247 lines
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247 lines
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<document id="994ADB6BD7317D6B5FA704E6420DCEC0" ID-DOI="10.24199/j.mmv.2015.73.07" ID-ISSN="1447-2554" ID-Zenodo-Dep="12213223" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718996479650" checkinUser="felipe" docAuthor="O'Loughlin, P. Mark, Skarbnik-López, Jessica, Mackenzie, Melanie & VandenSpiegel, Didier" docDate="2015" docId="039B87A1FFA2C74DFF27FA16FC71F8A1" docLanguage="en" docName="MemMusVictoria.73.59-93.pdf" docOrigin="Memoirs of Museum Victoria 73" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-73-2015/pages-59-93/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Psolus heardi O'Loughlin & Skarbnik-Lopez 2015, sp. nov." docType="treatment" docVersion="2" lastPageNumber="81" masterDocId="FFA2FFD9FFB4C75BFFABFFC2FF98FFCE" masterDocTitle="Sea cucumbers of the Kerguelen Plateau, with descriptions of new genus and species (Echinodermata: Holothuroidea)" masterLastPageNumber="93" masterPageNumber="59" pageNumber="81" updateTime="1719012382058" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B686C838E0B0CB06900AD65163D98251">Sea cucumbers of the Kerguelen Plateau, with descriptions of new genus and species (Echinodermata: Holothuroidea)</mods:title>
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<mods:namePart id="39F648F2F4E0A40254284D264BCC8789">O'Loughlin, P. Mark</mods:namePart>
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</mods:role>
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<mods:namePart id="D2B9F9D5CA3661D06472E45CC7D67442">Skarbnik-López, Jessica</mods:namePart>
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</mods:name>
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<mods:name id="3343C29B17AF5D1F8D9978A56A747284" type="personal">
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</mods:role>
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<mods:namePart id="0387CC66970DB8DAEDFDCB1E3F02A662">Mackenzie, Melanie</mods:namePart>
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</mods:name>
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<mods:name id="AABACA6454D12EDAB3DF3BAD4A471887" type="personal">
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</mods:role>
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<mods:namePart id="140CF3557E99361574F9D6668A3CFBCC">VandenSpiegel, Didier</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="49518953CBF97557E7B0050A328B19B6">text</mods:typeOfResource>
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<mods:title id="479E1848F3F232D9848B09634573E9DD">Memoirs of Museum Victoria</mods:title>
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</mods:titleInfo>
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<mods:part id="FA221C67C7AC4245C091CB60127C307C">
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<mods:date id="27B75BFF5EC6C8D42AE8B3DCED1EB483">2015</mods:date>
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<mods:detail id="4872CB2CF311115B2F7F0CDB69D096C4" type="pubDate">
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<mods:number id="54EE3E9EC610BDB9AF999AE559BAD9E1">2015-12-31</mods:number>
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</mods:detail>
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<mods:detail id="3AB6A1147ABA080F89A900D440E9601A" type="volume">
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<mods:number id="980E9B4EDE5C5633FB3E43EE031CDFDE">73</mods:number>
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<mods:start id="9EB9D34154D2F860629DF7C6406C4A93">59</mods:start>
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<mods:end id="F7F8B454138118F5BBCFB8BB41990B66">93</mods:end>
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<mods:url id="AF494AA92DD3834F0B01BC36F35BA2A2">https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-73-2015/pages-59-93/</mods:url>
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</mods:location>
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<mods:classification id="AD7A696B6EFB6A69067A42557F4A3690">journal article</mods:classification>
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<mods:identifier id="CC243EAB982BBB26E89DEA40E3ADF082" type="DOI">10.24199/j.mmv.2015.73.07</mods:identifier>
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<mods:identifier id="F01D6B5CB1593FEA226C6372F534C9D6" type="ISSN">1447-2554</mods:identifier>
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<mods:identifier id="F29B427628C1953C23702C2BE26EF20A" type="Zenodo-Dep">12213223</mods:identifier>
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</mods:mods>
