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<document id="DE1AD6FCF3187BCBE789DDAFD6DC052A" ID-DOI="10.5281/zenodo.274226" ID-GBIF-Dataset="f1310289-6972-4b5a-b391-92feea2e1434" ID-ISSN="1175-5326" ID-Zenodo-Dep="274226" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459838000819" checkinUser="plazi" docAuthor="Schöning, Caspar, Gotwald, William H., Kronauer, Daniel J. C. &amp; Vilhelmsen, Lars" docDate="2008" docId="B32C87BAFFF0DC38FF72F8FB999FFD59" docLanguage="en" docName="zt01749p052.pdf" docOrigin="Zootaxa 1749" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Dorylus gribodoi Emery 1892" docType="treatment" docVersion="10" lastPageNumber="48" masterDocId="4F15FFC2FFF3DC32FFE5FFFD997CFFBB" masterDocTitle="Taxonomy of the African army ant Dorylus gribodoi Emery, 1892 (Hymenoptera, Formicidae) — new insights from DNA sequence data and morphology" masterLastPageNumber="52" masterPageNumber="39" pageNumber="42" updateTime="1698223933840" updateUser="plazi">
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<mods:title id="DBDC6CF3ECAFBD594B9FFA95F911F24A">Taxonomy of the African army ant Dorylus gribodoi Emery, 1892 (Hymenoptera, Formicidae) — new insights from DNA sequence data and morphology</mods:title>
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<treatment id="B32C87BAFFF0DC38FF72F8FB999FFD59" ID-DOI="http://doi.org/10.5281/zenodo.3510522" ID-GBIF-Taxon="119359463" ID-Zenodo-Dep="3510522" LSID="urn:lsid:plazi:treatment:B32C87BAFFF0DC38FF72F8FB999FFD59" httpUri="http://treatment.plazi.org/id/B32C87BAFFF0DC38FF72F8FB999FFD59" lastPageId="10" lastPageNumber="48" pageId="3" pageNumber="42">
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<paragraph id="3B3A36ACFFF0DC31FF72F8FB9B7DF89B" blockId="3.[151,513,1798,1860]" box="[151,513,1798,1824]" pageId="3" pageNumber="42">
<heading id="607281C0FFF0DC31FF72F8FB9B7DF89B" bold="true" box="[151,513,1798,1824]" fontSize="11" level="1" pageId="3" pageNumber="42" reason="1">
<taxonomicName id="FC854D2FFFF0DC31FF72F8FB9B7DF89B" ID-CoL="37FCF" authority="Emery 1892" authorityName="Emery" authorityYear="1892" box="[151,513,1798,1824]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFF0DC31FF72F8FB9B7DF89B" bold="true" box="[151,513,1798,1824]" pageId="3" pageNumber="42">
<emphasis id="09F1EABEFFF0DC31FF72F8FB981FF89B" bold="true" box="[151,355,1798,1824]" italics="true" pageId="3" pageNumber="42">Dorylus gribodoi</emphasis>
<bibRefCitation id="5F144B5DFFF0DC31FE8FF8FB9B7DF89B" author="Emery" box="[362,513,1798,1824]" pageId="3" pageNumber="51" refString="Emery, C. (1892) Voyage de M. Ch. Alluaud dans le territoire d'Assinie (Afrique occidentale) en juillet et aout 1886. Formicides. Annales de la Societe Entomologique de France, 60, 553 - 574." type="journal article" year="1892">Emery 1892</bibRefCitation>
</emphasis>
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</paragraph>
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(
<figureCitation id="A3BE2A29FFF0DC31FF7AF8D79852F8FF" box="[159,302,1834,1860]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="4.[151,255,1561,1585]" captionStartId-1="5.[151,255,1113,1137]" captionStartId-2="5.[151,255,1859,1883]" captionStartId-3="6.[151,255,1015,1039]" captionTargetBox-0="[253,1337,700,1534]" captionTargetBox-1="[238,1338,183,1113]" captionTargetBox-2="[247,1337,1174,1859]" captionTargetBox-3="[264,1340,178,996]" captionTargetId-0="figure@4.[253,1338,696,1534]" captionTargetId-1="figure@5.[238,1339,179,1113]" captionTargetId-2="figure@5.[247,1340,1145,1875]" captionTargetId-3="figure@6.[238,1343,168,998]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 2. Lateral overview of a Dorylus gribodoi male from Taï, Ivory Coast." captionText-1="FIGURE 3. Frontal view of the head of a Dorylus gribodoi male from Taï, Ivory Coast." captionText-2="FIGURE 4. Dorylus gribodoi male: genital capsule and subgenital plate." captionText-3="FIGURE 5. Lateral overview of a large Dorylus gribodoi worker from Taï, Ivory Coast." httpUri-0="https://zenodo.org/record/274228/files/figure.png" httpUri-1="https://zenodo.org/record/274229/files/figure.png" httpUri-2="https://zenodo.org/record/274230/files/figure.png" httpUri-3="https://zenodo.org/record/274231/files/figure.png" pageId="3" pageNumber="42">Figures 2–5</figureCitation>
, 
<figureCitation id="A3BE2A29FFF0DC31FEDFF8D79837F8FF" box="[314,331,1834,1860]" captionStart="FIGURE 7" captionStartId="7.[151,255,763,787]" captionTargetBox="[468,1117,258,737]" captionTargetId="figure@7.[468,1118,250,738]" captionTargetPageId="7" captionText="FIGURE 7. Lateral view of the head of a large D. gribodoi worker (HW 2.79 mm), Taï, Ivory Coast." httpUri="https://zenodo.org/record/274233/files/figure.png" pageId="3" pageNumber="42">7</figureCitation>
, 
<figureCitation id="A3BE2A29FFF0DC31FEBDF8D79815F8FF" box="[344,361,1834,1860]" captionStart="FIGURE 9" captionStartId="8.[151,255,1031,1055]" captionTargetBox="[385,1200,354,1006]" captionTargetId="figure@8.[385,1202,354,1007]" captionTargetPageId="8" captionText="FIGURE 9. Dorsal view of the petiole of a large D. gribodoi worker (HW 2.85 mm) from Taï, Ivory Coast." httpUri="https://zenodo.org/record/274235/files/figure.png" pageId="3" pageNumber="42">9</figureCitation>
, 
<figureCitation id="A3BE2A29FFF0DC31FE90F8D798C2F8FF" box="[373,446,1834,1860]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="9.[151,255,1650,1674]" captionStartId-1="10.[151,255,1514,1538]" captionStartId-2="11.[151,255,913,937]" captionTargetBox-0="[389,1198,978,1626]" captionTargetBox-2="[363,1229,194,892]" captionTargetId-0="figure@9.[389,1198,978,1626]" captionTargetId-2="figure@11.[363,1231,194,892]" captionTargetPageId-0="9" captionTargetPageId-2="11" captionText-0="FIGURE 11. Lateral overview of the D. gribodoi queen collected from a nest at Lamto, Ivory Coast." captionText-1="FIGURE 12. Frontal view of the head of the D. gribodoi queen (HW 4.58 mm)." captionText-2="FIGURE 13. Dorsal view of the hypopygium of the D. gribodoi queen (HAD 2.53 mm)." httpUri-0="https://zenodo.org/record/274237/files/figure.png" httpUri-2="https://zenodo.org/record/274238/files/figure.png" pageId="3" pageNumber="42">11–13</figureCitation>
)
</paragraph>
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<paragraph id="3B3A36ACFFF0DC31FF72F887983CF84B" blockId="3.[151,1436,1913,2033]" pageId="3" pageNumber="42">
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<taxonomicName id="FC854D2FFFF0DC31FF72F8879B68F82B" ID-CoL="37FCF" authority="Emery 1892: 570" authorityName="Emery" authorityPageNumber="570" authorityYear="1892" box="[151,532,1913,1937]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFF0DC31FF72F887982CF82A" box="[151,336,1914,1937]" italics="true" pageId="3" pageNumber="42">Dorylus gribodoi</emphasis>
<treatmentCitation id="BA2410BDFFF0DC31FEB2F8849B68F82B" author="Emery" box="[343,532,1913,1936]" page="570" pageId="3" pageNumber="42" year="1892">
<bibRefCitation id="5F144B5DFFF0DC31FEB2F88498A2F82B" author="Emery" box="[343,478,1913,1936]" pageId="3" pageNumber="51" refString="Emery, C. (1892) Voyage de M. Ch. Alluaud dans le territoire d'Assinie (Afrique occidentale) en juillet et aout 1886. Formicides. Annales de la Societe Entomologique de France, 60, 553 - 574." type="journal article" year="1892">Emery 1892</bibRefCitation>
: 570
</treatmentCitation>
</taxonomicName>
. Two 
<typeStatus id="E43E880EFFF0DC31FDB0F8849BD4F82B" box="[597,680,1913,1936]" pageId="3" pageNumber="42" type="syntype">syntype</typeStatus>
males, West Africa [
<bibRefCitation id="5F144B5DFFF0DC31FC6DF8849D49F82B" author="Wheeler" box="[904,1077,1913,1936]" pageId="3" pageNumber="52" refString="Wheeler, W. M. (1922) Ants of the American Museum Congo Expedition. A Contribution to the Myrmecology of Africa. Bulletin of the American Museum of Natural History, 45, 1 - 1139." type="journal article" year="1922">Wheeler (1922)</bibRefCitation>
and 
<bibRefCitation id="5F144B5DFFF0DC31FB8CF8849C7CF82B" author="Bolton" box="[1129,1280,1913,1936]" pageId="3" pageNumber="51" refString="Bolton, B. (1995) A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, Massachusetts, 504 pp." type="book" year="1995">Bolton (1995)</bibRefCitation>
listed 
<collectingCountry id="4392763CFFF0DC31FAADF8849C02F82B" box="[1352,1406,1913,1936]" name="Togo" pageId="3" pageNumber="42">Togo</collectingCountry>
as country of the 
<typeStatus id="E43E880EFFF0DC31FE86F86498EDF80B" box="[355,401,1945,1968]" pageId="3" pageNumber="42">type</typeStatus>
location]: Amu, collected by M. Gribodo, MCSN (examined). 
<taxonomicName id="FC854D2FFFF0DC31FF72F847989FF86A" box="[151,483,1977,2001]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="gerstackeri" subGenus="Anomma">
<emphasis id="09F1EABEFFF0DC31FF72F8479991F86A" box="[151,237,1978,2001]" italics="true" pageId="3" pageNumber="42">Dorylus</emphasis>
(
<emphasis id="09F1EABEFFF0DC31FF19F8479827F86A" box="[252,347,1978,2001]" italics="true" pageId="3" pageNumber="42">Anomma</emphasis>
) 
<emphasis id="09F1EABEFFF0DC31FE8EF847989FF86A" box="[363,483,1978,2001]" italics="true" pageId="3" pageNumber="42">gerstäckeri</emphasis>
</taxonomicName>
(sic!) 
<treatmentCitation id="BA2410BDFFF0DC31FDCDF8449B9EF86B" author="Emery" box="[552,738,1977,2000]" page="713" pageId="3" pageNumber="42" year="1895">
<bibRefCitation id="5F144B5DFFF0DC31FDCDF8449BD3F86B" author="Emery" box="[552,687,1977,2000]" pageId="3" pageNumber="51" refString="Emery, C. (1895) Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbucher Abteilung fur Systematik Okologie und Geographie der Tiere, 8, 685 - 778." type="journal article" year="1895">Emery 1895</bibRefCitation>
: 713
</treatmentCitation>
– 714. 
