treatments-xml/data/03/FD/82/03FD8227FFE6E31AFF78FC42FC9B81E8.xml

<document id="C24E50F269C44722A5628A63429CD164" ID-DOI="10.11646/zootaxa.5450.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="11232500" ID-ZooBank="1B5D192F-1D5B-4460-9133-9AEAE9C920BF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716281048833" checkinUser="plazi" docAuthor="Johnson, Robert G." docDate="2024" docId="03FD8227FFE6E31AFF78FC42FC9B81E8" docLanguage="en" docName="zootaxa.5450.1.1.pdf" docOrigin="Zootaxa 5450 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5450.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Lioharpes ammari Johnson 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="59" masterDocId="FFC4FA5FFFDFE320FFEFF87CFFB58771" masterDocTitle="Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco" masterLastPageNumber="185" masterPageNumber="1" pageNumber="58" updateTime="1716282690877" updateUser="ExternalLinkService">
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<mods:title id="1F2691B497A69A7FBBFB0D5A14FD6B80">Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco</mods:title>
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<heading id="D0A3845DFFE6E319FF78FC42FE748328" bold="true" box="[151,449,1086,1113]" fontSize="11" level="2" pageId="57" pageNumber="58" reason="4">
<emphasis id="B920EF23FFE6E319FF78FC42FE748328" bold="true" box="[151,449,1086,1113]" pageId="57" pageNumber="58">
<taxonomicName id="4C5448B2FFE6E319FF78FC42FECD8328" authority="Johnson, 2024" authorityName="Johnson" authorityYear="2024" box="[151,376,1086,1113]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ammari" status="sp. nov.">
<emphasis id="B920EF23FFE6E319FF78FC42FECD8328" bold="true" box="[151,376,1086,1113]" italics="true" pageId="57" pageNumber="58">Lioharpes ammari</emphasis>
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</paragraph>
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<subSubSection id="C34E60BAFFE6E319FF78FC1FFE3A830C" box="[151,399,1123,1149]" pageId="57" pageNumber="58" type="description">
<paragraph id="8BEB3331FFE6E319FF78FC1FFE3A830C" blockId="57.[151,449,1086,1149]" box="[151,399,1123,1149]" pageId="57" pageNumber="58">
<figureCitation id="136F2FB4FFE6E319FF78FC1FFE8F830C" box="[151,314,1123,1149]" captionStart="PLATE 12" captionStartId="118.[151,234,1744,1768]" captionTargetBox="[213,1375,181,1720]" captionTargetId="figure-20@118.[213,1375,181,1720]" captionTargetPageId="118" captionText="PLATE 12. A–F Lioharpes cf. morocconensis, from a lower Emsian horizon, Boutiskaouine Formation, 28.3 km SSW of Rissani (Fig. 2B, Map 14, site 10), dorsal exoskeleton of partial cephalon, NHMUK It 29273. A, dorsal, B, anterior and C and D, lateral view. E, lateral view of eye lobe and F, dorsal view of brim perforation. G–M Lioharpes ammari n. sp., from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 9), dorsal exoskeleton with pygidium enrolled, holotype, NHMUK It 29263. G, dorsal, H, anterior and I, lateral view. J and K, dorsal and lateral views, respectively, of eye lobe. L and M, dorsal view of brim perforations and exterior rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232571" httpUri="https://zenodo.org/record/11232571/files/figure.png" pageId="57" pageNumber="58">Plates 12G–M</figureCitation>
, 
<figureCitation id="136F2FB4FFE6E319FEA9FC1FFE3A830C" box="[326,399,1123,1149]" captionStart="PLATE 13" captionStartId="119.[151,234,1650,1674]" captionTargetBox="[247,1340,181,1626]" captionTargetId="figure-15@119.[247,1340,181,1626]" captionTargetPageId="119" captionText="PLATE 13. A–J Lioharpes ammari n. sp., from horizon 2, Section 1 (Fig. 6), Tazoulait Formation, upper Emisian, South flank of Jbel Ou-Driss (Fig. 2B, Map 4). A–E dorsal exoskeleton, pygidium missing, paratype, NHMUK It 29264 (site 12).A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–H dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29265 (1) (site 13). F, dorsal, G, lateral and H, anterior view. I–J ventral exoskeleton of a partial upper lamella, paratype, NHMUK It 29265 (2) (site 13). I, ventral view and J, lateral view of ventral surface. K–O Pinnuloharpes igaouii n. sp., from a horizon 20 metres down slope from type horizon of Lioharpes ammari n. sp. Boutiskaouine Formation?, lower Emsian North flank of Jbel Ou-Driss (Fig. 2B Map 4 site 6). Dorsal exoskeleton of cephalon only, paratype NHMUK it 29267. K dorsal, L anterior and M lateral view. N lateral view of eye lobe and O dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232573" httpUri="https://zenodo.org/record/11232573/files/figure.png" pageId="57" pageNumber="58">13A–J</figureCitation>
</paragraph>
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<subSubSection id="C34E60BAFFE6E319FF78FCD6FD088240" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="8BEB3331FFE6E319FF78FCD6FD088240" blockId="57.[151,1437,1194,2013]" pageId="57" pageNumber="58">
<emphasis id="B920EF23FFE6E319FF78FCD6FEBF83B5" bold="true" box="[151,266,1194,1220]" pageId="57" pageNumber="58">Diagnosis</emphasis>
. Glabella is conical, carinate and with tectiform flanks and tubercles along crest. S1 short and shallow. Eye lobes inflated, tall, subcircular and with two eye lenses each. Brim is wide (sag.) and steepens strongly on prolongations. Brim width ratio is around 0.70. In lateral view, internal rim nearly horizontal with slightly wave-like profile. Pleural furrows narrow and well defined.