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<treatment id="039B87A1FFA2C74DFF27FA16FC71F8A1" LSID="urn:lsid:plazi:treatment:039B87A1FFA2C74DFF27FA16FC71F8A1" httpUri="http://treatment.plazi.org/id/039B87A1FFA2C74DFF27FA16FC71F8A1" lastPageNumber="81" pageId="22" pageNumber="81">
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<subSubSection id="C328653CFFA2C74DFF27FA16FEFBF9E3" pageId="22" pageNumber="81" type="nomenclature">
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<paragraph id="8B8D36B7FFA2C74DFF27FA16FD36FA24" blockId="22.[140,686,1492,1514]" box="[140,686,1492,1514]" pageId="22" pageNumber="81">
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<heading id="D0C581DBFFA2C74DFF27FA16FD36FA24" box="[140,686,1492,1514]" fontSize="9" level="2" pageId="22" pageNumber="81" reason="5">
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<taxonomicName id="4C324D34FFA2C74DFF27FA16FDC3FA24" ID-CoL="4PL97" authority="O'Loughlin & Skarbnik-Lopez" authorityName="O'Loughlin & Skarbnik-Lopez" authorityYear="2015" box="[140,603,1492,1514]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="species" species="heardi" status="sp. nov.">
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<emphasis id="B946EAA5FFA2C74DFF27FA16FE84FA24" bold="true" box="[140,284,1492,1514]" italics="true" pageId="22" pageNumber="81">Psolus heardi</emphasis>
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O’Loughlin & Skarbnik-López
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</taxonomicName>
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<taxonomicNameLabel id="A27557DEFFA2C74DFDCAFA17FD36FA24" box="[609,686,1493,1514]" pageId="22" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8B8D36B7FFA2C74DFF19FA3DFEFBF9E3" blockId="22.[140,770,1535,1607]" pageId="22" pageNumber="81">
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<emphasis id="B946EAA5FFA2C74DFF19FA3DFEDBF9DD" box="[178,323,1535,1555]" italics="true" pageId="22" pageNumber="81">Zoobank LSID.</emphasis>
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http://zoobank.org:act:
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<uuid id="FF940C62FFA2C74DFDB5FA3DFEFBF9E3" pageId="22" pageNumber="81">FF1FB240-6E7D-44BA- 9AF3-B1FADA9EE993</uuid>
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</paragraph>
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</subSubSection>
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<subSubSection id="C328653CFFA2C74DFF19F9F6FC71F8A1" pageId="22" pageNumber="81" type="description">
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<paragraph id="8B8D36B7FFA2C74DFF19F9F6FD27F989" blockId="22.[140,770,1535,1607]" box="[178,703,1588,1607]" pageId="22" pageNumber="81">
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<taxonomicName id="4C324D34FFA2C74DFF19F9F6FF76F989" authorityName="O'Loughlin & Skarbnik-López & Mackenzie & VandenSpiegel" authorityYear="2015" box="[178,238,1588,1607]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="genus" status="sp. nov.">
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<emphasis id="B946EAA5FFA2C74DFF19F9F6FF76F989" box="[178,238,1588,1607]" italics="true" pageId="22" pageNumber="81">Psolus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27557DEFFA2C74DFF5FF9F6FEA1F989" box="[244,313,1588,1607]" pageId="22" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
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(HOL 23).—Hibberd & Moore, 2009: 120,
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</paragraph>
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<paragraph id="8B8D36B7FFA2C74DFF27F99CFE2EF9BD" blockId="22.[140,438,1630,1652]" box="[140,438,1630,1652]" pageId="22" pageNumber="81">
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<tableCitation id="C6B0030CFFA2C74DFF27F99CFF61F9BD" box="[140,249,1630,1652]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStartId-0="4.[140,191,229,248]" captionStartId-1="4.[140,189,1179,1198]" captionStartId-2="5.[147,196,1313,1332]" captionText-0="Table 1. Voyages to HIMI that collected Holothuroidea specimens." captionText-1="Table 2. Holothuroidea species from HIMI (systematic identity established by Mark O’Loughlin)." captionText-2="Table 3. Holothuroidea specimens from HIMI held in NMV." pageId="22" pageNumber="81">Tables 1–3</tableCitation>
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,
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<tableCitation id="C6B0030CFFA2C74DFEA8F99CFE89F9BA" box="[259,273,1630,1652]" captionStart="Table 6" captionStartId="13.[147,196,229,248]" captionText="Table 6. Kerguelen Islands and HIMI holothuroid species distributed beyond the Kerguelen Plateau." pageId="22" pageNumber="81">6</tableCitation>
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; figures 1, 9, 10.