<typeStatus id="E43E880EFFF0DC31FCD7F8449AE9F86B" box="[818,917,1977,2000]" pageId="3" pageNumber="42" type="holotype">Holotype</typeStatus>
worker, 
<collectingCountry id="4392763CFFF0DC31FC11F8449D42F86B" box="[1012,1086,1977,2000]" name="Ghana" pageId="3" pageNumber="42">Ghana</collectingCountry>
: Accra, MCSN (examined), 
<emphasis id="09F1EABEFFF0DC31FA8BF8449845F84A" bold="true" pageId="3" pageNumber="42">new synonomy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
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<taxonomicName id="FC854D2FFFF7DC36FF72FF6598C1FF14" ID-CoL="37FCF" box="[151,445,152,175]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="gribodoi" subGenus="Dorylus">
<emphasis id="09F1EABEFFF7DC36FF72FF659991FF14" box="[151,237,152,175]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
(
<emphasis id="09F1EABEFFF7DC36FF19FF65982EFF14" box="[252,338,152,175]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
) 
<emphasis id="09F1EABEFFF7DC36FE84FF6598C1FF14" box="[353,445,152,175]" italics="true" pageId="4" pageNumber="43">gribodoi</emphasis>
</taxonomicName>
; 
<treatmentCitation id="BA2410BDFFF7DC36FE2EFF659BF8FF14" author="Emery" box="[459,644,152,175]" page="723" pageId="4" pageNumber="43" year="1895">
<bibRefCitation id="5F144B5DFFF7DC36FE2EFF659B2EFF14" author="Emery" box="[459,594,152,175]" pageId="4" pageNumber="51" refString="Emery, C. (1895) Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbucher Abteilung fur Systematik Okologie und Geographie der Tiere, 8, 685 - 778." type="journal article" year="1895">Emery 1895</bibRefCitation>
: 723
</treatmentCitation>
(combination in subgenus 
<taxonomicName id="FC854D2FFFF7DC36FC41FF659A86FF14" ID-CoL="47Y3" box="[932,1018,152,175]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="09F1EABEFFF7DC36FC41FF659A86FF14" box="[932,1018,152,175]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
</taxonomicName>
), 
<treatmentCitation id="BA2410BDFFF7DC36FBF5FF659DC2FF14" author="Emery" box="[1040,1214,152,175]" page="10" pageId="4" pageNumber="43" year="1910">
<bibRefCitation id="5F144B5DFFF7DC36FBF5FF659DEBFF14" author="Emery" box="[1040,1175,152,175]" pageId="4" pageNumber="51" refString="Emery, C. (1910) Hymenoptera. Fam. Formicidae. Subfam. Dorylinae. Genera Insectorum, 102, 1 - 35." type="journal article" year="1910">Emery 1910</bibRefCitation>
: 10
</treatmentCitation>
, 
<treatmentCitation id="BA2410BDFFF7DC36FB2CFF659CE4FF14" author="Wheeler" box="[1225,1432,152,175]" page="732" pageId="4" pageNumber="43" year="1922">
<bibRefCitation id="5F144B5DFFF7DC36FB2CFF659C1FFF14" author="Wheeler" box="[1225,1379,152,175]" pageId="4" pageNumber="52" refString="Wheeler, W. M. (1922) Ants of the American Museum Congo Expedition. A Contribution to the Myrmecology of Africa. Bulletin of the American Museum of Natural History, 45, 1 - 1139." type="journal article" year="1922">Wheeler 1922</bibRefCitation>
: 732
</treatmentCitation>
, 
<treatmentCitation id="BA2410BDFFF7DC36FF22FF4598EEFF74" ID-DOI="http://doi.org/10.5281/zenodo.3504511" ID-Zenodo-Dep="3504511" author="Bernard" box="[199,402,184,207]" httpUri="http://treatment.plazi.org/id/25BCAF19A5D5D7DEFF28706BE00A3DD5" page="217" pageId="4" pageNumber="43" year="1953">
<bibRefCitation id="5F144B5DFFF7DC36FF22FF459820FF74" author="Bernard" box="[199,348,184,207]" pageId="4" pageNumber="51" refString="Bernard, F. (1953) La reserve naturelle integrale du Mt Nimba. XI. Hymenopteres, Formicidae. Memoires de l'Institut Francais d'Afrique Noire, 19, 165 - 270." type="journal article" year="1953">Bernard 1953</bibRefCitation>
: 217
</treatmentCitation>
, 
<treatmentCitation id="BA2410BDFFF7DC36FE79FF459B25FF74" author="Bolton" box="[412,601,184,207]" page="179" pageId="4" pageNumber="43" year="1995">
<bibRefCitation id="5F144B5DFFF7DC36FE79FF459B58FF74" author="Bolton" box="[412,548,184,207]" pageId="4" pageNumber="51" refString="Bolton, B. (1995) A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, Massachusetts, 504 pp." type="book" year="1995">Bolton 1995</bibRefCitation>
: 179
</treatmentCitation>
, 
<treatmentCitation id="BA2410BDFFF7DC36FD81FF459A16FF74" author="Kronauer" box="[612,874,184,207]" page="56" pageId="4" pageNumber="43" year="2007">
<bibRefCitation id="5F144B5DFFF7DC36FD81FF459A38FF74" author="Kronauer" box="[612,836,184,207]" pageId="4" pageNumber="52" refString="Kronauer, D. J. C., Schoning, C., Vilhelmsen, L. B. &amp; Boomsma, J. J. (2007) A molecular phylogeny of Dorylus army ants provides evidence for multiple evolutionary transitions in foraging niche. BMC Evolutionary Biology, 7, 56." type="journal article" year="2007">
Kronauer 
<emphasis id="09F1EABEFFF7DC36FD2AFF459A79FF74" box="[719,773,184,207]" italics="true" pageId="4" pageNumber="43">et al.</emphasis>
2007
</bibRefCitation>
: 56
</treatmentCitation>
(examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="3B3A36ACFFF7DC36FF72FF259DE6FEF4" blockId="4.[151,1436,152,655]" pageId="4" pageNumber="43">
<treatmentCitationGroup id="1B951182FFF7DC36FF72FF259DE6FEF4" pageId="4" pageNumber="43">
<taxonomicName id="FC854D2FFFF7DC36FF72FF259889FF54" ID-CoL="37FCD" box="[151,501,216,239]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="gerstaeckeri" subGenus="Anomma">
<emphasis id="09F1EABEFFF7DC36FF72FF259991FF54" box="[151,237,216,239]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
(
<emphasis id="09F1EABEFFF7DC36FF1AFF259822FF54" box="[255,350,216,239]" italics="true" pageId="4" pageNumber="43">Anomma</emphasis>
) 
<emphasis id="09F1EABEFFF7DC36FE94FF259889FF54" box="[369,501,216,239]" italics="true" pageId="4" pageNumber="43">gerstaeckeri</emphasis>
</taxonomicName>
; 
<treatmentCitation id="BA2410BDFFF7DC36FDE3FF259BB9FF54" author="Emery" box="[518,709,216,239]" page="415" pageId="4" pageNumber="43" year="1901">
<bibRefCitation id="5F144B5DFFF7DC36FDE3FF259BECFF54" author="Emery" box="[518,656,216,239]" pageId="4" pageNumber="51" refString="Emery, C. (1901) Studi sul polimorfismo e la metamorfosi nel genere Dorylus. Memorie della R. Accademia delle Scienze dell ' Istituto di Bologna, 9, 183 - 201." type="journal article" year="1901">Emery 1901</bibRefCitation>
: 415
</treatmentCitation>
– 436, 
<treatmentCitation id="BA2410BDFFF7DC36FCFBFF259AADFF54" author="Emery" box="[798,977,216,239]" page="11" pageId="4" pageNumber="43" year="1910">
<bibRefCitation id="5F144B5DFFF7DC36FCFBFF259AD4FF54" author="Emery" box="[798,936,216,239]" pageId="4" pageNumber="51" refString="Emery, C. (1910) Hymenoptera. Fam. Formicidae. Subfam. Dorylinae. Genera Insectorum, 102, 1 - 35." type="journal article" year="1910">Emery 1910</bibRefCitation>
: 11
</treatmentCitation>
, 
<treatmentCitation id="BA2410BDFFF7DC36FC05FF259DC8FF54" author="Santschi" box="[992,1204,216,239]" page="154" pageId="4" pageNumber="43" year="1912">
<bibRefCitation id="5F144B5DFFF7DC36FC05FF259D01FF54" author="Santschi" box="[992,1149,216,239]" pageId="4" pageNumber="52" refString="Santschi, F. (1912) Fourmis d'Afrique et de Madagascar. Annales de la Societe Entomologique de Belgique, 56, 150 - 167." type="journal article" year="1912">Santschi 1912</bibRefCitation>
: 154
</treatmentCitation>
, Santschi 1914: 332, 
<treatmentCitation id="BA2410BDFFF7DC36FF22FF0598E6FEB4" author="Wheeler" box="[199,410,248,271]" page="736" pageId="4" pageNumber="43" year="1922">
<bibRefCitation id="5F144B5DFFF7DC36FF22FF05981FFEB4" author="Wheeler" box="[199,355,248,271]" pageId="4" pageNumber="52" refString="Wheeler, W. M. (1922) Ants of the American Museum Congo Expedition. A Contribution to the Myrmecology of Africa. Bulletin of the American Museum of Natural History, 45, 1 - 1139." type="journal article" year="1922">Wheeler 1922</bibRefCitation>
: 736
</treatmentCitation>
, Raignier and van 
<treatmentCitation id="BA2410BDFFF7DC36FD8CFF059A65FEB4" author="Boven" box="[617,793,248,271]" page="85" pageId="4" pageNumber="43" year="1955">
<bibRefCitation id="5F144B5DFFF7DC36FD8CFF059B8CFEB4" author="Boven" box="[617,752,248,271]" pageId="4" pageNumber="52" refString="Raignier, A. &amp; Boven, J. K. A. van (1955) Etude taxonomique, biologique et biometrique des Dorylus du sous-genre Anomma (Hymenoptera Formicidae). Annales du Musee Royal du Congo Belge. Tervuren. Sciences Zoologiques. Serie, 2, 1 - 359." type="journal article" year="1955">Boven 1955</bibRefCitation>
: 85
</treatmentCitation>
, 
<treatmentCitation id="BA2410BDFFF7DC36FCC3FF059D7FFEB4" author="Gotwald" box="[806,1027,248,271]" page="877" pageId="4" pageNumber="43" year="1974">
<bibRefCitation id="5F144B5DFFF7DC36FCC3FF059AB3FEB4" author="Gotwald" box="[806,975,248,271]" pageId="4" pageNumber="51" refString="Gotwald, W. H. Jr. (1974 a) Foraging behavior of Anomma driver ants in Ghana cocoa farms. Bulletin Institut Fond. Afrique Noire Serie A, 36, 705 - 713." type="journal article" year="1974" yearSuffix="a">Gotwald 1974a</bibRefCitation>
: 877
</treatmentCitation>
–886, 
<treatmentCitation id="BA2410BDFFF7DC36FBADFF059C54FEB4" author="Gotwald" box="[1096,1320,248,271]" page="705" pageId="4" pageNumber="43" year="1974">
<bibRefCitation id="5F144B5DFFF7DC36FBADFF059D8FFEB4" author="Gotwald" box="[1096,1267,248,271]" pageId="4" pageNumber="51" refString="Gotwald, W. H. Jr. (1974 b) Predatory behavior and food preferences of driver ants in selected African habitats. Annals of the Entomological Society of America, 67, 877 - 886." type="journal article" year="1974" yearSuffix="b">Gotwald 1974b</bibRefCitation>
: 705
</treatmentCitation>
–713, 
<treatmentCitation id="BA2410BDFFF7DC36FA8EFF059817FE94" author="Gotwald" page="161" pageId="4" pageNumber="43" year="1978">
<bibRefCitation id="5F144B5DFFF7DC36FA8EFF059846FE94" author="Gotwald" pageId="4" pageNumber="51" refString="Gotwald, W. H. Jr. (1978) Trophic ecology and adaptation in tropical Old World ants of the subfamily Dorylinae. Biotropica, 10, 161 - 169." type="journal article" year="1978">Gotwald 1978</bibRefCitation>
: 161
</treatmentCitation>
–169, 
<treatmentCitation id="BA2410BDFFF7DC36FE4BFEE59B15FE94" author="Bolton" box="[430,617,280,303]" page="179" pageId="4" pageNumber="43" year="1995">
<bibRefCitation id="5F144B5DFFF7DC36FE4BFEE59B49FE94" author="Bolton" box="[430,565,280,303]" pageId="4" pageNumber="51" refString="Bolton, B. (1995) A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, Massachusetts, 504 pp." type="book" year="1995">Bolton 1995</bibRefCitation>
: 179
</treatmentCitation>
, 
<treatmentCitation id="BA2410BDFFF7DC36FD91FEE59A04FE94" author="Kronauer" box="[628,888,280,303]" page="56" pageId="4" pageNumber="43" year="2007">
<bibRefCitation id="5F144B5DFFF7DC36FD91FEE59A2FFE94" author="Kronauer" box="[628,851,280,303]" pageId="4" pageNumber="52" refString="Kronauer, D. J. C., Schoning, C., Vilhelmsen, L. B. &amp; Boomsma, J. J. (2007) A molecular phylogeny of Dorylus army ants provides evidence for multiple evolutionary transitions in foraging niche. BMC Evolutionary Biology, 7, 56." type="journal article" year="2007">
Kronauer 
<emphasis id="09F1EABEFFF7DC36FD3AFEE59A69FE94" box="[735,789,280,303]" italics="true" pageId="4" pageNumber="43">et al.</emphasis>
2007
</bibRefCitation>
: 56
</treatmentCitation>
(examined), Schöning 
<emphasis id="09F1EABEFFF7DC36FB8AFEE59DD8FE94" box="[1135,1188,280,303]" italics="true" pageId="4" pageNumber="43">et al.</emphasis>
2007: 125–133 (examined), 
<treatmentCitation id="BA2410BDFFF7DC36FEEEFEC59B3DFEF4" author="Boesch" box="[267,577,312,335]" page="86" pageId="4" pageNumber="43" year="1990">
<bibRefCitation id="5F144B5DFFF7DC36FEEEFEC59B67FEF4" author="Boesch" box="[267,539,312,335]" pageId="4" pageNumber="51" refString="Boesch, C. &amp; Boesch, H. (1990) Tool use and tool making in wild chimpanzees. Folia Primatologica, 54, 86 - 99." type="journal article" year="1990">Boesch and Boesch 1990</bibRefCitation>
: 86
</treatmentCitation>
–99, 
<treatmentCitation id="BA2410BDFFF7DC36FD90FEC59AA0FEF4" author="Humle" box="[629,988,312,335]" page="133" pageId="4" pageNumber="43" year="2002">
<bibRefCitation id="5F144B5DFFF7DC36FD90FEC59AD7FEF4" author="Humle" box="[629,939,312,335]" pageId="4" pageNumber="52" refString="Humle, T. &amp; Matsuzawa, T. (2002) Ant-dipping among the chimpanzees of Bossou, Guinea, and some comparisons with other sites. American Journal of Primatology, 58, 133 - 148." type="journal article" year="2002">Humle and Matsuzawa 2002</bibRefCitation>
: 133
</treatmentCitation>
–148 (examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="3B3A36ACFFF7DC36FF72FEA59AACFE34" blockId="4.[151,1436,152,655]" pageId="4" pageNumber="43">
<treatmentCitationGroup id="1B951182FFF7DC36FF72FEA59AACFE34" pageId="4" pageNumber="43">
<taxonomicName id="FC854D2FFFF7DC36FF72FEA5982CFED4" ID-CoL="37FCF" box="[151,336,344,367]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFF7DC36FF72FEA5982CFED4" box="[151,336,344,367]" italics="true" pageId="4" pageNumber="43">Dorylus gribodoi</emphasis>
</taxonomicName>
<taxonomicName id="FC854D2FFFF7DC36FEBDFEA59BD3FED4" authority="Santschi 1915: 246" authorityName="Santschi" authorityPageNumber="246" authorityYear="1915" box="[344,687,344,367]" pageId="4" pageNumber="43" rank="variety" variety="confusa">
var. 