</paragraph>
</subSubSection>
<subSubSection id="C34E60BAFFE6E319FF28FD47FD1C8208" pageId="57" pageNumber="58" type="etymology">
<paragraph id="8BEB3331FFE6E319FF28FD47FD1C8208" blockId="57.[151,1437,1194,2013]" pageId="57" pageNumber="58">
<emphasis id="B920EF23FFE6E319FF28FD47FEFB8224" bold="true" box="[199,334,1339,1365]" pageId="57" pageNumber="58">Etymology.</emphasis>
For Ammar Ait Hssain, a trilobite worker and preparator who prepared, together with his brother Hammi, much of the material presented herein.
</paragraph>
</subSubSection>
<subSubSection id="C34E60BAFFE6E319FF28FDFEFC5F81CC" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="8BEB3331FFE6E319FF28FDFEFD8D8104" blockId="57.[151,1437,1194,2013]" pageId="57" pageNumber="58">
<emphasis id="B920EF23FFE6E319FF28FDFEFE4182EC" bold="true" box="[199,500,1410,1437]" pageId="57" pageNumber="58">Material and occurrence.</emphasis>
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29263, 
<figureCitation id="136F2FB4FFE6E319FCA4FDFFFC7C82EC" box="[843,969,1411,1437]" captionStart="PLATE 12" captionStartId="118.[151,234,1744,1768]" captionTargetBox="[213,1375,181,1720]" captionTargetId="figure-20@118.[213,1375,181,1720]" captionTargetPageId="118" captionText="PLATE 12. A–F Lioharpes cf. morocconensis, from a lower Emsian horizon, Boutiskaouine Formation, 28.3 km SSW of Rissani (Fig. 2B, Map 14, site 10), dorsal exoskeleton of partial cephalon, NHMUK It 29273. A, dorsal, B, anterior and C and D, lateral view. E, lateral view of eye lobe and F, dorsal view of brim perforation. G–M Lioharpes ammari n. sp., from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 9), dorsal exoskeleton with pygidium enrolled, holotype, NHMUK It 29263. G, dorsal, H, anterior and I, lateral view. J and K, dorsal and lateral views, respectively, of eye lobe. L and M, dorsal view of brim perforations and exterior rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232571" httpUri="https://zenodo.org/record/11232571/files/figure.png" pageId="57" pageNumber="58">Pl. 12G–M</figureCitation>
, from an 
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horizon, 
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, north flank of 
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(
<figureCitation id="136F2FB4FFE6E319FD90FDDBFD6F82B0" box="[639,730,1447,1473]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="57" pageNumber="58">Fig. 2B</figureCitation>
, Map 4, site 9), dorsal exoskeleton with pygidium enrolled
</materialsCitation>
. 
