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<document id="FD296AB8224AF163B292DD167FEFC774" ID-DOI="10.35885/ruthenica.2022.32(2).5" ID-ISSN="2307-7336" ID-Zenodo-Dep="11041109" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1713810364962" checkinUser="felipe" docAuthor="Hasegawa, Kazunori" docDate="2022" docId="03F65E35FFBE3D4CFF7B0EE1A27FF982" docLanguage="en" docName="Ruthenica.32.2.85-92.pdf" docOrigin="Ruthenica, Russian Malacological Journal 32 (2)" docSource="http://dx.doi.org/10.35885/ruthenica.2022.32(2).5" docStyle="DocumentStyle:3A4A24405A53639AD762166AE048D148.1:Ruthenica.2022-.journal_article.english.index" docStyleId="3A4A24405A53639AD762166AE048D148" docStyleName="Ruthenica.2022-.journal_article.english.index" docStyleVersion="1" docTitle="Frigidoalvania asura" docType="treatment" docVersion="3" lastPageNumber="88" masterDocId="FFCF264DFFBD3D4FFFBA0C61A34EFFEB" masterDocTitle="Bathyal Rissoidae (Gastropoda: Rissooidea) off the Russian Far East coast of the Sea of Japan, with redescription of Punctulum reticulatum Golikov, 1986" masterLastPageNumber="92" masterPageNumber="85" pageNumber="88" updateTime="1715157570046" updateUser="tatiana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="307C087BD16221F81B07D1ABF4C3E587">Bathyal Rissoidae (Gastropoda: Rissooidea) off the Russian Far East coast of the Sea of Japan, with redescription of Punctulum reticulatum Golikov, 1986</mods:title>
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<mods:namePart id="BBB91BE19A88755657A0D4E7F855A383">Hasegawa, Kazunori</mods:namePart>
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<mods:title id="FFE6A50AC47986EEDB2AF31515EC4C57">Ruthenica, Russian Malacological Journal</mods:title>
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<mods:date id="5177C1D608AC979D1723A0FBCAC3753D">2022</mods:date>
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<mods:number id="FB93206DBA267D22B4F22CB6ABAE6636">32</mods:number>
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<mods:classification id="BDC8E3445498314CBB53383E57744BEC">journal article</mods:classification>
<mods:identifier id="0BC04BE07B0A1EED2E6C325D641F9785" type="DOI">10.35885/ruthenica.2022.32(2).5</mods:identifier>
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<treatment id="03F65E35FFBE3D4CFF7B0EE1A27FF982" LSID="urn:lsid:plazi:treatment:03F65E35FFBE3D4CFF7B0EE1A27FF982" httpUri="http://treatment.plazi.org/id/03F65E35FFBE3D4CFF7B0EE1A27FF982" lastPageNumber="88" pageId="3" pageNumber="88">
<subSubSection id="C345BCA8FFBE3D4CFF7B0EE1A184FD76" box="[193,714,640,669]" pageId="3" pageNumber="88" type="nomenclature">
<paragraph id="8BE0EF23FFBE3D4CFF7B0EE1A184FD76" blockId="3.[193,714,640,700]" box="[193,714,640,669]" pageId="3" pageNumber="88">