<emphasis id="09F1EABEFFF7DC36FE63FEA598A5FED4" box="[390,473,344,367]" italics="true" pageId="4" pageNumber="43">confusa</emphasis>
<treatmentCitation id="BA2410BDFFF7DC36FE04FEA59BD3FED4" author="Santschi" box="[481,687,344,367]" page="246" pageId="4" pageNumber="43" year="1915">
<bibRefCitation id="5F144B5DFFF7DC36FE04FEA59B00FED4" author="Santschi" box="[481,636,344,367]" pageId="4" pageNumber="52" refString="Santschi, F. (1915) Nouvelles fourmis d'Afrique. Annales Societe Entomologique de France, 84, 244 - 282." type="journal article" year="1915">Santschi 1915</bibRefCitation>
: 246
</treatmentCitation>
</taxonomicName>
– 247. Five 
<typeStatus id="E43E880EFFF7DC36FCDCFEA59AF0FED4" box="[825,908,344,367]" pageId="4" pageNumber="43" type="syntype">syntype</typeStatus>
males, 
<collectingCountry id="4392763CFFF7DC36FC05FEA59D1EFED4" box="[992,1122,344,367]" name="Ivory Coast" pageId="4" pageNumber="43">Ivory Coast</collectingCountry>
: Grand Bassam and Imbokro collected by J.H. Lohier and Posth, MNHN (examined), 
<emphasis id="09F1EABEFFF7DC36FCC6FE8A9AB5FE34" bold="true" box="[803,969,375,399]" pageId="4" pageNumber="43">new synonomy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3B3A36ACFFF7DC36FF72FE659DE0FE14" blockId="4.[151,1436,152,655]" box="[151,1180,408,431]" pageId="4" pageNumber="43">
<treatmentCitationGroup id="1B951182FFF7DC36FF72FE659DE0FE14" box="[151,1180,408,431]" pageId="4" pageNumber="43">
<taxonomicName id="FC854D2FFFF7DC36FF72FE6598C2FE14" ID-CoL="37FCF" box="[151,446,408,431]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="gribodoi" subGenus="Dorylus">
<emphasis id="09F1EABEFFF7DC36FF72FE659991FE14" box="[151,237,408,431]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
(
<emphasis id="09F1EABEFFF7DC36FF19FE65982EFE14" box="[252,338,408,431]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
) 
<emphasis id="09F1EABEFFF7DC36FE87FE6598C2FE14" box="[354,446,408,431]" italics="true" pageId="4" pageNumber="43">gribodoi</emphasis>
</taxonomicName>
<taxonomicName id="FC854D2FFFF7DC36FE21FE659B90FE14" authority="Santschi 1915" authorityName="Santschi" authorityYear="1915" box="[452,748,408,431]" pageId="4" pageNumber="43" rank="variety" variety="confusus">
var. 
<emphasis id="09F1EABEFFF7DC36FE17FE659B33FE14" box="[498,591,408,431]" italics="true" pageId="4" pageNumber="43">confusus</emphasis>
<bibRefCitation id="5F144B5DFFF7DC36FDB3FE659B90FE14" author="Santschi" box="[598,748,408,431]" pageId="4" pageNumber="52" refString="Santschi, F. (1915) Nouvelles fourmis d'Afrique. Annales Societe Entomologique de France, 84, 244 - 282." type="journal article" year="1915">Santschi 1915</bibRefCitation>
</taxonomicName>
; 
<treatmentCitation id="BA2410BDFFF7DC36FCE4FE659AB3FE14" author="Wheeler" box="[769,975,408,431]" page="732" pageId="4" pageNumber="43" year="1922">
<bibRefCitation id="5F144B5DFFF7DC36FCE4FE659AE7FE14" author="Wheeler" box="[769,923,408,431]" pageId="4" pageNumber="52" refString="Wheeler, W. M. (1922) Ants of the American Museum Congo Expedition. A Contribution to the Myrmecology of Africa. Bulletin of the American Museum of Natural History, 45, 1 - 1139." type="journal article" year="1922">Wheeler 1922</bibRefCitation>
: 732
</treatmentCitation>
, 
<treatmentCitation id="BA2410BDFFF7DC36FC3EFE659DE4FE14" author="Bolton" box="[987,1176,408,431]" page="178" pageId="4" pageNumber="43" year="1995">
<bibRefCitation id="5F144B5DFFF7DC36FC3EFE659D1FFE14" author="Bolton" box="[987,1123,408,431]" pageId="4" pageNumber="51" refString="Bolton, B. (1995) A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, Massachusetts, 504 pp." type="book" year="1995">Bolton 1995</bibRefCitation>
: 178
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3B3A36ACFFF7DC36FF72FE459AD7FE54" blockId="4.[151,1436,152,655]" pageId="4" pageNumber="43">
<treatmentCitationGroup id="1B951182FFF7DC36FF72FE459AD7FE54" pageId="4" pageNumber="43">
<taxonomicName id="FC854D2FFFF7DC36FF72FE4598BBFE74" ID-CoL="37FCF" box="[151,455,440,463]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="gribodoi" subGenus="Anomma">
<emphasis id="09F1EABEFFF7DC36FF72FE459991FE74" box="[151,237,440,463]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
(
<emphasis id="09F1EABEFFF7DC36FF19FE459827FE74" box="[252,347,440,463]" italics="true" pageId="4" pageNumber="43">Anomma</emphasis>
) 
<emphasis id="09F1EABEFFF7DC36FE8EFE4598BBFE74" box="[363,455,440,463]" italics="true" pageId="4" pageNumber="43">gribodoi</emphasis>
</taxonomicName>
<taxonomicName id="FC854D2FFFF7DC36FE2BFE459A58FE74" authority="Santschi 1937: 98" authorityName="Santschi" authorityPageNumber="98" authorityYear="1937" box="[462,804,440,463]" pageId="4" pageNumber="43" rank="variety" variety="insularis">
var. 
<emphasis id="09F1EABEFFF7DC36FE19FE459B25FE74" box="[508,601,440,463]" italics="true" pageId="4" pageNumber="43">insularis</emphasis>
<treatmentCitation id="BA2410BDFFF7DC36FD84FE459A58FE74" author="Santschi" box="[609,804,440,463]" page="98" pageId="4" pageNumber="43" year="1937">
<bibRefCitation id="5F144B5DFFF7DC36FD84FE459B87FE74" author="Santschi" box="[609,763,440,463]" pageId="4" pageNumber="52" refString="Santschi, F. (1937) Resultats entomologiques d'un voyage au Cameroun. Formicides recoltes par Mr. le Dr. F. Zumpt. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 17, 93 - 104." type="journal article" year="1937">Santschi 1937</bibRefCitation>
: 98
</treatmentCitation>
</taxonomicName>
. Two 
<typeStatus id="E43E880EFFF7DC36FC81FE459ACBFE74" box="[868,951,440,463]" pageId="4" pageNumber="43" type="syntype">syntype</typeStatus>
males, 
<collectingCountry id="4392763CFFF7DC36FBEFFE459DADFE74" box="[1034,1233,440,463]" name="Equatorial Guinea" pageId="4" pageNumber="43">Equatorial Guinea</collectingCountry>
: Bioko Island, collected by Conradt and J. Moser, NHMB (examined), 
<emphasis id="09F1EABEFFF7DC36FD18FE2A9AD8FE54" bold="true" box="[765,932,471,495]" pageId="4" pageNumber="43">new synonymy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3B3A36ACFFF7DC36FF72FE059865FD94" blockId="4.[151,1436,152,655]" pageId="4" pageNumber="43">
<treatmentCitationGroup id="1B951182FFF7DC36FF72FE059865FD94" pageId="4" pageNumber="43">
<taxonomicName id="FC854D2FFFF7DC36FF72FE059BE9FDB4" ID-CoL="37FCM" authority="Bernard 1953: 219" authorityName="Bernard" authorityPageNumber="219" authorityYear="1953" box="[151,661,504,527]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="lamottei" subGenus="Anomma">
<emphasis id="09F1EABEFFF7DC36FF72FE059991FDB4" box="[151,237,504,527]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
(
<emphasis id="09F1EABEFFF7DC36FF19FE059827FDB4" box="[252,347,504,527]" italics="true" pageId="4" pageNumber="43">Anomma</emphasis>
) 
<emphasis id="09F1EABEFFF7DC36FE8EFE0598BEFDB4" box="[363,450,504,527]" italics="true" pageId="4" pageNumber="43">lamottei</emphasis>
<treatmentCitation id="BA2410BDFFF7DC36FE2FFE059BE9FDB4" ID-DOI="http://doi.org/10.5281/zenodo.3507855" ID-Zenodo-Dep="3507855" author="Bernard" box="[458,661,504,527]" httpUri="http://treatment.plazi.org/id/3513B1D61D7E18427E866271E8544F67" page="219" pageId="4" pageNumber="43" year="1953">
<bibRefCitation id="5F144B5DFFF7DC36FE2FFE059B23FDB4" author="Bernard" box="[458,607,504,527]" pageId="4" pageNumber="51" refString="Bernard, F. (1953) La reserve naturelle integrale du Mt Nimba. XI. Hymenopteres, Formicidae. Memoires de l'Institut Francais d'Afrique Noire, 19, 165 - 270." type="journal article" year="1953">Bernard 1953</bibRefCitation>
: 219
</treatmentCitation>
</taxonomicName>
. 12 
<typeStatus id="E43E880EFFF7DC36FD27FE059A69FDB4" box="[706,789,504,527]" pageId="4" pageNumber="43" type="syntype">syntype</typeStatus>
workers, 
<collectingCountry id="4392763CFFF7DC36FC9AFE059AACFDB4" box="[895,976,504,527]" name="Guinea-Bissau" pageId="4" pageNumber="43">Guinea</collectingCountry>
: Mt Nimba, MNHN (examined), 
<emphasis id="09F1EABEFFF7DC36FAD2FE0A986EFD94" bold="true" pageId="4" pageNumber="43">new synonomy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3B3A36ACFFF7DC36FF72FDC59BEDFDD4" blockId="4.[151,1436,152,655]" pageId="4" pageNumber="43">
<treatmentCitationGroup id="1B951182FFF7DC36FF72FDC59BEDFDD4" pageId="4" pageNumber="43">
<taxonomicName id="FC854D2FFFF7DC36FF72FDC59B15FDF4" ID-CoL="37FCM" authority="Bernard 1953" authorityName="Bernard" authorityYear="1953" box="[151,617,568,591]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="lamottei" subGenus="Anomma">
<emphasis id="09F1EABEFFF7DC36FF72FDC59991FDF4" box="[151,237,568,591]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
(
<emphasis id="09F1EABEFFF7DC36FF1AFDC59822FDF4" box="[255,350,568,591]" italics="true" pageId="4" pageNumber="43">Anomma</emphasis>
) 
<emphasis id="09F1EABEFFF7DC36FE94FDC598B4FDF4" box="[369,456,568,591]" italics="true" pageId="4" pageNumber="43">lamottei</emphasis>
<bibRefCitation id="5F144B5DFFF7DC36FE37FDC59B15FDF4" author="Bernard" box="[466,617,568,591]" pageId="4" pageNumber="51" refString="Bernard, F. (1953) La reserve naturelle integrale du Mt Nimba. XI. Hymenopteres, Formicidae. Memoires de l'Institut Francais d'Afrique Noire, 19, 165 - 270." type="journal article" year="1953">Bernard 1953</bibRefCitation>
</taxonomicName>
; van 
<treatmentCitation id="BA2410BDFFF7DC36FD4DFDC59A84FDF4" author="Boven" box="[680,1016,568,591]" page="351" pageId="4" pageNumber="43" year="1970">
<bibRefCitation id="5F144B5DFFF7DC36FD4DFDC59ABFFDF4" author="Boven" box="[680,963,568,591]" pageId="4" pageNumber="51" refString="Boven, J. K. A. van &amp; Levieux, J. (1970) Les Dorylinae de la savane de Lamto (Hymenoptera: Formicidae). Annales Universite Abidjan Serie E (Ecologie), 1, 351 - 358." type="journal article" year="1970">Boven and Levieux 1970</bibRefCitation>
: 351
</treatmentCitation>
–358. 
<taxonomicName id="FC854D2FFFF7DC36FBDBFDC59C17FDF4" ID-CoL="37FCF" box="[1086,1387,568,591]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="gribodoi" subGenus="Dorylus">
<emphasis id="09F1EABEFFF7DC36FBDBFDC59DE8FDF4" box="[1086,1172,568,591]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
(
<emphasis id="09F1EABEFFF7DC36FB43FDC59D80FDF4" box="[1190,1276,568,591]" italics="true" pageId="4" pageNumber="43">Dorylus</emphasis>
) 
<emphasis id="09F1EABEFFF7DC36FAEAFDC59C17FDF4" box="[1295,1387,568,591]" italics="true" pageId="4" pageNumber="43">gribodoi</emphasis>
</taxonomicName>
<taxonomicName id="FC854D2FFFF7DC36FA91FDC598B8FDD4" authority="Santschi 1937" authorityName="Santschi" authorityYear="1937" pageId="4" pageNumber="43" rank="variety" variety="insularis">
var. 