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<typeStatus id="54EF8D93FFE6E319FF78FDB7FF488294" box="[151,253,1483,1509]" pageId="57" pageNumber="58" type="paratype">Paratype</typeStatus>
: 
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<location id="8E8B65EAFFE6E319FE93FDB7FE388294" LSID="urn:lsid:plazi:treatment:03FD8227FFE6E31AFF78FC42FC9B81E8:8E8B65EAFFE6E319FE93FDB7FE388294" box="[380,397,1483,1509]" county="Jbel Ou-Driss" municipality="South" name="It" pageId="57" pageNumber="58">It</location>
29264, 
<figureCitation id="136F2FB4FFE6E319FE04FDB7FDD18294" box="[491,612,1483,1509]" captionStart="PLATE 13" captionStartId="119.[151,234,1650,1674]" captionTargetBox="[247,1340,181,1626]" captionTargetId="figure-15@119.[247,1340,181,1626]" captionTargetPageId="119" captionText="PLATE 13. A–J Lioharpes ammari n. sp., from horizon 2, Section 1 (Fig. 6), Tazoulait Formation, upper Emisian, South flank of Jbel Ou-Driss (Fig. 2B, Map 4). A–E dorsal exoskeleton, pygidium missing, paratype, NHMUK It 29264 (site 12).A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–H dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29265 (1) (site 13). F, dorsal, G, lateral and H, anterior view. I–J ventral exoskeleton of a partial upper lamella, paratype, NHMUK It 29265 (2) (site 13). I, ventral view and J, lateral view of ventral surface. K–O Pinnuloharpes igaouii n. sp., from a horizon 20 metres down slope from type horizon of Lioharpes ammari n. sp. Boutiskaouine Formation?, lower Emsian North flank of Jbel Ou-Driss (Fig. 2B Map 4 site 6). Dorsal exoskeleton of cephalon only, paratype NHMUK it 29267. K dorsal, L anterior and M lateral view. N lateral view of eye lobe and O dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232573" httpUri="https://zenodo.org/record/11232573/files/figure.png" pageId="57" pageNumber="58">Pl. 13A–E</figureCitation>
, from horizon 2, 
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1 (
<figureCitation id="136F2FB4FFE6E319FC47FDB7FC5B8294" box="[936,1006,1483,1509]" captionStart="FIGURE 6" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[154,1432,185,1829]" captionTargetId="figure-20@12.[151,1435,181,1833]" captionTargetPageId="12" captionText="FIGURE 6. Section 1 Jbel Ou-Driss" figureDoi="http://doi.org/10.5281/zenodo.11232515" httpUri="https://zenodo.org/record/11232515/files/figure.png" pageId="57" pageNumber="58">Fig. 6</figureCitation>
), 
<location id="8E8B65EAFFE6E319FBEBFDB7FB598294" LSID="urn:lsid:plazi:treatment:03FD8227FFE6E31AFF78FC42FC9B81E8:8E8B65EAFFE6E319FBEBFDB7FB598294" box="[1028,1260,1483,1509]" county="Jbel Ou-Driss" municipality="South" name="Tazoulait Formation" pageId="57" pageNumber="58">Tazoulait Formation</location>
, 
<location id="8E8B65EAFFE6E319FB16FDB7FA2D8294" LSID="urn:lsid:plazi:treatment:03FD8227FFE6E31AFF78FC42FC9B81E8:8E8B65EAFFE6E319FB16FDB7FA2D8294" box="[1273,1432,1483,1509]" county="Jbel Ou-Driss" municipality="South" name="upper Emsian" pageId="57" pageNumber="58">upper Emsian</location>
, 
<collectingMunicipality id="6B8FA94BFFE6E319FF78FD93FF6D8178" box="[151,216,1519,1545]" pageId="57" pageNumber="58">South</collectingMunicipality>
flank of 
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(
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, Map 4, site 12), dorsal exoskeleton with pygidium missing
</materialsCitation>
; 
<materialsCitation id="3B3C396CFFE6E319FAC8FD93FA99815C" collectionCode="NHMUK" pageId="57" pageNumber="58" specimenCount="1" typeStatus="paratype">
<typeStatus id="54EF8D93FFE6E319FAC8FD93FA2D8178" box="[1319,1432,1519,1545]" pageId="57" pageNumber="58" type="paratype">Paratypes</typeStatus>
: 
<collectionCode id="ED45ABF4FFE6E319FF78FE6FFEB4815C" box="[151,257,1555,1581]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="57" pageNumber="58" type="Museum">NHMUK</collectionCode>
It 29265 (1), 
<figureCitation id="136F2FB4FFE6E319FE79FE6FFDBE815C" box="[406,523,1555,1581]" captionStart="PLATE 13" captionStartId="119.[151,234,1650,1674]" captionTargetBox="[247,1340,181,1626]" captionTargetId="figure-15@119.[247,1340,181,1626]" captionTargetPageId="119" captionText="PLATE 13. A–J Lioharpes ammari n. sp., from horizon 2, Section 1 (Fig. 6), Tazoulait Formation, upper Emisian, South flank of Jbel Ou-Driss (Fig. 2B, Map 4). A–E dorsal exoskeleton, pygidium missing, paratype, NHMUK It 29264 (site 12).A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–H dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29265 (1) (site 13). F, dorsal, G, lateral and H, anterior view. I–J ventral exoskeleton of a partial upper lamella, paratype, NHMUK It 29265 (2) (site 13). I, ventral view and J, lateral view of ventral surface. K–O Pinnuloharpes igaouii n. sp., from a horizon 20 metres down slope from type horizon of Lioharpes ammari n. sp. Boutiskaouine Formation?, lower Emsian North flank of Jbel Ou-Driss (Fig. 2B Map 4 site 6). Dorsal exoskeleton of cephalon only, paratype NHMUK it 29267. K dorsal, L anterior and M lateral view. N lateral view of eye lobe and O dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232573" httpUri="https://zenodo.org/record/11232573/files/figure.png" pageId="57" pageNumber="58">Pl. 13F–H</figureCitation>