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</paragraph>
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<paragraph id="8B8D36B7FFA2C74DFF27F94BFE46F925" blockId="22.[139,772,1673,1902]" pageId="22" pageNumber="81">
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<emphasis id="B946EAA5FFA2C74DFF27F94BFEA2F952" box="[140,314,1673,1692]" italics="true" pageId="22" pageNumber="81">Material examined</emphasis>
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.
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<materialsCitation id="3B5A3CEAFFA2C74DFEEDF94BFE46F925" collectingDate="1993-09-28" collectionCode="ANARE, T, NMV" collectorName="Southern Ocean, S" country="Heard Island and McDonald Islands" elevation="521" location="Southern Ocean" municipality="Stranks" pageId="22" pageNumber="81" specimenCode="F84986" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54898815FFA2C74DFEEDF94BFE05F952" box="[326,413,1673,1692]" pageId="22" pageNumber="81" type="holotype">Holotype</typeStatus>
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.
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<collectorName id="26C75361FFA2C74DFE0DF94BFDC8F952" box="[422,592,1673,1692]" pageId="22" pageNumber="81">
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<location id="8EED606CFFA2C74DFE0DF94BFDA3F952" LSID="urn:lsid:plazi:treatment:039B87A1FFA2C74DFF27FA16FC71F8A1:8EED606CFFA2C74DFE0DF94BFDA3F952" box="[422,571,1673,1692]" country="Heard Island and McDonald Islands" municipality="Stranks" name="Southern Ocean" pageId="22" pageNumber="81">Southern Ocean</location>
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, S
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</collectorName>
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Kerguelen Plateau, NE
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<collectingCountry id="F3257627FFA2C74DFF04F961FEBDF978" box="[175,293,1699,1718]" name="Heard Island and McDonald Islands" pageId="22" pageNumber="81">Heard Island</collectingCountry>
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,
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<locationDeviation id="9A3F2F95FFA2C74DFE86F966FE48F978" box="[301,464,1699,1719]" location="Shell Bank" pageId="22" pageNumber="81">
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NW of
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<emphasis id="B946EAA5FFA2C74DFEC4F961FE48F978" box="[367,464,1699,1718]" italics="true" pageId="22" pageNumber="81">Shell Bank</emphasis>
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</locationDeviation>
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,
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<collectionCode id="ED23AE72FFA2C74DFE70F961FDB2F978" box="[475,554,1699,1718]" pageId="22" pageNumber="81">ANARE</collectionCode>
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<taxonomicName id="4C324D34FFA2C74DFD85F966FC9BF979" authority=", HRD" authorityName="HRD" box="[558,771,1699,1719]" class="Holothuroidea" family="Cucumariidae" genus="Aurora" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="species" species="australis">
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<emphasis id="B946EAA5FFA2C74DFD85F966FD5FF978" box="[558,711,1699,1719]" italics="true" pageId="22" pageNumber="81">Aurora Australis</emphasis>
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, HRD
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</taxonomicName>
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93, stn. 71–S, epibenthic sled, -50.72 75.07,
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<quantity id="4CCA9B52FFA2C74DFDEAF97FFD35F91F" box="[577,685,1725,1745]" metricMagnitude="2" metricUnit="m" metricValue="5.21" metricValueMax="5.279999999999999" metricValueMin="5.14" pageId="22" pageNumber="81" unit="m" value="521.0" valueMax="528.0" valueMin="514.0">
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<elevation id="001FD184FFA2C74DFDEAF97FFD35F91F" box="[577,685,1725,1745]" metricMagnitude="2" metricUnit="m" metricValue="5.21" metricValueMax="5.279999999999999" metricValueMin="5.14" pageId="22" pageNumber="81" unit="m" value="521.0" valueMax="528.0" valueMin="514.0">514–528 m</elevation>
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</quantity>
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, coll.