<heading id="D0A8584FFFBE3D4CFF7B0EE1A184FD76" box="[193,714,640,669]" centered="true" fontSize="12" level="2" pageId="3" pageNumber="88" reason="4">
<taxonomicName id="4C5F94A0FFBE3D4CFF7B0EE1A184FD76" ID-CoL="6K236" authority="(Yokoyama, 1926)" baseAuthorityName="Yokoyama" baseAuthorityYear="1926" box="[193,714,640,669]" class="Gastropoda" family="Rissoidae" genus="Frigidoalvania" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="88" phylum="Mollusca" rank="species" species="asura">
<emphasis id="B92B3331FFBE3D4CFF7B0EE1A29AFD76" box="[193,468,640,669]" italics="true" pageId="3" pageNumber="88">Frigidoalvania asura</emphasis>
(
<bibRefCitation id="EFCE92D2FFBE3D4CFE5E0EE1A18FFD76" author="Yokoyama M." box="[484,705,640,669]" pageId="3" pageNumber="88" pagination="249 - 312" refId="ref6457" refString="Yokoyama M. 1926. Fossil shells from Sado. Journal of the Faculty of Science, Imperial University of Tokyo, Section II Geology, Mineralogy, Geography, Seismology, 1: 249 - 312, pls. 32 - 37." type="journal article" year="1926">Yokoyama, 1926</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C345BCA8FFBE3D4CFEC00EC3A15FFD57" box="[378,529,674,700]" pageId="3" pageNumber="88" type="description">
<paragraph id="8BE0EF23FFBE3D4CFEC00EC3A15FFD57" blockId="3.[193,714,640,700]" box="[378,529,674,700]" pageId="3" pageNumber="88">
(
<figureCitation id="1364F3A6FFBE3D4CFE390EC3A146FD57" box="[387,520,674,700]" captionStart="FIG" captionStartId="4.[174,216,1724,1745]" captionTargetBox="[174,1383,189,1701]" captionTargetId="figure-10@4.[174,1384,189,1701]" captionTargetPageId="4" captionText="FIG. 2. Species of Rissoidae. A. Punctulum akibai (Yokoyama, 1926), SoJaBio stn. A2-10 (455-456 m), MIBM 39336. B. Punctulum nihonkaiensis (Hasegawa, 2014), SoJaBio stn. A2-10 (455-456 m), MIBM 39337. C-D. Frigidoalvania asura (Yokoyama, 1926), SoJaBio stn. B7-7 (470-528 m), MIBM 39340. E-F. Frigidoalvania tanseimaruae Hasegawa, 2014, SoJaBio stn. B6-7 (1,001-1,011 m). G-J. Punctulum flavum (Okutani, 1964); G-H, “shallow-water form”, SoJaBio stn. A2-10 (455-456 m), MIBM 14270; I-J, “deep-water form”, SoJaBio stn. A6-7 (2,511-2,534 m), MIBM 39334. All at the same scale; scale = 1 mm. РИС. 2.Виды Rissoidae.A. Punctulum akibai (Yokoyama,1926), SoJaBioст.A2-10 (455-456м), MIBM 39336.B. Punctulum nihonkaiensis (Hasegawa, 2014), SoJaBio ст. A2-10 (455-456 м), MIBM 39337. C-D. Frigidoalvania asura (Yokoyama, 1926), SoJaBio ст. B7-7 (470-528 м), MIBM 39340. E-F. Frigidoalvania tanseimaruae Hasegawa, 2014, SoJaBio ст. B6-7 (1001-1011 м). G-J. Punctulum flavum (Okutani, 1964); G-H, “мелководнаЯ форма”, SoJaBio ст. A2-10 (455-456 м), MIBM 14270; I-J, “глубоководнаЯ форма”, SoJaBio ст. A6-7 (2511-2534 м), MIBM 39334. Все раковины в одном масШтабе, масШтабнаЯ линейка = 1 мм." figureDoi="http://doi.org/10.5281/zenodo.11041111" httpUri="https://zenodo.org/record/11041111/files/figure.png" pageId="3" pageNumber="88">Fig. 2 C–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C345BCA8FFBE3D4CFF1C0EB3A1AAFCE8" pageId="3" pageNumber="88" type="reference_group">