, and 
<collectionCode id="ED45ABF4FFE6E319FDACFE6FFD18815C" box="[579,685,1555,1581]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="57" pageNumber="58" type="Museum">NHMUK</collectionCode>
It 29265 (2), 
<figureCitation id="136F2FB4FFE6E319FCADFE6FFC12815C" box="[834,935,1555,1581]" captionStart="PLATE 13" captionStartId="119.[151,234,1650,1674]" captionTargetBox="[247,1340,181,1626]" captionTargetId="figure-15@119.[247,1340,181,1626]" captionTargetPageId="119" captionText="PLATE 13. A–J Lioharpes ammari n. sp., from horizon 2, Section 1 (Fig. 6), Tazoulait Formation, upper Emisian, South flank of Jbel Ou-Driss (Fig. 2B, Map 4). A–E dorsal exoskeleton, pygidium missing, paratype, NHMUK It 29264 (site 12).A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–H dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29265 (1) (site 13). F, dorsal, G, lateral and H, anterior view. I–J ventral exoskeleton of a partial upper lamella, paratype, NHMUK It 29265 (2) (site 13). I, ventral view and J, lateral view of ventral surface. K–O Pinnuloharpes igaouii n. sp., from a horizon 20 metres down slope from type horizon of Lioharpes ammari n. sp. Boutiskaouine Formation?, lower Emsian North flank of Jbel Ou-Driss (Fig. 2B Map 4 site 6). Dorsal exoskeleton of cephalon only, paratype NHMUK it 29267. K dorsal, L anterior and M lateral view. N lateral view of eye lobe and O dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232573" httpUri="https://zenodo.org/record/11232573/files/figure.png" pageId="57" pageNumber="58">Pl. 13I–J</figureCitation>
, from same location and horizon as
</materialsCitation>
<collectionCode id="ED45ABF4FFE6E319FADDFE6FFA29815C" box="[1330,1436,1555,1581]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="57" pageNumber="58" type="Museum">NHMUK</collectionCode>
It 29264 (site 13). 
<collectionCode id="ED45ABF4FFE6E319FE80FE4BFE6C8120" box="[367,473,1591,1617]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="57" pageNumber="58" type="Museum">NHMUK</collectionCode>
It 29265 (1), dorsal exoskeleton of a partial cephalon, and 
<collectionCode id="ED45ABF4FFE6E319FB6CFE4BFB588120" box="[1155,1261,1591,1617]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="57" pageNumber="58" type="Museum">NHMUK</collectionCode>
It 29265 (2), a ventral exoskeleton of upper lamella.
</paragraph>
<paragraph id="8BEB3331FFE6E319FF28FE03FC5F81CC" blockId="57.[151,1437,1194,2013]" pageId="57" pageNumber="58">
<materialsCitation id="3B3C396CFFE6E319FF28FE03FC5381CC" collectionCode="NHMUK" pageId="57" pageNumber="58" specimenCount="2">
<typeStatus id="54EF8D93FFE6E319FF28FE03FEDA81E8" box="[199,367,1663,1689]" isOtherTypeStatus="true" pageId="57" pageNumber="58">Other material</typeStatus>
: 
<taxonomicName id="4C5448B2FFE6E319FE94FE03FDF981E9" authorityName="Johnson" authorityYear="2024" box="[379,588,1663,1689]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE6E319FE94FE03FDF981E9" box="[379,588,1663,1689]" italics="true" pageId="57" pageNumber="58">Lioharpes ammari</emphasis>
</taxonomicName>
? (meraspis / early holaspid), 
<collectionCode id="ED45ABF4FFE6E319FC70FE03FBBC81E8" box="[927,1033,1663,1689]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="57" pageNumber="58" type="Museum">NHMUK</collectionCode>
It 29266, 
<figureCitation id="136F2FB4FFE6E319FB6EFE03FB4381E8" box="[1153,1270,1662,1689]" captionStart="PLATE 53" captionStartId="159.[151,234,1654,1678]" captionTargetBox="[247,1340,181,1630]" captionTargetId="figure-15@159.[247,1340,181,1630]" captionTargetPageId="159" captionText="PLATE 53. A–F Lioharpes ammari n. sp. (?), a small holaspid or late meraspid from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 25), dorsal exoskeleton of partial cephalon, NHMUK It 29266. A, dorsal view, B anterior view and C and D lateral views. E, dorsal view of brim perforations and F, laterodorsal view of eye lobe. G–L Eoharpes beningensis (Barrande 1872), from Dobrotivá beds, Llandeilo, Lower Ordovician Svatá Dobrovtivá central Bohemia. G–H partial dorsal exoskeleton, Museum of Comparative Zoology Harvard University, IP 173331. G dorsal view and H, lateral view. I–L partial dorsal exoskeleton, Museum of Comparative Zoology Harvard University, IP 173316. I, dorsal view and L, lateral view. J, dorsal view of brim perforations and K, lateral view of eye lobe and caeca on genal area. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232695" httpUri="https://zenodo.org/record/11232695/files/figure.png" pageId="57" pageNumber="58">Pl. 53A–F</figureCitation>