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<collectionCode id="ED23AE72FFA2C74DFD44F97CFD66F91F" box="[751,766,1726,1745]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="81">T</collectionCode>
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.
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<collectingMunicipality id="6BE9ACCDFFA2C74DFF27F91AFF4CF925" box="[140,212,1752,1771]" pageId="22" pageNumber="81">Stranks</collectingMunicipality>
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,
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<date id="FF8C1077FFA2C74DFF77F91AFED6F925" box="[220,334,1752,1771]" pageId="22" pageNumber="81" value="1993-09-28">
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<collectingDate id="EFC8E99FFFA2C74DFF77F91AFED6F925" box="[220,334,1752,1771]" pageId="22" pageNumber="81" value="1993-09-28">28 Sep 1993</collectingDate>
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</date>
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,
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<collectionCode id="ED23AE72FFA2C74DFEFCF91AFE15F925" box="[343,397,1752,1771]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="22" pageNumber="81" type="Museum">NMV</collectionCode>
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<specimenCode id="DB949ECCFFA2C74DFE39F91AFE42F925" box="[402,474,1752,1771]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="22" pageNumber="81" type="Museum">F84986</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B8D36B7FFA2C74DFF19F930FD5AF8CB" blockId="22.[139,772,1673,1902]" box="[178,706,1778,1797]" pageId="22" pageNumber="81">
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<typeStatus id="54898815FFA2C74DFF19F930FE96F8CB" box="[178,270,1778,1797]" pageId="22" pageNumber="81" type="paratype">Paratypes</typeStatus>
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.
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<materialsCitation id="3B5A3CEAFFA2C74DFEBDF930FD26F8CB" box="[278,702,1778,1797]" collectionCode="NMV" pageId="22" pageNumber="81" specimenCode="F198493" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54898815FFA2C74DFEBDF930FEF2F8CB" box="[278,362,1778,1797]" pageId="22" pageNumber="81" type="holotype">Holotype</typeStatus>
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locality and date,
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<collectionCode id="ED23AE72FFA2C74DFDB9F930FDD0F8CB" box="[530,584,1778,1797]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="22" pageNumber="81" type="Museum">NMV</collectionCode>
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<specimenCode id="DB949ECCFFA2C74DFDE5F930FD04F8CB" box="[590,668,1778,1797]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="22" pageNumber="81" type="Museum">F198493</specimenCode>
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(7)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8D36B7FFA2C74DFF19F8CEFA0DFDB9" blockId="22.[139,772,1673,1902]" lastBlockId="22.[807,1442,229,1186]" pageId="22" pageNumber="81">
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Other material (AAD species code: HOL 23). Southern
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<emphasis id="B946EAA5FFA2C74DFD7EF8CEFF23F8F7" italics="true" pageId="22" pageNumber="81">Shell Bank</emphasis>
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, AAD
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<emphasis id="B946EAA5FFA2C74DFF51F8E4FE28F8F7" box="[250,432,1830,1849]" italics="true" pageId="22" pageNumber="81">Southern Champion</emphasis>
|
|||
|
|
cruise 46 haul 111, beam trawl, -51.81 76.31,
|
|||
|
|
<quantity id="4CCA9B52FFA2C74DFF6FF883FF66F89A" box="[196,254,1857,1876]" metricMagnitude="2" metricUnit="m" metricValue="2.4699999999999998" pageId="22" pageNumber="81" unit="m" value="247.0">247 m</quantity>
|
|||
|
|
, AAD; southern
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFE3EF883FE6EF89A" box="[405,502,1857,1876]" italics="true" pageId="22" pageNumber="81">Shell Bank</emphasis>
|
|||
|
|
, SC 46(114), -51.81 76.06,
|
|||
|
|
<quantity id="4CCA9B52FFA2C74DFD48F883FF07F8A0" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="22" pageNumber="81" unit="m" value="410.0">410 m</quantity>
|
|||
|
|
, AAD; southern
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFE93F899FE02F8A0" box="[312,410,1883,1902]" italics="true" pageId="22" pageNumber="81">Shell Bank</emphasis>
|
|||
|
|
, SC 46(115), -51.81 75.98,
|
|||
|
|
<quantity id="4CCA9B52FFA2C74DFD21F899FD5BF8A0" box="[650,707,1883,1902]" metricMagnitude="2" metricUnit="m" metricValue="5.57" pageId="22" pageNumber="81" unit="m" value="557.0">557 m</quantity>
|
|||
|
|
, AAD.