<emphasis id="09F1EABEFFF7DC36FF22FDA59858FDD4" box="[199,292,600,623]" italics="true" pageId="4" pageNumber="43">insularis</emphasis>
<bibRefCitation id="5F144B5DFFF7DC36FECEFDA598B8FDD4" author="Santschi" box="[299,452,600,623]" pageId="4" pageNumber="52" refString="Santschi, F. (1937) Resultats entomologiques d'un voyage au Cameroun. Formicides recoltes par Mr. le Dr. F. Zumpt. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 17, 93 - 104." type="journal article" year="1937">Santschi 1937</bibRefCitation>
</taxonomicName>
; 
<treatmentCitation id="BA2410BDFFF7DC36FE2AFDA59BF0FDD4" author="Bolton" box="[463,652,600,623]" page="179" pageId="4" pageNumber="43" year="1995">
<bibRefCitation id="5F144B5DFFF7DC36FE2AFDA59B2BFDD4" author="Bolton" box="[463,599,600,623]" pageId="4" pageNumber="51" refString="Bolton, B. (1995) A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, Massachusetts, 504 pp." type="book" year="1995">Bolton 1995</bibRefCitation>
: 179
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3B3A36ACFFF7DC36FF72FD859ADEFD34" blockId="4.[151,1436,152,655]" box="[151,930,632,655]" pageId="4" pageNumber="43">
<taxonomicName id="FC854D2FFFF7DC36FF72FD85980BFD34" ID-CoL="37FCD" box="[151,375,632,655]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="gerstaeckeri">
<emphasis id="09F1EABEFFF7DC36FF72FD85980BFD34" box="[151,375,632,655]" italics="true" pageId="4" pageNumber="43">Dorylus gerstaeckeri</emphasis>
</taxonomicName>
complex sp. 1; Schöning 
<emphasis id="09F1EABEFFF7DC36FD69FD859BBEFD34" box="[652,706,632,655]" italics="true" pageId="4" pageNumber="43">et al.</emphasis>
in press (examined).
</paragraph>
</subSubSection>
<caption id="6FFA6624FFF7DC36FF72F9E49A8EF98A" box="[151,1010,1561,1585]" httpUri="https://zenodo.org/record/274228/files/figure.png" pageId="4" pageNumber="43" targetBox="[253,1337,700,1534]" targetPageId="4">
<paragraph id="3B3A36ACFFF7DC36FF72F9E49A8EF98A" blockId="4.[151,1010,1561,1585]" box="[151,1010,1561,1585]" pageId="4" pageNumber="43">
<emphasis id="09F1EABEFFF7DC36FF72F9E49865F98A" bold="true" box="[151,281,1561,1585]" pageId="4" pageNumber="43">FIGURE 2.</emphasis>
Lateral overview of a 
<taxonomicName id="FC854D2FFFF7DC36FDEEF9E79BB8F98A" box="[523,708,1562,1585]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFF7DC36FDEEF9E79BB8F98A" box="[523,708,1562,1585]" italics="true" pageId="4" pageNumber="43">Dorylus gribodoi</emphasis>
</taxonomicName>
male from Taï, Ivory Coast.
</paragraph>
</caption>
<subSubSection id="739F6527FFF7DC36FF72F9999D13F85D" pageId="4" pageNumber="43" type="materials_examined">
<paragraph id="3B3A36ACFFF7DC36FF72F9999D13F85D" blockId="4.[151,1436,1636,2022]" pageId="4" pageNumber="43">
<emphasis id="09F1EABEFFF7DC36FF72F99998A1F9C5" bold="true" box="[151,477,1636,1662]" pageId="4" pageNumber="43">Other material examined:</emphasis>
<collectingCountry id="4392763CFFF7DC36FE00F9999B4AF9C5" box="[485,566,1636,1662]" name="Ghana" pageId="4" pageNumber="43">Ghana</collectingCountry>
: Kade ([
<geoCoordinate id="5EB1506BFFF7DC36FD7DF9999B9BF9C5" box="[664,743,1636,1662]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="925" value="6.0666666">N 6º4'</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FD16F9999A2BF9C5" box="[755,855,1636,1662]" direction="west" orientation="longitude" pageId="4" pageNumber="43" precision="925" value="-0.81666666">W 0º49'</geoCoordinate>
], 
<quantity id="FC7D9B49FFF7DC36FC8FF9999AC9F9C5" box="[874,949,1636,1662]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="43" unit="m" value="150.0">150 m</quantity>
asl, W.H. Gotwald Jr., 
<collectionCode id="5D94AE69FFF7DC36FB2DF9999C5AF9C5" box="[1224,1318,1636,1662]" pageId="4" pageNumber="43">WHGC</collectionCode>
), Osenase ([
<geoCoordinate id="5EB1506BFFF7DC36FF42F971987BF91D" box="[167,263,1676,1702]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="925" value="5.95">N 5º57'</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FEF1F971980CF91D" box="[276,368,1676,1702]" direction="east" orientation="longitude" pageId="4" pageNumber="43" precision="925" value="0.75">E 0º45'</geoCoordinate>
], 
<quantity id="FC7D9B49FFF7DC36FE60F97198ADF91D" box="[389,465,1676,1702]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="43" unit="m" value="180.0">180 m</quantity>
asl, D. Leston, 
<collectionCode id="5D94AE69FFF7DC36FD76F9719B8DF91D" box="[659,753,1676,1702]" pageId="4" pageNumber="43">WHGC</collectionCode>
), Tafo ([
<geoCoordinate id="5EB1506BFFF7DC36FCB3F9719ACAF91D" box="[854,950,1676,1702]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="925" value="6.7166667">N 6º43'</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FC26F9719D5AF91D" box="[963,1062,1676,1702]" direction="west" orientation="longitude" pageId="4" pageNumber="43" precision="925" value="-1.6166667">W 1º37'</geoCoordinate>
, 
<quantity id="FC7D9B49FFF7DC36FBD1F9719DFCF91D" box="[1076,1152,1676,1702]" metricMagnitude="2" metricUnit="m" metricValue="2.78" pageId="4" pageNumber="43" unit="m" value="278.0">278 m</quantity>
asl], W.H. Gotwald Jr., 
<collectionCode id="5D94AE69FFF7DC36FF72F949998AF975" box="[151,246,1716,1742]" pageId="4" pageNumber="43">WHGC</collectionCode>
). 
<collectingCountry id="4392763CFFF7DC36FEEEF9499818F975" box="[267,356,1716,1742]" name="Guinea-Bissau" pageId="4" pageNumber="43">Guinea</collectingCountry>
: Bossou (
<geoCoordinate id="5EB1506BFFF7DC36FE3FF9499B1CF975" box="[474,608,1716,1742]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="15" value="7.645">N 7º38'42''</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FD8BF9499B84F975" box="[622,760,1716,1742]" direction="west" orientation="longitude" pageId="4" pageNumber="43" precision="15" value="-8.509722">W 8º30'35''</geoCoordinate>
, 
<quantity id="FC7D9B49FFF7DC36FCE2F9499A2FF975" box="[775,851,1716,1742]" metricMagnitude="2" metricUnit="m" metricValue="5.72" pageId="4" pageNumber="43" unit="m" value="572.0">572 m</quantity>
asl, T. Humle, K. Koops, Y. Sugiyama, 
<collectionCode id="5D94AE69FFF7DC36FADBF9499CF3F975" box="[1342,1423,1716,1742]" pageId="4" pageNumber="43">CSAC</collectionCode>
), Seringbara (
<geoCoordinate id="5EB1506BFFF7DC36FECDF92198D2F94D" box="[296,430,1756,1782]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="15" value="7.6269445">N 7º37'37''</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FE58F9219B45F94D" box="[445,569,1756,1782]" direction="west" orientation="longitude" pageId="4" pageNumber="43" precision="15" value="-8.467222">W 8º28'2''</geoCoordinate>
, 
<quantity id="FC7D9B49FFF7DC36FDACF9219BEAF94D" box="[585,662,1756,1782]" metricMagnitude="2" metricUnit="m" metricValue="6.08" pageId="4" pageNumber="43" unit="m" value="608.0">608 m</quantity>
asl, T. Humle, K. Koops, 
<collectionCode id="5D94AE69FFF7DC36FC3EF9219D51F94D" box="[987,1069,1756,1782]" pageId="4" pageNumber="43">CSAC</collectionCode>
). 
<collectingCountry id="4392763CFFF7DC36FBA1F9219DA9F94D" box="[1092,1237,1756,1782]" name="Ivory Coast" pageId="4" pageNumber="43">Ivory Coast</collectingCountry>
: Taï ([
<geoCoordinate id="5EB1506BFFF7DC36FAC6F9219CF8F94D" box="[1315,1412,1756,1782]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="925" value="6.3333335">N 6°20'</geoCoordinate>
– 5°10', 
<geoCoordinate id="5EB1506BFFF7DC36FF01F8F9983AF8A5" box="[228,326,1796,1822]" direction="west" orientation="longitude" pageId="4" pageNumber="43" precision="925" value="-4.3333335">W 4°20'</geoCoordinate>
– 6°50'], 
<quantity id="FC7D9B49FFF7DC36FE59F8F99B74F8A5" box="[444,520,1796,1822]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="43" unit="m" value="150.0">150 m</quantity>
asl, Y. Möbius, T. Deschner, C. Schöning, 
<collectionCode id="5D94AE69FFF7DC36FBE9F8F99D21F8A5" box="[1036,1117,1796,1822]" pageId="4" pageNumber="43">CSAC</collectionCode>
), Lamto ([
<geoCoordinate id="5EB1506BFFF7DC36FB3DF8F99C4AF8A5" box="[1240,1334,1796,1822]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="925" value="6.2166667">N 6º13'</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FAA6F8F99CECF8A5" box="[1347,1424,1796,1822]" direction="west" orientation="longitude" pageId="4" pageNumber="43" precision="925" value="-5.0333333">W5º2'</geoCoordinate>
], 
<quantity id="FC7D9B49FFF7DC36FF72F8D199A6F8FD" box="[151,218,1836,1862]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="43" unit="m" value="100.0">100m</quantity>
asl, queen specimen collected from nest along with workers in 1974, sample label #740701, J.M. Leroux, 
<collectionCode id="5D94AE69FFF7DC36FF33F8A99848F8D5" box="[214,308,1876,1902]" pageId="4" pageNumber="43">WHGC</collectionCode>
), Orstrom Experiment Station, nr. Abidjan ([
<geoCoordinate id="5EB1506BFFF7DC36FCBFF8A99AA3F8D5" box="[858,991,1876,1902]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="15" value="5.341111">N 5º20'28''</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FC01F8A99D1AF8D5" box="[996,1126,1876,1902]" direction="west" orientation="longitude" pageId="4" pageNumber="43" precision="15" value="-4.0280557">W 4º1'41''</geoCoordinate>
], 
<quantity id="FC7D9B49FFF7DC36FB9AF8A99DBCF8D5" box="[1151,1216,1876,1902]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="43" unit="m" value="70.0">70 m</quantity>
asl, W.L. Brown, 
<collectionCode id="5D94AE69FFF7DC36FF72F881998AF82D" box="[151,246,1916,1942]" pageId="4" pageNumber="43">WHGC</collectionCode>
). 
<collectingCountry id="4392763CFFF7DC36FEEFF881981FF82D" box="[266,355,1916,1942]" name="Liberia" pageId="4" pageNumber="43">Liberia</collectingCountry>
: Charlesville ([
<geoCoordinate id="5EB1506BFFF7DC36FDF2F8819BE1F82D" box="[535,669,1916,1942]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="15" value="6.206111">N 6º12'22''</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FD4EF8819A39F82D" box="[683,837,1916,1942]" direction="west" orientation="longitude" pageId="4" pageNumber="43" precision="15" value="-10.3747225">W 10º22'29''</geoCoordinate>
], 
<quantity id="FC7D9B49FFF7DC36FCBFF8819AEBF82D" box="[858,919,1916,1942]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="43" unit="m" value="90.0">90 m</quantity>
asl, D. Kistner, 
<collectionCode id="5D94AE69FFF7DC36FBB9F8819DC4F82D" box="[1116,1208,1916,1942]" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="4" pageNumber="43">RMCA</collectionCode>
). 
