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159 KiB
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1513 lines
159 KiB
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<document id="C2624DAB8BBD7D3CA4BFC46E7CDDEE3A" ID-DOI="10.5281/zenodo.214038" ID-GBIF-Dataset="3f654113-7579-48f4-8b49-0a3eeac7d6fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="214038" ID-ZooBank="9FC2166A-A0B8-4844-9EF6-8E325AD8F531" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460536329573" checkinUser="plazi" docAuthor="David, Patrick, Nguyen, Truong Quang, Nguyen, Tao Thien, Jiang, Ke, Teynié, Tianbo Chen Alexandre & Ziegler, Thomas" docDate="2012" docId="03B0BD2A1159BB5CFF00FE47FAA72312" docLanguage="en" docName="zt03498p062.pdf" docOrigin="Zootaxa 3498" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Oligodon nagao David, Nguyen, Nguyen, Jiang, Teynié & Ziegler, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="56" masterDocId="FF89C552115BBB57FF97FF9DFFB12025" masterDocTitle="A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from northern Vietnam, southern China and central Laos" masterLastPageNumber="62" masterPageNumber="45" pageNumber="47" updateTime="1698316480961" updateUser="plazi">
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<mods:title id="3761A6F8CD722F170927E0FA6496394F">A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from northern Vietnam, southern China and central Laos</mods:title>
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<mods:namePart id="7ABA017422BCCC8F4B4788E2FB268537">David, Patrick</mods:namePart>
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</mods:name>
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<mods:namePart id="BB60F82D84616082DB5E163809280465">Nguyen, Truong Quang</mods:namePart>
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<mods:namePart id="3B734B3E855F356178CB7461694F4202">Nguyen, Tao Thien</mods:namePart>
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</mods:name>
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<mods:name id="F0B30B4D709E75D0D306C1C21F463977" type="personal">
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<mods:roleTerm id="FCEB4370D7A5579D85CEB265FA646B08">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="77AEE5FA564AEAF1F32F28D1A92B315F">Jiang, Ke</mods:namePart>
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</mods:name>
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</mods:role>
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<mods:namePart id="A0C2663A0F3A2AE49F64BD84C7B84AB3">Teynié, Tianbo Chen Alexandre</mods:namePart>
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<mods:namePart id="C9B0B4B136B25685D17A4C6A00E230E0">Ziegler, Thomas</mods:namePart>
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<mods:typeOfResource id="7CBEDE60A661561A27416CDAC4F23077">text</mods:typeOfResource>
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<mods:title id="4B212E400589AD4FDBAEABDB26136033">Zootaxa</mods:title>
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<mods:part id="7D33AABC70E5C0C6D74192075B747BD1">
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<mods:date id="1C458BAB0668725C69594F59152961F0">2012</mods:date>
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<mods:detail id="EEC66E3335243C2B98032ABAB5149AF2" type="volume">
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<mods:number id="DBC4DE43E7D76C991269BC0A03BD6627">3498</mods:number>
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<mods:start id="70F6C0B515A2D206215B353BCA826F78">45</mods:start>
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<mods:classification id="DDA349EABADF68315533557753E12229">journal article</mods:classification>
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<mods:identifier id="1EEA73EAA8DE36F1191B01FF4092BF28" type="DOI">10.5281/zenodo.214038</mods:identifier>
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<mods:identifier id="AE563A56FB2DEC9C140ABDC0E24735AF" type="GBIF-Dataset">3f654113-7579-48f4-8b49-0a3eeac7d6fa</mods:identifier>
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<mods:identifier id="FB56813CFC9DE130BF385A93B173E5FC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="60814C8247F59C029CB3585EBD9E8F55" type="Zenodo-Dep">214038</mods:identifier>
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<treatment id="03B0BD2A1159BB5CFF00FE47FAA72312" ID-DOI="http://doi.org/10.5281/zenodo.5679604" ID-GBIF-Taxon="119527730" ID-Zenodo-Dep="5679604" LSID="urn:lsid:plazi:treatment:03B0BD2A1159BB5CFF00FE47FAA72312" httpUri="http://treatment.plazi.org/id/03B0BD2A1159BB5CFF00FE47FAA72312" lastPageId="11" lastPageNumber="56" pageId="2" pageNumber="47">
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<subSubSection id="C3035FB71159BB55FF00FE47FEA32230" pageId="2" pageNumber="47" type="nomenclature">
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<paragraph id="8BA60C3C1159BB55FF00FE47FE0F21D1" blockId="2.[151,446,474,533]" box="[151,446,474,500]" pageId="2" pageNumber="47">
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<heading id="D0EEBB501159BB55FF00FE47FE0F21D1" bold="true" box="[151,446,474,500]" fontSize="11" level="1" pageId="2" pageNumber="47" reason="1">
|
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<emphasis id="B96DD02E1159BB55FF00FE47FE0F21D1" bold="true" box="[151,446,474,500]" pageId="2" pageNumber="47">
|
|||
|
|
<taxonomicName id="4C1977BF1159BB55FF00FE47FEE821D1" box="[151,345,474,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
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|
<emphasis id="B96DD02E1159BB55FF00FE47FEE821D1" bold="true" box="[151,345,474,500]" italics="true" pageId="2" pageNumber="47">Oligodon nagao</emphasis>
|
|||
|
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</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551159BB55FEF6FE47FE0F21D1" box="[353,446,474,500]" pageId="2" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
|
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</emphasis>
|
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</heading>
|
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</paragraph>
|
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|
<paragraph id="8BA60C3C1159BB55FF00FE60FEA32230" blockId="2.[151,446,474,533]" box="[151,274,509,533]" pageId="2" pageNumber="47">
|
|||
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(
|
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<figureCitation id="132210B91159BB55FF08FE60FEB92230" box="[159,264,509,533]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,249,1942,1965]" captionStartId-1="3.[151,249,995,1018]" captionStartId-2="4.[151,249,1440,1463]" captionStartId-3="5.[151,249,1981,2004]" captionStartId-4="7.[151,249,1897,1920]" captionTargetBox-0="[163,1423,1412,1920]" captionTargetBox-1="[163,1436,193,973]" captionTargetBox-2="[151,1435,553,1418]" captionTargetBox-3="[202,1370,193,1937]" captionTargetBox-4="[151,1435,945,1872]" captionTargetId-0="figure@2.[163,1424,1412,1921]" captionTargetId-1="figure@3.[151,1436,193,974]" captionTargetId-2="figure@4.[151,1436,553,1419]" captionTargetId-3="figure@5.[202,1370,193,1938]" captionTargetId-4="figure@7.[151,1436,945,1873]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="7" captionText-0="FIGURE 1. Holotype of Oligodon nagao sp. nov. (VNMN A. 2012.1, adult male): A. Dorsal view. B. Ventral view. Photographs by Patrick David." captionText-1="FIGURE 2. Holotype of Oligodon nagao sp. nov. (VNMN A. 2012.1, adult male), views of the head: A. Dorsal side. B. Ventral side. C. Left side. D. Right side. Photographs by Truong Quang Nguyen." captionText-2="FIGURE 3. Oligodon nagao sp. nov., in life (IEBR A. 2012.6, adult male, Vietnam). Photograph by Truong Quang Nguyen." captionText-3="FIGURE 4. Oligodon nagao sp. nov., in life (ZFMK 93281, adult male, Laos). A. General view. B. Specimen showing its threatening behaviour. Photographs by Alexandre Teynié." captionText-4="FIGURE 5. Biotope of Oligodon nagao sp. nov. in Khammouane Province, Laos. Photographs by Alexandre Teynié." httpUri-0="https://zenodo.org/record/214039/files/figure.png" httpUri-1="https://zenodo.org/record/214040/files/figure.png" httpUri-2="https://zenodo.org/record/214041/files/figure.png" httpUri-3="https://zenodo.org/record/214042/files/figure.png" httpUri-4="https://zenodo.org/record/214043/files/figure.png" pageId="2" pageNumber="47">Figs. 1–5</figureCitation>
|
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)
|
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|
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</paragraph>
|
|||
|
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</subSubSection>
|
|||
|
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<subSubSection id="C3035FB71159BB55FF00FDD8FC41225C" pageId="2" pageNumber="47" type="reference_group">
|
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|
|
<paragraph id="8BA60C3C1159BB55FF00FDD8FC41225C" blockId="2.[151,1436,581,633]" pageId="2" pageNumber="47">
|
|||
|
|
<treatmentCitationGroup id="AB092B121159BB55FF00FDD8FC41225C" pageId="2" pageNumber="47">
|
|||
|
|
<taxonomicName id="4C1977BF1159BB55FF00FDD8FEE7227E" box="[151,342,581,603]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1159BB55FF00FDD8FEE7227E" box="[151,342,581,603]" italics="true" pageId="2" pageNumber="47">Oligodon joynsoni</emphasis>
|
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|
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</taxonomicName>
|
|||
|
|
(nec
|
|||
|
|
<taxonomicName id="4C1977BF1159BB55FE02FDDBFCFA227F" authority="Smith, 1917" authorityName="Smith" authorityYear="1917" box="[405,843,581,603]" class="Squamata" family="Colubridae" genus="Simotes" higherTaxonomySource="CoL" kingdom="Animalia" pageId="2" pageNumber="47" phylum="Chordata" rank="subSpecies" species="longicauda" subSpecies="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1159BB55FE02FDDBFD71227E" box="[405,704,581,603]" italics="true" pageId="2" pageNumber="47">Simotes longicauda joynsoni</emphasis>
|
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|
Smith, 1917
|
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|
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</taxonomicName>
|
|||
|
|
, a valid species, now
|
|||
|
|
<taxonomicName id="4C1977BF1159BB55FBAEFDD8FB49227E" box="[1081,1272,581,603]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1159BB55FBAEFDD8FB49227E" box="[1081,1272,581,603]" italics="true" pageId="2" pageNumber="47">Oligodon joynsoni</emphasis>
|
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</taxonomicName>
|
|||
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).—Anonymous 2002: 22, front cover.—
|
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|
|
<treatmentCitation id="0AB82A2D1159BB55FE2DFDFEFDEA225C" author="Zhao" box="[442,603,611,633]" page="229" pageId="2" pageNumber="47" year="2006">
|
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|
|
<bibRefCitation id="EF8871CD1159BB55FE2DFDFEFD9A225C" author="Zhao" box="[442,555,611,633]" pageId="2" pageNumber="47" refString="Zhao, E. (2006) Zhongguo Shelei [Snakes of China]. Anhui Science and Technology Publishing House Publ., Hefei, Vol. I, 372 pp.; Vol. II, 279 pp. (in Chinese)." type="book" year="2006">Zhao 2006</bibRefCitation>
|
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: 229
|
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</treatmentCitation>
|
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|
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.—
|
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<treatmentCitation id="0AB82A2D1159BB55FDEEFDFEFCAF225C" author="Zhang" box="[633,798,611,633]" page="97" pageId="2" pageNumber="47" year="2009">
|
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|
|
<bibRefCitation id="EF8871CD1159BB55FDEEFDFEFD49225C" author="Zhang" box="[633,760,611,633]" pageId="2" pageNumber="47" refString="Zhang, Y. (2009) Herpetology in Guangxi. Guangxi Biodiversity Studies, Guangxi Normal University Press, Guilin, 1 - 8 (col. plates), (2 unnumbered pages), 170 pp." type="book" year="2009">Zhang 2009</bibRefCitation>
|
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: 97
|
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</treatmentCitation>
|
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, 99 & 100: Fig. 100.