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFC82FF24FC3BFF35" box="[809,931,230,251]" italics="true" pageId="22" pageNumber="81">Description</emphasis>
|
|||
|
|
. Preserved body up to
|
|||
|
|
<quantity id="4CCA9B52FFA2C74DFB0CFF27FB6BFF35" box="[1191,1267,229,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="81" unit="mm" value="12.0">12 mm</quantity>
|
|||
|
|
long (excluding tentacles), body up to
|
|||
|
|
<quantity id="4CCA9B52FFA2C74DFBA4FEC0FBD4FED6" box="[1039,1100,258,280]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="81" unit="mm" value="5.0">5 mm</quantity>
|
|||
|
|
high, body flat ventrally, domed dorsally, lacking oral and anal cones, mouth with anterior orientation, dorsal and lateral body with conspicuous imbricating scales, lacking tube feet, ventro-lateral scales projecting slightly over a soft sole. Discrete sole, lacking scales, peripheral series of large tube feet, series not continuous anteriorly, rare mid-ventral tube feet. Tentacles dendritic, 10, ventral pair small. Calcareous ring solid, cucumariid-like, digitiform anterior projection and deep posterior notch on both radial and inter-radial plates, lacking posterior prolongations. Longitudinal muscles narrow, flat. Gonad comprises 2 tufts of sac-like tubules, one on each side of the dorsal mesentery, hermaphrodite, coelomic bood-protecting.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8D36B7FFA2C74DFCE4FDBCFBD6FB6C" blockId="22.[807,1442,229,1186]" pageId="22" pageNumber="81">
|
|||
|
|
Dorsal and lateral ossicles imbricating scales and cups; scales irregularly oval, up to about
|
|||
|
|
<quantity id="4CCA9B52FFA2C74DFB04FD59FB77FD7F" box="[1199,1263,667,689]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="81" unit="mm" value="1.0">1 mm</quantity>
|
|||
|
|
long, scales are perforated plates with mesh-like secondary layering, finely spinous dorsal surface layer; cups irregular, shallow to deep, predominantly four perforations in cross form, sometimes small corner perforations, cups variably knobbed or with short digitiform projections, some incipiently bridged, cups typically 140
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFC60FC89FC41FCAE" box="[971,985,843,864]" italics="true" pageId="22" pageNumber="81">µ</emphasis>
|
|||
|
|
m long. Sole with irregular, marginally knobbed, perforated, shallow, concave plates, up to about 8 perforations, up to about 360
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFBFCFC44FBFDFC55" box="[1111,1125,902,923]" italics="true" pageId="22" pageNumber="81">µ</emphasis>
|
|||
|
|
m long; plates inter-grade with shallow cups, similar to dorsal cups, predominantly 4 large perforations, sometimes smaller corner ones, margin knobbed, some secondary knobs and developments, cups about 140
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFC30FC39FC31FBDE" box="[923,937,1019,1040]" italics="true" pageId="22" pageNumber="81">µ</emphasis>
|
|||
|
|
m long. Tentacle ossicles thick, perforated rods and plates, irregular, curved, variably knobbed and digitate margins, some surface knobs, plates and rods up to 320
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFA2BFBF7FA14FB84" box="[1408,1420,1077,1098]" italics="true" pageId="22" pageNumber="81">µ</emphasis>
|
|||
|
|
m long; small thin curved plates, about 50
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFB75FB90FB74FBA9" box="[1246,1260,1106,1127]" italics="true" pageId="22" pageNumber="81">µ</emphasis>
|
|||
|
|
m long; rare fine irregular crosses, about 50
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFBC0FBADFBE1FB4A" box="[1131,1145,1135,1156]" italics="true" pageId="22" pageNumber="81">µ</emphasis>
|
|||
|
|
m long; small rosette-like ossicles, about 100
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFC59FB4FFB98FB6C" box="[1010,1024,1165,1186]" italics="true" pageId="22" pageNumber="81">µ</emphasis>
