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<document id="65362E6C2549DD87F01BE263CCCE410D" ID="10.11646/zootaxa.4235.1.1SLASH10244" ID-CLB-Dataset="36483" ID-DOI="10.5281/zenodo.322062" ID-GBIF-Dataset="29139646-2af5-4d4f-a7a4-b03c54cb4af8" ID-ISSN="1175-5326" ID-Zenodo-Dep="322062" ID-ZooBank="AEA565CC-1A14-4987-977A-1C269B0FE2A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487671437888" checkinUser="plazi" docAuthor="Garrison, Rosser W. &amp; Ellenrieder, Natalia Von" docDate="2017" docId="A15040439A2AFFD2FF39F98AFBDFFD9B" docLanguage="en" docName="zootaxa.4235.1.1SLASH10244.pdf" docOrigin="Zootaxa 4235 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Argia schneideri Garrison &amp; von Ellenrieder, n. sp." docType="treatment" docVersion="8" lastPageNumber="27" masterDocId="5D69383B9A32FFC8FFAEFF9EFF8AFFA4" masterDocTitle="New species of the damselfly genus Argia from Mexico, Central America and Ecuador with an emphasis on Costa Rica (Insecta: Odonata: Coenagrionidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="25" updateTime="1698437136683" updateUser="ExternalLinkService">
<mods:mods id="F2C377D1DB0AF18E6FB9C7BBA617D92F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8BF4481B8B4EDDBBFA5CC065BC174B2E">
<mods:title id="53F1C496D98BC6931554C90AB7DCE2C8">New species of the damselfly genus Argia from Mexico, Central America and Ecuador with an emphasis on Costa Rica (Insecta: Odonata: Coenagrionidae)</mods:title>
</mods:titleInfo>
<mods:name id="EC28876D15BEBAC88A6D0092E7D65B75" type="personal">
<mods:role id="30DEE5F9A425DC5567EC73C5E67E9043">
<mods:roleTerm id="A5370096439E267C998FB6E13E942E5A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="964BF2AE0A328E5E5F93BA00286ECE8F">Garrison, Rosser W.</mods:namePart>
</mods:name>
<mods:name id="214445099222D84E5CD1D0FCFCC417AB" type="personal">
<mods:role id="14A63BA117EDE52D453792C1637E4866">
<mods:roleTerm id="B943B79118118C734E410726B783AD53">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB8C2CE1518950BA633595CA92A1142F">Ellenrieder, Natalia Von</mods:namePart>
</mods:name>
<mods:typeOfResource id="92209B3CF04D5DE7C7404B99F6E1435E">text</mods:typeOfResource>
<mods:relatedItem id="6AE08FD00958494B3FB557D3150CD972" type="host">
<mods:titleInfo id="5BF0F5FC7FECFD52191B14E04216BE1B">
<mods:title id="F255E375E22213A259BE8C3747BC7ADD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7C269C00C902A1BC3CE9281EDE98674F">
<mods:date id="3885C00DCEA1257BA218311BCA5E91E6">2017</mods:date>
<mods:detail id="6BC9F8B33B0FD01373A9C84AC31AB376" type="volume">
<mods:number id="C2907AD21F15E75C28EE4B01109C48F3">4235</mods:number>
</mods:detail>
<mods:detail id="52B16A80B204568D6FA66F647F66241F" type="issue">
<mods:number id="B5327EE86E0DD0BFB934F4DA20CE6124">1</mods:number>
</mods:detail>
<mods:extent id="28C4464094FD18D3E02476951A23B45B" unit="page">
<mods:start id="686810917D431B8C5AD2C97EA10F70A8">1</mods:start>
<mods:end id="E63CD75F45948A0A7847524E257B7C3E">93</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="66986F8353AA8BE7F494EBA393B7A6AA">journal article</mods:classification>
<mods:identifier id="8BF0F1B150DBF1ABCA4A47518BA77875" type="CLB-Dataset">36483</mods:identifier>
<mods:identifier id="C0358D812D96095486D931466437A1CE" type="DOI">10.5281/zenodo.322062</mods:identifier>
<mods:identifier id="0C9AEA0212FAD82D8FEBBDB6D45DA2EF" type="GBIF-Dataset">29139646-2af5-4d4f-a7a4-b03c54cb4af8</mods:identifier>
<mods:identifier id="DCF3CE1D1011062BB80C3780255C2649" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="602B0A56BE72388A392D96DCE424C99C" type="Zenodo-Dep">322062</mods:identifier>
<mods:identifier id="337284B41048F268ABE063DD21F0C086" type="ZooBank">AEA565CC-1A14-4987-977A-1C269B0FE2A8</mods:identifier>
</mods:mods>
<treatment id="A15040439A2AFFD2FF39F98AFBDFFD9B" ID-DOI="http://doi.org/10.5281/zenodo.6030017" ID-GBIF-Taxon="127697284" ID-Zenodo-Dep="6030017" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B" httpUri="http://treatment.plazi.org/id/A15040439A2AFFD2FF39F98AFBDFFD9B" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
<subSubSection id="61E3A2DE9A2AFFD0FF39F98AFBBBF9D7" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="2946F1559A2AFFD0FF39F98AFC9BF98A" blockId="24.[151,1436,1556,1651]" box="[151,785,1556,1582]" pageId="24" pageNumber="25">
<heading id="720E46399A2AFFD0FF39F98AFC9BF98A" bold="true" box="[151,785,1556,1582]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="1B8D2D479A2AFFD0FF39F98AFC9BF98A" bold="true" box="[151,785,1556,1582]" pageId="24" pageNumber="25">
<taxonomicName id="EEF98AD69A2AFFD0FF39F98AFD48F98A" ID-CoL="9X234" authority="Garrison &amp; von Ellenrieder" authorityName="Garrison &amp; von Ellenrieder" box="[151,706,1556,1582]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="schneideri" status="sp. nov.">
<emphasis id="1B8D2D479A2AFFD0FF39F98AFEEAF98A" bold="true" box="[151,352,1556,1582]" italics="true" pageId="24" pageNumber="25">Argia schneideri</emphasis>
Garrison &amp; von Ellenrieder
</taxonomicName>
,
<taxonomicNameLabel id="00BE903C9A2AFFD0FD61F98AFC9BF98A" box="[719,785,1556,1582]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2946F1559A2AFFD0FF39F9A8FBBBF9D7" blockId="24.[151,1436,1556,1651]" pageId="24" pageNumber="25">
<figureCitation id="B1C2EDD09A2AFFD0FF39F9A8FF7BF9EB" box="[151,241,1590,1615]" captionStart="FIGURES 24 – 26" captionStartId="45.[151,264,1836,1859]" captionTargetBox="[160,1408,215,1680]" captionTargetId="figure@45.[151,1436,193,1815]" captionTargetPageId="45" captionText="FIGURES 24 – 26. Color pattern of male head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322068/files/figure.png" pageId="24" pageNumber="25">Figs. 25</figureCitation>
(head, thorax, S1–
<specimenCount id="3FFF3ADC9A2AFFD0FE6AF9A9FE78F9EB" box="[452,498,1591,1615]" pageId="24" pageNumber="25" type="male">4 ♂</specimenCount>
); 50 (head, thorax, S1–
<specimenCount id="3FFF3ADC9A2AFFD0FD57F9A9FCA8F9EB" box="[761,802,1591,1615]" pageId="24" pageNumber="25" type="female">3 ♀</specimenCount>
); 74 (S7–
<specimenCount id="3FFF3ADC9A2AFFD0FC3DF9A9FC44F9EB" box="[915,974,1591,1615]" pageId="24" pageNumber="25" type="male">10 ♂</specimenCount>
), 99 (S7–
<specimenCount id="3FFF3ADC9A2AFFD0FB95F9A9FBF8F9EB" box="[1083,1138,1591,1615]" pageId="24" pageNumber="25" type="female">10 ♀</specimenCount>
); 120 (mesostigmal plates
<specimenCount id="3FFF3ADC9A2AFFD0FF39F9C5FF21F9D7" box="[151,171,1627,1651]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
); 139 (genital ligula); 159 (appendages
<specimenCount id="3FFF3ADC9A2AFFD0FDC3F9C5FD0EF9D7" box="[621,644,1627,1651]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
); 169 (map);
<tableCitation id="647BC4EE9A2AFFD0FCB3F9C4FCFAF9D7" box="[797,880,1626,1651]" captionStart="TABLE 4" captionStartId="20.[119,197,507,527]" captionTargetBox="[119,1761,656,1074]" captionTargetPageId="20" captionText="TABLE 4. Wing counts and measurements (average and standard deviation followed by range in square brackets) for various Argia species treated in this study." httpUri="http://table.plazi.org/id/7D86A1DD9A26FFDCFFD9FE65F9B0FDAB" pageId="24" pageNumber="25" tableUuid="7D86A1DD9A26FFDCFFD9FE65F9B0FDAB">Table 4</tableCitation>
(measurements).
