue="Tacuarambó">Tacuarambo</normalizedToken>
, Uruguay (
<geoCoordinate degrees="31" direction="south" minutes="38" orientation="latitude" precision="925" value="-31.633333">31°38'S</geoCoordinate>
; 
<geoCoordinate degrees="56" direction="west" minutes="19" orientation="longitude" precision="925" value="-56.316666">56°19'W</geoCoordinate>
, 240 m of elevation; AMNH 205541 ♀, AMNH 205545 ♀, AMNH 205546 ♂, AMNH 205552 ♂, AMNH 205553 ♀, AMNH 205562 ♀, AMNH 205563 ♂, AMNH 205564 ♀, AMNH 205565 ♂).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="711" type="distribution">
<paragraph pageId="0" pageNumber="711">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="711">
Known from only two localities in the provinces of Artigas and 
<normalizedToken originalValue="Tacuarembó">Tacuarembo</normalizedToken>
, Uruguay (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Map with localities of occurrence for Myotis pampa, including the type locality (red star)." figureDoi="10.3897/vz.71.e73146.figure6" httpUri="https://binary.pensoft.net/fig/615062" pageId="0" pageNumber="711">6</figureCitation>
). The records come from the Uruguayan Pampas, between 30 and 240 m in altitude. The region is characterized by grassland plains with shrubby vegetation patches typical of subtropical landscapes. However, the margins of rivers can present denser riparian forests, with arborescent ferns, orchids, lianas and several trees with more than 10 m in height (
<bibRefCitation author="Sganga, JC" journalOrPublisher="Privately published, Washington, DC" pageId="0" pageNumber="711" refId="B37" refString="Sganga, JC, 1994. Caracterizacion de la vegetacion de la Republica Oriental del Uruguay. In: Ministerio de Ganaderia, Agricultura y Pesca (Org) Contribucion de los estudios edafologicos al conocimiento de la vegetacion en la Republica Oriental del Uruguay. Boletin Tecnico no. 13, Direccion de Suelos y Aguas, Montevideo, 5-14." title="Caracterizacion de la vegetacion de la Republica Oriental del Uruguay. In: Ministerio de Ganaderia, Agricultura y Pesca (Org) Contribucion de los estudios edafologicos al conocimiento de la vegetacion en la Republica Oriental del Uruguay. Boletin Tecnico no. 13, Direccion de Suelos y Aguas, Montevideo, 5 - 14." year="1994">Sganga 1994</bibRefCitation>
; 
<bibRefCitation author="Panario, D" journalOrPublisher="Convenio MGAP / PPR - Facultad de Ciencias / Vida Silvestre / Sociedad Zoologica del Uruguay / CIEDUR, Montevideo" pageId="0" pageNumber="711" refId="B30" refString="Panario, D, Gutierrez, O, Achkar, M, Bartesaghi, L, Ceroni, M, 2011. Clasificacion y mapeo de ambientes de Uruguay. Convenio MGAP/PPR - Facultad de Ciencias/Vida Silvestre/Sociedad Zoologica del Uruguay/CIEDUR, Montevideo" title="Clasificacion y mapeo de ambientes de Uruguay." year="2011">Panario et al. 2011</bibRefCitation>
). According to information available on museum labels, most individuals were captured with mist-nets near watercourses.