|
|||
|
|
m long.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8D36B7FFA2C74DFC82FB7AFBFFFB25" blockId="22.[809,1440,1208,1259]" pageId="22" pageNumber="81">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFC82FB7AFC30FB03" box="[809,936,1208,1229]" italics="true" pageId="22" pageNumber="81">Distribution</emphasis>
|
|||
|
|
. Southern Ocean, Kerguelen Plateau, NE
|
|||
|
|
<collectingCountry id="F3257627FFA2C74DFAC9FB7AFCF3FB25" name="Heard Island and McDonald Islands" pageId="22" pageNumber="81">Heard Island</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFCDFFB17FC7BFB24" box="[884,995,1237,1258]" italics="true" pageId="22" pageNumber="81">Shell Bank</emphasis>
|
|||
|
|
,
|
|||
|
|
<quantity id="4CCA9B52FFA2C74DFC5AFB17FBFBFB25" box="[1009,1123,1237,1259]" metricMagnitude="2" metricUnit="m" metricValue="4.02" metricValueMax="5.57" metricValueMin="2.4699999999999998" pageId="22" pageNumber="81" unit="m" value="402.0" valueMax="557.0" valueMin="247.0">247–557 m</quantity>
|
|||
|
|
,
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8D36B7FFA2C74DFC82FAC0FAE2FAD6" blockId="22.[809,1402,1282,1304]" box="[809,1402,1282,1304]" pageId="22" pageNumber="81">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFC82FAC0FC0FFAD9" box="[809,919,1282,1303]" italics="true" pageId="22" pageNumber="81">Etymology</emphasis>
|
|||
|
|
. Named
|
|||
|
|
<taxonomicName id="4C324D34FFA2C74DFC5FFAC0FBAFFAD9" authorityName="O'Loughlin & Skarbnik-Lopez" authorityYear="2015" box="[1012,1079,1282,1303]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="species" species="heardi">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFC5FFAC0FBAFFAD9" box="[1012,1079,1282,1303]" italics="true" pageId="22" pageNumber="81">heardi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
with reference to
|
|||
|
|
<collectingCountry id="F3257627FFA2C74DFB5BFAC0FAEFFAD6" box="[1264,1399,1282,1304]" name="Heard Island and McDonald Islands" pageId="22" pageNumber="81">Heard Island</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8D36B7FFA2C74DFC82FAECFC71F8A1" blockId="22.[809,1441,1326,1903]" pageId="22" pageNumber="81">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFC82FAECFC1BFA8D" box="[809,899,1326,1347]" italics="true" pageId="22" pageNumber="81">Remarks</emphasis>
|
|||
|
|
. We provisionally assign this new species to the currently suppressed
|
|||
|
|
<taxonomicName id="4C324D34FFA2C74DFBA8FA89FB5AFAAF" authority="Oken, 1815" authorityName="Oken" authorityYear="1815" box="[1027,1218,1355,1377]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="genus">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFBA8FA89FBDFFAAE" box="[1027,1095,1355,1376]" italics="true" pageId="22" pageNumber="81">Psolus</emphasis>
|
|||
|
|
<bibRefCitation id="EFA34B46FFA2C74DFBE6FA89FB5AFAAF" author="Oken, L." box="[1101,1218,1355,1377]" pageId="22" pageNumber="81" refId="ref19431" refString="Oken, L. 1815. Okens Lehrbuch der Naturgeschichte. Teil 3. Zoologie. 850 pp. C. H. Reclam, Jena." type="book" year="1815">Oken, 1815</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, pending an appeal to the ICZN (see Introduction). While
|
|||
|
|
<taxonomicName id="4C324D34FFA2C74DFB3EFAAAFC76FA52" authority="O'Loughlin & Skarbnik-Lopez" authorityName="O'Loughlin & Skarbnik-Lopez" authorityYear="2015" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="species" species="heardi" status="sp. nov.">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFB3EFAAAFAB9FAB3" box="[1173,1313,1384,1405]" italics="true" pageId="22" pageNumber="81">Psolus heardi</emphasis>
|
|||
|
|
O’Loughlin & Skarbnik-López
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A27557DEFFA2C74DFC51FA44FBD5FA55" box="[1018,1101,1414,1435]" pageId="22" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