<paragraph id="8BE0EF23FFBE3D4CFF1C0EB3A1AAFCE8" blockId="3.[166,740,721,771]" pageId="3" pageNumber="88">
<treatmentCitationGroup id="AB4FC80DFFBE3D4CFF1C0EB3A1AAFCE8" pageId="3" pageNumber="88">
<taxonomicName id="4C5F94A0FFBE3D4CFF1C0EB3A132FD0D" ID-CoL="7PFV3" authority="Yokoyama, 1926: 273" authorityName="Desmarest" authorityPageNumber="273" authorityYear="1814" box="[166,636,721,743]" class="Gastropoda" family="Rissoidae" genus="Rissoa" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="88" phylum="Mollusca" rank="genus">
<emphasis id="B92B3331FFBE3D4CFF1C0EB3A222FD0D" box="[166,364,721,743]" italics="true" pageId="3" pageNumber="88">Rissoa (Apicularia?)</emphasis>
<treatmentCitation id="0AFEC932FFBE3D4CFECB0EB3A132FD0D" author="Yokoyama M." box="[369,636,721,743]" page="273" pageId="3" pageNumber="88" year="1926">
<emphasis id="B92B3331FFBE3D4CFECB0EB3A2E9FD0C" box="[369,423,722,743]" italics="true" pageId="3" pageNumber="88">asura</emphasis>
<bibRefCitation id="EFCE92D2FFBE3D4CFE100EB0A132FD0D" author="Yokoyama M." box="[426,636,721,743]" pageId="3" pageNumber="88" pagination="249 - 312" refId="ref6457" refString="Yokoyama M. 1926. Fossil shells from Sado. Journal of the Faculty of Science, Imperial University of Tokyo, Section II Geology, Mineralogy, Geography, Seismology, 1: 249 - 312, pls. 32 - 37." type="journal article" year="1926">Yokoyama, 1926: 273</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, pl. 33, fig. 15 (see
<bibRefCitation id="EFCE92D2FFBE3D4CFEAD0E8FA2F1FCE9" author="Hasegawa K." box="[279,447,748,771]" pageId="3" pageNumber="88" pagination="75 - 148" refId="ref5887" refString="Hasegawa K. 2014. A review of bathyal Rissoidae in the Sea of Japan and adjacent waters (Gastropoda: Rissooidea). In: Fujita T. (Ed.). Deep-sea Fauna of the Sea of Japan. National Museum of Nature and Science Monographs, (44): 75 - 148." type="journal article" year="2014">Hasegawa [2014]</bibRefCitation>
for more detailed synonymy).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C345BCA8FFBE3D4CFF760F76A2A7FB5C" pageId="3" pageNumber="88" type="materials_examined">
<paragraph id="8BE0EF23FFBE3D4CFF760F76A263FC8E" blockId="3.[165,742,791,1208]" pageId="3" pageNumber="88">
<emphasis id="B92B3331FFBE3D4CFF760F76A2D7FCC6" bold="true" box="[204,409,791,813]" pageId="3" pageNumber="88">Material examined</emphasis>
. SoJaBio B 6-7, MIBM 14992; SoJaBio B 7-7, MIBM 39340 (