</paragraph>
<caption doi="10.3897/vz.71.e73146.figure6" httpUri="https://binary.pensoft.net/fig/615062" pageId="0" pageNumber="711" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="711">
<emphasis bold="true" pageId="0" pageNumber="711">Figure 6.</emphasis>
Map with localities of occurrence for 
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
, including the type locality (red star).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="711" type="etymology">
<paragraph pageId="0" pageNumber="711">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="711">
We name 
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
after the Pampas ecoregion, the main habitat occurrence for the species. Pampa is a Quechua word for 
<normalizedToken originalValue="“plain”">&quot;plain&quot;</normalizedToken>
, and in Latin languages it is used in the masculine gender, which agrees with the generic name 
<taxonomicName authorityName="Kaup" authorityYear="1829" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis</emphasis>
</taxonomicName>
, also masculine (
<bibRefCitation author="ICZN" journalOrPublisher="Journal of Statistical Software" pageId="0" pageNumber="711" refId="B15" refString="ICZN, 1958. Direction 98 on the Interpretation under the Plenary Powers of the nominal species Vespertilio murinus, pp. 144-145. In Opinions and declarations rendered by the International Commission on Zoological Nomenclature. Vol 1. Section F. Part F.9." title="Direction 98 on the Interpretation under the Plenary Powers of the nominal species Vespertilio murinus, pp. 144 - 145. In Opinions and declarations rendered by the International Commission on Zoological Nomenclature. Vol 1. Section F. Part F. 9." year="1958">ICZN 1958</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.2307/1382580" author="Pritchard, PCH" journalOrPublisher="Journal of Mammalogy" pageId="0" pageNumber="711" pagination="549 - 550" refId="B33" refString="Pritchard, PCH, 1994. Comment on gender and declension of generic names. Journal of Mammalogy 75: 549 - 550, DOI: https://doi.org/10.2307/1382580" title="Comment on gender and declension of generic names." url="https://doi.org/10.2307/1382580" volume="75" year="1994">Pritchard 1994</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="711" type="diagnosis">
<paragraph pageId="0" pageNumber="711">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="711">
The tricolored ventral fur, with a Drab basal band, Smoke Gray middle band, and yellowish (generally Naples Yellow) tip is unique among South American 
<taxonomicName authorityName="Kaup" authorityYear="1829" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis</emphasis>
</taxonomicName>
. In addition, the following set of traits distinguishes 
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
from all its South American congeners: small to medium sized species (FA 33.8-36.3 mm; sagittal crest absent or very low; elongated skull (GLS 13.4-14.4 mm); braincase not inflated and low in profile; braincase roof formed by the parietal bone strongly straight and not inclined forward; frontal bone steeply sloped towards the rostrum; posterior region of the braincase rounded and projected beyond the limit of the occipital condyles; mastoid processes narrower and practically not visible in dorsal view; dorsal fur notably long (7-9 mm), woolly and bicolored, with tips Dresden Brown or paler Ochraceous-Tawny, and bases brownish-gray (giving a fuliginous aspect); ventral fur moderately long (6-7 mm); legs and dorsal surface of the uropatagium covered by scattered fur that extends up to the knees; plagiopatagium inserted into the foot by a broad band of membrane.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="711" type="description">
<paragraph pageId="0" pageNumber="711">Morphological description and comparisons.</paragraph>
<paragraph pageId="0" pageNumber="711">
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
is a small to medium species of 
<taxonomicName authorityName="Kaup" authorityYear="1829" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis</emphasis>
</taxonomicName>
(FA 33.8-36.3 mm; other measurements in Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. External and craniodental measurements of Myotis sp. nov. from Uruguay (including holotype abd paratypes), M. riparius and M. ruber including morphometric variation and sample size (N). The measurements are in millimeters. Acronyms and descriptions are available in Table 1." httpUri="http://table.plazi.org/id/95B4C1DE5310088F352A8C2E3B5A1919" pageId="0" pageNumber="711" tableUuid="95B4C1DE5310088F352A8C2E3B5A1919">2</tableCitation>
) and the fur texture (wooly) and cranial morphology (broad skull with sagittal and lambdoidal crests presents) resembles species allocated to the 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis ruber" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="ruber">Myotis ruber</taxonomicName>
-group (
<emphasis italics="true" pageId="0" pageNumber="711">q.v</emphasis>
., 
<bibRefCitation DOI="https://doi.org/10.1206/3780.2" author="Moratelli, R" journalOrPublisher="American Museum Novitates" pageId="0" pageNumber="711" pagination="1 - 36" refId="B22" refString="Moratelli, R, Gardner, AL, Oliveira, JA, Wilson, DE, 2013. Review of Myotis (Chiroptera, Vespertilionidae) from northern South America, including description of a new species. American Museum Novitates 3780: 1 - 36, DOI: https://doi.org/10.1206/3780.2" title="Review of Myotis (Chiroptera, Vespertilionidae) from northern South America, including description of a new species." url="https://doi.org/10.1206/3780.2" volume="3780" year="2013">Moratelli et al. 2013</bibRefCitation>
, 
<bibRefCitation author="Moratelli, R" editor="Wilson, DE" journalOrPublisher="9. Bats. Lynx Editions, Barcelona" pageId="0" pageNumber="711" pagination="716 - 981" refId="B24" refString="Moratelli, R, Burgin, C, Claudio, VC, Novaes, RLM, Lopez-Baucells, A, Haslauer, R, 2019a. Family Vespertilionidae (Vesper Bats). In: Wilson, DE, Mittermeier, AR, Eds., Handbook of the mammals of the world, vol. 9. Bats. Lynx Editions, Barcelona: 716 - 981" title="Family Vespertilionidae (Vesper Bats)." volumeTitle="Handbook of the mammals of the world, vol." year="2019 a">2019a</bibRefCitation>
). Ears are Mummy Brown and comparatively medium sized (EL 11-14 mm), reaching the portion of the rostrum between the eyes and nostrils when extended forward. Tragus is long and slender, with a wide base and a narrower spear-shaped terminal half, almost straight anterior edge, and rounded tip. Membranes are Mummy Brown; the dorsal surface of tibia, and uropatagium has scattered hairs extending to the level of the knees. The uropatagium lacks the fringe of hairs along the trailing edge. The plagiopatagium is attached to the foot at the level of the toes by a broad band of membrane.