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A2AFFD1FF39F93CFB3BFDC4" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="etymology">
<paragraph id="2946F1559A2AFFD0FF39F93CFA02F8A7" blockId="24.[151,1436,1698,2011]" pageId="24" pageNumber="25">
<emphasis id="1B8D2D479A2AFFD0FF39F93CFE9CF91F" bold="true" box="[151,278,1698,1723]" pageId="24" pageNumber="25">Etymology</emphasis>
. Named
<taxonomicName id="EEF98AD69A2AFFD0FE2AF93CFE76F91F" box="[388,508,1698,1723]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="1B8D2D479A2AFFD0FE2AF93CFE76F91F" bold="true" box="[388,508,1698,1723]" italics="true" pageId="24" pageNumber="25">schneideri</emphasis>
</taxonomicName>
(Latinized name) in honor of our friend and colleague Wolfgang Schneider, in recognition of his valuable contributions to the study of
<taxonomicName id="EEF98AD69A2AFFD0FCA8F958FCEFF97B" box="[774,869,1734,1759]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
and his active part in establishing the International Dragonfly Fund, including personal donations, which have helped further our knowledge of
<taxonomicName id="EEF98AD69A2AFFD0FB0DF974FA88F8A7" box="[1187,1282,1770,1795]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
worldwide.
</paragraph>
<paragraph id="2946F1559A2AFFD1FF69F890FAE3FD98" blockId="24.[151,1436,1698,2011]" lastBlockId="25.[151,1437,151,2016]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="1B8D2D479A2AFFD0FF69F890FE35F883" bold="true" box="[199,447,1806,1831]" pageId="24" pageNumber="25">Specimens examined</emphasis>
. 35 ♂, 5 ♀.
<emphasis id="1B8D2D479A2AFFD0FDE1F890FD19F883" bold="true" box="[591,659,1806,1831]" pageId="24" pageNumber="25">Types</emphasis>
.
<materialsCitation id="9991FB089A2AFFD0FD0DF890FC39F8EE" ID-GBIF-Occurrence="1434659049" collectingDate="1936-12-11" collectionCode="UMMZ" collectorName="W. Clark-Macintyre" country="Ecuador" location="Rio Anzu" municipality="Las Palmas" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="holotype">
<emphasis id="1B8D2D479A2AFFD0FD0DF890FC9AF883" bold="true" box="[675,784,1806,1831]" pageId="24" pageNumber="25">
<typeStatus id="F6424FF79A2AFFD0FD0DF890FC9AF883" box="[675,784,1806,1831]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="3FFF3ADC9A2AFFD0FCB9F891FCA5F882" box="[791,815,1807,1830]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
:
<collectingCountry id="51EEB1C59A2AFFD0FC92F891FC4CF883" box="[828,966,1807,1831]" name="Ecuador" pageId="24" pageNumber="25">ECUADOR</collectingCountry>
,
<collectingRegion id="EB3D3FB79A2AFFD0FC7CF891FB85F882" box="[978,1039,1807,1830]" country="Ecuador" name="Napo" pageId="24" pageNumber="25">Napo</collectingRegion>
Prov.,
<collectingMunicipality id="C9226B2F9A2AFFD0FBF1F891FB69F883" box="[1119,1251,1806,1831]" pageId="24" pageNumber="25">Las Palmas</collectingMunicipality>
, on
<location id="2C26A78E9A2AFFD0FABCF891FAF4F883" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B:2C26A78E9A2AFFD0FABCF891FAF4F883" box="[1298,1406,1807,1831]" country="Ecuador" municipality="Las Palmas" name="Rio Anzu" pageId="24" pageNumber="25" stateProvince="Napo">Río Anzu</location>
in Río
<collectingRegion id="EB3D3FB79A2AFFD0FF67F8ADFE8CF8EE" box="[201,262,1843,1866]" country="Ecuador" name="Napo" pageId="24" pageNumber="25">Napo</collectingRegion>
watershed,
<date id="5D47D7959A2AFFD0FE3FF8ADFD92F8EF" box="[401,536,1842,1867]" pageId="24" pageNumber="25" value="1936-12-11">
<collectingDate id="4D032E7D9A2AFFD0FE3FF8ADFD92F8EF" box="[401,536,1842,1867]" pageId="24" pageNumber="25" value="1936-12-11">11 xii 1936</collectingDate>
</date>
,
<collectorName id="840C94839A2AFFD0FD88F8ADFC80F8EF" box="[550,778,1842,1867]" pageId="24" pageNumber="25">W. Clark-Macintyre</collectorName>
leg. [
<collectionCode id="4FE869909A2AFFD0FCE2F8ADFC26F8EE" box="[844,940,1843,1866]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="24" pageNumber="25">UMMZ</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="9991FB089A2AFFD0FC6EF8ADFAEAF8CA" ID-GBIF-Occurrence="1434659032" collectingDate="1942-09-18" collectionCode="UMMZ" country="Ecuador" elevation="500" latitude="-0.98333335" location="Ishpinga-yacu" longLatPrecision="1308" longitude="-77.833336" pageId="24" pageNumber="25" specimenCount="3" specimenCount-male="3" stateProvince="Napo" typeStatus="paratype">
<typeStatus id="F6424FF79A2AFFD0FC6EF8ADFBB8F8EF" box="[960,1074,1843,1867]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="51EEB1C59A2AFFD0FB91F8ADFB43F8EF" box="[1087,1225,1843,1867]" name="Ecuador" pageId="24" pageNumber="25">ECUADOR</collectingCountry>
,
<collectingRegion id="EB3D3FB79A2AFFD0FB79F8ADFA9EF8EE" box="[1239,1300,1843,1866]" country="Ecuador" name="Napo" pageId="24" pageNumber="25">Napo</collectingRegion>
Prov.:
<specimenCount id="3FFF3ADC9A2AFFD0FAC7F8ADFA1DF8EE" box="[1385,1431,1843,1867]" pageId="24" pageNumber="25" type="male">3 ♂</specimenCount>
, upper E branch of
<location id="2C26A78E9A2AFFD0FEC6F8C8FD8DF8CB" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B:2C26A78E9A2AFFD0FEC6F8C8FD8DF8CB" box="[360,519,1878,1903]" country="Ecuador" latitude="-0.98333335" longLatPrecision="1308" longitude="-77.833336" name="Ishpinga-yacu" pageId="24" pageNumber="25" stateProvince="Napo">Ishpinga-yacu</location>
{
<geoCoordinate id="4CCD97929A2AFFD0FDB8F8C8FDE4F8CB" box="[534,622,1878,1903]" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-0.98333335">0°59' S</geoCoordinate>
,
<geoCoordinate id="4CCD97929A2AFFD0FDD4F8C8FD6FF8CA" box="[634,741,1878,1903]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="-77.833336">77°50' W</geoCoordinate>
,
<quantity id="EE015CB09A2AFFD0FD5EF8C8FCB7F8CB" box="[752,829,1878,1903]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="500.0">
<elevation id="A2D416669A2AFFD0FD5EF8C8FCB7F8CB" box="[752,829,1878,1903]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="500.0">500 m</elevation>
</quantity>
},
<date id="5D47D7959A2AFFD0FCFCF8C9FC47F8CB" box="[850,973,1878,1903]" pageId="24" pageNumber="25" value="1942-09-18">
<collectingDate id="4D032E7D9A2AFFD0FCFCF8C9FC47F8CB" box="[850,973,1878,1903]" pageId="24" pageNumber="25" value="1942-09-18">18 ix 1942</collectingDate>
</date>
, W. Clarke-McIntyre leg. [
<collectionCode id="4FE869909A2AFFD0FB57F8C9FAD3F8CA" box="[1273,1369,1879,1902]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="24" pageNumber="25">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2AFFD0FAC5F8C9FE2BF813" ID-GBIF-Occurrence="1434659047" collectingDate="1935-07-27" collectionCode="CSCA" collectorName="El Partidero &amp; Macintyre" country="Ecuador" elevation="700" latitude="-1.0166667" location="El Partidero" longLatPrecision="1308" longitude="-77.833336" municipality="Cushillu-Yacu" pageId="24" pageNumber="25" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2AFFD0FAC5F8C9FA1DF8CA" box="[1387,1431,1879,1903]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
,
<specimenCount id="3FFF3ADC9A2AFFD0FF39F8E5FF4CF836" box="[151,198,1915,1939]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="C9226B2F9A2AFFD0FF74F8E4FEF4F837" box="[218,382,1914,1939]" pageId="24" pageNumber="25">Cushillu-Yacu</collectingMunicipality>
, Río
<collectingRegion id="EB3D3FB79A2AFFD0FE6DF8E5FD8AF836" box="[451,512,1915,1938]" country="Ecuador" name="Napo" pageId="24" pageNumber="25">Napo</collectingRegion>
watershed,
<location id="2C26A78E9A2AFFD0FD3FF8E4FC97F837" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B:2C26A78E9A2AFFD0FD3FF8E4FC97F837" box="[657,797,1914,1939]" country="Ecuador" latitude="-1.0166667" longLatPrecision="1308" longitude="-77.833336" municipality="Cushillu-Yacu" name="El Partidero" pageId="24" pageNumber="25" stateProvince="Napo">
<collectorName id="840C94839A2AFFD0FD3FF8E4FC97F837" box="[657,797,1914,1939]" pageId="24" pageNumber="25">El Partidero</collectorName>
</location>
{
<geoCoordinate id="4CCD97929A2AFFD0FC9BF8E4FC0AF837" box="[821,896,1914,1939]" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-1.0166667">1°1' S</geoCoordinate>
,