|
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</treatmentCitationGroup>
|
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</paragraph>
|
|||
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</subSubSection>
|
|||
|
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<subSubSection id="C3035FB71159BB55FF00FD3AFE942409" pageId="2" pageNumber="47" type="materials_examined">
|
|||
|
|
<paragraph id="8BA60C3C1159BB55FF00FD3AFB9122C7" blockId="2.[151,1436,678,1359]" pageId="2" pageNumber="47">
|
|||
|
|
<typeStatus id="54A2B29E1159BB55FF00FD3AFEB5229A" box="[151,260,679,703]" pageId="2" pageNumber="47" type="holotype">
|
|||
|
|
<emphasis id="B96DD02E1159BB55FF00FD3AFEB5229A" bold="true" box="[151,260,679,703]" pageId="2" pageNumber="47">Holotype</emphasis>
|
|||
|
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</typeStatus>
|
|||
|
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.
|
|||
|
|
<collectionCode id="ED0894F91159BB55FE85FD3AFED9229B" box="[274,360,679,702]" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="2" pageNumber="47">VNMN</collectionCode>
|
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A.2012.1, adult male, from Huu Lien forest, Huu Lung District, Lang Son Province,
|
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|
|
<collectingCountry id="F30E4CAC1159BB55FA8CFD3BFACC229B" box="[1307,1405,678,702]" name="Vietnam" pageId="2" pageNumber="47">Vietnam</collectingCountry>
|
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|
|
, at elevation of about
|
|||
|
|
<quantity id="4CE1A1D91159BB55FEFFFD57FE1E22C7" box="[360,431,714,738]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="47" unit="m" value="300.0">300 m</quantity>
|
|||
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|
a.s.l.; collected by Tao Thien Nguyen on
|
|||
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|
<date id="FFA72AFC1159BB55FC14FD57FBAA22C7" box="[899,1051,714,738]" pageId="2" pageNumber="47" value="2009-06-30">30 June 2009</date>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1159BB55FF52FD72FE942409" blockId="2.[151,1436,678,1359]" pageId="2" pageNumber="47">
|
|||
|
|
<typeStatus id="54A2B29E1159BB55FF52FD72FE8C2322" box="[197,317,751,775]" pageId="2" pageNumber="47" type="paratype">
|
|||
|
|
<emphasis id="B96DD02E1159BB55FF52FD72FE8C2322" bold="true" box="[197,317,751,775]" pageId="2" pageNumber="47">Paratypes</emphasis>
|
|||
|
|
</typeStatus>
|
|||
|
|
. Four specimens, all adult males:
|
|||
|
|
<collectionCode id="ED0894F91159BB55FD58FD72FC942323" LSID="urn:lsid:biocol.org:col:34988" box="[719,805,751,774]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="47">MNHN</collectionCode>
|
|||
|
|
2012.0216, from Huu Lien forest, Huu Lung District, Lang Son Province,
|
|||
|
|
<collectingCountry id="F30E4CAC1159BB55FE03FC8FFE47230F" box="[404,502,786,810]" name="Vietnam" pageId="2" pageNumber="47">Vietnam</collectingCountry>
|
|||
|
|
, at elevation of about
|
|||
|
|
<quantity id="4CE1A1D91159BB55FC83FC8FFCD2230F" box="[788,867,786,810]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="47" unit="m" value="300.0">300 m</quantity>
|
|||
|
|
a.s.l.; collected by Tao Thien Nguyen,
|
|||
|
|
<date id="FFA72AFC1159BB55FAC6FC8FFF63236B" pageId="2" pageNumber="47" value="2009-07-02">2 July 2009</date>
|
|||
|
|
.—
|
|||
|
|
<collectionCode id="ED0894F91159BB55FF65FCABFE82236B" box="[242,307,822,846]" pageId="2" pageNumber="47">IEBR</collectionCode>
|
|||
|
|
A.2012.6, from
|
|||
|
|
<collectingCountry id="F30E4CAC1159BB55FE79FCAAFDAC236B" box="[494,541,823,846]" name="Germany" pageId="2" pageNumber="47">Duc</collectingCountry>
|
|||
|
|
Quang Commune, Ha Lang District, Cao Bang Province,
|
|||
|
|
<collectingCountry id="F30E4CAC1159BB55FB30FCABFABB236B" box="[1191,1290,822,846]" name="Vietnam" pageId="2" pageNumber="47">Vietnam</collectingCountry>
|
|||
|
|
; collected by Truong Quang Nguyen
|
|||
|
|
<emphasis id="B96DD02E1159BB55FE39FCC3FE5A2350" box="[430,491,861,885]" italics="true" pageId="2" pageNumber="47">et al.</emphasis>
|
|||
|
|
,
|
|||
|
|
<date id="FFA72AFC1159BB55FE6BFCC0FD0E2350" box="[508,703,861,885]" pageId="2" pageNumber="47" value="2011-10-14">14 October 2011</date>
|
|||
|
|
(
|
|||
|
|
<geoCoordinate id="EE2D6AFB1159BB55FD45FCC0FCD42350" box="[722,869,858,885]" direction="north" orientation="latitude" pageId="2" pageNumber="47" precision="1" value="22.714317">22o42.859’N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE2D6AFB1159BB55FCE3FCC0FBBE2350" box="[884,1039,858,885]" direction="east" orientation="longitude" pageId="2" pageNumber="47" precision="1" value="106.664215">106o39.853’E</geoCoordinate>
|
|||
|
|
, elevation
|
|||
|
|
<quantity id="4CE1A1D91159BB55FB07FCC0FB6D2350" box="[1168,1244,861,885]" metricMagnitude="2" metricUnit="m" metricValue="4.76" pageId="2" pageNumber="47" unit="m" value="476.0">476 m</quantity>
|
|||
|
|
).—
|
|||
|
|
<collectionCode id="ED0894F91159BB55FA92FCC0FA872350" box="[1285,1334,861,885]" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="2" pageNumber="47">KIZ</collectionCode>
|
|||
|
|
0 14591, from Nonggang National Nature Reserve, Longzhou County, Guangxi Autonomous Region,
|
|||
|
|
<collectingCountry id="F30E4CAC1159BB55FB26FC1CFF512398" name="China" pageId="2" pageNumber="47">People’s Republic of China</collectingCountry>
|
|||
|
|
; collected by Tianbo Chen, October, 2011.—
|
|||
|
|
<collectionCode id="ED0894F91159BB55FC99FC38FCEE2398" LSID="urn:lsid:biocol.org:col:34613" box="[782,863,933,957]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="2" pageNumber="47">ZFMK</collectionCode>
|
|||
|
|
93281, from Ban Nathan, Hin Boun District, Khammouane Province,
|
|||
|
|
<collectingCountry id="F30E4CAC1159BB55FE38FC50FE5723C1" box="[431,486,973,996]" name="Laos" pageId="2" pageNumber="47">Laos</collectingCountry>
|
|||
|
|
PDR (
|
|||
|
|
<geoCoordinate id="EE2D6AFB1159BB55FDA4FC51FD7523C1" box="[563,708,969,996]" direction="north" orientation="latitude" pageId="2" pageNumber="47" precision="1" value="17.9809">17o58.854’N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE2D6AFB1159BB55FD47FC51FCDA23C1" box="[720,875,969,996]" direction="east" orientation="longitude" pageId="2" pageNumber="47" precision="1" value="104.82529">104o49.517’E</geoCoordinate>
|
|||
|
|
, elevation
|
|||
|
|
<quantity id="4CE1A1D91159BB55FC72FC51FB8123C1" box="[997,1072,972,996]" metricMagnitude="2" metricUnit="m" metricValue="1.72" pageId="2" pageNumber="47" unit="m" value="172.0">172 m</quantity>
|
|||
|
|
); collected by Alexandre Teynié
|
|||
|
|
<emphasis id="B96DD02E1159BB55FF0EFC6CFF65242D" box="[153,212,1008,1032]" italics="true" pageId="2" pageNumber="47">et al.</emphasis>
|
|||
|
|
,
|
|||
|
|
<date id="FFA72AFC1159BB55FF72FC6DFE35242D" box="[229,388,1008,1033]" pageId="2" pageNumber="47" value="2012-05-17">17 May 2012</date>
|
|||
|
|
, during the mission “
|
|||
|
|
<emphasis id="B96DD02E1159BB55FDE8FC6CFEA32409" italics="true" pageId="2" pageNumber="47">
|
|||
|
|
Opération Canopée, Inventaire de la Biodiversité Forestière du
|
|||
|
|
<collectingCountry id="F30E4CAC1159BB55FAF1FC6CFA2D242D" box="[1382,1436,1009,1032]" name="Laos" pageId="2" pageNumber="47">Laos</collectingCountry>
|
|||
|
|
<date id="FFA72AFC1159BB55FF0EFB89FEA32409" box="[153,274,1044,1068]" pageId="2" pageNumber="47" value="2012" valueMax="2015">2012-2015</date>
|
|||
|
|
</emphasis>
|
|||
|
|
”.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3035FB71159BB55FF52FBAAFB14256A" pageId="2" pageNumber="47" type="diagnosis">
|
|||
|
|
<paragraph id="8BA60C3C1159BB55FF52FBAAFB14256A" blockId="2.[151,1436,678,1359]" pageId="2" pageNumber="47">
|
|||
|
|
<emphasis id="B96DD02E1159BB55FF52FBAAFE892475" bold="true" box="[197,312,1079,1104]" pageId="2" pageNumber="47">Diagnosis</emphasis>
|
|||
|
|
. A large species (TL up to at least
|
|||
|
|
<quantity id="4CE1A1D91159BB55FD2AFBA5FCAB2475" box="[701,794,1080,1104]" metricMagnitude="-1" metricUnit="m" metricValue="7.859999999999999" pageId="2" pageNumber="47" unit="mm" value="786.0">786 mm</quantity>
|
|||
|
|
) of the genus
|
|||
|
|
<taxonomicName id="4C1977BF1159BB55FC2FFBA5FB932475" box="[952,1058,1080,1104]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="47" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B96DD02E1159BB55FC2FFBA5FB932475" box="[952,1058,1080,1104]" italics="true" pageId="2" pageNumber="47">Oligodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
characterized by the combination of (1) 9 or 10 maxillary teeth, the last three or four strongly enlarged, (2) hemipenes not forked but divided into two lobes, thick and bulbous, reaching in situ the 16th SC, smooth, each lobe with a papilla, (3) 17–17–15 (or 17–15–15 DSR in one specimen); (4) cloacal plate entire, (5) complete complement of head scales, including 1 loreal on each side, (6) 8 supralabials (
|
|||
|
|
<quantity id="4CE1A1D91159BB55FE3DFB56FE6624C6" box="[426,471,1227,1251]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="2" pageNumber="47" unit="in" value="7.0">7 in</quantity>
|
|||
|
|