</paragraph>
<paragraph pageId="0" pageNumber="711">
Dorsal and ventral fur are wooly and long (LDF 7-9 mm, LVF 6-7 mm). The dorsal pelage is bicolored with brownish-gray bases (3/4 of the total fur length) and reddish tips (1/4 of the total fur length), ranging from Dresden Brown to paler Ochraceous-Tawny. The dorsal fur has a coloration that looks like the bat is partially covered by soot and with the tip of the hairs burned. The ventral fur is tricolored, with Drab bases (1/4 of the total fur length) progressively turning Smoke Gray in the middle band (2/4 length) and light-yellow tips (1/4 [ranging from Naples Yellow to Cream-Buff]). The tricolored ventral fur is unique among all South American 
<taxonomicName authorityName="Kaup" authorityYear="1829" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis</emphasis>
</taxonomicName>
species (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal and ventral views of the skins of of Myotis sp. nov. from Uruguay (AMNH 205471 [holotype]), Myotis riparius from Panama (USNM 310255 [holotype]), and Myotis ruber from Paraguay (USNM 115097 [neotype])." figureDoi="10.3897/vz.71.e73146.figure2" httpUri="https://binary.pensoft.net/fig/615058" pageId="0" pageNumber="711">2</figureCitation>
). However, this pattern may be less evident in some lighter specimens, where the coloration of the middle band may be confused with the tip band of the hairs.
</paragraph>
<paragraph pageId="0" pageNumber="711">
Dental formula is I 2/3, C 1/1, PM 3/3, M 3/3 = 38, typical of all New World species of 
<taxonomicName authorityName="Kaup" authorityYear="1829" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis</emphasis>
</taxonomicName>
. Skull delicate and medium-sized in length (GLS 13.1-14.1 mm), resembling 
<taxonomicName authorityName="Kaup" authorityYear="1829" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis</emphasis>
</taxonomicName>
of the 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis ruber" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="ruber">Myotis ruber</taxonomicName>
-group species. The 2nd upper premolar (P3) is in the toothrow, not displaced to the lingual side and smaller than first upper premolar (P2). The 1st lower molar (m1) is myotodont, with the postcristid connecting hypoconid and entoconid. Braincase delicate and elongated; sagittal crest and lambdoidal crests are absent or very low; the occipital region is rounded and projects beyond the posterior limits of the occipital condyles; mastoid processes weakly developed. Frontal bone slightly sloping; rostrum comparatively short and narrow.