, from the 
<typeStatus id="54EF8D93FFE6E319FA83FE03FA2981E8" box="[1388,1436,1663,1689]" pageId="57" pageNumber="58">type</typeStatus>
horizon and location, north flank of Jbel Ou-Driss (
<figureCitation id="136F2FB4FFE6E319FD39FEDFFC9B81CC" box="[726,814,1699,1725]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="57" pageNumber="58">Fig. 2B</figureCitation>
, Map 4, site 25)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E60BAFFE6E31AFF28FEBAFF79842C" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="description">
<paragraph id="8BEB3331FFE6E31AFF28FEBAFA9A87A4" blockId="57.[151,1437,1194,2013]" lastBlockId="58.[151,1437,150,1689]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<emphasis id="B920EF23FFE6E319FF28FEBAFEEC8191" bold="true" box="[199,345,1734,1760]" pageId="57" pageNumber="58">Description.</emphasis>
Cephalon widest at midpoint between eyes and alae, genicranium broad (tr.) with width about 1.4 x length (sag.) and widest at posterior border. Glabella anterior to S0 carinate, with pronounced tubercles along crest and broad with width (tr.) over 75% of length (sag,). S1 slanting posteriorly, not reaching halfway up flank of glabella, no muscle insertion furrows at S2, S3, S4. Axial furrows convex, converging at around 
<superScript id="7C219E79FFE6E319FB19FF4FFAAF8031" attach="left" box="[1270,1306,1842,1869]" fontSize="6" pageId="57" pageNumber="58">170</superScript>
to axis and not crossing junction between F1 and ala. Preglabellar furrow shallow and straight (tr.) in anterior view. Occipital furrow deep, straight (tr.) and medially broader (sag.) than posterior border (exsag.). Occipital ring same width (sag.) as posterior border (exsag.), higher than crest of glabella in lateral view and with pronounced tubercles on dorsal surface. Occipital node large and strongly inflated. L1 inflated, alae laterally directed, large at over 30% of width (tr.) of glabella at S1 and over 85% of length (exsag.). Alar furrow shallow and alar depression present. Posterior border short width (tr.), just under 30% of width of occipital ring (tr.) and preglabellar genal area narrow.
</paragraph>
<paragraph id="8BEB3331FFE5E31AFF28F8A3FB4B86A0" blockId="58.[151,1437,150,1689]" pageId="58" pageNumber="59">
Genal area moderately broad with width (tr.) over 60% length (exsag.). Eye lobes strongly inflated, tall, oval in dorsal view, tubercles on dorsal surface and with two eye lenses. Eye lenses of about equal size and elongated, recumbent rectangles with rounded corners in shape (
<figureCitation id="136F2FB4FFE5E31AFD12F95BFCEF8630" box="[765,858,295,321]" captionStart="PLATE 12" captionStartId="118.[151,234,1744,1768]" captionTargetBox="[213,1375,181,1720]" captionTargetId="figure-20@118.[213,1375,181,1720]" captionTargetPageId="118" captionText="PLATE 12. A–F Lioharpes cf. morocconensis, from a lower Emsian horizon, Boutiskaouine Formation, 28.3 km SSW of Rissani (Fig. 2B, Map 14, site 10), dorsal exoskeleton of partial cephalon, NHMUK It 29273. A, dorsal, B, anterior and C and D, lateral view. E, lateral view of eye lobe and F, dorsal view of brim perforation. G–M Lioharpes ammari n. sp., from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 9), dorsal exoskeleton with pygidium enrolled, holotype, NHMUK It 29263. G, dorsal, H, anterior and I, lateral view. J and K, dorsal and lateral views, respectively, of eye lobe. L and M, dorsal view of brim perforations and exterior rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232571" httpUri="https://zenodo.org/record/11232571/files/figure.png" pageId="58" pageNumber="59">Pl. 12K</figureCitation>
). Eye lobes close to internal margin of fringe; eye location ratio over 0.80. No eye ridge or genal ridge. Anterior boss barely inflated but reaching down to weak girder kink. Genal roll sloping gently, at just under 
<superScript id="7C219E79FFE5E31AFD63F912FD05860D" attach="left" box="[652,688,366,393]" fontSize="6" pageId="58" pageNumber="59">500</superScript>
anteriorly and about 
<superScript id="7C219E79FFE5E31AFC70F912FC76860D" attach="left" box="[927,963,366,393]" fontSize="6" pageId="58" pageNumber="59">550</superScript>
laterally. Row of large perforations above girder and across anterior boss at internal margin of fringe. Above row of large perforations at girder, perforations smaller than those on brim and decrease in size upwards away from girder. Genal roll without caeca.