one specimen), fourth and fifth (third and fourth in one specimen) entering orbit, (7) 184–193 ventrals in five males (females unknown), (8) tail relatively short in males (ratio TaL/TL: 0.135–0.146), (9) dorsal pattern made of a dark background colour with 27–37 darker, pale centered, butterfly-shaped blotches on the body and 5–8 on the tail, and (10) venter cream, heavily marked with dark pigmentation.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="DF665CB41159BB55FF00F80BFE7F27EE" httpUri="https://zenodo.org/record/214039/files/figure.png" pageId="2" pageNumber="47" targetBox="[163,1423,1412,1920]" targetPageId="2">
|
|||
|
|
<paragraph id="8BA60C3C1159BB55FF00F80BFE7F27EE" blockId="2.[151,1437,1942,1995]" pageId="2" pageNumber="47">
|
|||
|
|
<emphasis id="B96DD02E1159BB55FF00F80BFEA82789" bold="true" box="[151,281,1942,1965]" pageId="2" pageNumber="47">FIGURE 1.</emphasis>
|
|||
|
|
Holotype of
|
|||
|
|
<taxonomicName id="4C1977BF1159BB55FE24F80AFDEE2788" box="[435,607,1943,1965]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1159BB55FE24F80AFDEE2788" box="[435,607,1943,1965]" italics="true" pageId="2" pageNumber="47">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B96DD02E1159BB55FDFBF80AFD722789" bold="true" box="[620,707,1943,1964]" pageId="2" pageNumber="47">
|
|||
|
|
<taxonomicNameLabel id="A25E6D551159BB55FDFBF80AFD722789" box="[620,707,1943,1964]" pageId="2" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(VNMN A.2012.1, adult male): A. Dorsal view. B. Ventral view. Photographs by Patrick David.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF665CB41158BB54FF00FC7EFCCB243D" httpUri="https://zenodo.org/record/214040/files/figure.png" pageId="3" pageNumber="48" targetBox="[163,1436,193,973]" targetPageId="3">
|
|||
|
|
<paragraph id="8BA60C3C1158BB54FF00FC7EFCCB243D" blockId="3.[151,1436,995,1048]" pageId="3" pageNumber="48">
|
|||
|
|
<emphasis id="B96DD02E1158BB54FF00FC7EFEA323DC" bold="true" box="[151,274,995,1018]" pageId="3" pageNumber="48">FIGURE 2.</emphasis>
|
|||
|
|
Holotype of
|
|||
|
|
<taxonomicName id="4C1977BF1158BB54FE01FC79FD8A23DF" box="[406,571,996,1018]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="48" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1158BB54FE01FC79FD8A23DF" box="[406,571,996,1018]" italics="true" pageId="3" pageNumber="48">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551158BB54FDD6FC79FD2023DC" box="[577,657,996,1017]" pageId="3" pageNumber="48" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1158BB54FDD6FC79FD2023DC" bold="true" box="[577,657,996,1017]" pageId="3" pageNumber="48">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
(VNMN A.2012.1, adult male), views of the head: A. Dorsal side. B. Ventral side. C. Left side. D. Right side. Photographs by Truong Quang Nguyen.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C3035FB71158BB5FFF52FBD8FAAB21D1" lastPageId="8" lastPageNumber="53" pageId="3" pageNumber="48" type="description">
|
|||
|
|
<paragraph id="8BA60C3C1158BB54FF52FBD8FC4424C8" blockId="3.[151,1436,1093,1588]" pageId="3" pageNumber="48">
|
|||
|
|
<emphasis id="B96DD02E1158BB54FF52FBD8FE522478" bold="true" box="[197,483,1093,1118]" pageId="3" pageNumber="48">
|
|||
|
|
Description of
|
|||
|
|
<typeStatus id="54A2B29E1158BB54FEEAFBD8FE522478" box="[381,483,1093,1117]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
. Body elongate, cylindrical and robust; head short (3.7 % of SVL), ovoid, rather broad and distinct from thick neck, thick but depressed anteriorly; snout long, narrowing anteriorly, slightly rounded, extending well beyond lower jaw, about 1.9 times as long as eye diameter; large, oval nostril piercing laterally the central part of nasal; eye rather small, its diameter about 0.9 times the distance between eye and lip, round pupil; tail average, robust at its base, tapering progressively to a point.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1158BB54FF52FB67FA2D2534" blockId="3.[151,1436,1093,1588]" box="[197,1436,1273,1297]" pageId="3" pageNumber="48">
|
|||
|
|
<emphasis id="B96DD02E1158BB54FF52FB67FED62534" box="[197,359,1274,1297]" italics="true" pageId="3" pageNumber="48">Measurements</emphasis>
|
|||
|
|
.SVL:
|
|||
|
|
<quantity id="4CE1A1D91158BB54FE25FB64FDA32534" box="[434,530,1273,1297]" metricMagnitude="-1" metricUnit="m" metricValue="5.61" pageId="3" pageNumber="48" unit="mm" value="561.0">561 mm</quantity>
|
|||
|
|
; TaL:
|
|||
|
|
<quantity id="4CE1A1D91158BB54FDCFFB64FD1A2534" box="[600,683,1273,1297]" metricMagnitude="-2" metricUnit="m" metricValue="9.2" pageId="3" pageNumber="48" unit="mm" value="92.0">92 mm</quantity>
|
|||
|
|
; TL:
|
|||
|
|
<quantity id="4CE1A1D91158BB54FD70FB64FCF62534" box="[743,839,1273,1297]" metricMagnitude="-1" metricUnit="m" metricValue="6.529999999999999" pageId="3" pageNumber="48" unit="mm" value="653.0">653 mm</quantity>
|
|||
|
|
; ratio TaL/TL: 0.141; HL:
|
|||
|
|
<quantity id="4CE1A1D91158BB54FBE4FB64FB592534" box="[1139,1256,1273,1297]" metricMagnitude="-2" metricUnit="m" metricValue="2.0949999999999998" pageId="3" pageNumber="48" unit="mm" value="20.95">20.95 mm</quantity>
|
|||
|
|
; SnL:
|
|||
|
|
<quantity id="4CE1A1D91158BB54FAA6FB64FA292534" box="[1329,1432,1273,1297]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" pageId="3" pageNumber="48" unit="mm" value="6.35">6.35 mm</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1158BB54FF52FA83FDAD257C" blockId="3.[151,1436,1093,1588]" pageId="3" pageNumber="48">
|
|||
|
|
<emphasis id="B96DD02E1158BB54FF52FA83FE9E2510" box="[197,303,1310,1333]" italics="true" pageId="3" pageNumber="48">Dentition</emphasis>
|
|||
|
|
. Maxillary teeth: left maxilla with 9 teeth under the formula: 6 subequal teeth + 3 strongly enlarged, blade-like teeth, without diastema.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1158BB54FF52FAF4FF0F25ED" blockId="3.[151,1436,1093,1588]" pageId="3" pageNumber="48">
|
|||
|
|
<emphasis id="B96DD02E1158BB54FF52FAF4FE8C25A5" box="[197,317,1385,1408]" italics="true" pageId="3" pageNumber="48">Hemipenis</emphasis>
|
|||
|
|
.In situ, the hemipenis is massive, unforked, with simple sulcus, and reaches the 16th SC; first basal quarter of the hemipenis covered with calyces; no spines; a long papilla each, about 5 SC long, emerging from the tip.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1158BB54FF52FA49FD172611" blockId="3.[151,1436,1093,1588]" pageId="3" pageNumber="48">
|
|||
|
|
<emphasis id="B96DD02E1158BB54FF52FA49FEDD25C9" box="[197,364,1492,1516]" italics="true" pageId="3" pageNumber="48">Body scalation</emphasis>
|
|||
|
|
. DSR: 17–17–15, all smooth; scales of the outer dorsal scale row distinctly enlarged. 189 VEN (+ 1 preventral), strongly angulated; 45 SC, all paired; cloacal entire; terminal caudal scale pointing. The dorsal scale row reductions are as follows:
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1158BB54FF6FF9E8FDA126A9" blockId="3.[185,571,1653,1749]" box="[248,528,1653,1676]" pageId="3" pageNumber="48">5+6 ↔ 5 (VEN 115) (left)</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1158BB54FF2EF905FD8A268A" blockId="3.[185,571,1653,1749]" box="[185,571,1688,1711]" pageId="3" pageNumber="48">17 ————————— 15</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1158BB54FF66F923FDA926F0" blockId="3.[185,571,1653,1749]" box="[241,536,1726,1749]" pageId="3" pageNumber="48">5+6 ↔ 5 (VEN 119) (right)</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1158BB53FF50F88FFB1F21D1" blockId="3.[151,1436,1809,2013]" lastBlockId="4.[151,1436,151,500]" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
|
|||
|
|
<emphasis id="B96DD02E1158BB54FF50F88FFEC0270F" box="[199,369,1810,1834]" italics="true" pageId="3" pageNumber="48">Head scalation</emphasis>
|
|||
|
|
. Head scalation complement complete, including 2 internasals, 2 prefrontals, 2 supraoculars, 1 frontal, and 2 parietals. Rostral large, wider than high, well visible from above, inserting deeply and broadly between internasals on about one half of their total length; 1 / 1 large, elongate, pentagonal nasal, 1.9 times as long as high, entire, most of scale area being occupied by the nostril; internasals subrectangular, narrow, separated by a short suture, much wider than long, 0.7 times as long as the suture between prefrontals and only 0.4 times as long as prefrontal; prefrontals large, pentagonal, much wider than long, 0.5 times as long as frontal but separated by a suture only 0.25 times as long as frontal; supraoculars subrectangular, relatively large, about 1.6 times as long as broad, 0.4 times as wide as frontal; frontal large, hexagonal, wide, rather squat, 1.2 times longer than wide; parietals moderate, longer than wide, extending on about 30 % of HL, about 1.05 times longer than frontal, abruptly truncated posteriorly with a straight posterior margin; no nuchal scale behind parietal; 1 / 1 small loreal scale, subrectangular, 1.4 times longer than high; 8 / 8 SL, first and second in contact with nasal, second and third in contact with loreal, fourth and fifth entering orbit, sixth and seventh SL largest; 1 / 1 narrow preocular, in contact with prefrontal but not reaching the frontal; 1 / 1 minute presubocular; 2 / 2 small, narrow postoculars, lowest one slightly largest; 1 + 3 temporals on each side, anterior one large, long and high, rather squat, posterior ones rather small; 8 / 8