</paragraph>
<paragraph pageId="0" pageNumber="711">
Considering species from the 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis ruber" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="ruber">Myotis ruber</taxonomicName>
-group (
<taxonomicName authorityName="Carrion-Bonilla &amp; Cook" authorityYear="2020" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis armiensis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="armiensis">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis armiensis</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Hall" authorityYear="1962" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis elegans" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis elegans</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="J. A. Allen" authorityYear="1914" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis keaysi" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="keaysi">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis keaysi</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Moratelli &amp; Wilson" authorityYear="2014" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis midastactus" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="midastactus">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis midastactus</emphasis>
</taxonomicName>
, 
<emphasis italics="true" pageId="0" pageNumber="711">moratellii</emphasis>
, 
<taxonomicName authorityName="LaVal" authorityYear="1973" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pilosatibialis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pilosatibialis">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pilosatibialis</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Handley" authorityYear="1960" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis riparius" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="riparius">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis riparius</emphasis>
</taxonomicName>
, 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis ruber" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="ruber">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis ruber</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Thomas" authorityYear="1901" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis simus" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="simus">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis simus</emphasis>
</taxonomicName>
), and species from the 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis albescens" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="albescens">Myotis albescens</taxonomicName>
-group in the South American southern cone (
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis albescens" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="albescens">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis albescens</emphasis>
</taxonomicName>
, 
<taxonomicName baseAuthorityName="Waterhouse" baseAuthorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis chiloensis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="chiloensis">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis chiloensis</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Thomas" authorityYear="1902" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis dinellii" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="dinellii">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis dinellii</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Moratelli, Peracchi, Dias &amp; de Oliveira" authorityYear="2011" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis izecksohni" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="izecksohni">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis izecksohni</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Moratelli, Peracchi, Dias &amp; de Oliveira" authorityYear="2011" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis lavali" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="lavali">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis lavali</emphasis>
</taxonomicName>
, 
<taxonomicName baseAuthorityName="I. Geoffroy" baseAuthorityYear="1824" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis levis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="levis">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis levis</emphasis>
</taxonomicName>
, 
<taxonomicName baseAuthorityName="Schinz" baseAuthorityYear="1821" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis nigricans" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="nigricans">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis nigricans</emphasis>
</taxonomicName>
), 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">M. pampa</emphasis>
</taxonomicName>
can be distinguished from all by the set of diagnostic traits reported above.
</paragraph>
<paragraph pageId="0" pageNumber="711">
These specimens of 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">M. pampa</emphasis>
</taxonomicName>
were originally identified as 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. riparius" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="riparius">
<emphasis italics="true" pageId="0" pageNumber="711">M. riparius</emphasis>
</taxonomicName>
and this identification was maintained by 
<bibRefCitation author="LaVal, RK" journalOrPublisher="Natural History Museum of Los Angeles County, Science Bulletin" pageId="0" pageNumber="711" pagination="1 - 54" refId="B17" refString="LaVal, RK, 1973. A revision of the Neotropical bats of the genus Myotis. Natural History Museum of Los Angeles County, Science Bulletin 15: 1 - 54" title="A revision of the Neotropical bats of the genus Myotis." volume="15" year="1973">LaVal (1973)</bibRefCitation>
and 
<bibRefCitation DOI="https://doi.org/10.1644/1545-1542(2001)082%3C0138:TSOMCV%3E2.0.CO;2" author="Lopez-Gonzalez, C" journalOrPublisher="Journal of Mammalogy" pageId="0" pageNumber="711" pagination="138 - 160" refId="B18" refString="Lopez-Gonzalez, C, Presley, SJ, Owen, RD, Willig, MR, 2001. Taxonomic status of Myotis (Chiroptera: Vespertilionidae) in Paraguay. Journal of Mammalogy 82: 138 - 160, DOI: https://doi.org/10.1644/1545-1542(2001)082%3C0138:TSOMCV%3E2.0.CO;2" title="Taxonomic status of Myotis (Chiroptera: Vespertilionidae) in Paraguay." url="https://doi.org/10.1644/1545-1542(2001)082%3C0138:TSOMCV%3E2.0.CO;2" volume="82" year="2001">
<normalizedToken originalValue="López-González">Lopez-Gonzalez</normalizedToken>
et al. (2001)
</bibRefCitation>
. In fact, 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">M. pampa</emphasis>
</taxonomicName>
resembles 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. riparius" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="riparius">
<emphasis italics="true" pageId="0" pageNumber="711">M. riparius</emphasis>
</taxonomicName>
, but it can be distinguished by its smaller skull (GLS &lt;14.4 in 
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
and generally&gt; 14.5 mm in 
<taxonomicName authorityName="Handley" authorityYear="1960" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis riparius" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="riparius">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis riparius</emphasis>
</taxonomicName>
), braincase lower in lateral view, sagittal crest lower (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lateral, dorsal and ventral views of skulls of Myotis sp. nov. from Uruguay (AMNH 205471 [holotype]), Myotis riparius from Panama (USNM 310255 [holotype]), and Myotis ruber from Paraguay (USNM 115097 [neotype]). The images of the mandibles of M. riparius and M. ruber are inverted." figureDoi="10.3897/vz.71.e73146.figure1" httpUri="https://binary.pensoft.net/fig/615057" pageId="0" pageNumber="711">1</figureCitation>
), longer and bicolored dorsal fur with grayish base, tricolored ventral fur (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal and ventral views of the skins of of Myotis sp. nov. from Uruguay (AMNH 205471 [holotype]), Myotis riparius from Panama (USNM 310255 [holotype]), and Myotis ruber from Paraguay (USNM 115097 [neotype])." figureDoi="10.3897/vz.71.e73146.figure2" httpUri="https://binary.pensoft.net/fig/615058" pageId="0" pageNumber="711">2</figureCitation>
), and presence of scattered hairs along the leg and dorsal surface of the uropatagium. 