<collectingCountry id="4392763CFFF7DC36FB29F8819C54F82D" box="[1228,1320,1916,1942]" name="Nigeria" pageId="4" pageNumber="43">Nigeria</collectingCountry>
: Gambari ([
<geoCoordinate id="5EB1506BFFF7DC36FF42F8599851F805" box="[167,301,1956,1982]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="15" value="8.206111">N 8º12'22''</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FEDEF85998DEF805" box="[315,418,1956,1982]" direction="west" orientation="longitude" pageId="4" pageNumber="43" precision="925" value="-4.3166666">W 4º19’</geoCoordinate>
], 
<quantity id="FC7D9B49FFF7DC36FE5CF8599B79F805" box="[441,517,1956,1982]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="4" pageNumber="43" unit="m" value="308.0">308 m</quantity>
asl, B. Bolton, 
<collectionCode id="5D94AE69FFF7DC36FD21F8599A5EF805" box="[708,802,1956,1982]" pageId="4" pageNumber="43">WHGC</collectionCode>
), Gashaka (
<geoCoordinate id="5EB1506BFFF7DC36FC4BF8599D35F805" box="[942,1097,1956,1982]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="1" value="7.329472">N 7º19'46.1''</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FBB3F8599D97F805" box="[1110,1259,1956,1982]" direction="east" orientation="longitude" pageId="4" pageNumber="43" precision="1" value="11.58386">E 11º35'1.9''</geoCoordinate>
, 
<quantity id="FC7D9B49FFF7DC36FB1CF8599C39F805" box="[1273,1349,1956,1982]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="43" unit="m" value="560.0">560 m</quantity>
asl, D. Ellis, 
<collectionCode id="5D94AE69FFF7DC36FF3EF8319850F85D" box="[219,300,1996,2022]" pageId="4" pageNumber="43">CSAC</collectionCode>
), Ibadan ([
<geoCoordinate id="5EB1506BFFF7DC36FE42F8319B57F85D" box="[423,555,1996,2022]" direction="north" orientation="latitude" pageId="4" pageNumber="43" precision="15" value="7.387778">N 7º23'16''</geoCoordinate>
, 
<geoCoordinate id="5EB1506BFFF7DC36FDDDF8319BCBF85D" box="[568,695,1996,2022]" direction="east" orientation="longitude" pageId="4" pageNumber="43" precision="15" value="3.8963888">E 3º53'47''</geoCoordinate>
], 
<quantity id="FC7D9B49FFF7DC36FD2EF8319A6AF85D" box="[715,790,1996,2022]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="43" unit="m" value="230.0">230 m</quantity>
asl, W.H. Gotwald, 
<collectionCode id="5D94AE69FFF7DC36FBE6F8319D1EF85D" box="[1027,1122,1996,2022]" pageId="4" pageNumber="43">WHGC</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="6FFA6624FFF6DC37FF72FBA49D3EFBCB" box="[151,1090,1113,1137]" httpUri="https://zenodo.org/record/274229/files/figure.png" pageId="5" pageNumber="44" targetBox="[238,1338,183,1113]" targetPageId="5">
<paragraph id="3B3A36ACFFF6DC37FF72FBA49D3EFBCB" blockId="5.[151,1090,1113,1137]" box="[151,1090,1113,1137]" pageId="5" pageNumber="44">
<emphasis id="09F1EABEFFF6DC37FF72FBA49865FBCA" bold="true" box="[151,281,1113,1137]" pageId="5" pageNumber="44">FIGURE 3.</emphasis>
Frontal view of the head of a 
<taxonomicName id="FC854D2FFFF6DC37FDBFFBA49A6FFBCB" box="[602,787,1113,1136]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFF6DC37FDBFFBA49A6FFBCB" box="[602,787,1113,1136]" italics="true" pageId="5" pageNumber="44">Dorylus gribodoi</emphasis>
</taxonomicName>
male from Taï, Ivory Coast.
</paragraph>
</caption>
<caption id="6FFA6624FFF6DC37FF72F8BE9ADFF8E0" box="[151,931,1859,1883]" httpUri="https://zenodo.org/record/274230/files/figure.png" pageId="5" pageNumber="44" targetBox="[247,1337,1174,1859]" targetPageId="5">
<paragraph id="3B3A36ACFFF6DC37FF72F8BE9ADFF8E0" blockId="5.[151,931,1859,1883]" box="[151,931,1859,1883]" pageId="5" pageNumber="44">
<emphasis id="09F1EABEFFF6DC37FF72F8BE9865F8E0" bold="true" box="[151,281,1859,1883]" pageId="5" pageNumber="44">FIGURE 4.</emphasis>
<taxonomicName id="FC854D2FFFF6DC37FEC5F8B998A4F8E0" box="[288,472,1860,1883]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFF6DC37FEC5F8B998A4F8E0" box="[288,472,1860,1883]" italics="true" pageId="5" pageNumber="44">Dorylus gribodoi</emphasis>
</taxonomicName>
male: genital capsule and subgenital plate.
</paragraph>
</caption>
<subSubSection id="739F6527FFF6DC35FF20F8769DACF998" lastPageId="7" lastPageNumber="46" pageId="5" pageNumber="44" type="description">
<paragraph id="3B3A36ACFFF6DC37FF20F8769AB2F84D" blockId="5.[151,1436,1931,2038]" pageId="5" pageNumber="44">
<emphasis id="09F1EABEFFF6DC37FF20F8769892F81E" bold="true" box="[197,494,1931,1957]" pageId="5" pageNumber="44">Worker measurements:</emphasis>
Worker caste highly polymorphic, among samples from Taï (
<collectingCountry id="4392763CFFF6DC37FB29F8719C27F81D" box="[1228,1371,1932,1958]" name="Ivory Coast" pageId="5" pageNumber="44">Ivory Coast</collectingCountry>
) HW varies between 
<quantity id="FC7D9B49FFF6DC37FEA9F84998C8F875" box="[332,436,1972,1998]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="5" pageNumber="44" unit="mm" value="0.78">0.78 mm</quantity>
and 
<quantity id="FC7D9B49FFF6DC37FE09F8499B35F875" box="[492,585,1972,1998]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="44" unit="mm" value="2.9">2.9 mm</quantity>
. SL, hind leg length and mandible length notably shorter than in workers of the driver ant clade (
<bibRefCitation id="5F144B5DFFF6DC37FE42F8219BE3F84D" author="Kronauer" box="[423,671,2012,2038]" pageId="5" pageNumber="52" refString="Kronauer, D. J. C., Schoning, C., Vilhelmsen, L. B. &amp; Boomsma, J. J. (2007) A molecular phylogeny of Dorylus army ants provides evidence for multiple evolutionary transitions in foraging niche. BMC Evolutionary Biology, 7, 56." type="journal article" year="2007">
Kronauer 
<emphasis id="09F1EABEFFF6DC37FDFAF8219B25F84E" box="[543,601,2012,2037]" italics="true" pageId="5" pageNumber="44">et al.</emphasis>
2007
</bibRefCitation>
, Schöning 
<emphasis id="09F1EABEFFF6DC37FCC5F8219A27F84E" box="[800,859,2012,2037]" italics="true" pageId="5" pageNumber="44">et al.</emphasis>
in press).
</paragraph>
<caption id="6FFA6624FFF5DC34FF72FC0A9D3AFBB4" box="[151,1094,1015,1039]" httpUri="https://zenodo.org/record/274231/files/figure.png" pageId="6" pageNumber="45" targetBox="[264,1340,178,996]" targetPageId="6">
<paragraph id="3B3A36ACFFF5DC34FF72FC0A9D3AFBB4" blockId="6.[151,1094,1015,1039]" box="[151,1094,1015,1039]" pageId="6" pageNumber="45">
<emphasis id="09F1EABEFFF5DC34FF72FC0A9865FBB4" bold="true" box="[151,281,1015,1039]" pageId="6" pageNumber="45">FIGURE 5.</emphasis>
Lateral overview of a large 
<taxonomicName id="FC854D2FFFF5DC34FDA3FC059B83FBB4" box="[582,767,1016,1039]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFF5DC34FDA3FC059B83FBB4" box="[582,767,1016,1039]" italics="true" pageId="6" pageNumber="45">Dorylus gribodoi</emphasis>
</taxonomicName>
worker from Taï, Ivory Coast.
</paragraph>
</caption>
<caption id="6FFA6624FFF5DC34FF72F8919A8BF838" box="[151,1015,1900,1924]" httpUri="https://zenodo.org/record/274232/files/figure.png" pageId="6" pageNumber="45" targetBox="[240,1348,1074,1873]" targetPageId="6">
<paragraph id="3B3A36ACFFF5DC34FF72F8919A8BF838" blockId="6.[151,1015,1900,1924]" box="[151,1015,1900,1924]" pageId="6" pageNumber="45">
<emphasis id="09F1EABEFFF5DC34FF72F8919865F83F" bold="true" box="[151,281,1900,1924]" pageId="6" pageNumber="45">FIGURE 6.</emphasis>
Lateral overview of a large 
<taxonomicName id="FC854D2FFFF5DC34FDA3F8919BCCF838" box="[582,688,1900,1923]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="09F1EABEFFF5DC34FDA3F8919BCCF838" box="[582,688,1900,1923]" italics="true" pageId="6" pageNumber="45">D. emeryi</emphasis>
</taxonomicName>
worker from Taï, Ivory Coast.
</paragraph>
</caption>
<paragraph id="3B3A36ACFFF5DC35FF20F84C9B71FF61" blockId="6.[151,1437,1969,2036]" lastBlockId="7.[151,1436,152,218]" lastPageId="7" lastPageNumber="46" pageId="6" pageNumber="45">
<emphasis id="09F1EABEFFF5DC34FF20F84C98BEF870" bold="true" box="[197,450,1969,1995]" pageId="6" pageNumber="45">Male measurements</emphasis>
(n = 12, mean ± SD, range): HW 4.38 ± 0.11, 4.19 – 4.55; SL 1.57 ± 0.05, 1.48 – 1.67; HL 2.24 ± 0.07, 2.12 – 2.33; EL 1.78 ± 0.08, 1.61 – 1.88; DPF 2.77 ± 0.12, 2.57 – 2.95; PeW 3.70 ± 0.14, 3.52 – 3.90; HTL 2.29 ± 0.08, 2.07 – 2.42; PL 2.10 ± 0.08, 1.99 – 2.24; PD 1.01 ± 0.05, 0.96 – 1.09; SPWB 1.05 ± 0.07, 0.93 – 1.14.
</paragraph>
<caption id="6FFA6624FFF4DC35FF72FD069DB2FCA8" box="[151,1230,763,787]" httpUri="https://zenodo.org/record/274233/files/figure.png" pageId="7" pageNumber="46" targetBox="[468,1117,258,737]" targetPageId="7">
<paragraph id="3B3A36ACFFF4DC35FF72FD069DB2FCA8" blockId="7.[151,1230,763,787]" box="[151,1230,763,787]" pageId="7" pageNumber="46">
<emphasis id="09F1EABEFFF4DC35FF72FD069865FCA8" bold="true" box="[151,281,763,787]" pageId="7" pageNumber="46">FIGURE 7.</emphasis>
Lateral view of the head of a large 
<taxonomicName id="FC854D2FFFF4DC35FD71FD019A6DFCA8" box="[660,785,764,787]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFF4DC35FD71FD019A6DFCA8" box="[660,785,764,787]" italics="true" pageId="7" pageNumber="46">D. gribodoi</emphasis>
</taxonomicName>
worker (HW 2.79 mm), Taï, Ivory Coast.
</paragraph>
</caption>
<caption id="6FFA6624FFF4DC35FF72FA6B9D92FA16" box="[151,1262,1430,1454]" httpUri="https://zenodo.org/record/274234/files/figure.png" pageId="7" pageNumber="46" targetBox="[409,1159,844,1407]" targetPageId="7">
<paragraph id="3B3A36ACFFF4DC35FF72FA6B9D92FA16" blockId="7.[151,1262,1430,1454]" box="[151,1262,1430,1454]" pageId="7" pageNumber="46">
<emphasis id="09F1EABEFFF4DC35FF72FA6B9864FA15" bold="true" box="[151,280,1430,1454]" pageId="7" pageNumber="46">FIGURE 8.</emphasis>
Lateral view of the head of a large 
<taxonomicName id="FC854D2FFFF4DC35FD71FA6A9B82FA15" box="[660,766,1431,1454]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="09F1EABEFFF4DC35FD71FA6A9B82FA15" box="[660,766,1431,1454]" italics="true" pageId="7" pageNumber="46">D. emeryi</emphasis>
</taxonomicName>
worker (HW 3.62 mm) from Taï, Ivory Coast.
</paragraph>
</caption>
<paragraph id="3B3A36ACFFF4DC35FF20FA1D9DACF998" blockId="7.[151,1436,1504,2011]" pageId="7" pageNumber="46">
<emphasis id="09F1EABEFFF4DC35FF20FA1D98AFFA41" bold="true" box="[197,467,1504,1530]" pageId="7" pageNumber="46">Queen measurements</emphasis>
(n = 1): HW 4.58, SL 1.44, HL 3.32, PeW 4.54, HFL 2.73, HAD 2.53, maximum total length, in lateral view, from anterior surface of head to tips of hypopygial apices 40.45.