satisfies the general diagnostic criteria for referral to
|
|||
|
|
<taxonomicName id="4C324D34FFA2C74DFBB6FA61FBF9FA76" authorityName="Oken" authorityYear="1815" box="[1053,1121,1443,1464]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="genus">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFBB6FA61FBF9FA76" box="[1053,1121,1443,1464]" italics="true" pageId="22" pageNumber="81">Psolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, we are aware that emerging phylogenetic data do not support the family
|
|||
|
|
<taxonomicName id="4C324D34FFA2C74DFAB3FA02FAF7FA18" authorityName="Burmeister" authorityYear="1837" box="[1304,1391,1472,1494]" class="Holothuroidea" family="Psolidae" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="family">Psolidae</taxonomicName>
|
|||
|
|
and provide evidence that the genera
|
|||
|
|
<taxonomicName id="4C324D34FFA2C74DFB29FA1FFA9CFA3D" authority="Oken" authorityName="Oken" authorityYear="1815" box="[1154,1284,1501,1523]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="genus">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFB29FA1FFB5EFA3C" box="[1154,1222,1501,1522]" italics="true" pageId="22" pageNumber="81">Psolus</emphasis>
|
|||
|
|
Oken
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C324D34FFA2C74DFA93FA1FFC2FF9DE" authority="Ludwig, 1887" authorityName="Ludwig" authorityYear="1887" class="Holothuroidea" family="Psolidae" genus="Psolidium" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="genus">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFA93FA1FFA39FA3C" box="[1336,1441,1501,1522]" italics="true" pageId="22" pageNumber="81">Psolidium</emphasis>
|
|||
|
|
<bibRefCitation id="EFA34B46FFA2C74DFC82FA38FC2FF9DE" author="Ludwig, H." box="[809,951,1530,1552]" pageId="22" pageNumber="81" pagination="1 - 36" refId="ref19018" refString="Ludwig, H. 1887. Die von G. Chierchia auf der Fahrt der Kgl. Italianische Corvette Vettor Pisani gesammelten Holothurien. Zoologische Jahrbucher 2: 1 - 36, 2 pls." type="journal article" year="1887">Ludwig, 1887</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are polyphyletic (Gustav Paulay pers. comm.).
|
|||
|
|
<taxonomicName id="4C324D34FFA2C74DFC82F9DAFA99F9E0" authority="O'Loughlin & Skarbnik-Lopez" authorityName="O'Loughlin & Skarbnik-Lopez" authorityYear="2015" box="[809,1281,1560,1582]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="81" phylum="Echinodermata" rank="species" species="heardi" status="sp. nov.">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFC82F9DAFC21F9E3" box="[809,953,1560,1581]" italics="true" pageId="22" pageNumber="81">Psolus heardi</emphasis>
|
|||
|
|
O’Loughlin & Skarbnik-López
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A27557DEFFA2C74DFAACF9DAFACEF9E3" box="[1287,1366,1560,1581]" pageId="22" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
differs from all other
|
|||
|
|
<taxonomicName id="4C324D34FFA2C74DFC60F9F7FBFDF985" box="[971,1125,1589,1611]" pageId="22" pageNumber="81">
|
|||
|
|
<emphasis id="B946EAA5FFA2C74DFC60F9F7FB97F984" box="[971,1039,1589,1610]" italics="true" pageId="22" pageNumber="81">Psolus</emphasis>
|
|||
|
|
species
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having a combination of: coelomic brood-protecting habit; poorly defined ventrolateral margin overlapping the sole; dorsal scales that are single-layered with mesh-like secondary layering that has fine dorsal surface spines; shallow, irregular, marginally knobbed cups that are similar dorsally and in sole. “Other material” refers to lots that were identified in the AAD by comparison with voucher specimens that were identified by Mark O’Loughlin. These lots are held (unregistered) in the AAD and the determinations not confirmed by Mark O’Loughlin.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|