<geoCoordinate id="4CCD97929A2AFFD0FC3CF8E4FB88F836" box="[914,1026,1914,1939]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="-77.833336">77°50' W</geoCoordinate>
,
<quantity id="EE015CB09A2AFFD0FBBCF8E5FBEFF837" box="[1042,1125,1914,1939]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="25" unit="m" value="700.0">
<elevation id="A2D416669A2AFFD0FBBCF8E5FBEFF837" box="[1042,1125,1914,1939]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="25" unit="m" value="700.0">700 m</elevation>
</quantity>
},
<date id="5D47D7959A2AFFD0FBD1F8E5FA87F837" box="[1151,1293,1914,1939]" pageId="24" pageNumber="25" value="1935-07-27">
<collectingDate id="4D032E7D9A2AFFD0FBD1F8E5FA87F837" box="[1151,1293,1914,1939]" pageId="24" pageNumber="25" value="1935-07-27">27 vii 1935</collectingDate>
</date>
, W. Clarke-
<collectorName id="840C94839A2AFFD0FF39F800FE80F813" box="[151,266,1950,1975]" pageId="24" pageNumber="25">Macintyre</collectorName>
leg. [
<collectionCode id="4FE869909A2AFFD0FEE6F801FE10F813" box="[328,410,1951,1975]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="24" pageNumber="25">CSCA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2AFFD0FE02F801FD0FF87E" ID-GBIF-Occurrence="1434659034" collectingDate="1936-12-11" collectionCode="UMMZ" collectorName="W. Clark-Macintyre" country="Ecuador" elevation="900" latitude="-1.0666667" location="Rio Anzu" longLatPrecision="1308" longitude="-77.8" municipality="Las Palmas" pageId="24" pageNumber="25" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2AFFD0FE02F801FE52F812" box="[428,472,1951,1975]" pageId="24" pageNumber="25" type="male">2 ♂</specimenCount>
,
<specimenCount id="3FFF3ADC9A2AFFD0FE4AF801FD84F812" box="[484,526,1951,1975]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="C9226B2F9A2AFFD0FDB3F801FD2AF813" box="[541,672,1950,1975]" pageId="24" pageNumber="25">Las Palmas</collectingMunicipality>
, on
<location id="2C26A78E9A2AFFD0FD61F801FCB1F813" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B:2C26A78E9A2AFFD0FD61F801FCB1F813" box="[719,827,1951,1975]" country="Ecuador" latitude="-1.0666667" longLatPrecision="1308" longitude="-77.8" municipality="Las Palmas" name="Rio Anzu" pageId="24" pageNumber="25" stateProvince="Napo">Río Anzu</location>
in Río
<collectingRegion id="EB3D3FB79A2AFFD0FC20F801FC41F812" box="[910,971,1951,1974]" country="Ecuador" name="Napo" pageId="24" pageNumber="25">Napo</collectingRegion>
watershed {
<geoCoordinate id="4CCD97929A2AFFD0FBFBF800FB17F813" box="[1109,1181,1950,1975]" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-1.0666667">1°4' S</geoCoordinate>
,
<geoCoordinate id="4CCD97929A2AFFD0FB07F801FA9EF812" box="[1193,1300,1951,1975]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="-77.8">77°48' W</geoCoordinate>
,
<quantity id="EE015CB09A2AFFD0FAB1F801FAE6F813" box="[1311,1388,1950,1975]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="25" unit="m" value="900.0">
<elevation id="A2D416669A2AFFD0FAB1F801FAE6F813" box="[1311,1388,1950,1975]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="25" unit="m" value="900.0">900 m</elevation>
</quantity>
},
<date id="5D47D7959A2AFFD0FA2FF801FF7CF87F" pageId="24" pageNumber="25" value="1936-12-11">
<collectingDate id="4D032E7D9A2AFFD0FA2FF801FF7CF87F" pageId="24" pageNumber="25" value="1936-12-11">11 xii 1936</collectingDate>
</date>
,
<collectorName id="840C94839A2AFFD0FEAFF85DFE6BF87F" box="[257,481,1986,2011]" pageId="24" pageNumber="25">W. Clark-Macintyre</collectorName>
leg. [
<collectionCode id="4FE869909A2AFFD0FDB0F85DFDF4F87E" box="[542,638,1987,2010]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="24" pageNumber="25">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2AFFD1FD21F85DFC24FF0B" ID-GBIF-Occurrence="1434659031" collectingDate="1941-04-27" collectionCode="UMMZ" collectorName="W. Clarke-Macintyre" country="Ecuador" elevation="1000" lastPageId="25" lastPageNumber="26" latitude="-1.0833334" location="Rio Arajuno" longLatPrecision="1308" longitude="-77.53333" municipality="Headwaters" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2AFFD0FD21F85DFD31F87E" box="[655,699,1987,2011]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="C9226B2F9A2AFFD0FD69F85CFCC1F87F" box="[711,843,1986,2011]" pageId="24" pageNumber="25">Headwaters</collectingMunicipality>
of
<location id="2C26A78E9A2AFFD0FCDEF85DFC74F87F" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B:2C26A78E9A2AFFD0FCDEF85DFC74F87F" box="[880,1022,1986,2011]" country="Ecuador" latitude="-1.0833334" longLatPrecision="1308" longitude="-77.53333" municipality="Headwaters" name="Rio Arajuno" pageId="24" pageNumber="25" stateProvince="Napo">Río Arajuno</location>
,
<collectingRegion id="EB3D3FB79A2AFFD0FBA6F85DFBCFF87E" box="[1032,1093,1987,2010]" country="Ecuador" name="Napo" pageId="24" pageNumber="25">Napo</collectingRegion>
watershed, stream # 3 {
<geoCoordinate id="4CCD97929A2AFFD0FAFEF85CFA1DF87F" box="[1360,1431,1986,2011]" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-1.0833334">1°5' S</geoCoordinate>
,
<geoCoordinate id="4CCD97929A2BFFD1FF39FF06FE89FF0B" box="[151,259,152,176]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="-77.53333">77°32' W</geoCoordinate>
,
<quantity id="EE015CB09A2BFFD1FEBEFF06FEFEFF14" box="[272,372,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="m" value="1000.0">
<elevation id="A2D416669A2BFFD1FEBEFF06FEFEFF14" box="[272,372,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="m" value="1000.0">1,000 m</elevation>
</quantity>
},
<date id="5D47D7959A2BFFD1FE25FF06FD82FF14" box="[395,520,151,176]" pageId="25" pageNumber="26" value="1941-04-27">
<collectingDate id="4D032E7D9A2BFFD1FE25FF06FD82FF14" box="[395,520,151,176]" pageId="25" pageNumber="26" value="1941-04-27">27 iv 1941</collectingDate>
</date>
,
<collectorName id="840C94839A2BFFD1FDB8FF06FC8FFF14" box="[534,773,151,176]" pageId="25" pageNumber="26">W. Clarke-Macintyre</collectorName>
leg. [
<collectionCode id="4FE869909A2BFFD1FCE9FF06FC2DFF0B" box="[839,935,152,175]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FC15FF06FECEFF70" ID-GBIF-Occurrence="1434659020" collectingDate="1941-04-26" collectionCode="UMMZ" collectorName="W. Clarke-Macintyre" country="Ecuador" elevation="1000" latitude="-1.0833334" location="Rio Arajuno" longLatPrecision="1308" longitude="-77.53333" municipality="Headwaters" pageId="25" pageNumber="26" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FC15FF06FC62FF0B" box="[955,1000,152,176]" pageId="25" pageNumber="26" type="male">4 ♂</specimenCount>
,
<specimenCount id="3FFF3ADC9A2BFFD1FC59FF06FBA8FF0B" box="[1015,1058,152,176]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
, same data but stream # 4,
<date id="5D47D7959A2BFFD1FACCFF09FF45FF70" pageId="25" pageNumber="26" value="1941-04-26">
<collectingDate id="4D032E7D9A2BFFD1FACCFF09FF45FF70" pageId="25" pageNumber="26" value="1941-04-26">26 iv 1941</collectingDate>
</date>
[
<collectionCode id="4FE869909A2BFFD1FF73FF23FEB7FF70" box="[221,317,189,212]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FEFEFF22FB2EFF70" ID-GBIF-Occurrence="1434659059" box="[336,1188,188,213]" collectionCode="UMMZ" collectorName="Staundinger" country="Ecuador" elevation="700" latitude="-1.2" location="Napo" longLatPrecision="1308" longitude="-77.833336" pageId="25" pageNumber="26" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FEFEFF22FEF7FF70" box="[336,381,188,212]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
,
<specimenCount id="3FFF3ADC9A2BFFD1FE24FF22FE3FFF70" box="[394,437,188,212]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
,
<collectingRegion id="EB3D3FB79A2BFFD1FE6AFF23FD8BFF70" box="[452,513,189,212]" country="Ecuador" name="Napo" pageId="25" pageNumber="26">Napo</collectingRegion>
{
<geoCoordinate id="4CCD97929A2BFFD1FDBDFF22FDE3FF70" box="[531,617,188,213]" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="-1.2">1°12' S</geoCoordinate>
,
<geoCoordinate id="4CCD97929A2BFFD1FDD9FF22FD68FF70" box="[631,738,188,213]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="-77.833336">77°50' W</geoCoordinate>
,