|
|||
|
|
<collectingCountry id="F30E4CAC115FBB53FEB5FE25FE8E21EA" box="[290,319,440,463]" name="Iceland" pageId="4" pageNumber="49">IL</collectingCountry>
|
|||
|
|
, first in contact with each other, first to fifth in contact with anterior chin shields (fifth in contact only punctually); mental small; anterior chin shields 2.2 times as long as short posterior ones.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF665CB4115FBB53FF00FA3DFA352593" box="[151,1412,1440,1463]" httpUri="https://zenodo.org/record/214041/files/figure.png" pageId="4" pageNumber="49" targetBox="[151,1435,553,1418]" targetPageId="4">
|
|||
|
|
<paragraph id="8BA60C3C115FBB53FF00FA3DFA352593" blockId="4.[151,1412,1440,1463]" box="[151,1412,1440,1463]" pageId="4" pageNumber="49">
|
|||
|
|
<emphasis id="B96DD02E115FBB53FF00FA3DFEA32593" bold="true" box="[151,274,1440,1463]" pageId="4" pageNumber="49">FIGURE 3.</emphasis>
|
|||
|
|
<taxonomicName id="4C1977BF115FBB53FE8FFA3DFE0C2592" box="[280,445,1440,1463]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="49" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E115FBB53FE8FFA3DFE0C2592" box="[280,445,1440,1463]" italics="true" pageId="4" pageNumber="49">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D55115FBB53FE53FA3CFDA52593" box="[452,532,1441,1462]" pageId="4" pageNumber="49" rank="species">
|
|||
|
|
<emphasis id="B96DD02E115FBB53FE53FA3CFDA52593" bold="true" box="[452,532,1441,1462]" pageId="4" pageNumber="49">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
, in life (IEBR A.2012.6, adult male, Vietnam). Photograph by Truong Quang Nguyen.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA60C3C115FBB53FF52FA43FABA27F0" blockId="4.[153,1436,1502,2005]" pageId="4" pageNumber="49">
|
|||
|
|
<emphasis id="B96DD02E115FBB53FF52FA43FE1025D3" box="[197,417,1502,1526]" italics="true" pageId="4" pageNumber="49">Colour and pattern</emphasis>
|
|||
|
|
. Body is dark greyish-brown, somewhat darker or more grey on the lower sides, with all scales finely but densely dotted with blackish-brown; outer margins of scales of the vertebral row and upper margins of each adjacent row beige or pale greyish-brown, and central part of scales of the vertebral row densely dotted with brown, producing an irregular paler brown vertebral stripe more visible on the neck; a series of 27 blackish-brown dorsal blotches, somewhat paler greyish-brown in their centre, edged by darker pigmentation, about 3 DSR long and a total of 5 to 7 DSR broad, i.e. covering the vertebral scale row and the 2 or 3 adjacent DSR (DSR 6th or 7th); first blotch located at 13 scales behind parietals; first 11 blotches butterfly-shaped, bisected along the vertebral line although in narrow contact, posterior ones rather “bat-like”; on each side, a faint, irregular dark blotch on each side of the paler vertebral on 7th and 8th DSR in between each dorsal blotch; an elongate, irregularly shaped lateral, blackish-brown blotch on 3rd and 4th and sometimes also on the lower edge of 5th DSR, just below each dorsal blotch, about 1 or 1.5 DSR long. The tail is as the body, with a much better defined vertebral stripe due to the pale greyish-brown colour of the two upper tail rows; 8 blackish-brown dorsal blotches, somewhat paler in their centre, edged with darker pigmentation, progressively more bisected towards the end of the tail, no blotch on the lateral side of the tail; 1st scale row of the tail and lower half of 2nd row uniformly blackish-brown.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF665CB4115EBB52FF00F820FD6E27D7" httpUri="https://zenodo.org/record/214042/files/figure.png" pageId="5" pageNumber="50" targetBox="[202,1370,193,1937]" targetPageId="5">
|
|||
|
|
<paragraph id="8BA60C3C115EBB52FF00F820FD6E27D7" blockId="5.[151,1436,1981,2034]" pageId="5" pageNumber="50">
|
|||
|
|
<emphasis id="B96DD02E115EBB52FF00F820FEA527F6" bold="true" box="[151,276,1981,2004]" pageId="5" pageNumber="50">FIGURE 4.</emphasis>
|
|||
|
|
<taxonomicName id="4C1977BF115EBB52FE8AF823FE7527F1" box="[285,452,1982,2004]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="50" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E115EBB52FE8AF823FE7527F1" box="[285,452,1982,2004]" italics="true" pageId="5" pageNumber="50">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D55115EBB52FE5AF823FD9127F6" box="[461,544,1982,2003]" pageId="5" pageNumber="50" rank="species">
|
|||
|
|
<emphasis id="B96DD02E115EBB52FE5AF823FD9127F6" bold="true" box="[461,544,1982,2003]" pageId="5" pageNumber="50">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
, in life (ZFMK 93281, adult male, Laos). A. General view. B. Specimen showing its threatening behaviour. Photographs by Alexandre Teynié.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA60C3C115DBB51FF52FF0AFCB3224E" blockId="6.[153,1437,151,1965]" pageId="6" pageNumber="51">The head is greyish-brown as the body, much paler on the sides of the snout and rostral; a dark brown crossband across the anterior part of the head extends on internasals, prefrontals and the anterior part of the frontal, then obliquely downwards on each side through the eyes down to 5th–6th SL; a strong diagonal dark brown streak, extends obliquely downwards from the parietals across the temporal region down to the corner of the mouth, then onto the lower side of the neck, where it borders the 3rd to 5th ventral; a rounded dark brown blotch in the middle of frontal; another rounded dark brown blotch on the distal part of frontal and anterior part of parietals; a large and conspicuous dark brown chevron on the occiput, its apex reaching forward the posterior central part of parietals, and its branches extending straight posteriorly on 6th, 7th and lower half of 8th DSR of the neck, each branch contrasting sharply with the pale, brown vertebral stripe on the neck; on each side of the neck, an oblique, elongate dark brown streak in short contact with each branch of the nuchal chevron; nasal, loreal, outer margins of prefrontals and supralabials pale yellow with irregular greyish-brown spots; 5th and 6th SL, and anterior part of 7th SL with an oblique dark brown streak. The chin and throat are creamish-yellow with faint greyish-brown spots on infralabials; one such spot on each anterior chin shield.</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115DBB51FF52FDEAFE9023AE" blockId="6.[153,1437,151,1965]" pageId="6" pageNumber="51">The venter is pale yellow, uniform on the first 6 ventrals, then with rectangular dark greyish-brown blotches irregularly arranged on the outer part of each or one out of two ventrals, then on each ventral; the blotches become progressively wider and, after midbody, the dark pigmentation covers much of ventrals, in leaving only narrow streaks in the middle of each ventral; tips of ventrals dark greyish-brown but separated throughout the venter from the greyish-brown blotches by a narrow, yellow streak which produces on each side a discontinuous ventrolateral stripe. The ventral surface of the tail is pale yellow, with dark brown rectangular blotches on the outer parts of subcaudals; these blotches disappear progressively and the ventral surface of the last quarter of the tail is uniformly pale yellow.</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115DBB51FF52FC0BFE48241A" blockId="6.[153,1437,151,1965]" pageId="6" pageNumber="51">
|
|||
|
|
<emphasis id="B96DD02E115DBB51FF52FC0BFE82238A" bold="true" box="[197,307,918,943]" pageId="6" pageNumber="51">Variation</emphasis>
|
|||
|
|
. Besides the
|
|||
|
|
<typeStatus id="54A2B29E115DBB51FE5DFC0AFD9E238A" box="[458,559,919,943]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
|
|||
|
|
, four other males are known. The main morphological characters of all known specimens are summarized in
|
|||
|
|
<tableCitation id="C69B3987115DBB51FE7EFC26FDF723F6" box="[489,582,955,979]" captionStart="TABLE 1" captionStartId="7.[151,235,152,173]" captionText="TABLE 1. Main variation in the type series of Oligodon nagao sp. nov. Part 1: body and tail scalation." pageId="6" pageNumber="51">Tables 1</tableCitation>
|
|||
|
|
and
|
|||
|
|
<tableCitation id="C69B3987115DBB51FDEBFC26FD3C23F6" box="[636,653,955,979]" captionStart="TABLE 2" captionStartId="7.[151,235,560,581]" captionText="TABLE 2. Main variation in the type series of Oligodon nagao sp. nov. Part 2: head scalation. See Materials and Methods for abbreviations." pageId="6" pageNumber="51">2</tableCitation>
|
|||
|
|
. The great homogeneity in variation is noteworthy, at the exception of the number of supralabials. All other external morphological characters, either in morphometry or scalation, agree with those described for the
|
|||
|
|
<typeStatus id="54A2B29E115DBB51FE4EFB9EFD8F243E" box="[473,574,1027,1051]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