</paragraph>
<paragraph id="8BEB3331FFE5E31AFF28F9A7FDBE85F4" blockId="58.[151,1437,150,1689]" pageId="58" pageNumber="59">
Brim sloping moderately at over 
<superScript id="7C219E79FFE5E31AFDAAF9A7FDDC8699" attach="none" box="[581,617,474,501]" fontSize="6" pageId="58" pageNumber="59">100</superScript>
, concave (sag.) straight laterally and width (sag.) over 35% of cephalic length (sag.). Brim width ratio 0.70 and standardised brim perforations moderately fine at around 140μm with larger row of perforations at distal and proximal edges. Light caeca reaching across brim. External rim stout with a few scattered fine tubercles on dorsal surface. Marginal band near vertical and without tubercles. Brim steepens quickly to near vertical on prolongations.
</paragraph>
<paragraph id="8BEB3331FFE5E31AFF28FAF3FCB68464" blockId="58.[151,1437,150,1689]" pageId="58" pageNumber="59">Prolongations moderately short (exsag.), around 80% of cephalic length (sag.). Internal and external rims curving adaxially posteriorly and internal rims with weak wave-like profile in lateral view. Extension of girder meeting internal rim about 45% of way down prolongation. Tubercles on dorsal surface of internal rim and genal spine short, under 10% length (exsag.) of prolongation.</paragraph>
<paragraph id="8BEB3331FFE5E31AFF28FB63FF79842C" blockId="58.[151,1437,150,1689]" pageId="58" pageNumber="59">Thorax with 20 segments and axis tapering posteriorly with width (tr.) of last axial ring being under 40% of first.</paragraph>
</subSubSection>
<subSubSection id="C34E60BAFFE5E31AFF28FB1BFC9B81E8" pageId="58" pageNumber="59" type="discussion">
<paragraph id="8BEB3331FFE5E31AFF28FB1BFDA784D4" blockId="58.[151,1437,150,1689]" pageId="58" pageNumber="59">
<emphasis id="B920EF23FFE5E31AFF28FB1BFE8E84F0" bold="true" box="[199,315,871,897]" pageId="58" pageNumber="59">Remarks.</emphasis>
<taxonomicName id="4C5448B2FFE5E31AFEAEFB1BFDBA84F1" authorityName="Johnson" authorityYear="2024" box="[321,527,871,897]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE5E31AFEAEFB1BFDBA84F1" box="[321,527,871,897]" italics="true" pageId="58" pageNumber="59">Lioharpes ammari</emphasis>
</taxonomicName>
lacks one of the diagnostic characters of its genus as shown in 
<tableCitation id="C6D6068AFFE5E31AFB29FB1BFA9284F0" box="[1222,1319,870,897]" captionStart="TABLE 5" captionStartId="182.[151,239,503,529]" captionText="TABLE 5b. Proposed diagnostic characters for Devonian harpetid genera (continued)" pageId="58" pageNumber="59">Table 5a</tableCitation>
: it has two eye lenses per eye lobe, not three.