<figureCitation id="1364F3A6FFBE3D4CFE5A0F52A11AFCA2" box="[480,596,819,841]" captionStart="FIG" captionStartId="4.[174,216,1724,1745]" captionTargetBox="[174,1383,189,1701]" captionTargetId="figure-10@4.[174,1384,189,1701]" captionTargetPageId="4" captionText="FIG. 2. Species of Rissoidae. A. Punctulum akibai (Yokoyama, 1926), SoJaBio stn. A2-10 (455-456 m), MIBM 39336. B. Punctulum nihonkaiensis (Hasegawa, 2014), SoJaBio stn. A2-10 (455-456 m), MIBM 39337. C-D. Frigidoalvania asura (Yokoyama, 1926), SoJaBio stn. B7-7 (470-528 m), MIBM 39340. E-F. Frigidoalvania tanseimaruae Hasegawa, 2014, SoJaBio stn. B6-7 (1,001-1,011 m). G-J. Punctulum flavum (Okutani, 1964); G-H, “shallow-water form”, SoJaBio stn. A2-10 (455-456 m), MIBM 14270; I-J, “deep-water form”, SoJaBio stn. A6-7 (2,511-2,534 m), MIBM 39334. All at the same scale; scale = 1 mm. РИС. 2.Виды Rissoidae.A. Punctulum akibai (Yokoyama,1926), SoJaBioст.A2-10 (455-456м), MIBM 39336.B. Punctulum nihonkaiensis (Hasegawa, 2014), SoJaBio ст. A2-10 (455-456 м), MIBM 39337. C-D. Frigidoalvania asura (Yokoyama, 1926), SoJaBio ст. B7-7 (470-528 м), MIBM 39340. E-F. Frigidoalvania tanseimaruae Hasegawa, 2014, SoJaBio ст. B6-7 (1001-1011 м). G-J. Punctulum flavum (Okutani, 1964); G-H, “мелководнаЯ форма”, SoJaBio ст. A2-10 (455-456 м), MIBM 14270; I-J, “глубоководнаЯ форма”, SoJaBio ст. A6-7 (2511-2534 м), MIBM 39334. Все раковины в одном масШтабе, масШтабнаЯ линейка = 1 мм." figureDoi="http://doi.org/10.5281/zenodo.11041111" httpUri="https://zenodo.org/record/11041111/files/figure.png" pageId="3" pageNumber="88">Fig. 2 C–D</figureCitation>
) [depth range:
<quantity id="4CA742C6FFBE3D4CFF1C0F2EA26CFC8E" box="[166,290,847,869]" metricMagnitude="2" metricUnit="m" metricValue="7.404999999999999" metricValueMax="10.11" metricValueMin="4.7" pageId="3" pageNumber="88" unit="m" value="740.5" valueMax="1011.0" valueMin="470.0">470–1011 m</quantity>
].
</paragraph>
<paragraph id="8BE0EF23FFBE3D4CFF760F0AA2A7FB5C" blockId="3.[165,742,791,1208]" pageId="3" pageNumber="88">
<emphasis id="B92B3331FFBE3D4CFF760F0AA14FFC6A" bold="true" box="[204,513,875,897]" pageId="3" pageNumber="88">Additional material examined</emphasis>
(cited from
<bibRefCitation id="EFCE92D2FFBE3D4CFDC60F0DA394FC75" author="Hasegawa K." pageId="3" pageNumber="88" pagination="75 - 148" refId="ref5887" refString="Hasegawa K. 2014. A review of bathyal Rissoidae in the Sea of Japan and adjacent waters (Gastropoda: Rissooidea). In: Fujita T. (Ed.). Deep-sea Fauna of the Sea of Japan. National Museum of Nature and Science Monographs, (44): 75 - 148." type="journal article" year="2014">Hasegawa, 2014</bibRefCitation>
).
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<specimenCode id="DBF94758FFBE3D4CFF520FE9A21FFC75" box="[232,337,904,926]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="88" type="Museum">ZIN 33623</specimenCode>
, off
<collectorName id="26AA8AF5FFBE3D4CFEC20FE9A141FC76" box="[376,527,904,926]" pageId="3" pageNumber="88">SW Sakhalin, N</collectorName>
.