</paragraph>
</subSubSection>
<subSubSection id="739F6527FFF4DC35FF20F9CD9A9CF9F0" box="[197,992,1584,1611]" pageId="7" pageNumber="46" type="distribution">
<paragraph id="3B3A36ACFFF4DC35FF20F9CD9A9CF9F0" blockId="7.[151,1436,1504,2011]" box="[197,992,1584,1611]" pageId="7" pageNumber="46">
<emphasis id="09F1EABEFFF4DC35FF20F9CD9814F9F1" bold="true" box="[197,360,1584,1610]" pageId="7" pageNumber="46">Distribution:</emphasis>
West Africa (from 
<collectingCountry id="4392763CFFF4DC35FDA8F9CC9BDEF9F0" box="[589,674,1585,1611]" name="Guinea-Bissau" pageId="7" pageNumber="46">Guinea</collectingCountry>
and 
<collectingCountry id="4392763CFFF4DC35FD3EF9CC9A4CF9F0" box="[731,816,1585,1611]" name="Liberia" pageId="7" pageNumber="46">Liberia</collectingCountry>
to 
<collectingCountry id="4392763CFFF4DC35FCB0F9CC9AA9F9F0" box="[853,981,1585,1611]" name="Cameroon" pageId="7" pageNumber="46">Cameroon</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="739F6527FFF4DC3AFF20F9A598C7FE91" lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46" type="biology_ecology">
<paragraph id="3B3A36ACFFF4DC3AFF20F9A598C7FE91" blockId="7.[151,1436,1504,2011]" lastBlockId="8.[151,1436,152,298]" lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46">
<emphasis id="09F1EABEFFF4DC35FF20F9A59851F9C9" bold="true" box="[197,301,1624,1650]" pageId="7" pageNumber="46">Biology:</emphasis>
Restricted to humid habitats such as rainforest and gallery forest in mixed savanna – forest landscapes. Occurs at least up to 
<quantity id="FC7D9B49FFF4DC35FE11F97C9B32F920" box="[500,590,1665,1691]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="46" unit="m" value="1600.0">1600 m</quantity>
asl. Raids are conducted in the leaf-litter stratum (
<bibRefCitation id="5F144B5DFFF4DC35FB4AF97C9C15F920" author="Gotwald" box="[1199,1385,1665,1691]" pageId="7" pageNumber="51" refString="Gotwald, W. H. Jr. (1974 a) Foraging behavior of Anomma driver ants in Ghana cocoa farms. Bulletin Institut Fond. Afrique Noire Serie A, 36, 705 - 713." type="journal article" year="1974" yearSuffix="a">Gotwald 1974a</bibRefCitation>
, see also 
<bibRefCitation id="5F144B5DFFF4DC35FF28F95498BFF978" author="Kronauer" box="[205,451,1705,1731]" pageId="7" pageNumber="52" refString="Kronauer, D. J. C., Schoning, C., Vilhelmsen, L. B. &amp; Boomsma, J. J. (2007) A molecular phylogeny of Dorylus army ants provides evidence for multiple evolutionary transitions in foraging niche. BMC Evolutionary Biology, 7, 56." type="journal article" year="2007">
Kronauer 
<emphasis id="09F1EABEFFF4DC35FEA6F9549801F979" box="[323,381,1705,1730]" italics="true" pageId="7" pageNumber="46">et al.</emphasis>
2007
</bibRefCitation>
). Workers retreat in tunnels when disturbed or exposed to sun (
<bibRefCitation id="5F144B5DFFF4DC35FB44F9549C31F978" author="Gotwald" box="[1185,1357,1705,1731]" pageId="7" pageNumber="51" refString="Gotwald, W. H. Jr. (1978) Trophic ecology and adaptation in tropical Old World ants of the subfamily Dorylinae. Biotropica, 10, 161 - 169." type="journal article" year="1978">Gotwald 1978</bibRefCitation>
). Limited observations indicate that the species seems to prey almost exclusively on earthworms (
<bibRefCitation id="5F144B5DFFF4DC35FB39F92C98B5F8A8" author="Gotwald" pageId="7" pageNumber="51" refString="Gotwald, W. H. Jr. (1974 b) Predatory behavior and food preferences of driver ants in selected African habitats. Annals of the Entomological Society of America, 67, 877 - 886." type="journal article" year="1974" yearSuffix="b">Gotwald 1974b, C.S. personal observation</bibRefCitation>
), but this information has to be viewed with caution since variation in prey composition can be considerable in other 
<taxonomicName id="FC854D2FFFF4DC35FDDDF8DC9BEAF881" box="[568,662,1825,1850]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="09F1EABEFFF4DC35FDDDF8DC9BEAF881" box="[568,662,1825,1850]" italics="true" pageId="7" pageNumber="46">Dorylus</emphasis>
</taxonomicName>
species (
<bibRefCitation id="5F144B5DFFF4DC35FCE1F8DC9D7DF880" author="Schoning" box="[772,1025,1825,1851]" pageId="7" pageNumber="52" refString="Schoning, C., Njagi, W. &amp; Kinuthia, W. (2008) Prey spectra of two swarm-raiding army ant species in East Africa. Journal of Zoology, 274, 85 - 93." type="journal article" year="2008">
Schöning 
<emphasis id="09F1EABEFFF4DC35FC98F8DC9AC6F881" box="[893,954,1825,1850]" italics="true" pageId="7" pageNumber="46">et al.</emphasis>
2008
</bibRefCitation>
). Analysis of more extensive samples is therefore needed. Chimpanzees are known to feed on 
<taxonomicName id="FC854D2FFFF4DC35FC8EF8B49A89F8D9" box="[875,1013,1865,1890]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFF4DC35FC8EF8B49A89F8D9" box="[875,1013,1865,1890]" italics="true" pageId="7" pageNumber="46">D. gribodoi</emphasis>
</taxonomicName>
at two sites (Bossou in 
<collectingCountry id="4392763CFFF4DC35FAF1F8B49C15F8D8" box="[1300,1385,1865,1891]" name="Guinea-Bissau" pageId="7" pageNumber="46">Guinea</collectingCountry>
and Taï in 
<collectingCountry id="4392763CFFF4DC35FF02F88C9804F830" box="[231,376,1905,1931]" name="Ivory Coast" pageId="7" pageNumber="46">Ivory Coast</collectingCountry>
); they either use sticks to “dip” for ant workers at trails or nests or open nests and take brood and workers with their hands (
<bibRefCitation id="5F144B5DFFF4DC35FDAEF8649A06F808" author="Boesch" box="[587,890,1945,1971]" pageId="7" pageNumber="51" refString="Boesch, C. &amp; Boesch, H. (1990) Tool use and tool making in wild chimpanzees. Folia Primatologica, 54, 86 - 99." type="journal article" year="1990">Boesch and Boesch 1990</bibRefCitation>
, 
<bibRefCitation id="5F144B5DFFF4DC35FC62F8649DA1F808" author="Humle" box="[903,1245,1945,1971]" pageId="7" pageNumber="52" refString="Humle, T. &amp; Matsuzawa, T. (2002) Ant-dipping among the chimpanzees of Bossou, Guinea, and some comparisons with other sites. American Journal of Primatology, 58, 133 - 148." type="journal article" year="2002">Humle and Matsuzawa 2002</bibRefCitation>
, Schöning 
<emphasis id="09F1EABEFFF4DC35FA85F8649CE0F809" box="[1376,1436,1945,1970]" italics="true" pageId="7" pageNumber="46">et al.</emphasis>
in press). Nests are much less conspicuous than those of driver ant species and extremely difficult to find (for humans, chimpanzees seem to do much better). It is interesting to note that 
<taxonomicName id="FC854D2FFFFBDC3AFBCCFF659DE3FF0A" box="[1065,1183,152,177]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="09F1EABEFFFBDC3AFBCCFF659DE3FF0A" box="[1065,1183,152,177]" italics="true" pageId="8" pageNumber="47">D. emeryi</emphasis>
</taxonomicName>
is broadly sympatric with 
<taxonomicName id="FC854D2FFFFBDC3AFF37FF3D9827FF62" box="[210,347,192,217]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFBDC3AFF37FF3D9827FF62" box="[210,347,192,217]" italics="true" pageId="8" pageNumber="47">D. gribodoi</emphasis>
</taxonomicName>
and has nearly identical hunting behavior and habitat requirements, such that the two species may be easily confused in the field. The largest workers of 
<taxonomicName id="FC854D2FFFFBDC3AFCBAFF159AA9FEBA" box="[863,981,232,257]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="09F1EABEFFFBDC3AFCBAFF159AA9FEBA" box="[863,981,232,257]" italics="true" pageId="8" pageNumber="47">D. emeryi</emphasis>
</taxonomicName>
are much larger (HW up to 
<quantity id="FC7D9B49FFFBDC3AFACFFF159CE4FEB9" box="[1322,1432,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="3.92" pageId="8" pageNumber="47" unit="mm" value="3.92">3.92 mm</quantity>
, Schöning 
<emphasis id="09F1EABEFFFBDC3AFEE8FEED9834FE92" box="[269,328,272,297]" italics="true" pageId="8" pageNumber="47">et al.</emphasis>
in press).
</paragraph>
</subSubSection>
<caption id="6FFA6624FFFBDC3AFF72FBFA9C6FFBA5" box="[151,1299,1031,1055]" httpUri="https://zenodo.org/record/274235/files/figure.png" pageId="8" pageNumber="47" targetBox="[385,1200,354,1006]" targetPageId="8">
<paragraph id="3B3A36ACFFFBDC3AFF72FBFA9C6FFBA5" blockId="8.[151,1299,1031,1055]" box="[151,1299,1031,1055]" pageId="8" pageNumber="47">
<emphasis id="09F1EABEFFFBDC3AFF72FBFA9865FBA4" bold="true" box="[151,281,1031,1055]" pageId="8" pageNumber="47">FIGURE 9.</emphasis>
Dorsal view of the petiole of a large 
<taxonomicName id="FC854D2FFFFBDC3AFD43FBFA9A5FFBA5" box="[678,803,1031,1054]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFBDC3AFD43FBFA9A5FFBA5" box="[678,803,1031,1054]" italics="true" pageId="8" pageNumber="47">D. gribodoi</emphasis>
</taxonomicName>
worker (HW 2.85 mm) from Taï, Ivory Coast.
</paragraph>
</caption>
<caption id="6FFA6624FFFBDC3AFF72F9049C72F8AB" box="[151,1294,1785,1809]" httpUri="https://zenodo.org/record/274236/files/figure.png" pageId="8" pageNumber="47" targetBox="[385,1200,1108,1761]" targetPageId="8">
<paragraph id="3B3A36ACFFFBDC3AFF72F9049C72F8AB" blockId="8.[151,1294,1785,1809]" box="[151,1294,1785,1809]" pageId="8" pageNumber="47">
<emphasis id="09F1EABEFFFBDC3AFF72F904985AF8AA" bold="true" box="[151,294,1785,1809]" pageId="8" pageNumber="47">FIGURE 10.</emphasis>
Dorsal view of the petiole of a large 
<taxonomicName id="FC854D2FFFFBDC3AFD51F9079A61F8AA" box="[692,797,1786,1809]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="09F1EABEFFFBDC3AFD51F9079A61F8AA" box="[692,797,1786,1809]" italics="true" pageId="8" pageNumber="47">D. emeryi</emphasis>
</taxonomicName>
worker (HW 3.74 mm) from Taï, Ivory Coast.