|
|||
|
|
, including the single cloacal, 1 loreal and 1 small presubocular on each side. Other variations are as follows:
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115DBB51FF52FBD6FBEE24F2" blockId="6.[153,1437,151,1965]" pageId="6" pageNumber="51">
|
|||
|
|
<emphasis id="B96DD02E115DBB51FF52FBD6FE3E2446" box="[197,399,1099,1123]" italics="true" pageId="6" pageNumber="51">Dorsal scale rows</emphasis>
|
|||
|
|
. Four of the five specimens have 17 DSR at midbody at the exception of MNHN 2012.0216. In this latter specimen, the first reduction (17 ® 16 DSR) occurs at the level of 91st VEN; the second reduction (16 ® 15 DSR) occurs at the level of the 93rd VEN, so just before midbody, here considered to be located at the level of the 94th VEN. We consider this value of 15 DSR exactly at midbody to be an anomaly.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115DBB51FF52FB79FC752566" blockId="6.[153,1437,151,1965]" pageId="6" pageNumber="51">
|
|||
|
|
<emphasis id="B96DD02E115DBB51FF52FB79FEC224D9" box="[197,371,1252,1276]" italics="true" pageId="6" pageNumber="51">Head scalation</emphasis>
|
|||
|
|
. Four of the five available specimens have 8 supralabials on each side. The specimen from
|
|||
|
|
<collectingCountry id="F30E4CAC115DBB51FF0EFA95FF63253A" box="[153,210,1288,1311]" name="Laos" pageId="6" pageNumber="51">Laos</collectingCountry>
|
|||
|
|
, ZFMK 93281 has 7 / 7 supralabials, third and fourth entering orbit. All specimens have 8 infralabials on each side at the exception of specimen KIZ 0 14591 which has 7 scales at left.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115DBB51FF52FACEFC0A25D8" blockId="6.[153,1437,151,1965]" pageId="6" pageNumber="51">
|
|||
|
|
<emphasis id="B96DD02E115DBB51FF52FACEFE8C254F" box="[197,317,1363,1386]" italics="true" pageId="6" pageNumber="51">Hemipenis</emphasis>
|
|||
|
|
. In MNHN 2012.0216, the organ also reaches 16th SC. In KIZ 0 14591, in situ, hemipenes reach the 13th SC with the origin of
|
|||
|
|
<emphasis id="B96DD02E115DBB51FE2FFAE4FC9B25B4" box="[440,810,1401,1425]" italics="true" pageId="6" pageNumber="51">musculus retractor penis magnus</emphasis>
|
|||
|
|
at 31st SC. The everted organ may be described as such: hemipenis bulbous, not forked but partly divided into two large lobes; sulcus simple; a papilla present at the tip of each lobe, the papilla of the left side of the snake longer and distinctly thicker than the papilla of the right lobe; calyces present on the first basal quarter of the copulatory organ length.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115DBB51FF52F997FB172660" blockId="6.[153,1437,151,1965]" pageId="6" pageNumber="51">
|
|||
|
|
<emphasis id="B96DD02E115DBB51FF52F997FE9E2604" box="[197,303,1546,1569]" italics="true" pageId="6" pageNumber="51">Dentition</emphasis>
|
|||
|
|
. There are 9 or 10 maxillary teeth. The formula of maxillary teeth of specimen MNHN 2012.0216 is 6 + 4 enlarged teeth, especially the last two ones; in IEBR 2012.6, the formula is 6 + 3 teeth.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115DBB51FF52F9CCFD082738" blockId="6.[153,1437,151,1965]" pageId="6" pageNumber="51">
|
|||
|
|
<emphasis id="B96DD02E115DBB51FF52F9CCFEDB264C" box="[197,362,1617,1641]" italics="true" pageId="6" pageNumber="51">Dorsal pattern</emphasis>
|
|||
|
|
. It is quite similar in all these specimens. The number of dorsal blotches varies from 27 to 37 on the body and 5 to 8 on the tail for a total ranging from 35 to 42. Specimen MNHN 2012.0216 is more distinctly brown than the
|
|||
|
|
<typeStatus id="54A2B29E115DBB51FEDBF904FE1F2694" box="[332,430,1689,1713]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
|
|||
|
|
and lateral blotches are smaller and more irregular. Other elements of the dorsal pattern are similar. In life (
|
|||
|
|
<figureCitation id="132210B9115DBB51FEE2F920FE0826F0" box="[373,441,1725,1749]" captionStart="FIGURE 5" captionStartId="7.[151,249,1897,1920]" captionTargetBox="[151,1435,945,1872]" captionTargetId="figure@7.[151,1436,945,1873]" captionTargetPageId="7" captionText="FIGURE 5. Biotope of Oligodon nagao sp. nov. in Khammouane Province, Laos. Photographs by Alexandre Teynié." httpUri="https://zenodo.org/record/214043/files/figure.png" pageId="6" pageNumber="51">Fig. 5</figureCitation>
|
|||
|
|
), upper dorsal surfaces of specimens IEBR A.2012.6 and ZFMK 93281 were greenishbrown, with dark dorsal blotches, golden in their centre and edged with blackish-brown. In alcohol, the paler center is not as visible and turns to pale greyish-brown.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115DBB51FF52F8B5FD052788" blockId="6.[153,1437,151,1965]" pageId="6" pageNumber="51">
|
|||
|
|
<emphasis id="B96DD02E115DBB51FF52F8B5FED52764" bold="true" box="[197,356,1832,1857]" pageId="6" pageNumber="51">Comparisons</emphasis>
|
|||
|
|
. Our comparisons of
|
|||
|
|
<taxonomicName id="4C1977BF115DBB51FDCBF8B4FCA22764" box="[604,787,1833,1857]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="51" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E115DBB51FDCBF8B4FCA22764" box="[604,787,1833,1857]" italics="true" pageId="6" pageNumber="51">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D55115DBB51FC8BF8B4FCC72764" box="[796,886,1833,1857]" pageId="6" pageNumber="51" rank="species">
|
|||
|
|
<emphasis id="B96DD02E115DBB51FC8BF8B4FCC72764" bold="true" box="[796,886,1833,1857]" pageId="6" pageNumber="51">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
with other species of the genus is based on data available in
|
|||
|
|
<bibRefCitation id="EF8871CD115DBB51FEB3F8D0FE0D2740" author="Smith" box="[292,444,1869,1893]" pageId="6" pageNumber="51" refString="Smith, M. A. (1943) The fauna of British India, Ceylon and Burma, including the whole of the Indo-chinese subregion. Reptilia and Amphibia. Vol. III, Serpentes. Taylor & Francis, London, xii + 583 pp." type="book" year="1943">Smith (1943)</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EF8871CD115DBB51FE5EF8D3FDC52740" author="Wagner" box="[457,628,1869,1894]" pageId="6" pageNumber="51" refString="Wagner, F. W. (1975) A revision of the Asian colubrid snakes Oligodon cinereus (Gunther), Oligodon joynsoni (Smith), and Oligodon cyclurus (Cantor). Unpublished MS thesis, Louisiana State University, Baton Rouge (USA), 97 pp." type="book" year="1975">Wagner (1975)</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EF8871CD115DBB51FD16F8D0FCC72740" box="[641,886,1869,1893]" pageId="6" pageNumber="51" refString="Pauwels, O. S. G., Wallach, V., David, P. & Chanhome, L. (2002) A new species of Oligodon Fitzinger, 1826 (Serpentes, Colubridae) from southern Peninsular Thailand. Natural History Journal of Chulalongkorn University, Bangkok, 2 (2), 7 - 18." type="journal article">
|
|||
|
|
Pauwels
|
|||
|
|
<emphasis id="B96DD02E115DBB51FD70F8D3FCAB2740" box="[743,794,1869,1893]" italics="true" pageId="6" pageNumber="51">et al</emphasis>
|
|||
|
|
. (2002)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EF8871CD115DBB51FC14F8D0FBA32740" author="Zhao" box="[899,1042,1869,1893]" pageId="6" pageNumber="51" refString="Zhao, E. (2006) Zhongguo Shelei [Snakes of China]. Anhui Science and Technology Publishing House Publ., Hefei, Vol. I, 372 pp.; Vol. II, 279 pp. (in Chinese)." type="book" year="2006">Zhao (2006)</bibRefCitation>
|
|||
|
|
, David
|
|||
|
|
<emphasis id="B96DD02E115DBB51FBFCF8D3FB2C2740" box="[1131,1181,1869,1893]" italics="true" pageId="6" pageNumber="51">et al</emphasis>
|
|||
|
|
. (2008a-b, 2011),
|
|||
|
|
<bibRefCitation id="EF8871CD115DBB51FAE7F8D3FF5727AC" author="Das" pageId="6" pageNumber="51" refString="Das, I. (2010) A field guide to the Reptiles of South-east Asia. New Holland Publishers (UK) Ltd, London-Cape Town-Sydney- Auckland, 376 pp." type="book" year="2010">Das (2010)</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EF8871CD115DBB51FF64F8ECFDAB27AC" author="Tillack" box="[243,538,1905,1929]" pageId="6" pageNumber="51" refString="Tillack, F. & Gunther, R. (2010) Revision of the species of Oligodon from Sumatra and adjacent islands, with comments on the taxonomic status of Oligodon subcarinatus (Gunther, 1872) and Oligodon annulifer (Boulenger, 1893) from Borneo (Reptilia, Squamata, Colubridae). Russian Journal of Herpetology, 16 [2009] (4), 265 - 294." type="journal article" year="2010">Tillack & Günther (2010)</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EF8871CD115DBB51FDB0F8ECFCB427AC" box="[551,773,1905,1929]" pageId="6" pageNumber="51" refString="Zhang, J., Jang, K., Li, P., Hou, M. & Rao, D. (2011) Taxonomic revisions on genus Oligodon of China (Serpentes, Colubridae). Acta Zootaxonomica Sinica, 36 (2), 423 - 430." type="journal article">
|
|||
|
|
Zhang
|
|||
|
|
<emphasis id="B96DD02E115DBB51FDE0F8EFFD1B27AC" box="[631,682,1905,1929]" italics="true" pageId="6" pageNumber="51">et al</emphasis>
|
|||
|
|