<collectorName id="26AA8AF5FFBE3D4CFDAC0FE9A176FC76" box="[534,568,904,925]" pageId="3" pageNumber="88">Sea</collectorName>
of
<collectingCountry id="F348AFB3FFBE3D4CFDEE0FE9A1C2FC76" box="[596,652,904,925]" name="Japan" pageId="3" pageNumber="88">Japan</collectingCountry>
,
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,
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,
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,
<quantity id="4CA742C6FFBE3D4CFE1E0FC5A2AFFC52" box="[420,481,932,953]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="88" unit="m" value="100.0">100 m</quantity>
;
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<specimenCode id="DBF94758FFBE3D4CFE530FC5A11FFC51" box="[489,593,932,954]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="88" type="Museum">ZIN 33625</specimenCode>
,
<location id="8E80B9F8FFBE3D4CFDE30FC5A1C7FC52" LSID="urn:lsid:plazi:treatment:03F65E35FFBE3D4CFF7B0EE1A27FF982:8E80B9F8FFBE3D4CFDE30FC5A1C7FC52" box="[601,649,932,953]" country="Japan" latitude="42.31333" longLatPrecision="114" longitude="131.15834" municipality="Great Bay" name="Peter" pageId="3" pageNumber="88">Peter</location>
the
<collectingMunicipality id="6B847559FFBE3D4CFD150FC5A383FC3D" pageId="3" pageNumber="88">Great Bay</collectingMunicipality>
,
<geoCoordinate id="EE6B89E4FFBE3D4CFF630FA1A20EFC3E" box="[217,320,960,981]" degrees="42" direction="north" minutes="18.8" orientation="latitude" pageId="3" pageNumber="88" precision="92" value="42.31333">42°18.8’N</geoCoordinate>
,
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,
<quantity id="4CA742C6FFBE3D4CFE720FA1A146FC3D" box="[456,520,960,982]" metricMagnitude="2" metricUnit="m" metricValue="2.4699999999999998" pageId="3" pageNumber="88" unit="m" value="247.0">
<elevation id="00720810FFBE3D4CFE720FA1A146FC3D" box="[456,520,960,982]" metricMagnitude="2" metricUnit="m" metricValue="2.4699999999999998" pageId="3" pageNumber="88" unit="m" value="247.0">247 m</elevation>
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</materialsCitation>
;
<materialsCitation id="3B37E57EFFBE3D4CFDAF0FA0A179FC19" collectionCode="ZIN" country="Japan" elevation="140" latitude="42.256668" location="Peter" longLatPrecision="114" longitude="130.74333" municipality="Great Bay" pageId="3" pageNumber="88" specimenCode="ZIN 36609" specimenCount="1">
<specimenCode id="DBF94758FFBE3D4CFDAF0FA0A1CCFC3D" box="[533,642,960,982]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="88" type="Museum">ZIN 36609</specimenCode>
,
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the
<collectingMunicipality id="6B847559FFBE3D4CFF1C0FBCA249FC19" box="[166,263,989,1010]" pageId="3" pageNumber="88">Great Bay</collectingMunicipality>
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,
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,
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;
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the
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the
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, 157–
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<specimenCode id="DBF94758FFBE3D4CFD070874A3AAFBAC" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="88" type="Museum">ZIN 36612</specimenCode>
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<location id="8E80B9F8FFBE3D4CFF540853A251FBAC" LSID="urn:lsid:plazi:treatment:03F65E35FFBE3D4CFF7B0EE1A27FF982:8E80B9F8FFBE3D4CFF540853A251FBAC" box="[238,287,1074,1095]" country="Japan" latitude="42.55" longLatPrecision="114" longitude="132.85" municipality="Great Bay" name="Peter" pageId="3" pageNumber="88">Peter</location>
the
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<specimenCode id="DBF94758FFBE3D4CFF1C082FA25FFB88" box="[166,273,1101,1123]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="88" type="Museum">ZIN 36613</specimenCode>
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the
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<specimenCode id="DBF94758FFBE3D4CFF7E080BA261FB94" box="[196,303,1129,1151]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="88" type="Museum">ZIN 36614</specimenCode>
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the
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the
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<geoCoordinate id="EE6B89E4FFBE3D4CFDAD08E7A124FB70" box="[535,618,1158,1179]" degrees="42" direction="north" minutes="38" orientation="latitude" pageId="3" pageNumber="88" precision="925" value="42.633335">42°38’N</geoCoordinate>
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,
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<materialsCitation id="3B37E57EFFBE3D4CFF5308C3A2ABFB5C" box="[233,485,1185,1208]" collectionCode="ZIN" country="Japan" elevation="174" isTailEnumeration="true" latitude="42.633335" location="Peter" longLatPrecision="927" longitude="132.72" municipality="Great Bay" pageId="3" pageNumber="88" specimenCount="1">
[depth range:
<quantity id="4CA742C6FFBE3D4CFED408C3A291FB5C" box="[366,479,1185,1207]" metricMagnitude="2" metricUnit="m" metricValue="1.735" metricValueMax="2.4699999999999998" metricValueMin="1.0" pageId="3" pageNumber="88" unit="m" value="173.5" valueMax="247.0" valueMin="100.0">
<elevation id="00720810FFBE3D4CFED408C3A291FB5C" box="[366,479,1185,1207]" metricMagnitude="2" metricUnit="m" metricValue="1.735" metricValueMax="2.4699999999999998" metricValueMin="1.0" pageId="3" pageNumber="88" unit="m" value="173.5" valueMax="247.0" valueMin="100.0">100–247 m</elevation>
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]
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.