<elevation id="A2D416669A2BFFD1FD40FF22FCB6FF71" box="[750,828,188,213]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="26" unit="m" value="700.0">
<quantity id="EE015CB09A2BFFD1FD40FF22FCB6FF71" box="[750,828,188,213]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="26" unit="m" value="700.0">700 m</quantity>
</elevation>
},
<collectorName id="840C94839A2BFFD1FCFCFF22FC53FF71" box="[850,985,188,213]" pageId="25" pageNumber="26">Staundinger</collectorName>
vendor [
<collectionCode id="4FE869909A2BFFD1FB90FF23FB14FF70" box="[1086,1182,189,212]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FB1EFF22FEC9FF53" ID-GBIF-Occurrence="1434659061" collectingDate="1940-05" collectionCode="UMMZ" collectorName="Staundinger" country="Ecuador" elevation="700" latitude="-1.2" location="Napo" longLatPrecision="1308" longitude="-77.833336" pageId="25" pageNumber="26" specimenCount="4" specimenCount-male="4" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FB1EFF22FB57FF70" box="[1200,1245,188,212]" pageId="25" pageNumber="26" type="male">4 ♂</specimenCount>
, same data but
<date id="5D47D7959A2BFFD1FA20FF23FF45FF5C" pageId="25" pageNumber="26" value="1940-05">
<collectingDate id="4D032E7D9A2BFFD1FA20FF23FF45FF5C" pageId="25" pageNumber="26" value="1940-05">v 1940</collectingDate>
</date>
[
<collectionCode id="4FE869909A2BFFD1FF72FF7EFEB6FF53" box="[220,316,224,247]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FEE0FF7EFB95FF5C" ID-GBIF-Occurrence="1434659067" box="[334,1055,223,248]" collectingDate="1935-12-24" collectionCode="UMMZ" collectorName="W. Clark-Macintyre" country="Ecuador" elevation="700" latitude="-1.2" location="Napo" longLatPrecision="1308" longitude="-77.833336" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FEE0FF7EFEF0FF53" box="[334,378,224,248]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, same data but
<date id="5D47D7959A2BFFD1FD8BFF7EFD2DFF5C" box="[549,679,223,248]" pageId="25" pageNumber="26" value="1935-12-24">
<collectingDate id="4D032E7D9A2BFFD1FD8BFF7EFD2DFF5C" box="[549,679,223,248]" pageId="25" pageNumber="26" value="1935-12-24">24 xii 1935</collectingDate>
</date>
,
<collectorName id="840C94839A2BFFD1FD1CFF7EFC18FF5C" box="[690,914,223,248]" pageId="25" pageNumber="26">W. Clark-Macintyre</collectorName>
leg. [RWG]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FB84FF7EFAEAFF5C" ID-GBIF-Occurrence="1434659045" box="[1066,1376,223,248]" collectingDate="1935-12-24" collectionCode="CSCA" collectorName="W. Clark-Macintyre" country="Ecuador" elevation="700" latitude="-1.2" location="Napo" longLatPrecision="1308" longitude="-77.833336" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FB84FF7EFBDFFF53" box="[1066,1109,224,248]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, same data but [
<collectionCode id="4FE869909A2BFFD1FAA6FF7EFAD3FF5C" box="[1288,1369,224,248]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="25" pageNumber="26">CSCA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FAC5FF7EFB6CFEB8" ID-GBIF-Occurrence="1434659063" collectingDate="1939-12" collectionCode="UMMZ" country="Ecuador" elevation="400" latitude="-1.6666667" location="Napo" longLatPrecision="1308" longitude="-77.416664" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FAC5FF7EFA1DFF53" box="[1387,1431,224,248]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, Concepción-Río
<collectingRegion id="EB3D3FB79A2BFFD1FEF8FE9BFE19FEB8" box="[342,403,261,284]" country="Ecuador" name="Napo" pageId="25" pageNumber="26">Napo</collectingRegion>
{
<geoCoordinate id="4CCD97929A2BFFD1FE0BFE9AFE71FEB8" box="[421,507,260,285]" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="-1.6666667">1°40' S</geoCoordinate>
,
<geoCoordinate id="4CCD97929A2BFFD1FDA7FE9AFDFEFEB8" box="[521,628,260,285]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="-77.416664">77°25' W</geoCoordinate>
,
<quantity id="EE015CB09A2BFFD1FD2EFE9AFD44FEB9" box="[640,718,260,285]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="m" value="400.0">
<elevation id="A2D416669A2BFFD1FD2EFE9AFD44FEB9" box="[640,718,260,285]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="m" value="400.0">400 m</elevation>
</quantity>
},
<date id="5D47D7959A2BFFD1FD4AFE9AFCCEFEB8" box="[740,836,260,285]" pageId="25" pageNumber="26" value="1939-12">
<collectingDate id="4D032E7D9A2BFFD1FD4AFE9AFCCEFEB8" box="[740,836,260,285]" pageId="25" pageNumber="26" value="1939-12">xii 1939</collectingDate>
</date>
, W. Clarke-MacIntyre leg. [
<collectionCode id="4FE869909A2BFFD1FBD1FE9BFB55FEB8" box="[1151,1247,261,284]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FB5CFE9AFE54FE9B" ID-GBIF-Occurrence="1434659060" collectingDate="1939-12-28" collectingDateMax="1939-12-29" collectingDateMin="1939-12-28" collectionCode="UMMZ" country="Ecuador" elevation="400" latitude="-1.6666667" location="Napo" longLatPrecision="1308" longitude="-77.416664" pageId="25" pageNumber="26" specimenCount="3" specimenCount-male="3" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FB5CFE9AFA95FEB8" box="[1266,1311,260,284]" pageId="25" pageNumber="26" type="male">3 ♂</specimenCount>
, same data but
<date id="5D47D7959A2BFFD1FF6FFEB6FEE0FEE4" box="[193,362,295,320]" pageId="25" pageNumber="26" value="1939-12-28" valueMax="1939-12-29" valueMin="1939-12-28">
<collectingDate id="4D032E7D9A2BFFD1FF6FFEB6FEE0FEE4" box="[193,362,295,320]" pageId="25" pageNumber="26" value="1939-12-28" valueMax="1939-12-29" valueMin="1939-12-28">28–29 xii 1939</collectingDate>
</date>
[
<collectionCode id="4FE869909A2BFFD1FED9FEB6FE5DFE9B" box="[375,471,296,319]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FE46FEB6FA12FE9B" ID-GBIF-Occurrence="1434659052" box="[488,1432,295,320]" collectingDate="1939-12-29" collectingDateMax="1939-12-29" collectingDateMin="1939-12-28" collectionCode="UMMZ" country="Ecuador" elevation="400" latitude="-1.6666667" location="Rio Pacai-Yacu" longLatPrecision="1308" longitude="-77.416664" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FE46FEB6FD9EFE9B" box="[488,532,296,320]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, same data but
<location id="2C26A78E9A2BFFD1FD11FEB6FCE5FEE4" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B:2C26A78E9A2BFFD1FD11FEB6FCE5FEE4" box="[703,879,295,320]" name="Rio Pacai-Yacu" pageId="25" pageNumber="26">Río Pacai-Yacu</location>
,
<date id="5D47D7959A2BFFD1FCD4FEB6FC76FEE4" box="[890,1020,295,320]" pageId="25" pageNumber="26" value="1939-12-29">
<collectingDate id="4D032E7D9A2BFFD1FCD4FEB6FC76FEE4" box="[890,1020,295,320]" pageId="25" pageNumber="26" value="1939-12-29">29 xii 1939</collectingDate>
</date>
, W. Clarke-MacIntyre leg. [
<collectionCode id="4FE869909A2BFFD1FA9FFEB6FA1BFE9B" box="[1329,1425,296,319]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FF39FED3FB6FFE23" ID-GBIF-Occurrence="1434659058" collectingDate="1935-09-29" collectionCode="UMMZ" collectorName="El Partidero &amp; W. Clarke-Macintyre" country="Ecuador" county="El Partidero" elevation="1000" latitude="-1.3666667" location="Mera" longLatPrecision="1308" longitude="-78.0" municipality="Rio Anzu" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
<collectingRegion id="EB3D3FB79A2BFFD1FF39FED3FF61FEC0" box="[151,235,333,356]" country="Ecuador" name="Pastaza" pageId="25" pageNumber="26">Pastaza</collectingRegion>
Prov.:
<specimenCount id="3FFF3ADC9A2BFFD1FEEFFED2FEE5FEC0" box="[321,367,332,356]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
,
<collectingCounty id="C02789D99A2BFFD1FED0FED2FD80FEC1" box="[382,522,332,357]" pageId="25" pageNumber="26">
<collectorName id="840C94839A2BFFD1FED0FED2FD80FEC1" box="[382,522,332,357]" pageId="25" pageNumber="26">El Partidero</collectorName>
</collectingCounty>
, indian village on E bank of
<collectingMunicipality id="C9226B2F9A2BFFD1FCCCFED2FC5EFEC0" box="[866,980,332,356]" pageId="25" pageNumber="26">Río Anzu</collectingMunicipality>
, junction of trails from