</paragraph>
<paragraph id="8BEB3331FFE5E31AFF28FBD3FCDD8344" blockId="58.[151,1437,150,1689]" pageId="58" pageNumber="59">
<taxonomicName id="4C5448B2FFE5E31AFF28FBD3FE2D84B9" authorityName="Johnson" authorityYear="2024" box="[199,408,943,969]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE5E31AFF28FBD3FE2D84B9" box="[199,408,943,969]" italics="true" pageId="58" pageNumber="59">Lioharpes ammari</emphasis>
</taxonomicName>
sister species is 
<taxonomicName id="4C5448B2FFE5E31AFDB8FBD3FD5484B8" authorityName="Johnson" authorityYear="2024" box="[599,737,943,969]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="scopulum">
<emphasis id="B920EF23FFE5E31AFDB8FBD3FD5484B8" box="[599,737,943,969]" italics="true" pageId="58" pageNumber="59">L. scopulum</emphasis>
</taxonomicName>
and differs in that 
<taxonomicName id="4C5448B2FFE5E31AFC55FBD3FB8584B9" authorityName="Johnson" authorityYear="2024" box="[954,1072,943,968]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE5E31AFC55FBD3FB8584B9" box="[954,1072,943,968]" italics="true" pageId="58" pageNumber="59">L. ammari</emphasis>
</taxonomicName>
has: a broader genicranium and glabella; axial furrows that converge more strongly; a broader occipital furrow; larger alae (tr. and exsag.); eye lobes that are more strongly inflated and have two lenses each; no caeca on the genal roll; and internal rims on the prolongations that have a weak wave-like profile in lateral view.
</paragraph>
<paragraph id="8BEB3331FFE5E31AFF28FC43FAF98294" blockId="58.[151,1437,150,1689]" pageId="58" pageNumber="59">
<taxonomicName id="4C5448B2FFE5E31AFF28FC43FE238329" authorityName="Johnson" authorityYear="2024" box="[199,406,1087,1113]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE5E31AFF28FC43FE238329" box="[199,406,1087,1113]" italics="true" pageId="58" pageNumber="59">Lioharpes ammari</emphasis>
</taxonomicName>
is similar to certain 
<taxonomicName id="4C5448B2FFE5E31AFD98FC43FCC68328" box="[631,883,1087,1113]" pageId="58" pageNumber="59">
<emphasis id="B920EF23FFE5E31AFD98FC43FCA98328" box="[631,796,1087,1113]" italics="true" pageId="58" pageNumber="59">Pinnuloharpes</emphasis>
species
</taxonomicName>
and in particular 
<taxonomicName id="4C5448B2FFE5E31AFBDAFC43FB0C8328" authorityName="Johnson" authorityYear="2024" box="[1077,1209,1087,1113]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="haustrum">
<emphasis id="B920EF23FFE5E31AFBDAFC43FB0C8328" box="[1077,1209,1087,1113]" italics="true" pageId="58" pageNumber="59">P. haustrum</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C5448B2FFE5E31AFB29FC43FA208329" authorityName="Johnson" authorityYear="2024" box="[1222,1429,1087,1113]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE5E31AFB29FC43FA208329" box="[1222,1429,1087,1113]" italics="true" pageId="58" pageNumber="59">Lioharpes ammari</emphasis>
</taxonomicName>
, however, has a number of differences, the most important of which are: a wider (tr.) genicranium and glabella; a glabella that is more tapered anteriorly and has tubercles along the crest; a wider occipital furrow and an occipital ring that is higher than the glabella anterior to S0. 
<taxonomicName id="4C5448B2FFE5E31AFD38FCD7FC1D83B5" authorityName="Johnson" authorityYear="2024" box="[727,936,1195,1221]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE5E31AFD38FCD7FC1D83B5" box="[727,936,1195,1221]" italics="true" pageId="58" pageNumber="59">Lioharpes ammari</emphasis>
</taxonomicName>
also has an axial furrow that does not cross the junction of L1 with ala; deeper alar furrows; taller eye lobes that are set much closer to the inner margin of fringe; there is no genal ridge or eye ridge, both of which are just visible on 
<taxonomicName id="4C5448B2FFE5E31AFC06FC8FFB4A827C" authorityName="Johnson" authorityYear="2024" box="[1001,1279,1267,1293]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="haustrum">
<emphasis id="B920EF23FFE5E31AFC06FC8FFB4A827C" box="[1001,1279,1267,1293]" italics="true" pageId="58" pageNumber="59">Pinnuloharpes haustrum</emphasis>
</taxonomicName>
; inner margin of fringe across the anterior of genal area is straight, not concave; and the genal roll is not so steeply-sloping. Brim of 
<taxonomicName id="4C5448B2FFE5E31AFF59FD47FE308225" authorityName="Johnson" authorityYear="2024" box="[182,389,1339,1365]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE5E31AFF59FD47FE308225" box="[182,389,1339,1365]" italics="true" pageId="58" pageNumber="59">Lioharpes ammari</emphasis>
</taxonomicName>
is wider (sag.), steeper-sloping and with a higher brim width ratio at 0.70, rather than 
<quantity id="4CAC9ED4FFE5E31AFAA1FD46FA298224" box="[1358,1436,1338,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.4224" pageId="58" pageNumber="59" unit="in" value="0.56">0.56 in</quantity>
the case of 
<emphasis id="B920EF23FFE5E31AFEFBFD23FD848208" box="[276,561,1375,1401]" italics="true" pageId="58" pageNumber="59">
<taxonomicName id="4C5448B2FFE5E31AFEFBFD23FD988208" authorityName="Johnson" authorityYear="2024" box="[276,557,1375,1401]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="haustrum">Pinnuloharpes haustrum</taxonomicName>
.