<taxonomicName authorityName="Handley" authorityYear="1960" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis riparius" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="riparius">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis riparius</emphasis>
</taxonomicName>
have a higher braincase in lateral view, sagittal crest well-developed, shorter and unicolored dorsal fur, bicolored ventral fur and absence of scattered hairs along the leg and dorsal surface of the uropatagium.
</paragraph>
<paragraph pageId="0" pageNumber="711">
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
can also be confused with 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. ruber" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="ruber">
<emphasis italics="true" pageId="0" pageNumber="711">M. ruber</emphasis>
</taxonomicName>
, but it can be easly distinguished by its smaller size (both external and cranial; FA&gt; 37.5 mm and GLS&gt; 14.0 mm in 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. ruber" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="ruber">
<emphasis italics="true" pageId="0" pageNumber="711">M. ruber</emphasis>
</taxonomicName>
), skull more delicate, braincase lower in lateral view, mastoid processes less developed, dorsal fur longer and clearly bicolored with grayish base (unicolored in 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis ruber" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="ruber">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis ruber</emphasis>
</taxonomicName>
), tricolored ventral fur (bicolored in 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis ruber" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="ruber">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis ruber</emphasis>
</taxonomicName>
), and presence of scattered hairs along the leg and dorsal surface of the uropatagium (absent in 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis ruber" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="ruber">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis ruber</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="711">
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
differs from 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. armiensis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="armiensis">
<emphasis italics="true" pageId="0" pageNumber="711">M. armiensis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. keaysi" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="keaysi">
<emphasis italics="true" pageId="0" pageNumber="711">M. keaysi</emphasis>
</taxonomicName>
, and 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. pilosatibialis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pilosatibialis">
<emphasis italics="true" pageId="0" pageNumber="711">M. pilosatibialis</emphasis>
</taxonomicName>
by its external smaller size (FA&gt; 36 mm in 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. armiensis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="armiensis">
<emphasis italics="true" pageId="0" pageNumber="711">M. armiensis</emphasis>
</taxonomicName>
and 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. keaysi" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="keaysi">
<emphasis italics="true" pageId="0" pageNumber="711">M. keaysi</emphasis>
</taxonomicName>
), more delicate skull, braincase lower in lateral view, sagittal crest lower, mastoid processes less developed, dorsal fur longer and clearly bicolored with grayish base, tricolored ventral fur, and pelage on the dorsal surface of the uropatagium composed of scattered hairs not extending beyond the knees. It differs from 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. elegans" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="711">M. elegans</emphasis>
</taxonomicName>
by larger size in general (both external and cranial; FA &lt;34.5 mm, GLS &lt;13.0 mm, and BCB &lt;6.3 mm in 
<taxonomicName authorityName="Hall" authorityYear="1962" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis elegans" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis elegans</emphasis>
</taxonomicName>
), skull more robust in general, parietal bone not inclined forward, rostrum longer, dorsal fur clearly bicolored with grayish base, ventral fur tricolored. 