<location id="2C26A78E9A2BFFD1FB45FED3FAACFEC0" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B:2C26A78E9A2BFFD1FB45FED3FAACFEC0" box="[1259,1318,333,356]" country="Ecuador" county="El Partidero" latitude="-1.3666667" longLatPrecision="1308" longitude="-78.0" municipality="Rio Anzu" name="Mera" pageId="25" pageNumber="26" stateProvince="Pastaza">Mera</location>
and
<location id="2C26A78E9A2BFFD1FACCFED3FA16FEC1" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B:2C26A78E9A2BFFD1FACCFED3FA16FEC1" box="[1378,1436,333,357]" country="Ecuador" county="El Partidero" latitude="-1.3666667" longLatPrecision="1308" longitude="-78.0" municipality="Rio Anzu" name="Puyo" pageId="25" pageNumber="26" stateProvince="Pastaza">Puyo</location>
northward to
<collectingRegion id="EB3D3FB79A2BFFD1FE9EFEEEFEE7FE23" box="[304,365,368,391]" country="Ecuador" name="Napo" pageId="25" pageNumber="26">Napo</collectingRegion>
{
<geoCoordinate id="4CCD97929A2BFFD1FED1FEF1FE5CFE2C" box="[383,470,367,392]" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="-1.3666667">1°22' S</geoCoordinate>
,
<geoCoordinate id="4CCD97929A2BFFD1FE4AFEEEFDCBFE23" box="[484,577,368,392]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="-78.0">78°0' W</geoCoordinate>
,
<quantity id="EE015CB09A2BFFD1FDE3FEEEFD3AFE2C" box="[589,688,367,392]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="m" value="1000.0">
<elevation id="A2D416669A2BFFD1FDE3FEEEFD3AFE2C" box="[589,688,367,392]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="m" value="1000.0">1,000 m</elevation>
</quantity>
},
<date id="5D47D7959A2BFFD1FD69FEEEFCCEFE2C" box="[711,836,367,392]" pageId="25" pageNumber="26" value="1935-09-29">
<collectingDate id="4D032E7D9A2BFFD1FD69FEEEFCCEFE2C" box="[711,836,367,392]" pageId="25" pageNumber="26" value="1935-09-29">29 ix 1935</collectingDate>
</date>
,
<collectorName id="840C94839A2BFFD1FCFEFEEEFBB4FE2C" box="[848,1086,367,392]" pageId="25" pageNumber="26">W. Clarke-Macintyre</collectorName>
leg. [
<collectionCode id="4FE869909A2BFFD1FBD0FEEEFB54FE23" box="[1150,1246,368,391]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FB5CFEEEFE39FE08" ID-GBIF-Occurrence="1434659030" collectingDate="1935-11-12" collectionCode="UMMZ" collectorName="El Partidero &amp; W. Clarke-Macintyre" country="Ecuador" county="El Partidero" elevation="1000" latitude="-1.3666667" location="Mera" longLatPrecision="1308" longitude="-78.0" municipality="Rio Anzu" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FB5CFEEEFA95FE23" box="[1266,1311,368,392]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, same data but
<date id="5D47D7959A2BFFD1FF6DFE0AFEB7FE09" box="[195,317,404,429]" pageId="25" pageNumber="26" value="1935-11-12">
<collectingDate id="4D032E7D9A2BFFD1FF6DFE0AFEB7FE09" box="[195,317,404,429]" pageId="25" pageNumber="26" value="1935-11-12">12 xi 1935</collectingDate>
</date>
[
<collectionCode id="4FE869909A2BFFD1FEE2FE0BFE26FE08" box="[332,428,405,428]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FE11FE0AFB32FE08" ID-GBIF-Occurrence="1434659051" box="[447,1208,404,429]" collectingDate="1935-07-27" collectionCode="CSCA" collectorName="W. Clarke-Macintyre" country="Ecuador" county="El Partidero" elevation="1000" latitude="-1.3666667" location="Mera" longLatPrecision="1308" longitude="-78.0" municipality="Rio Anzu" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FE11FE0AFE67FE08" box="[447,493,404,428]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, same data but
<date id="5D47D7959A2BFFD1FD31FE0AFCAEFE09" box="[671,804,404,429]" pageId="25" pageNumber="26" value="1935-07-27">
<collectingDate id="4D032E7D9A2BFFD1FD31FE0AFCAEFE09" box="[671,804,404,429]" pageId="25" pageNumber="26" value="1935-07-27">27 vii 1935</collectingDate>
</date>
,
<collectorName id="840C94839A2BFFD1FC9FFE0BFB95FE09" box="[817,1055,404,429]" pageId="25" pageNumber="26">W. Clarke-Macintyre</collectorName>
leg. [
<collectionCode id="4FE869909A2BFFD1FBCEFE0AFB38FE08" box="[1120,1202,404,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="25" pageNumber="26">CSCA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FB6BFE0AFE8EFE6B" ID-GBIF-Occurrence="1434659046" collectingDate="1935-07-27" collectionCode="UMMZ" collectorName="W. Clarke-Macintyre" country="Ecuador" county="El Partidero" elevation="1000" latitude="-1.3666667" location="Mera" longLatPrecision="1308" longitude="-78.0" municipality="Rio Anzu" pageId="25" pageNumber="26" specimenCount="2" specimenCount-male="2" stateProvince="Pastaza" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FB6BFE0AFB78FE08" box="[1221,1266,404,428]" pageId="25" pageNumber="26" type="male">2 ♂</specimenCount>
, same data but [
<collectionCode id="4FE869909A2BFFD1FF33FE26FF77FE6B" box="[157,253,440,463]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FEBFFE26FD4EFE74" ID-GBIF-Occurrence="1434659069" box="[273,708,439,464]" collectingDate="1935-05-27" collectionCode="UMMZ" collectorName="W. Clarke-Macintyre" country="Ecuador" county="El Partidero" elevation="1000" latitude="-1.3666667" location="Mera" longLatPrecision="1308" longitude="-78.0" municipality="Rio Anzu" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FEBFFE26FEB5FE6B" box="[273,319,440,464]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, same data but
<date id="5D47D7959A2BFFD1FE5CFE26FDECFE74" box="[498,614,439,464]" pageId="25" pageNumber="26" value="1935-05-27">
<collectingDate id="4D032E7D9A2BFFD1FE5CFE26FDECFE74" box="[498,614,439,464]" pageId="25" pageNumber="26" value="1935-05-27">27 v 1935</collectingDate>
</date>
[RWG]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FD7FFE26FDF8FE50" ID-GBIF-Occurrence="1434659071" collectingDate="1936-11-28" collectionCode="UMMZ" country="Ecuador" elevation="900" latitude="-1.6333333" location="Pastaza" longLatPrecision="1308" longitude="-77.833336" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FD7FFE26FD75FE6B" box="[721,767,440,464]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, Río
<collectingRegion id="EB3D3FB79A2BFFD1FC90FE26FC18FE6B" box="[830,914,440,463]" country="Ecuador" name="Pastaza" pageId="25" pageNumber="26">Pastaza</collectingRegion>
watershed {
<geoCoordinate id="4CCD97929A2BFFD1FBB0FE29FBFCFE74" box="[1054,1142,439,464]" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="-1.6333333">1°38' S</geoCoordinate>
,
<geoCoordinate id="4CCD97929A2BFFD1FB2AFE29FB7AFE6B" box="[1156,1264,439,464]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="-77.833336">77°50' W</geoCoordinate>
,
<quantity id="EE015CB09A2BFFD1FB52FE26FAC1FE74" box="[1276,1355,439,464]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="25" pageNumber="26" unit="m" value="900.0">
<elevation id="A2D416669A2BFFD1FB52FE26FAC1FE74" box="[1276,1355,439,464]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="25" pageNumber="26" unit="m" value="900.0">900 m</elevation>
</quantity>
},
<date id="5D47D7959A2BFFD1FACCFE26FF58FE51" pageId="25" pageNumber="26" value="1936-11-28">
<collectingDate id="4D032E7D9A2BFFD1FACCFE26FF58FE51" pageId="25" pageNumber="26" value="1936-11-28">28 xi 1936</collectingDate>
</date>
, W. Clarke-MacIntyre leg. [
<collectionCode id="4FE869909A2BFFD1FDA5FE43FDE1FE50" box="[523,619,477,500]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FDD3FE42FB9EFDB3" ID-GBIF-Occurrence="1434659038" collectingDate="1935-11-09" collectionCode="UMMZ" collectorName="W. Clarke-Macintyre" country="Ecuador" elevation="1700" latitude="-1.4" location="Guamo Yacu" longLatPrecision="1308" longitude="-78.36667" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Tungurahua" typeStatus="paratype">
<collectingRegion id="EB3D3FB79A2BFFD1FDD3FE42FC89FE51" box="[637,771,476,501]" country="Ecuador" name="Tungurahua" pageId="25" pageNumber="26">Tungurahua</collectingRegion>
Prov.:
<specimenCount id="3FFF3ADC9A2BFFD1FCFAFE42FC0AFE50" box="[852,896,476,500]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
,
<location id="2C26A78E9A2BFFD1FC22FE42FB94FE50" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B:2C26A78E9A2BFFD1FC22FE42FB94FE50" box="[908,1054,476,500]" country="Ecuador" latitude="-1.4" longLatPrecision="1308" longitude="-78.36667" name="Guamo Yacu" pageId="25" pageNumber="26" stateProvince="Tungurahua">Guamo Yacu</location>