. (2011)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EF8871CD115DBB51FC84F8ECFBA527AC" author="David" box="[787,1044,1905,1929]" pageId="6" pageNumber="51" refString="David, P. & Vogel, G. (2012) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from Pulau Nias, Indonesia. Zootaxa, 3201, 58 - 68." type="journal article" year="2012">David & Vogel (2012)</bibRefCitation>
|
|||
|
|
, and
|
|||
|
|
<bibRefCitation id="EF8871CD115DBB51FBC5F8ECFAF727AC" box="[1106,1350,1905,1929]" pageId="6" pageNumber="51" refString="Geissler, P., Krohn, A. R. & Rennert, A. (2012) Herpetofaunal records in coastal dune areas, Binh Thuan Province, Southern Vietnam, with the rediscovery of Oligodon macrurus Angel, 1927. Russian Journal of Herpetology, 18 [2011] (4), 317 - 324." type="journal article">
|
|||
|
|
Geissler
|
|||
|
|
<emphasis id="B96DD02E115DBB51FB20F8EFFB5B27AC" box="[1207,1258,1905,1929]" italics="true" pageId="6" pageNumber="51">et al</emphasis>
|
|||
|
|
. (2012)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
, and as well as on examined specimens (see Appendix).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF665CB4115CBB50FF00FF05FB1A208B" box="[151,1195,152,174]" pageId="7" pageNumber="52">
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FF00FF05FB1A208B" blockId="7.[151,1394,152,491]" box="[151,1195,152,174]" pageId="7" pageNumber="52">
|
|||
|
|
<emphasis id="B96DD02E115CBB50FF00FF05FEB52088" bold="true" box="[151,260,152,173]" pageId="7" pageNumber="52">TABLE 1.</emphasis>
|
|||
|
|
Main variation in the type series of
|
|||
|
|
<taxonomicName id="4C1977BF115CBB50FDE4FF05FCA9208B" box="[627,792,152,174]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E115CBB50FDE4FF05FCA9208B" box="[627,792,152,174]" italics="true" pageId="7" pageNumber="52">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D55115CBB50FC89FF05FCDE2088" box="[798,879,152,173]" pageId="7" pageNumber="52" rank="species">
|
|||
|
|
<emphasis id="B96DD02E115CBB50FC89FF05FCDE2088" bold="true" box="[798,879,152,173]" pageId="7" pageNumber="52">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
Part 1: body and tail scalation.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FF00FF2AFCD920E9" blockId="7.[151,1394,152,491]" box="[151,872,182,204]" pageId="7" pageNumber="52">See Materials and Methods for abbreviations. All specimens are males.</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FF08FF75FAC320D8" blockId="7.[151,1394,152,491]" box="[159,1394,232,254]" pageId="7" pageNumber="52">Number Body + tail SVL TaL TL Ratio TaL/ DSR VEN SC</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FE32FE9BFC642139" blockId="7.[151,1394,152,491]" box="[421,981,262,285]" pageId="7" pageNumber="52">blotches (mm) (mm) (mm) TL</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FF08FEACFADD2163" blockId="7.[151,1394,152,491]" box="[159,1388,304,326]" pageId="7" pageNumber="52">VNMN A.2012.1 27 + 8 561 92 653 0.141 17–17–15 189 45</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FF08FEC7FADD214A" blockId="7.[151,1394,152,491]" box="[159,1388,345,367]" pageId="7" pageNumber="52">MNHN 2012.0216 30 + 6 628 107 735 0.146 17–15–15 188 47</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FF08FE1EFADD21BD" blockId="7.[151,1394,152,491]" box="[159,1388,386,408]" pageId="7" pageNumber="52">IEBR A.2012.6 32 + 6 680 106 786 0.135 17–17–15 191 44</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FF08FE30FADD21E4" blockId="7.[151,1394,152,491]" box="[159,1388,427,450]" pageId="7" pageNumber="52">KIZ 0 14591 37+5 623 104 727 0.143 17–17–15 193 46</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FF08FE4BFB402341" blockId="7.[151,1394,152,491]" lastBlockId="7.[151,1436,560,868]" pageId="7" pageNumber="52">
|
|||
|
|
ZFMK 93281 31+6 609 98 707 0.139 17–17–15 184 43 Number SL SL / orbit PreOc PosOc Temporals
|
|||
|
|
<collectingCountry id="F30E4CAC115CBB50FBC1FD1CFB5322B3" box="[1110,1250,641,662]" name="Iceland" pageId="7" pageNumber="52">IL IL</collectingCountry>
|
|||
|
|
/ chin shields VNMN A.2012.1 8 / 8 4–5 / 4–5 1 / 1 2 / 2 1+3 / 1+3 8 / 8 1–5 MNHN 2012.0216 8 / 8 4–5 / 4–5 1 / 1 2 / 2 1+2 / 1+2 8 / 8 1–4 IEBR A.2012.6 8 / 8 4–5 / 4–5 1 / 1 2 / 2 1+2 / 1+2 8 / 8 1–4 KIZ 0 14591 8 / 8 4–5 / 4–5 1 / 1 2 / 2 1+2 / 1+2 7 / 8 1–4 ZFMK 93281 7 / 7 3–4 / 3–4 1 / 1 2 / 2 1+2 / 1+2 8 / 8 1–4
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF665CB4115CBB50FF00FDADFE972241" pageId="7" pageNumber="52">
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FF00FDADFE972241" blockId="7.[151,1436,560,868]" pageId="7" pageNumber="52">
|
|||
|
|
<emphasis id="B96DD02E115CBB50FF00FDADFEB52260" bold="true" box="[151,260,560,581]" pageId="7" pageNumber="52">TABLE 2.</emphasis>
|
|||
|
|
Main variation in the type series of
|
|||
|
|
<taxonomicName id="4C1977BF115CBB50FDE1FDADFCAD2263" box="[630,796,560,582]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E115CBB50FDE1FDADFCAD2263" box="[630,796,560,582]" italics="true" pageId="7" pageNumber="52">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D55115CBB50FCB4FDADFCC52260" box="[803,884,560,581]" pageId="7" pageNumber="52" rank="species">
|
|||
|
|
<emphasis id="B96DD02E115CBB50FCB4FDADFCC52260" bold="true" box="[803,884,560,581]" pageId="7" pageNumber="52">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
Part 2: head scalation. See Materials and Methods for abbreviations.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF665CB4115CBB50FF00F8F4FAF1275A" box="[151,1344,1897,1920]" httpUri="https://zenodo.org/record/214043/files/figure.png" pageId="7" pageNumber="52" targetBox="[151,1435,945,1872]" targetPageId="7">
|
|||
|
|
<paragraph id="8BA60C3C115CBB50FF00F8F4FAF1275A" blockId="7.[151,1344,1897,1920]" box="[151,1344,1897,1920]" pageId="7" pageNumber="52">
|
|||
|
|
<emphasis id="B96DD02E115CBB50FF00F8F4FEA3275A" bold="true" box="[151,274,1897,1920]" pageId="7" pageNumber="52">FIGURE 5.</emphasis>
|
|||
|
|
Biotope of
|
|||
|
|
<taxonomicName id="4C1977BF115CBB50FE1DF8F4FD9E275A" box="[394,559,1897,1919]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E115CBB50FE1DF8F4FD9E275A" box="[394,559,1897,1919]" italics="true" pageId="7" pageNumber="52">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D55115CBB50FDA1F8F7FD37275A" box="[566,646,1898,1919]" pageId="7" pageNumber="52" rank="species">
|
|||
|
|
<emphasis id="B96DD02E115CBB50FDA1F8F7FD37275A" bold="true" box="[566,646,1898,1919]" pageId="7" pageNumber="52">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
in Khammouane Province, Laos. Photographs by Alexandre Teynié.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA60C3C1153BB5FFF52FF0AFAAB21D1" blockId="8.[153,1436,151,2011]" pageId="8" pageNumber="53">
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFF52FF0AFEEB208A" author="Smith" box="[197,346,151,175]" pageId="8" pageNumber="53" refString="Smith, M. A. (1943) The fauna of British India, Ceylon and Burma, including the whole of the Indo-chinese subregion. Reptilia and Amphibia. Vol. III, Serpentes. Taylor & Francis, London, xii + 583 pp." type="book" year="1943">Smith (1943)</bibRefCitation>
|
|||
|
|
defined informal groups in the genus
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC9BFF05FCC72095" box="[780,886,152,176]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC9BFF05FCC72095" box="[780,886,152,176]" italics="true" pageId="8" pageNumber="53">Oligodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
on the basis of the hemipenial morphology. The distinctiveness of these groups was confirmed by
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFD5FFF21FC1020F1" box="[712,929,188,212]" pageId="8" pageNumber="53" refString="Green, M. D., Orlov, N. L. & Murphy, R. W. (2010) Toward a phylogeny of the Kukri snakes, genus Oligodon. Asian Herpetological Research, 1 (1), 1 - 21." type="journal article">
|
|||
|
|
Green
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC83FF20FCF420F1" box="[788,837,188,212]" italics="true" pageId="8" pageNumber="53">et al</emphasis>
|
|||
|
|
. (2010)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
. Among the 50 species or so of the genus for which the hemipenial morphology is known according to
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFCD9FF42FB8620D2" box="[846,1079,223,248]" pageId="8" pageNumber="53" refString="Green, M. D., Orlov, N. L. & Murphy, R. W. (2010) Toward a phylogeny of the Kukri snakes, genus Oligodon. Asian Herpetological Research, 1 (1), 1 - 21." type="journal article">
|
|||
|
|
Green
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC08FF7CFC6720DD" box="[927,982,224,248]" italics="true" pageId="8" pageNumber="53">et al</emphasis>
|
|||
|
|
. (2010)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
, only the groups of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFAA5FF7DFEB12139" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="taeniatus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFAA5FF7DFEB12139" italics="true" pageId="8" pageNumber="53">Oligodon taeniatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFEA8FE99FDA52139" box="[319,532,260,284]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cyclurus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFEA8FE99FDA52139" box="[319,532,260,284]" italics="true" pageId="8" pageNumber="53">Oligodon cyclurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