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
can be easily distinguished from 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. simus" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="simus">
<emphasis italics="true" pageId="0" pageNumber="711">M. simus</emphasis>
</taxonomicName>
and 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. midastactus" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="midastactus">
<emphasis italics="true" pageId="0" pageNumber="711">M. midastactus</emphasis>
</taxonomicName>
by its smaller size (generally FA&gt; 36 mm and GLS&gt; 13.5 mm in 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. simus" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="simus">
<emphasis italics="true" pageId="0" pageNumber="711">M. simus</emphasis>
</taxonomicName>
and 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. midastactus" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="midastactus">
<emphasis italics="true" pageId="0" pageNumber="711">M. midastactus</emphasis>
</taxonomicName>
), more delicate and narrower skull (BCB &lt;6.8 in 
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
and&gt; 6.8 in 
<taxonomicName authorityName="Thomas" authorityYear="1901" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis simus" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="simus">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis simus</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Moratelli &amp; Wilson" authorityYear="2014" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis midastactus" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="midastactus">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis midastactus</emphasis>
</taxonomicName>
), dorsal fur longer and clearly bicolored, ventral fur tricolored, dorsal surface of the uropatagium covered by scattered hairs that extend up to the knees; plagiopatagium inserted into the foot by a broad band of membrane.
</paragraph>
<paragraph pageId="0" pageNumber="711">
In comparison to species from the 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis albescens" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="albescens">Myotis albescens</taxonomicName>
-group, 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">M. pampa</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. albescens" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="albescens">
<emphasis italics="true" pageId="0" pageNumber="711">M. albescens</emphasis>
</taxonomicName>
, 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. dinellii" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="dinellii">
<emphasis italics="true" pageId="0" pageNumber="711">M. dinellii</emphasis>
</taxonomicName>
and 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. levis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="levis">
<emphasis italics="true" pageId="0" pageNumber="711">M. levis</emphasis>
</taxonomicName>
by the absence of a fringe of hairs on the posterior margin of the uropatagium, reddish dorsal fur bicolored with grayish base, and braincase less inflated. In addition, 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">M. pampa</emphasis>
</taxonomicName>
differs from 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. dinellii" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="dinellii">
<emphasis italics="true" pageId="0" pageNumber="711">M. dinellii</emphasis>
</taxonomicName>
and 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. levis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="levis">
<emphasis italics="true" pageId="0" pageNumber="711">M. levis</emphasis>
</taxonomicName>
by its generally smaller size and comparatively shorter ears. It differs from 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. izecksohni" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="izecksohni">
<emphasis italics="true" pageId="0" pageNumber="711">M. izecksohni</emphasis>
</taxonomicName>
and 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. nigricans" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="nigricans">
<emphasis italics="true" pageId="0" pageNumber="711">M. nigricans</emphasis>
</taxonomicName>
by the braincase less inflated and reddish dorsal fur clearly bicolored with grayish base. 
<taxonomicName authorityName="Novaes &amp; Wilson &amp; Moratelli" authorityYear="2021" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">Myotis pampa</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. chiloensis" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="chiloensis">
<emphasis italics="true" pageId="0" pageNumber="711">M. chiloensis</emphasis>
</taxonomicName>
from its smaller size in general, skull lower in lateral view, braincase less inflated, dorsal fur shorter and bicolored, with grayish bases and reddish tips. It differs from 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. lavali" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="lavali">
<emphasis italics="true" pageId="0" pageNumber="711">M. lavali</emphasis>
</taxonomicName>
by its second upper premolar (P3) smaller than first upper premolar (P2), and by its reddish fur with grayish bases (being blackish bases in 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. lavali" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="lavali">
<emphasis italics="true" pageId="0" pageNumber="711">M. lavali</emphasis>
</taxonomicName>
). In addition, 
<taxonomicName class="Mammalia" genus="M." kingdom="Animalia" lsidName="M. pampa" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="pampa">
<emphasis italics="true" pageId="0" pageNumber="711">M. pampa</emphasis>
</taxonomicName>
can be distinguished of all species from the 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis albescens" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="albescens">Myotis albescens</taxonomicName>
-group by its wooly fur (silky in 
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" class="Mammalia" family="Vespertilionidae" genus="Myotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myotis albescens" order="Chiroptera" pageId="0" pageNumber="711" phylum="Chordata" rank="species" species="albescens">Myotis albescens</taxonomicName>
-group species), ventral fur tricolored, and fur composed of scattered hairs on dorsal surface of the uropatagium.
</paragraph>
</subSubSection>
</treatment>
</document>