(Agoyan), Río
<collectingRegion id="EB3D3FB79A2BFFD1FB61FE43FAA9FE50" box="[1231,1315,477,500]" country="Ecuador" name="Pastaza" pageId="25" pageNumber="26">Pastaza</collectingRegion>
watershed {
<geoCoordinate id="4CCD97929A2BFFD1FF0EFE61FF73FDBC" box="[160,249,511,536]" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="-1.4">1°24' S</geoCoordinate>
,
<geoCoordinate id="4CCD97929A2BFFD1FEA6FD9EFEFEFDB3" box="[264,372,512,536]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="-78.36667">78°22' W</geoCoordinate>
,
<quantity id="EE015CB09A2BFFD1FE2FFD9EFE6FFDBC" box="[385,485,511,536]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="25" pageNumber="26" unit="m" value="1700.0">
<elevation id="A2D416669A2BFFD1FE2FFD9EFE6FFDBC" box="[385,485,511,536]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="25" pageNumber="26" unit="m" value="1700.0">1,700 m</elevation>
</quantity>
},
<date id="5D47D7959A2BFFD1FE53FD9EFDE4FDBC" box="[509,622,511,536]" pageId="25" pageNumber="26" value="1935-11-09">
<collectingDate id="4D032E7D9A2BFFD1FE53FD9EFDE4FDBC" box="[509,622,511,536]" pageId="25" pageNumber="26" value="1935-11-09">9 xi 1935</collectingDate>
</date>
,
<collectorName id="840C94839A2BFFD1FDD5FD9EFCE1FDBC" box="[635,875,511,536]" pageId="25" pageNumber="26">W. Clarke-Macintyre</collectorName>
leg. [
<collectionCode id="4FE869909A2BFFD1FC03FD9EFB87FDB3" box="[941,1037,512,535]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FB8FFD9EFA12FDBC" ID-GBIF-Occurrence="1434659062" box="[1057,1432,511,536]" collectingDate="1935-11-09" collectionCode="UMMZ" collectorName="W. Clarke-Macintyre" country="Ecuador" elevation="1700" latitude="-1.4" location="Guamo Yacu" longLatPrecision="1308" longitude="-78.36667" pageId="25" pageNumber="26" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tungurahua" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A2BFFD1FB8FFD9EFBC5FDB3" box="[1057,1103,512,536]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
,
<specimenCount id="3FFF3ADC9A2BFFD1FBF0FD9EFB00FDB3" box="[1118,1162,512,536]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
, same data but [RWG]
</materialsCitation>
;
<materialsCitation id="9991FB089A2BFFD1FF39FDBBFAECFD98" ID-GBIF-Occurrence="1434659072" box="[151,1382,548,573]" collectionCode="UMMZ" collectorName="L. Gomez" country="Ecuador" elevation="708" latitude="-2.5166667" location="Rio Upano" longLatPrecision="1307" longitude="-77.88333" municipality="Mangosisa" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Morona Santiago" typeStatus="paratype">
<collectingRegion id="EB3D3FB79A2BFFD1FF39FDBBFED2FD99" box="[151,344,548,573]" country="Ecuador" name="Morona-Santiago" pageId="25" pageNumber="26">
Morona
<collectingRegion id="EB3D3FB79A2BFFD1FF58FDBAFED2FD99" box="[246,344,548,573]" country="Philippines" name="Santiago" pageId="25" pageNumber="26">Santiago</collectingRegion>
</collectingRegion>
Prov.:
<specimenCount id="3FFF3ADC9A2BFFD1FE06FDBAFE5EFD98" box="[424,468,548,572]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="C9226B2F9A2BFFD1FE4EFDBAFDD6FD99" box="[480,604,548,573]" pageId="25" pageNumber="26">Mangosisa</collectingMunicipality>
,
<location id="2C26A78E9A2BFFD1FDC6FDBAFD68FD98" LSID="urn:lsid:plazi:treatment:A15040439A2AFFD2FF39F98AFBDFFD9B:2C26A78E9A2BFFD1FDC6FDBAFD68FD98" box="[616,738,548,572]" country="Ecuador" latitude="-2.5166667" longLatPrecision="1307" longitude="-77.88333" municipality="Mangosisa" name="Rio Upano" pageId="25" pageNumber="26" stateProvince="Morona Santiago">Río Upano</location>
{
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,
<geoCoordinate id="4CCD97929A2BFFD1FCFBFDBAFC35FD98" box="[853,959,548,573]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="-77.88333">77°53' W</geoCoordinate>
,
<quantity id="EE015CB09A2BFFD1FC64FDBAFB9DFD99" box="[970,1047,548,573]" metricMagnitude="2" metricUnit="m" metricValue="7.08" pageId="25" pageNumber="26" unit="m" value="708.0">
<elevation id="A2D416669A2BFFD1FC64FDBAFB9DFD99" box="[970,1047,548,573]" metricMagnitude="2" metricUnit="m" metricValue="7.08" pageId="25" pageNumber="26" unit="m" value="708.0">708 m</elevation>
</quantity>
},
<collectorName id="840C94839A2BFFD1FB82FDBBFB16FD98" box="[1068,1180,548,572]" pageId="25" pageNumber="26">L. Gomez</collectorName>
vendor [
<collectionCode id="4FE869909A2BFFD1FB51FDBBFAD5FD98" box="[1279,1375,549,572]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="2946F1559A2BFFD1FF69FDD6FB3BFDC4" blockId="25.[151,1437,151,2016]" box="[199,1201,583,608]" pageId="25" pageNumber="26">
A medium-sized largely dark species with strongly forcipate male appendages (
<figureCitation id="B1C2EDD09A2BFFD1FBECFDD9FB2EFDC4" box="[1090,1188,583,608]" captionStart="FIGURES 158 – 160" captionStartId="80.[151,264,1826,1849]" captionTargetBox="[189,1398,197,1790]" captionTargetId="figure@80.[188,1398,193,1805]" captionTargetPageId="80" captionText="FIGURES 158 – 160. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view." httpUri="https://zenodo.org/record/322103/files/figure.png" pageId="25" pageNumber="26">Fig. 159</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A2BFFD2FF69FDF2FCCEFEC3" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
<paragraph id="2946F1559A2BFFD1FF69FDF2FCA6FD54" blockId="25.[151,1437,151,2016]" pageId="25" pageNumber="26">
<emphasis id="1B8D2D479A2BFFD1FF69FDF2FDA3FD21" bold="true" box="[199,553,620,645]" pageId="25" pageNumber="26">
Description of male
<typeStatus id="F6424FF79A2BFFD1FE6CFDF2FDA3FD21" box="[450,553,620,645]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
</emphasis>
. Head: labrum pale rimmed with narrow line of black apically and basally, anteclypeus dark brown, base of mandibles, genae, postclypeus pale (purple), remainder of head black except for postocular spots and small pale spot anterolateral to lateral ocellus (as in
<figureCitation id="B1C2EDD09A2BFFD1FC47FD2AFBB5FD69" box="[1001,1087,692,717]" captionStart="FIGURES 24 – 26" captionStartId="45.[151,264,1836,1859]" captionTargetBox="[160,1408,215,1680]" captionTargetId="figure@45.[151,1436,193,1815]" captionTargetPageId="45" captionText="FIGURES 24 – 26. Color pattern of male head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322068/files/figure.png" pageId="25" pageNumber="26">Fig. 25</figureCitation>
); antennae black, rear of head black except for pale narrow margin bordering eye margin.
</paragraph>
<paragraph id="2946F1559A2BFFD1FF69FD62FE72FBFF" blockId="25.[151,1437,151,2016]" pageId="25" pageNumber="26">
Prothorax black with following areas pale: obscure pale rim on anterior lobe, small dorsolateral spot on middle lobe, lateral 0.30 of posterior margin of propleuron. Pale areas of pterothorax purple, with broad black middorsal stripe about four times as wide as pale antehumeral stripe, the latter slightly narrowing dorsally; broad parallel black humeral stripe extending from base of mesinfraepisternum and connecting below antealar crest with middorsal stripe above and with dorsal portion of obsolete interpleural suture; metapleural stripe narrow; pale colors on side of thorax purple (as in
<figureCitation id="B1C2EDD09A2BFFD1FDCFFC31FD33FC6C" box="[609,697,943,968]" captionStart="FIGURES 24 – 26" captionStartId="45.[151,264,1836,1859]" captionTargetBox="[160,1408,215,1680]" captionTargetId="figure@45.[151,1436,193,1815]" captionTargetPageId="45" captionText="FIGURES 24 – 26. Color pattern of male head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322068/files/figure.png" pageId="25" pageNumber="26">Fig. 25</figureCitation>
). Wings hyaline with venation black; pterostigma dark brown, surmounting 1.5 cells in right Fw, 2 cells in left Fw; postnodals Fw; 2 cells in left Hw, 1.5 cells in right Hw; postnodals Fw 17/18, Hw 16/15; postquadrangular cells Fw 4/4, Hw 4/3; RP2 at Fw 8.5/8.5, Hw 7/7. Coxae and trochanters pale except for black on ventral and anterior portions of coxae and dorsum of trochanters; femora, tibiae, tarsi and armature black.