</paragraph>
</caption>
<subSubSection id="739F6527FFFBDC3BFF23F8B99AD6F89C" lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47" type="discussion">
<paragraph id="3B3A36ACFFFBDC3BFF23F8B99BBEFC27" blockId="8.[151,1436,1860,2035]" lastBlockId="9.[151,1437,152,924]" lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47">
<emphasis id="09F1EABEFFFBDC3AFF23F8B9982BF8E5" bold="true" box="[198,343,1860,1886]" pageId="8" pageNumber="47">Comments:</emphasis>
The diagnostic features of the 
<taxonomicName id="FC854D2FFFFBDC3AFD2CF8B99A2FF8E6" box="[713,851,1860,1885]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFBDC3AFD2CF8B99A2FF8E6" box="[713,851,1860,1885]" italics="true" pageId="8" pageNumber="47">D. gribodoi</emphasis>
</taxonomicName>
male are its mandible shape (the anterior quarter notably flexed inward, 
<figureCitation id="A3BE2A29FFFBDC3AFE42F8949891F838" box="[423,493,1897,1923]" captionStart="FIGURE 3" captionStartId="5.[151,255,1113,1137]" captionTargetBox="[238,1338,183,1113]" captionTargetId="figure@5.[238,1339,179,1113]" captionTargetPageId="5" captionText="FIGURE 3. Frontal view of the head of a Dorylus gribodoi male from Taï, Ivory Coast." httpUri="https://zenodo.org/record/274229/files/figure.png" pageId="8" pageNumber="47">Fig. 3</figureCitation>
) (
<bibRefCitation id="5F144B5DFFFBDC3AFDE0F8949BE5F838" author="Emery" box="[517,665,1897,1923]" pageId="8" pageNumber="51" refString="Emery, C. (1892) Voyage de M. Ch. Alluaud dans le territoire d'Assinie (Afrique occidentale) en juillet et aout 1886. Formicides. Annales de la Societe Entomologique de France, 60, 553 - 574." type="journal article" year="1892">Emery 1892</bibRefCitation>
) and the nearly flat posterior head margin (
<figureCitation id="A3BE2A29FFFBDC3AFB72F8949DA2F838" box="[1175,1246,1897,1923]" captionStart="FIGURE 3" captionStartId="5.[151,255,1113,1137]" captionTargetBox="[238,1338,183,1113]" captionTargetId="figure@5.[238,1339,179,1113]" captionTargetPageId="5" captionText="FIGURE 3. Frontal view of the head of a Dorylus gribodoi male from Taï, Ivory Coast." httpUri="https://zenodo.org/record/274229/files/figure.png" pageId="8" pageNumber="47">Fig. 3</figureCitation>
). In the original description Emery compared 
<taxonomicName id="FC854D2FFFFBDC3AFE12F8729BFEF813" box="[503,642,1935,1960]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFBDC3AFE12F8729BFEF813" box="[503,642,1935,1960]" italics="true" pageId="8" pageNumber="47">D. gribodoi</emphasis>
</taxonomicName>
with 
<taxonomicName id="FC854D2FFFFBDC3AFD2DF8729A3DF813" box="[712,833,1935,1960]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="09F1EABEFFFBDC3AFD2DF8729A3DF813" box="[712,833,1935,1960]" italics="true" pageId="8" pageNumber="47">D. atratus</emphasis>
</taxonomicName>
and stated that 
<taxonomicName id="FC854D2FFFFBDC3AFBE4F8729D06F813" box="[1025,1146,1935,1960]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="09F1EABEFFFBDC3AFBE4F8729D06F813" box="[1025,1146,1935,1960]" italics="true" pageId="8" pageNumber="47">D. atratus</emphasis>
</taxonomicName>
was darker than 
<taxonomicName id="FC854D2FFFFBDC3AFAAFF87299A6F876" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFBDC3AFAAFF87299A6F876" italics="true" pageId="8" pageNumber="47">D. gribodoi</emphasis>
</taxonomicName>
and that its scape was longer than half the length of the funiculus. Both 
<bibRefCitation id="5F144B5DFFFBDC3AFBA9F8499D95F875" author="Smith" box="[1100,1257,1972,1998]" pageId="8" pageNumber="52" refString="Smith, F. (1859) Catalogue of Hymenopterous insects in the collections of the British Museum. Part VII. Dorylidae and Thynnidae. British Museum, London. 76 pp." type="book" year="1859">Smith (1859)</bibRefCitation>
in the original description and later Santschi (1914) mentioned that the posterior head margin of the 
<taxonomicName id="FC854D2FFFFBDC3AFB42F8249C5DF849" box="[1191,1313,2009,2034]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="09F1EABEFFFBDC3AFB42F8249C5DF849" box="[1191,1313,2009,2034]" italics="true" pageId="8" pageNumber="47">D. atratus</emphasis>
</taxonomicName>
male was convex. While the colouration of 
<taxonomicName id="FC854D2FFFFADC3BFDC8FF659BC4FF0A" box="[557,696,152,177]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFDC8FF659BC4FF0A" box="[557,696,152,177]" italics="true" pageId="9" pageNumber="48">D. gribodoi</emphasis>
</taxonomicName>
is variable, it is not uniformly black in any of the specimens examined by us and the head shape is nearly rectangular with a flat posterior margin. Unfortunately, the 
<taxonomicName id="FC854D2FFFFADC3BFA9AFF409996FF40" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="09F1EABEFFFADC3BFA9AFF409996FF40" italics="true" pageId="9" pageNumber="48">D. atratus</emphasis>
</taxonomicName>
<typeStatus id="E43E880EFFFADC3BFF14FF1F985FFF47" box="[241,291,226,252]" pageId="9" pageNumber="48">type</typeStatus>
specimen appears to be lost (George Else pers. comm., Natural History Museum London), so that we could not directly examine the relationship between 
<taxonomicName id="FC854D2FFFFADC3BFCC2FEF59ACCFE9A" box="[807,944,264,289]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFCC2FEF59ACCFE9A" box="[807,944,264,289]" italics="true" pageId="9" pageNumber="48">D. gribodoi</emphasis>
</taxonomicName>
and 
<taxonomicName id="FC854D2FFFFADC3BFC0CFEF59D1CFE9A" box="[1001,1120,264,289]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="09F1EABEFFFADC3BFC0CFEF59D1CFE9A" box="[1001,1120,264,289]" italics="true" pageId="9" pageNumber="48">D. atratus</emphasis>
</taxonomicName>
. However, the distinctness of the taxa appears beyond doubt. Workers of 
<taxonomicName id="FC854D2FFFFADC3BFD4AFED09A4BFEFD" box="[687,823,301,326]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFD4AFED09A4BFEFD" box="[687,823,301,326]" italics="true" pageId="9" pageNumber="48">D. gribodoi</emphasis>
</taxonomicName>
are unique (and easily identifiable) among all 
<taxonomicName id="FC854D2FFFFADC3BFAB3FED099C5FED0" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="09F1EABEFFFADC3BFAB3FED099C5FED0" italics="true" pageId="9" pageNumber="48">Dorylus</emphasis>
</taxonomicName>
species in having a nearly round petiole in dorsal view (
<figureCitation id="A3BE2A29FFFADC3BFCB8FEAF9AD8FED7" box="[861,932,338,364]" captionStart="FIGURE 9" captionStartId="8.[151,255,1031,1055]" captionTargetBox="[385,1200,354,1006]" captionTargetId="figure@8.[385,1202,354,1007]" captionTargetPageId="8" captionText="FIGURE 9. Dorsal view of the petiole of a large D. gribodoi worker (HW 2.85 mm) from Taï, Ivory Coast." httpUri="https://zenodo.org/record/274235/files/figure.png" pageId="9" pageNumber="48">Fig. 9</figureCitation>
) and a distinct ridge on the posterior head margin that is laterally developed into tiny horns in larger specimens (&gt; 
<quantity id="FC7D9B49FFFADC3BFC38FE859D49FE29" box="[989,1077,376,402]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="48" unit="mm" value="1.5">1.5 mm</quantity>
HW). 
<taxonomicName id="FC854D2FFFFADC3BFB60FE859C33FE2A" box="[1157,1359,376,401]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFB60FE859C33FE2A" box="[1157,1359,376,401]" italics="true" pageId="9" pageNumber="48">Dorylus gribodoi</emphasis>
</taxonomicName>
workers of the same size class can be separated from workers of the 
<taxonomicName id="FC854D2FFFFADC3BFC66FE609A9EFE0D" box="[899,994,413,438]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="kohli">
<emphasis id="09F1EABEFFFADC3BFC66FE609A9EFE0D" box="[899,994,413,438]" italics="true" pageId="9" pageNumber="48">D. kohli</emphasis>
</taxonomicName>
/ 
<taxonomicName id="FC854D2FFFFADC3BFC1CFE609DD4FE0D" box="[1017,1192,413,438]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="congolensis">
<emphasis id="09F1EABEFFFADC3BFC1CFE609DD4FE0D" box="[1017,1192,413,438]" italics="true" pageId="9" pageNumber="48">D. congolensis</emphasis>
</taxonomicName>
complex also by the posterior margin of the head being angular in lateral view and not smoothly rounded (
<figureCitation id="A3BE2A29FFFADC3BFB6CFE3F9DAEFE67" box="[1161,1234,450,476]" captionStart="FIGURE 7" captionStartId="7.[151,255,763,787]" captionTargetBox="[468,1117,258,737]" captionTargetId="figure@7.[468,1118,250,738]" captionTargetPageId="7" captionText="FIGURE 7. Lateral view of the head of a large D. gribodoi worker (HW 2.79 mm), Taï, Ivory Coast." httpUri="https://zenodo.org/record/274233/files/figure.png" pageId="9" pageNumber="48">Fig. 7</figureCitation>
). Larger 
<taxonomicName id="FC854D2FFFFADC3BFAD8FE3F999BFDBA" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="09F1EABEFFFADC3BFAD8FE3F999BFDBA" italics="true" pageId="9" pageNumber="48">Dorylus emeryi</emphasis>
</taxonomicName>
workers can be separated from 
<taxonomicName id="FC854D2FFFFADC3BFD84FE159A51FDBA" box="[609,813,488,513]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFD84FE159A51FDBA" box="[609,813,488,513]" italics="true" pageId="9" pageNumber="48">Dorylus gribodoi</emphasis>
</taxonomicName>
workers of the same size class by the following features: posterior angles of head drawn out backwards and ventrally [“Die ziemlich dick-kegeligen, am 
<collectingCountry id="4392763CFFFADC3BFA85FDF09CE0FD9C" box="[1376,1436,525,551]" name="India" pageId="9" pageNumber="48">Ende</collectingCountry>
stark abgerundeten Hinterecken des Kopfes (den Kopf von der Seite gesehen und nach vorne gestreckt gedacht) ziemlich stark hinabgebogen und zwar schief nach hinten und unten gerichtet.” (
<bibRefCitation id="5F144B5DFFFADC3BFB26FDA59C37FDC9" author="Mayr" box="[1219,1355,600,626]" pageId="9" pageNumber="52" refString="Mayr, G. (1896) Beitrage zur Kenntniss der Insektenfauna von Kamerun. 5. Formiciden gesammelt von Herrn Yngve Sjostedt. Entomologisk Tidskrift, 17, 225 - 252." type="journal article" year="1896">Mayr 1896</bibRefCitation>
)] (
<figureCitation id="A3BE2A29FFFADC3BFA88FDA599DBFD2C" captionStart="FIGURE 8" captionStartId="7.[151,255,1430,1454]" captionTargetBox="[409,1159,844,1407]" captionTargetId="figure@7.[409,1160,844,1407]" captionTargetPageId="7" captionText="FIGURE 8. Lateral view of the head of a large D. emeryi worker (HW 3.62 mm) from Taï, Ivory Coast." httpUri="https://zenodo.org/record/274234/files/figure.png" pageId="9" pageNumber="48">Fig. 8</figureCitation>
), petiole in dorsal view angled at widest point (
<figureCitation id="A3BE2A29FFFADC3BFD2AFD809A59FD2C" box="[719,805,637,663]" captionStart="FIGURE 10" captionStartId="8.[151,255,1785,1809]" captionTargetBox="[385,1200,1108,1761]" captionTargetId="figure@8.[385,1202,1108,1761]" captionTargetPageId="8" captionText="FIGURE 10. Dorsal view of the petiole of a large D. emeryi worker (HW 3.74 mm) from Taï, Ivory Coast." httpUri="https://zenodo.org/record/274236/files/figure.png" pageId="9" pageNumber="48">Fig. 10</figureCitation>
), petiole has several conspicuous erect setae on dorsal surface (which are lacking in 
<taxonomicName id="FC854D2FFFFADC3BFDEBFD5F9BE1FD00" box="[526,669,674,699]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFDEBFD5F9BE1FD00" box="[526,669,674,699]" italics="true" pageId="9" pageNumber="48">D. gribodoi</emphasis>
</taxonomicName>
), largest workers have massive heads with width 
<emphasis id="09F1EABEFFFADC3BFAF4FD5F9C5DFD07" bold="true" box="[1297,1313,674,700]" pageId="9" pageNumber="48">≤</emphasis>
<quantity id="FC7D9B49FFFADC3BFACBFD5F9CE7FD07" box="[1326,1435,674,700]" metricMagnitude="-3" metricUnit="m" metricValue="3.92" pageId="9" pageNumber="48" unit="mm" value="3.92">3.92 mm</quantity>
(HWmax of largest 
<taxonomicName id="FC854D2FFFFADC3BFE9BFD359B7AFD5A" box="[382,518,712,737]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFE9BFD359B7AFD5A" box="[382,518,712,737]" italics="true" pageId="9" pageNumber="48">D. gribodoi</emphasis>
</taxonomicName>
workers 
<quantity id="FC7D9B49FFFADC3BFD97FD359BB2FD59" box="[626,718,712,738]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="48" unit="mm" value="2.9">2.9 mm</quantity>
, see above). The unique petiole shape of 
<taxonomicName id="FC854D2FFFFADC3BFB49FD359C48FD5A" box="[1196,1332,712,737]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFB49FD359C48FD5A" box="[1196,1332,712,737]" italics="true" pageId="9" pageNumber="48">D. gribodoi</emphasis>
</taxonomicName>
was recognized by Emery in his original description of 
<taxonomicName id="FC854D2FFFFADC3BFD3BFD109D3AFCBC" authority="Emery 1895" authorityName="Emery" authorityYear="1895" box="[734,1094,749,775]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gerstaeckeri">
<emphasis id="09F1EABEFFFADC3BFD3BFD109AEAFCBD" box="[734,918,749,774]" italics="true" pageId="9" pageNumber="48">D. gerstaeckeri</emphasis>
(
<bibRefCitation id="5F144B5DFFFADC3BFC4DFD109D42FCBC" author="Emery" box="[936,1086,749,775]" pageId="9" pageNumber="51" refString="Emery, C. (1895) Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbucher Abteilung fur Systematik Okologie und Geographie der Tiere, 8, 685 - 778." type="journal article" year="1895">Emery 1895</bibRefCitation>
)
</taxonomicName>
and also by Bernard (1952) who described 
<taxonomicName id="FC854D2FFFFADC3BFEA8FCEF98AFFC90" box="[333,467,786,811]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="lamottei">
<emphasis id="09F1EABEFFFADC3BFEA8FCEF98AFFC90" box="[333,467,786,811]" italics="true" pageId="9" pageNumber="48">D. lamottei</emphasis>
</taxonomicName>
based on workers collected at Mount Nimba in 
<collectingCountry id="4392763CFFFADC3BFBF8FCEF9D0EFC97" box="[1053,1138,786,812]" name="Guinea-Bissau" pageId="9" pageNumber="48">Guinea</collectingCountry>
but failed to realize that these specimens were in fact conspecific with Emery’s 
<taxonomicName id="FC854D2FFFFADC3BFCC0FCC59AA7FCEA" box="[805,987,824,849]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gerstaeckeri">
<emphasis id="09F1EABEFFFADC3BFCC0FCC59AA7FCEA" box="[805,987,824,849]" italics="true" pageId="9" pageNumber="48">D. gerstaeckeri</emphasis>
</taxonomicName>
. 
<bibRefCitation id="5F144B5DFFFADC3BFC0EFCC59DDCFCE9" author="Bernard" box="[1003,1184,824,850]" pageId="9" pageNumber="51" refString="Bernard, F. (1953) La reserve naturelle integrale du Mt Nimba. XI. Hymenopteres, Formicidae. Memoires de l'Institut Francais d'Afrique Noire, 19, 165 - 270." type="journal article" year="1953">Bernard (1953)</bibRefCitation>
also identified males taken from the same location as 
<taxonomicName id="FC854D2FFFFADC3BFDC3FCA09BCFFCCD" box="[550,691,861,886]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFDC3FCA09BCFFCCD" box="[550,691,861,886]" italics="true" pageId="9" pageNumber="48">D. gribodoi</emphasis>
</taxonomicName>
, but the association between these males and his 
<taxonomicName id="FC854D2FFFFADC3BFAF1FCA09CE1FCCD" box="[1300,1437,861,886]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="lamottei">
<emphasis id="09F1EABEFFFADC3BFAF1FCA09CE1FCCD" box="[1300,1437,861,886]" italics="true" pageId="9" pageNumber="48">D. lamottei</emphasis>
</taxonomicName>
workers necessarily remained unknown to him.