</emphasis>
Internal rim and external meet at the end of prolongations, whereas those of 
<taxonomicName id="4C5448B2FFE5E31AFA67FD23FEB482EC" authorityName="Johnson" authorityYear="2024" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="haustrum">
<emphasis id="B920EF23FFE5E31AFA67FD23FEB482EC" italics="true" pageId="58" pageNumber="59">P. haustrum</emphasis>
</taxonomicName>
meet and run together for a short distance posteriorly before joining. The genal spine of 
<taxonomicName id="4C5448B2FFE5E31AFB21FDFFFA2982ED" authorityName="Johnson" authorityYear="2024" box="[1230,1436,1411,1437]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE5E31AFB21FDFFFA2982ED" box="[1230,1436,1411,1437]" italics="true" pageId="58" pageNumber="59">Lioharpes ammari</emphasis>
</taxonomicName>
is shorter and there are no larger perforations under the internal rim. The thorax of 
<taxonomicName id="4C5448B2FFE5E31AFBA7FDDBFB0B82B1" authorityName="Johnson" authorityYear="2024" box="[1096,1214,1447,1472]" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE5E31AFBA7FDDBFB0B82B1" box="[1096,1214,1447,1472]" italics="true" pageId="58" pageNumber="59">L. ammari</emphasis>
</taxonomicName>
has a more tapered axis, lacks pitting on the axial rings and pleurae, and the distal ends of the pleurae are rounded, not pointed.
</paragraph>
<paragraph id="8BEB3331FFE5E31AFF28FD93FC9B81E8" blockId="58.[151,1437,150,1689]" pageId="58" pageNumber="59">
The 
<emphasis id="B920EF23FFE5E31AFF15FD93FED78178" box="[250,354,1519,1545]" italics="true" pageId="58" pageNumber="59">meraspid</emphasis>
or perhaps early holaspid (
<figureCitation id="136F2FB4FFE5E31AFD7DFD93FCBF8178" box="[658,778,1518,1545]" captionStart="PLATE 53" captionStartId="159.[151,234,1654,1678]" captionTargetBox="[247,1340,181,1630]" captionTargetId="figure-15@159.[247,1340,181,1630]" captionTargetPageId="159" captionText="PLATE 53. A–F Lioharpes ammari n. sp. (?), a small holaspid or late meraspid from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 25), dorsal exoskeleton of partial cephalon, NHMUK It 29266. A, dorsal view, B anterior view and C and D lateral views. E, dorsal view of brim perforations and F, laterodorsal view of eye lobe. G–L Eoharpes beningensis (Barrande 1872), from Dobrotivá beds, Llandeilo, Lower Ordovician Svatá Dobrovtivá central Bohemia. G–H partial dorsal exoskeleton, Museum of Comparative Zoology Harvard University, IP 173331. G dorsal view and H, lateral view. I–L partial dorsal exoskeleton, Museum of Comparative Zoology Harvard University, IP 173316. I, dorsal view and L, lateral view. J, dorsal view of brim perforations and K, lateral view of eye lobe and caeca on genal area. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232695" httpUri="https://zenodo.org/record/11232695/files/figure.png" pageId="58" pageNumber="59">Pl. 53A–F</figureCitation>
), coming as it does from the 
<typeStatus id="54EF8D93FFE5E31AFBA2FD93FBC88178" box="[1101,1149,1519,1545]" pageId="58" pageNumber="59">type</typeStatus>
horizon, is most likely 
<taxonomicName id="4C5448B2FFE5E31AFA69FD93FF5B815D" authorityName="Johnson" authorityYear="2024" class="Trilobita" family="Harpetidae" genus="Lioharpes" kingdom="Animalia" order="Harpetida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ammari">
<emphasis id="B920EF23FFE5E31AFA69FD93FF5B815D" italics="true" pageId="58" pageNumber="59">L. ammari</emphasis>
</taxonomicName>
. It has the same brim width ratio and other morphological features. However, the eye lobes are not as tall, the glabella is not carinated and the tubercles on the glabella and eye lobes appear not to be so well developed. It may be that some of these features have been lost in the preparation of such a small specimen (cephalon sag. length = 
<quantity id="4CAC9ED4FFE5E31AFF41FE02FEB081E8" box="[174,261,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="58" pageNumber="59" unit="mm" value="7.5">7.5 mm</quantity>
) or that they may have developed in later instars.
</paragraph>
</subSubSection>
</treatment>
</document>