</paragraph>
<paragraph id="2946F1559A2BFFD1FF69FBF8FAB2FAF3" blockId="25.[151,1437,151,2016]" pageId="25" pageNumber="26">
Abdomen (as in
<figureCitation id="B1C2EDD09A2BFFD1FE2CFBF8FE55FBDB" box="[386,479,1126,1151]" captionStart="FIGURES 24 – 26" captionStartId="45.[151,264,1836,1859]" captionTargetBox="[160,1408,215,1680]" captionTargetId="figure@45.[151,1436,193,1815]" captionTargetPageId="45" captionText="FIGURES 24 – 26. Color pattern of male head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322068/files/figure.png" pageId="25" pageNumber="26">Figs. 25</figureCitation>
,
<figureCitation id="B1C2EDD09A2BFFD1FE44FBF9FD8DFBDB" box="[490,519,1127,1151]" captionStart="FIGURES 73 – 79" captionStartId="55.[151,264,1620,1643]" captionTargetBox="[157,1423,211,1533]" captionTargetId="figure@55.[151,1436,193,1600]" captionTargetPageId="55" captionText="FIGURES 73 – 79. Color pattern of male S 7 – 10 of other species, lateral view." httpUri="https://zenodo.org/record/322078/files/figure.png" pageId="25" pageNumber="26">74</figureCitation>
) mostly black; S1 with a black basal ring, remainder purple; S2 black with a pale (purple) dorsal campanulate spot occupying basal 0.80 of segment, abruptly narrowing to a thin line almost extending to black apical annulus, laterally black with a pale lateral stripe; S3 black except for narrow basal dorsal inverted triangular spot at basal 0.10; S4–5 similar to S3 but with only a small obscure pale dorsal spot at basal 0.05; S5–7 black; S8 with a dorsolateral pale (purple) spot interrupted basally by a narrow ring of black and posteriorly by black lateral stripe extending dorsally at apical 0.10; S9 pale dorsally, black laterally; S10 black with a pair of purple spots dorsally interrupted by narrow middorsal black stripe; torus pale, appendages black.
</paragraph>
<paragraph id="2946F1559A2BFFD1FF69FAFCFE80FA67" blockId="25.[151,1437,151,2016]" pageId="25" pageNumber="26">
Genital ligula (
<figureCitation id="B1C2EDD09A2BFFD1FEDBFAFCFE5CFADF" box="[373,470,1378,1403]" captionStart="FIGURES 138 – 139" captionStartId="72.[151,264,1855,1878]" captionTargetBox="[166,1436,218,1754]" captionTargetId="figure@72.[151,1436,193,1834]" captionTargetPageId="72" captionText="FIGURES 138 – 139. Genital ligula of other species. (a) lateral view; (b) ectal view; (c) entolateral view; (d) detail of field of michrothrichia on distal segment’s base." httpUri="https://zenodo.org/record/322095/files/figure.png" pageId="25" pageNumber="26">Fig. 139</figureCitation>
) with a large elongate microspinulate patch on ental surface proximal to flexure on sclerotized area (
<figureCitation id="B1C2EDD09A2BFFD1FEFBFA18FE3FFA3B" box="[341,437,1414,1439]" captionStart="FIGURES 138 – 139" captionStartId="72.[151,264,1855,1878]" captionTargetBox="[166,1436,218,1754]" captionTargetId="figure@72.[151,1436,193,1834]" captionTargetPageId="72" captionText="FIGURES 138 – 139. Genital ligula of other species. (a) lateral view; (b) ectal view; (c) entolateral view; (d) detail of field of michrothrichia on distal segment’s base." httpUri="https://zenodo.org/record/322095/files/figure.png" pageId="25" pageNumber="26">Fig. 139</figureCitation>
d); distal segment lacking lateral sclerotized lobes and terminating in a single long curved flagellum.
</paragraph>
<paragraph id="2946F1559A2BFFD1FF69FA4EFBA3F9F7" blockId="25.[151,1437,151,2016]" pageId="25" pageNumber="26">
Torus small, transversely oval, swollen, occupying entire ventral margin of torifer but not overlapping bilobed epiproct (
<figureCitation id="B1C2EDD09A2BFFD1FEACFA6CFEEBF9AF" box="[258,353,1522,1547]" captionStart="FIGURES 158 – 160" captionStartId="80.[151,264,1826,1849]" captionTargetBox="[189,1398,197,1790]" captionTargetId="figure@80.[188,1398,193,1805]" captionTargetPageId="80" captionText="FIGURES 158 – 160. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view." httpUri="https://zenodo.org/record/322103/files/figure.png" pageId="25" pageNumber="26">Fig. 159</figureCitation>
a, c); area around epiproct and base of same black; cercus (
<figureCitation id="B1C2EDD09A2BFFD1FC5FFA6CFBDBF9AF" box="[1009,1105,1522,1547]" captionStart="FIGURES 158 – 160" captionStartId="80.[151,264,1826,1849]" captionTargetBox="[189,1398,197,1790]" captionTargetId="figure@80.[188,1398,193,1805]" captionTargetPageId="80" captionText="FIGURES 158 – 160. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view." httpUri="https://zenodo.org/record/322103/files/figure.png" pageId="25" pageNumber="26">Fig. 159</figureCitation>
a, c) long, about twice as long as paraproct (
<figureCitation id="B1C2EDD09A2BFFD1FE96F988FE17F98B" box="[312,413,1558,1583]" captionStart="FIGURES 158 – 160" captionStartId="80.[151,264,1826,1849]" captionTargetBox="[189,1398,197,1790]" captionTargetId="figure@80.[188,1398,193,1805]" captionTargetPageId="80" captionText="FIGURES 158 – 160. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view." httpUri="https://zenodo.org/record/322103/files/figure.png" pageId="25" pageNumber="26">Fig. 159</figureCitation>
b), forcipate and ending in a capitate tooth; paraproct bilobed, its ventral branch much smaller than larger dorsal branch, in lateral view, dorsal branch bluntly triangular.
</paragraph>
<paragraph id="2946F1559A2BFFD1FF69F9C0FCB3F9D3" blockId="25.[151,1437,151,2016]" box="[199,825,1630,1655]" pageId="25" pageNumber="26">
<emphasis id="1B8D2D479A2BFFD1FF69F9C0FED8F9D3" bold="true" box="[199,338,1630,1655]" pageId="25" pageNumber="26">Dimensions</emphasis>
. Hw 21.5, abdomen 28.4, total length 36.7.
</paragraph>
<paragraph id="2946F1559A2BFFD1FF69F91CFD13F8EB" blockId="25.[151,1437,151,2016]" pageId="25" pageNumber="26">
<emphasis id="1B8D2D479A2BFFD1FF69F91CFDCBF93F" bold="true" box="[199,577,1666,1691]" pageId="25" pageNumber="26">
Description of female
<typeStatus id="F6424FF79A2BFFD1FE78F91CFDCBF93F" box="[470,577,1666,1691]" pageId="25" pageNumber="26" type="paratype">paratype</typeStatus>
</emphasis>
(
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:
<collectingRegion id="EB3D3FB79A2BFFD1FD6AF91DFC8BF93E" box="[708,769,1667,1690]" country="Ecuador" name="Napo" pageId="25" pageNumber="26">Napo</collectingRegion>
Prov., Cushillu-Yacu, Río
<collectingRegion id="EB3D3FB79A2BFFD1FB97F91DFBFCF93E" box="[1081,1142,1667,1690]" country="Ecuador" name="Napo" pageId="25" pageNumber="26">Napo</collectingRegion>
watershed, El Partidero). Head, pro- and pterothorax and S1–2 as in male (
<figureCitation id="B1C2EDD09A2BFFD1FD68F938FC90F91B" box="[710,794,1702,1727]" captionStart="FIGURES 50 – 52" captionStartId="52.[151,264,1842,1865]" captionTargetBox="[186,1393,242,1658]" captionTargetId="figure@52.[151,1436,193,1821]" captionTargetPageId="52" captionText="FIGURES 50 – 52. Color pattern of female head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322075/files/figure.png" pageId="25" pageNumber="26">Fig. 50</figureCitation>
), but postocular spots and antehumeral stripe dull purple and pale colors on side of thorax olive brown; S1 as in male; S2 black dorsally with a narrow ventral pale stripe; S3–7 black with vestiges of narrow pale lateral stripe; S8 black with blue distal spot occupying apical 0.40; S9 black with posterior 0.50 blue extending laterally and projecting anteriorly to basal 0.30; S10 blue dorsally, black ventrally, cerci and ovipositor black (
<figureCitation id="B1C2EDD09A2BFFD1FD96F8A8FD06F8EB" box="[568,652,1846,1871]" captionStart="FIGURES 96 – 102" captionStartId="58.[151,264,1799,1822]" captionTargetBox="[159,1428,206,1751]" captionTargetId="figure@58.[151,1436,193,1778]" captionTargetPageId="58" captionText="FIGURES 96 – 102. Color pattern of female S 7 – 10 of other species, lateral view (93 – 100, 102); S 1 – 5, dorsolateral view (101)." httpUri="https://zenodo.org/record/322081/files/figure.png" pageId="25" pageNumber="26">Fig. 97</figureCitation>
).