have deeply forked hemipenes. Species of all other groups have non-forked copulatory organs. Among these groups, only the groups of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFCF8FEB5FBBC2165" box="[879,1037,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cruentatus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFCF8FEB5FBBC2165" box="[879,1037,296,320]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFCF8FEB5FC37211A" box="[879,902,296,319]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cruentatus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFBB6FEB5FB162165" box="[1057,1191,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cinereus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFBB6FEB5FB162165" box="[1057,1191,296,320]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFBB6FEB5FB89211A" box="[1057,1080,296,319]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cinereus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFB2BFEB5FADC2165" box="[1212,1389,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="octolineatus">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFB2BFEB5FB61211A" box="[1212,1232,296,319]" name="Iceland" pageId="8" pageNumber="53">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFB2BFEB5FB61211A" box="[1212,1232,296,319]" italics="true" pageId="8" pageNumber="53">O</emphasis>
|
|||
|
|
</collectingCountry>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFB73FEB5FADC2165" box="[1252,1389,296,320]" italics="true" pageId="8" pageNumber="53">octolineatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFA16FEB5FE832141" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="purpurascens">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFA16FEB5FE832141" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFA16FEB5FA29211A" box="[1409,1432,296,319]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. purpurascens
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFEF1FED0FE412141" box="[358,496,332,356]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="modestus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFEF1FED0FE412141" box="[358,496,332,356]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFEF1FED0FECC2141" box="[358,381,333,356]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. modestus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
have organs with long papillae but spines are absent only in species of the groups of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFF0EFEEDFEAA21AD" box="[153,283,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cinereus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF0EFEEDFEAA21AD" box="[153,283,368,392]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFF0EFEEDFF0121A2" box="[153,176,368,391]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cinereus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFEBBFEEDFE6821AD" box="[300,473,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="octolineatus">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFEBBFEEDFEF121A2" box="[300,320,368,391]" name="Iceland" pageId="8" pageNumber="53">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFEBBFEEDFEF121A2" box="[300,320,368,391]" italics="true" pageId="8" pageNumber="53">O</emphasis>
|
|||
|
|
</collectingCountry>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFEC7FEEDFE6821AD" box="[336,473,368,392]" italics="true" pageId="8" pageNumber="53">octolineatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFE7EFEEDFDC721AD" box="[489,630,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="modestus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFE7EFEEDFDC721AD" box="[489,630,368,392]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFE7EFEEDFDB121A2" box="[489,512,368,391]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. modestus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFD2FFEEDFCC421AD" box="[696,885,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="purpurascens">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD2FFEEDFCC421AD" box="[696,885,368,392]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD2FFEEDFD7E21A2" box="[696,719,368,391]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. purpurascens
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. We consider that
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFBD8FEEDFAB921AD" box="[1103,1288,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFBD8FEEDFAB921AD" box="[1103,1288,368,392]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFA86FEEDFADC21AD" box="[1297,1389,368,392]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFA86FEEDFADC21AD" bold="true" box="[1297,1389,368,392]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
has affinities with only one of these groups.
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFDF9FE09FD3C2189" box="[622,653,404,428]" name="American Samoa" pageId="8" pageNumber="53">As</collectingCountry>
|
|||
|
|
members of the groups of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC5DFE08FB362189" box="[970,1159,405,428]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="purpurascens">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC5DFE08FB362189" box="[970,1159,405,428]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFC5DFE08FC502189" box="[970,993,405,428]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. purpurascens
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFB0FFE08FAF42189" box="[1176,1349,404,428]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="octolineatus">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFB0FFE08FB1D2189" box="[1176,1196,405,428]" name="Iceland" pageId="8" pageNumber="53">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFB0FFE08FB1D2189" box="[1176,1196,405,428]" italics="true" pageId="8" pageNumber="53">O</emphasis>
|
|||
|
|
</collectingCountry>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFB2BFE09FAF42189" box="[1212,1349,404,428]" italics="true" pageId="8" pageNumber="53">octolineatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFA16FE08FEB021F5" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="modestus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFA16FE08FEB021F5" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFA16FE08FA292189" box="[1409,1432,405,428]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. modestus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
include species inhabiting the Malay Peninsula, Indo-Malayan Archipelago and the
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFB5DFE2AFA2621EA" box="[1226,1431,439,463]" name="Philippines" pageId="8" pageNumber="53">Philippine Islands</collectingCountry>
|
|||
|
|
, affinities are rather with the group of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFDAAFE40FD0C21D1" box="[573,701,477,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cinereus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFDAAFE40FD0C21D1" box="[573,701,477,500]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFDAAFE40FDE521D1" box="[573,596,477,500]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cinereus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
for both zoogeographical and morphological reasons.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3035FB71153BB5EFF52FD9DFC542743" lastPageId="9" lastPageNumber="54" pageId="8" pageNumber="53" type="reference_group">
|
|||
|
|
<paragraph id="8BA60C3C1153BB5FFF52FD9DFBE0242A" blockId="8.[153,1436,151,2011]" pageId="8" pageNumber="53">
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFF52FD9DFECC223D" box="[197,381,512,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF52FD9DFECC223D" box="[197,381,512,536]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFE12FD9DFE51223D" box="[389,480,512,536]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFE12FD9DFE51223D" bold="true" box="[389,480,512,536]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
differs from members of the groups of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC33FD9DFB94223D" box="[932,1061,512,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cinereus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC33FD9DFB94223D" box="[932,1061,512,536]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFC33FD9DFC0A2232" box="[932,955,512,535]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cinereus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFBCAFD9DFAA8223D" box="[1117,1305,512,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="purpurascens">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFBCAFD9DFAA8223D" box="[1117,1305,512,536]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFBCAFD9DFBC52232" box="[1117,1140,512,535]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. purpurascens
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as follows.