</paragraph>
</subSubSection>
<subSubSection id="C345BCA8FFBE3D4CFF7608ADA15FFA8C" pageId="3" pageNumber="88" type="distribution">
<paragraph id="8BE0EF23FFBE3D4CFF7608ADA15FFA8C" blockId="3.[164,743,1228,1641]" pageId="3" pageNumber="88">
<emphasis id="B92B3331FFBE3D4CFF7608ADA214FB0D" bold="true" box="[204,346,1228,1254]" pageId="3" pageNumber="88">Distribution</emphasis>
. Widely distributed in the Northwest Pacific, from the eastern part of the Okhotsk Sea to the southernmost part of the Sea of
<collectingCountry id="F348AFB3FFBE3D4CFDF1096CA1C0FACC" box="[587,654,1293,1319]" name="Japan" pageId="3" pageNumber="88">Japan</collectingCountry>
, and to Sagami Bay along the Pacific coast. Depth range in the Sea of
<collectingCountry id="F348AFB3FFBE3D4CFEA7092CA213FA8C" box="[285,349,1357,1383]" name="Japan" pageId="3" pageNumber="88">Japan</collectingCountry>
is
<quantity id="4CA742C6FFBE3D4CFEC4092CA143FA8C" box="[382,525,1357,1383]" metricMagnitude="2" metricUnit="m" metricValue="5.555" metricValueMax="10.11" metricValueMin="1.0" pageId="3" pageNumber="88" unit="m" value="555.5" valueMax="1011.0" valueMin="100.0">100–1011 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C345BCA8FFBE3D4CFF76090CA27FF982" pageId="3" pageNumber="88" type="discussion">
<paragraph id="8BE0EF23FFBE3D4CFF76090CA27FF982" blockId="3.[164,743,1228,1641]" pageId="3" pageNumber="88">
<emphasis id="B92B3331FFBE3D4CFF76090CA277FA6C" bold="true" box="[204,313,1389,1415]" pageId="3" pageNumber="88">Remarks</emphasis>
. Although this species was considered by
<bibRefCitation id="EFCE92D2FFBE3D4CFF7209EFA2C0FA43" author="Hasegawa K." box="[200,398,1421,1448]" pageId="3" pageNumber="88" pagination="75 - 148" refId="ref5887" refString="Hasegawa K. 2014. A review of bathyal Rissoidae in the Sea of Japan and adjacent waters (Gastropoda: Rissooidea). In: Fujita T. (Ed.). Deep-sea Fauna of the Sea of Japan. National Museum of Nature and Science Monographs, (44): 75 - 148." type="journal article" year="2014">Hasegawa [2014]</bibRefCitation>
to be widely distributed in the North Pacific and its marginal seas, there are considerable differences in the shape and sculpture of the shell between geographic areas, even within the Sea of
<collectingCountry id="F348AFB3FFBE3D4CFF550A6EA27CF9C2" box="[239,306,1551,1577]" name="Japan" pageId="3" pageNumber="88">Japan</collectingCountry>
. More detailed analysis using molecular techniques may reveal genetic and thus taxonomic distinctions.
</paragraph>
</subSubSection>
</treatment>
</document>
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