</paragraph>
<paragraph id="2946F1559A2BFFD1FF69F8C4FAF8F81F" blockId="25.[151,1437,151,2016]" pageId="25" pageNumber="26">
Mesostigmal lobe as in
<figureCitation id="B1C2EDD09A2BFFD1FE73F8C4FDB5F8D7" box="[477,575,1882,1907]" captionStart="FIGURES 119 – 120" captionStartId="65.[151,264,1887,1910]" captionTargetBox="[159,1420,209,1840]" captionTargetId="figure@65.[151,1436,193,1866]" captionTargetPageId="65" captionText="FIGURES 119 – 120. Female mesostigmal plates of other species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view." httpUri="https://zenodo.org/record/322088/files/figure.png" pageId="25" pageNumber="26">Fig. 120</figureCitation>
, well developed but small, forming a flat medially directed digit-like lobe elevated above mesepisternum and not overlying branch of middorsal carina; in posterior view (as in
<figureCitation id="B1C2EDD09A2BFFD1FA89F8E0FA0DF833" box="[1319,1415,1918,1943]" captionStart="FIGURES 119 – 120" captionStartId="65.[151,264,1887,1910]" captionTargetBox="[159,1420,209,1840]" captionTargetId="figure@65.[151,1436,193,1866]" captionTargetPageId="65" captionText="FIGURES 119 – 120. Female mesostigmal plates of other species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view." httpUri="https://zenodo.org/record/322088/files/figure.png" pageId="25" pageNumber="26">Fig. 120</figureCitation>
e) lobe thickened externally but lacking a tubercle at juncture with mesepisternum; mesepisternal tubercle absent.
</paragraph>
<paragraph id="2946F1559A2BFFD2FF69F858FCCEFEC3" blockId="25.[151,1437,151,2016]" lastBlockId="26.[151,1437,151,575]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="1B8D2D479A2BFFD1FF69F858FE5FF87B" bold="true" box="[199,469,1990,2015]" pageId="25" pageNumber="26">
Variation in
<typeStatus id="F6424FF79A2BFFD1FEF1F858FE5FF87B" box="[351,469,1990,2015]" pageId="25" pageNumber="26" type="paratype">paratypes</typeStatus>
</emphasis>
. Little variation was observed in the
<typeStatus id="F6424FF79A2BFFD1FCD1F856FC55F844" box="[895,991,1992,2016]" pageId="25" pageNumber="26" type="paratype">paratype</typeStatus>
series. The dorsolateral spot on S
<quantity id="EE015CB09A2BFFD1FACFF859FA16F87B" box="[1377,1436,1990,2015]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="10.0">10 in</quantity>
male can range from present as in
<typeStatus id="F6424FF79A28FFD2FDE7FF09FD21FF14" box="[585,683,151,176]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
to very small (El Partidero) or absent (Río
<collectingRegion id="EB3D3FB79A28FFD2FB61FF06FA86FF0B" box="[1231,1292,152,175]" country="Ecuador" name="Napo" pageId="26" pageNumber="27">Napo</collectingRegion>
watershed). Pterostigma surmounting 1–2 cells in males,
<quantity id="EE015CB09A28FFD2FD38FF22FD79FF71" box="[662,755,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="4.445" metricValueMax="5.08" metricValueMin="3.81" pageId="26" pageNumber="27" unit="in" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 in</quantity>
females; postnodals: Fw
<quantity id="EE015CB09A28FFD2FBB8FF22FBF0FF71" box="[1046,1146,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="26" pageNumber="27" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
males,
<quantity id="EE015CB09A28FFD2FB7CFF22FABCFF71" box="[1234,1334,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="26" pageNumber="27" unit="in" value="16.5" valueMax="17.0" valueMin="16.0">16–17 in</quantity>
females, Hw
<quantity id="EE015CB09A28FFD2FF64FF41FEBAFF5C" box="[202,304,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.556" pageId="26" pageNumber="27" unit="in" value="15.0" valueMax="16.0" valueMin="14.0">14–16 in</quantity>
males,
<quantity id="EE015CB09A28FFD2FE23FF41FE7EFF5C" box="[397,500,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="3.81" metricValueMin="3.556" pageId="26" pageNumber="27" unit="in" value="14.5" valueMax="15.0" valueMin="14.0">14–15 in</quantity>
females; postquadrangular cells Fw 3–5, Hw
<quantity id="EE015CB09A28FFD2FBA0FF7EFBD3FF5C" box="[1038,1113,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="26" pageNumber="27" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
males, Fw 4–5, Hw
<quantity id="EE015CB09A28FFD2FAFCFF7EFA17FF5C" box="[1362,1437,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="26" pageNumber="27" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
females; RP2 at Fw 7–9, Hw
<quantity id="EE015CB09A28FFD2FE77FE9AFD95FEB9" box="[473,543,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="26" pageNumber="27" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">6–7 in</quantity>
males, Fw 7–8, Hw
<quantity id="EE015CB09A28FFD2FCA8FE9AFCC7FEB9" box="[774,845,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="26" pageNumber="27" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">6–7 in</quantity>
females.
<emphasis id="1B8D2D479A28FFD2FC19FE9AFBC8FEB9" bold="true" box="[951,1090,260,285]" pageId="26" pageNumber="27">Dimensions</emphasis>
.
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: Hw 20.9 ± 1.09 [19–22.3], abdomen 27.3 ± 1.22 [25.7–29.3], total length 35.2 ± 1.51 [33.3–37.8];
<specimenCount id="3FFF3ADC9A28FFD2FC6BFEB5FC56FEE6" box="[965,988,299,322]" pageId="26" pageNumber="27" type="female">♀</specimenCount>
: Hw 22.1 ± 1.45 [20.2–23.7], abdomen 27.2 ± 1.84 [25.3–29.4], total length 35.2 ± 2.08 [33.1–37.7].
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A28FFD2FF69FEECFD2DFDBF" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="2946F1559A28FFD2FF69FEECFC12FE0B" blockId="26.[151,1437,151,575]" pageId="26" pageNumber="27">
<emphasis id="1B8D2D479A28FFD2FF69FEECFEB0FE2F" bold="true" box="[199,314,370,395]" pageId="26" pageNumber="27">Diagnosis</emphasis>
. This species is superficially similar to
<taxonomicName id="EEF98AD69A28FFD2FCA9FEEDFCF1FE2F" box="[775,891,371,395]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="carolus">
<emphasis id="1B8D2D479A28FFD2FCA9FEEDFCF1FE2F" box="[775,891,371,395]" italics="true" pageId="26" pageNumber="27">A. carolus</emphasis>
</taxonomicName>
and is diagnosed under that species. Within its range (
<collectingCountry id="51EEB1C59A28FFD2FF4AFE08FECDFE0B" box="[228,327,406,431]" name="Ecuador" pageId="26" pageNumber="27">Ecuador</collectingCountry>
), it is unlikely to be confused with any other species.
</paragraph>
<paragraph id="2946F1559A28FFD2FF69FE24FD2DFDBF" blockId="26.[151,1437,151,575]" pageId="26" pageNumber="27">
<emphasis id="1B8D2D479A28FFD2FF69FE24FEA9FE77" bold="true" box="[199,291,442,467]" pageId="26" pageNumber="27">Habitat</emphasis>
. Streams and rivers in forest. Specimens were taken at elevations ranging from about
<quantity id="EE015CB09A28FFD2FAE0FE25FA16FE76" box="[1358,1436,443,467]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="m" value="400.0">400 m</quantity>
(Concepción-Río
<collectingRegion id="EB3D3FB79A28FFD2FECFFE41FE15FE53" box="[353,415,479,503]" country="Ecuador" name="Napo" pageId="26" pageNumber="27">Napo</collectingRegion>
) to
<quantity id="EE015CB09A28FFD2FE7EFE41FDA5FE53" box="[464,559,479,503]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="26" pageNumber="27" unit="m" value="1700.0">1,700 m</quantity>
(Guamo Yacu). Flight dates range from April (Headwaters of Río Arajuno) through December (Las Palmas, on Río Anzu).
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A28FFD2FF69FDB8FBDFFD9B" box="[199,1109,550,575]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="2946F1559A28FFD2FF69FDB8FBDFFD9B" blockId="26.[151,1437,151,575]" box="[199,1109,550,575]" pageId="26" pageNumber="27">
<emphasis id="1B8D2D479A28FFD2FF69FDB8FED3FD9B" bold="true" box="[199,345,550,575]" pageId="26" pageNumber="27">Distribution</emphasis>
. As far as known restricted to
<collectingCountry id="51EEB1C59A28FFD2FD1FFDB8FC84FD9B" box="[689,782,550,575]" name="Ecuador" pageId="26" pageNumber="27">Ecuador</collectingCountry>
east of the Andes (
<figureCitation id="B1C2EDD09A28FFD2FC48FDB8FBC2FD9B" box="[998,1096,550,575]" captionStart="FIGURES 169 – 170" captionStartId="84.[151,264,1812,1835]" captionTargetBox="[154,1366,212,1759]" captionTargetId="figure@84.[151,1436,193,1791]" captionTargetPageId="84" captionText="FIGURES 169 – 170. Distribution of other species." httpUri="https://zenodo.org/record/322107/files/figure.png" pageId="26" pageNumber="27">Fig. 169</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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