</paragraph>
<caption id="6FFA6624FFFADC3BFF72F98F9DB0F931" box="[151,1228,1650,1674]" httpUri="https://zenodo.org/record/274237/files/figure.png" pageId="9" pageNumber="48" targetBox="[389,1198,978,1626]" targetPageId="9">
<paragraph id="3B3A36ACFFFADC3BFF72F98F9DB0F931" blockId="9.[151,1228,1650,1674]" box="[151,1228,1650,1674]" pageId="9" pageNumber="48">
<emphasis id="09F1EABEFFFADC3BFF72F98F9859F931" bold="true" box="[151,293,1650,1674]" pageId="9" pageNumber="48">FIGURE 11.</emphasis>
Lateral overview of the 
<taxonomicName id="FC854D2FFFFADC3BFDC9F98E9BD4F931" box="[556,680,1651,1674]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFDC9F98E9BD4F931" box="[556,680,1651,1674]" italics="true" pageId="9" pageNumber="48">D. gribodoi</emphasis>
</taxonomicName>
queen collected from a nest at Lamto, Ivory Coast.
</paragraph>
</caption>
<paragraph id="3B3A36ACFFFADC3BFF20F9409AD6F89C" blockId="9.[151,1436,1725,2031]" pageId="9" pageNumber="48">
A 
<taxonomicName id="FC854D2FFFFADC3BFF07F9439810F96C" box="[226,364,1726,1751]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFF07F9439810F96C" box="[226,364,1726,1751]" italics="true" pageId="9" pageNumber="48">D. gribodoi</emphasis>
</taxonomicName>
queen was collected along with workers from a nest at Lamto (
<collectingCountry id="4392763CFFFADC3BFBB3F9409D98F96C" box="[1110,1252,1725,1751]" name="Ivory Coast" pageId="9" pageNumber="48">Ivory Coast</collectingCountry>
) by Jean-Marie Leroux and below we provide the queen description. 
<taxonomicName id="FC854D2FFFFADC3BFCFFF91B9A9BF944" box="[794,999,1766,1791]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFFADC3BFCFFF91B9A9BF944" box="[794,999,1766,1791]" italics="true" pageId="9" pageNumber="48">Dorylus gribodoi</emphasis>
</taxonomicName>
is thus now one of the few 
<taxonomicName id="FC854D2FFFFADC3BFAD8F91B9CE7F944" box="[1341,1435,1766,1791]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="09F1EABEFFFADC3BFAD8F91B9CE7F944" box="[1341,1435,1766,1791]" italics="true" pageId="9" pageNumber="48">Dorylus</emphasis>
</taxonomicName>
species for which workers, the male and the queen are now known.
</paragraph>
</subSubSection>
<subSubSection id="739F6527FFFADC38FF20F8C8999FFD59" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" type="description">
<paragraph id="3B3A36ACFFFADC38FF20F8C8999FFD59" blockId="9.[151,1436,1725,2031]" lastBlockId="10.[151,1436,152,738]" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48">
<emphasis id="09F1EABEFFFADC3BFF20F8C898CFF8F4" bold="true" box="[197,435,1845,1871]" pageId="9" pageNumber="48">Queen description.</emphasis>
Habitus as in 
<figureCitation id="A3BE2A29FFFADC3BFDBAF8C89BAFF8F4" box="[607,723,1845,1871]" captionStart="FIGURE 11" captionStartId="9.[151,255,1650,1674]" captionTargetBox="[389,1198,978,1626]" captionTargetId="figure@9.[389,1198,978,1626]" captionTargetPageId="9" captionText="FIGURE 11. Lateral overview of the D. gribodoi queen collected from a nest at Lamto, Ivory Coast." httpUri="https://zenodo.org/record/274237/files/figure.png" pageId="9" pageNumber="48">Figure 11</figureCitation>
. Head: Antenna 11-segmented (
<figureCitation id="A3BE2A29FFFADC3BFBABF8C89DDAF8F4" box="[1102,1190,1845,1871]" captionStart="FIGURE 12" captionStartId="10.[151,255,1514,1538]" captionText="FIGURE 12. Frontal view of the head of the D. gribodoi queen (HW 4.58 mm)." pageId="9" pageNumber="48">Fig. 12</figureCitation>
), funicular segments and scape yellow; head glossy, finely punctate, without conspicuous pilosity, antennal fossae deeply impressed; small (
<quantity id="FC7D9B49FFFADC3BFE95F87898A1F824" box="[368,477,1925,1951]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="9" pageNumber="48" unit="mm" value="0.18">0.18 mm</quantity>
diameter) ocellus-like body located on the midline of head in frontal view 
<quantity id="FC7D9B49FFFADC3BFA8DF87899B9F87C" metricMagnitude="-3" metricUnit="m" metricValue="1.98" pageId="9" pageNumber="48" unit="mm" value="1.98">1.98 mm</quantity>
dorsad of the anterior clypeal border (
<figureCitation id="A3BE2A29FFFADC3BFD60F8509BA0F87C" box="[645,732,1965,1991]" captionStart="FIGURE 12" captionStartId="10.[151,255,1514,1538]" captionText="FIGURE 12. Frontal view of the head of the D. gribodoi queen (HW 4.58 mm)." pageId="9" pageNumber="48">Fig. 12</figureCitation>
); head impressed along the midline; mandibles with smooth medial (masticatory) border (i.e., without subapical teeth), sharply pointed apical tooth directed medially; clypeal margin straight with a row of short setae; labrum smoothly emarginate medially and weakly bilobed, reminiscent of the labrum in queens of the New World genus 
<taxonomicName id="FC854D2FFFF9DC38FC80FF3D9ACDFF62" box="[869,945,192,217]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="09F1EABEFFF9DC38FC80FF3D9ACDFF62" box="[869,945,192,217]" italics="true" pageId="10" pageNumber="49">Eciton</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="5F144B5DFFF9DC38FC0AFF3D9DE6FF61" author="Gotwald" box="[1007,1178,192,218]" pageId="10" pageNumber="51" refString="Gotwald, W. H., Jr. (1969) Comparative morphological studies of the ants, with particular reference to the mouthparts (Hymenoptera: Formicidae). Cornell University Agricultural Experiment Station Memoirs, 408, 1 - 150." type="journal article" year="1969">Gotwald 1969</bibRefCitation>
), not of other 
<taxonomicName id="FC854D2FFFF9DC38FAD8FF3D9CE7FF62" box="[1341,1435,192,217]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="09F1EABEFFF9DC38FAD8FF3D9CE7FF62" box="[1341,1435,192,217]" italics="true" pageId="10" pageNumber="49">Dorylus</emphasis>
</taxonomicName>
queens; head deeply impressed posteriorly at pronotal insertion, posterior angles rounded, directed posteriorly (
<figureCitation id="A3BE2A29FFF9DC38FF7AFEED9985FE91" box="[159,249,272,298]" captionStart="FIGURE 14" captionStartId="11.[151,255,1667,1691]" captionText="FIGURE 14. Dorsal view of D. gribodoi queen mesosoma (PeW 4.54 mm)." pageId="10" pageNumber="49">Fig. 14</figureCitation>
). Head and mandibles orange brown. Mesosoma: Similar in colour to head except for dark reddishbrown patches on dorsum of pronotum, mesonotum, and propodeum. Pronotal suture deeply impressed; mesonotum, metanotum and propodeum separated by shallow but well-defined sutures (
<figureCitation id="A3BE2A29FFF9DC38FB4DFE9D9D83FEC1" box="[1192,1279,352,378]" captionStart="FIGURE 14" captionStartId="11.[151,255,1667,1691]" captionText="FIGURE 14. Dorsal view of D. gribodoi queen mesosoma (PeW 4.54 mm)." pageId="10" pageNumber="49">Fig. 14</figureCitation>
); propodeum in dorsal view rectilinear (
<figureCitation id="A3BE2A29FFF9DC38FE2CFE759B5CFE19" box="[457,544,392,418]" captionStart="FIGURE 14" captionStartId="11.[151,255,1667,1691]" captionText="FIGURE 14. Dorsal view of D. gribodoi queen mesosoma (PeW 4.54 mm)." pageId="10" pageNumber="49">Fig. 14</figureCitation>
); mesosoma finely punctate without conspicuous pilosity. Petiole: Dark reddish-brown, finely punctate, without conspicuous setae or pilosity; posterior lateral lobes conspicuous in dorsal view (
<figureCitation id="A3BE2A29FFF9DC38FEE8FE25981BFE49" box="[269,359,472,498]" captionStart="FIGURE 14" captionStartId="11.[151,255,1667,1691]" captionText="FIGURE 14. Dorsal view of D. gribodoi queen mesosoma (PeW 4.54 mm)." pageId="10" pageNumber="49">Fig. 14</figureCitation>
). Gaster: Segments reddish-brown to orange-brown, finely punctate, without conspicuous pilosity; posterior margin of pygidium in dorsal view with deep semicircular notch medially, sharp, downward pointed horns laterally (
<figureCitation id="A3BE2A29FFF9DC38FE4AFDD59B73FDF9" box="[431,527,552,578]" captionStart="FIGURE 11" captionStartId="9.[151,255,1650,1674]" captionTargetBox="[389,1198,978,1626]" captionTargetId="figure@9.[389,1198,978,1626]" captionTargetPageId="9" captionText="FIGURE 11. Lateral overview of the D. gribodoi queen collected from a nest at Lamto, Ivory Coast." httpUri="https://zenodo.org/record/274237/files/figure.png" pageId="10" pageNumber="49">Figs. 11</figureCitation>
, 
<figureCitation id="A3BE2A29FFF9DC38FDF9FDD59B40FDF9" box="[540,572,552,578]" captionStart="FIGURE 13" captionStartId="11.[151,255,913,937]" captionTargetBox="[363,1229,194,892]" captionTargetId="figure@11.[363,1231,194,892]" captionTargetPageId="11" captionText="FIGURE 13. Dorsal view of the hypopygium of the D. gribodoi queen (HAD 2.53 mm)." httpUri="https://zenodo.org/record/274238/files/figure.png" pageId="10" pageNumber="49">13</figureCitation>
). The two lobes of the hypopygium of the 
<taxonomicName id="FC854D2FFFF9DC38FBD4FDD59DC6FDFA" box="[1073,1210,552,577]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="gribodoi">
<emphasis id="09F1EABEFFF9DC38FBD4FDD59DC6FDFA" box="[1073,1210,552,577]" italics="true" pageId="10" pageNumber="49">D. gribodoi</emphasis>
</taxonomicName>
queen terminate in apically pointed processes (
<figureCitation id="A3BE2A29FFF9DC38FE3DFDAD9B50FDD1" box="[472,556,592,618]" captionStart="FIGURE 13" captionStartId="11.[151,255,913,937]" captionTargetBox="[363,1229,194,892]" captionTargetId="figure@11.[363,1231,194,892]" captionTargetPageId="11" captionText="FIGURE 13. Dorsal view of the hypopygium of the D. gribodoi queen (HAD 2.53 mm)." httpUri="https://zenodo.org/record/274238/files/figure.png" pageId="10" pageNumber="49">Fig. 13</figureCitation>
) as in 
<taxonomicName id="FC854D2FFFF9DC38FD9CFDAD9BABFDD2" box="[633,727,592,617]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="kohli">
<emphasis id="09F1EABEFFF9DC38FD9CFDAD9BABFDD2" box="[633,727,592,617]" italics="true" pageId="10" pageNumber="49">D. kohli</emphasis>
</taxonomicName>
(currently classified in 
<taxonomicName id="FC854D2FFFF9DC38FC0FFDAD9D2EFDD2" box="[1002,1106,592,617]" class="Insecta" family="Formicidae" genus="Anomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="09F1EABEFFF9DC38FC0FFDAD9D2EFDD2" box="[1002,1106,592,617]" italics="true" pageId="10" pageNumber="49">Anomma</emphasis>
</taxonomicName>
) (van Boven 1968), 
<taxonomicName id="FC854D2FFFF9DC38FAA6FDAD999CFD2A" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="ghanensis">
<emphasis id="09F1EABEFFF9DC38FAA6FDAD999CFD2A" italics="true" pageId="10" pageNumber="49">D. ghanensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="FC854D2FFFF9DC38FEFCFD8598B8FD2A" box="[281,452,632,657]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="09F1EABEFFF9DC38FEFCFD8598B8FD2A" box="[281,452,632,657]" italics="true" pageId="10" pageNumber="49">D. brevipennis</emphasis>
</taxonomicName>
(both classified in the subgenus 
<taxonomicName id="FC854D2FFFF9DC38FCA3FD859AD8FD2A" box="[838,932,632,657]" class="Insecta" family="Formicidae" genus="Dorylus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="09F1EABEFFF9DC38FCA3FD859AD8FD2A" box="[838,932,632,657]" italics="true" pageId="10" pageNumber="49">Dorylus</emphasis>
</taxonomicName>
) (van Boven 1975), whereas the two lobes are very broad posteriorly in driver ant queen hypopygia (see e.g. Figs. 20a and 20b in Raignier &amp; van 
<bibRefCitation id="5F144B5DFFF9DC38FAAAFD5D99AAFD59" author="Boven" pageId="10" pageNumber="52" refString="Raignier, A. &amp; Boven, J. K. A. van (1955) Etude taxonomique, biologique et biometrique des Dorylus du sous-genre Anomma (Hymenoptera Formicidae). Annales du Musee Royal du Congo Belge. Tervuren. Sciences Zoologiques. Serie, 2, 1 - 359." type="journal article" year="1955">Boven 1955</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>