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFF0EFDB9FEE12219" box="[153,336,548,572]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF0EFDB9FEE12219" box="[153,336,548,572]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFECFFDB9FE032219" box="[344,434,548,572]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFECFFDB9FE032219" bold="true" box="[344,434,548,572]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
has a dorsal pattern made of 27–37 dark, butterfly-shaped blotches vs. a pattern uniform in
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFF21FDD5FF7B227A" box="[182,202,584,607]" name="Iceland" pageId="8" pageNumber="53">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF21FDD5FF7B227A" box="[182,202,584,607]" italics="true" pageId="8" pageNumber="53">O</emphasis>
|
|||
|
|
</collectingCountry>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF4FFDD4FF552245" box="[216,228,585,608]" italics="true" pageId="8" pageNumber="53">c</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFF64FDD4FEE02245" box="[243,337,585,608]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cinereus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF64FDD4FEE02245" box="[243,337,585,608]" italics="true" pageId="8" pageNumber="53">cinereus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFE1FFDD5FDA72245" box="[392,534,584,608]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="inornatus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFE1FFDD5FDA72245" box="[392,534,584,608]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFE1FFDD5FE2E227A" box="[392,415,584,607]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. inornatus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, or reticulate in
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD58FDD5FD52227A" box="[719,739,584,607]" name="Iceland" pageId="8" pageNumber="53">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD58FDD5FD52227A" box="[719,739,584,607]" italics="true" pageId="8" pageNumber="53">O</emphasis>
|
|||
|
|
</collectingCountry>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD65FDD4FD4F2245" box="[754,766,585,608]" italics="true" pageId="8" pageNumber="53">c</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC9BFDD4FCDB2245" box="[780,874,585,608]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cinereus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC9BFDD4FCDB2245" box="[780,874,585,608]" italics="true" pageId="8" pageNumber="53">cinereus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC36FDD5FB922245" box="[929,1059,584,608]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC36FDD5FB922245" box="[929,1059,584,608]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFC36FDD5FC09227A" box="[929,952,584,607]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. joynsoni
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, or made of solid black crossbars in
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFF2BFDF0FD1022A1" authority="Bourret, 1935" authorityName="Bourret" authorityYear="1935" box="[188,673,620,644]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="subSpecies" species="cinereus" subSpecies="tamdaoensis">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF2BFDF0FE6C22A1" box="[188,477,620,644]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFF2BFDF0FF6222A1" box="[188,211,621,644]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cinereus tamdaoensis
|
|||
|
|
</emphasis>
|
|||
|
|
(Bourret, 1935)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, or of black-edged pale crossbars in
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFBFEFDF0FEFE2282" authority="Bourret, 1934" authorityName="Bourret" authorityYear="1934" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="subSpecies" species="cinereus" subSpecies="pallidocinctus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFBFEFDF0FA2D22A1" box="[1129,1436,620,644]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFBFEFDF0FB3122A1" box="[1129,1152,621,644]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cinereus pallidocinctus
|
|||
|
|
</emphasis>
|
|||
|
|
(Bourret, 1934)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, one morph of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFD9AFD0DFCDA2282" authority="Cantor, 1839" authorityName="Cantor" authorityYear="1839" box="[525,875,655,680]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="albocinctus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD9AFD0DFD07228D" box="[525,694,656,680]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD9AFD0DFD952282" box="[525,548,656,679]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. albocinctus
|
|||
|
|
</emphasis>
|
|||
|
|
(Cantor, 1839)
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC3BFD0DFBCA228D" box="[940,1147,656,680]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="melanozonatus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC3BFD0DFBCA228D" box="[940,1147,656,680]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFC3BFD0DFC722282" box="[940,963,656,679]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. melanozonatus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, or, lastly, striped in
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFA16FD0DFE9722E9" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="woodmasoni">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFA16FD0DFE9722E9" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFA16FD0DFA292282" box="[1409,1432,656,679]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. woodmasoni
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Additional differences are given in
|
|||
|
|
<tableCitation id="C69B39871153BB5FFD5EFD29FC9622E9" box="[713,807,692,716]" captionStart="TABLE 3" captionStartId="9.[151,235,356,377]" captionTargetBox="[151,1436,464,1022]" captionTargetPageId="9" captionText="TABLE 3. Main characters of mainland Oligodon species with 15 dorsal scale rows at midbody. Exceptional values are placed in brackets. See Materials and Methods for abbreviations, with the addition of: Cloacal plate: D—divided; E—entire;? — data unavailable." httpUri="http://table.plazi.org/id/DF665CB41152BB5EFF00FEF9FEA22192" pageId="8" pageNumber="53" tableUuid="DF665CB41152BB5EFF00FEF9FEA22192">Tables 3</tableCitation>
|
|||
|
|
and 4 below. In contrast, one morph of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFB67FD28FA2422E9" box="[1264,1429,692,716]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="albocinctus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFB67FD28FA2422E9" box="[1264,1429,692,716]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFB67FD28FAB622E9" box="[1264,1287,693,716]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. albocinctus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFF0EFD45FEE922D5" box="[153,344,728,752]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="purpurascens">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF0EFD45FEE922D5" box="[153,344,728,752]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFF0EFD45FF0122CA" box="[153,176,728,751]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. purpurascens
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFE08FD45FD8E22D5" box="[415,575,728,752]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="splendidus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFE08FD45FD8E22D5" box="[415,575,728,752]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFE08FD45FE0722CA" box="[415,438,728,751]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. splendidus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has a blotched dorsal pattern.
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC26FD45FBDC22D5" box="[945,1133,728,752]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC26FD45FBDC22D5" box="[945,1133,728,752]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFBEEFD45FB6622D5" box="[1145,1239,728,752]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFBEEFD45FB6622D5" bold="true" box="[1145,1239,728,752]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
differs from the blotched morph (Form II of
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFE44FD61FDE42331" author="Smith" box="[467,597,764,788]" pageId="8" pageNumber="53" refString="Smith, M. A. (1943) The fauna of British India, Ceylon and Burma, including the whole of the Indo-chinese subregion. Reptilia and Amphibia. Vol. III, Serpentes. Taylor & Francis, London, xii + 583 pp." type="book" year="1943">Smith 1943</bibRefCitation>
|
|||
|
|
) of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFD15FD60FC942331" box="[642,805,764,788]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="albocinctus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD15FD60FC942331" box="[642,805,764,788]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD15FD60FD282331" box="[642,665,765,788]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. albocinctus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by (1) 17 vs. 19 or 21 DSR, (2) a much shorter tail, ratio Tal/TL 0.135–0.146 vs. 0.171–0.200 in males of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFD74FCBDFC3A231D" box="[739,907,800,824]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="albocinctus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD74FCBDFC3A231D" box="[739,907,800,824]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD74FCBDFD4B2312" box="[739,762,800,823]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. albocinctus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and (3) a largely dark venter vs. pale with rectangular blotches.
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFE19FCD9FDF92379" box="[398,584,836,860]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFE19FCD9FDF92379" box="[398,584,836,860]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFDC5FCD9FD1F2379" box="[594,686,836,860]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFDC5FCD9FD1F2379" bold="true" box="[594,686,836,860]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFCDDFCD8FBB62379" box="[842,1031,837,860]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="purpurascens">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFCDDFCD8FBB62379" box="[842,1031,837,860]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFCDDFCD8FCD02379" box="[842,865,837,860]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. purpurascens
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by (1) 17 vs. 19 or 21 DSR, (2) a shorter tail, ratio TaL / TL 0.135–0.146 vs.
|
|||
|
|
<quantity id="4CE1A1D91153BB5FFD0FFCFAFCF5235A" box="[664,836,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="4.318" metricValueMax="4.7752" metricValueMin="3.8608" pageId="8" pageNumber="53" unit="in" value="0.16999999999999998" valueMax="0.188" valueMin="0.152">0.152–0.188 in</quantity>
|
|||
|
|
males of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC2CFCF5FBC823A5" box="[955,1145,872,896]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="purpurascens">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC2CFCF5FBC823A5" box="[955,1145,872,896]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFC2CFCF5FC63235A" box="[955,978,872,895]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. purpurascens
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and a higher number of dorsal blotches, 27–37 vs.
|
|||
|
|
<quantity id="4CE1A1D91153BB5FFE4AFC11FDF62381" box="[477,583,908,932]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="4.572" metricValueMin="2.54" pageId="8" pageNumber="53" unit="in" value="14.0" valueMax="18.0" valueMin="10.0">10–18 in</quantity>
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFDC3FC10FCA42381" box="[596,789,909,932]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="purpurascens">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFDC3FC10FCA42381" box="[596,789,909,932]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFDC3FC10FDDA2381" box="[596,619,909,932]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. purpurascens
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFCBEFC11FC572381" box="[809,998,908,932]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFCBEFC11FC572381" box="[809,998,908,932]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFC63FC11FBE52381" box="[1012,1108,908,932]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC63FC11FBE52381" bold="true" box="[1012,1108,908,932]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFB6BFC10FEFE23E2" authority="Gunther, 1875" authorityName="Gunther" authorityYear="1875" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="splendidus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFB6BFC10FA2D2381" box="[1276,1436,908,932]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFB6BFC10FAA22381" box="[1276,1299,909,932]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. splendidus
|
|||
|
|
</emphasis>
|
|||
|
|
(Günther, 1875)
|
|||
|
|
</taxonomicName>
|
|||
|
|
by 17 vs. 21 DSR, (2) 2 prefrontals vs. 4, and (3) 27–37 vs. 14–17 dorsal blotches. Lastly,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFA16FC2DFEBD23C9" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="maculatus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFA16FC2DFEBD23C9" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFA16FC2DFA2923E2" box="[1409,1432,944,967]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. maculatus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, endemic to the
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFE5FFC49FDF723C9" box="[456,582,980,1004]" name="Philippines" pageId="8" pageNumber="53">Philippines</collectingCountry>
|
|||
|
|
has also 17 DSR and a blotched dorsal pattern but this latter species has no more than 164 ventrals (
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFE3EFC6AFD87242A" author="Alcala" box="[425,566,1015,1039]" pageId="8" pageNumber="53" refString="Alcala, A. C. (1986) Guide to Philippine flora and fauna. Vol. X. Amphibians and reptiles. Natural Resources Management Center, Ministry of Natural Resources and University of the Philippines, Manila, xiv + 195 pp." type="book" year="1986">Alcala 1986</bibRefCitation>
|
|||
|
|
), 7 supralabials and only 20–24 dorsal blotches.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1153BB5FFF52FB81FCB82529" blockId="8.[153,1436,151,2011]" pageId="8" pageNumber="53">
|
|||
|
|
When it was discovered in
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD95FB81FDF42411" box="[514,581,1052,1076]" name="China" pageId="8" pageNumber="53">China</collectingCountry>
|
|||
|
|
in 1998 (Anonymous 2002),
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC0AFB81FBE62411" box="[925,1111,1052,1076]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC0AFB81FBE62411" box="[925,1111,1052,1076]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFBF7FB81FB0C2411" box="[1120,1213,1052,1076]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFBF7FB81FB0C2411" bold="true" box="[1120,1213,1052,1076]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
was confused with
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFF0EFBDDFEDB247D" box="[153,362,1088,1112]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF0EFBDDFEDB247D" box="[153,362,1088,1112]" italics="true" pageId="8" pageNumber="53">Oligodon joynsoni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. This rare species is currently known from three provinces of northern
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFB0EFBA2FB4D2472" box="[1177,1276,1087,1111]" name="Thailand" pageId="8" pageNumber="53">Thailand</collectingCountry>
|
|||
|
|
(Chiang Mai, Lampang and Loei) and from an unknown locality in
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD61FBF8FC9C2459" box="[758,813,1125,1148]" name="Laos" pageId="8" pageNumber="53">Laos</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFCABFBF9FBB42459" author="David" box="[828,1029,1124,1148]" pageId="8" pageNumber="53" refString="David, P., Das, I. & Vogel, G. (2011) On some taxonomic and nomenclatural problems in Indian species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae). Zootaxa, 2799, 1 - 14." type="journal article" year="2011">
|
|||
|
|
David
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC1DFBF8FC0A2459" box="[906,955,1124,1148]" italics="true" pageId="8" pageNumber="53">et al</emphasis>
|
|||
|
|
. 2011
|
|||
|
|
</bibRefCitation>
|
|||
|
|
). On the basis of the five specimens examined by us, of four specimens cited by
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFD1FFB1AFCAC24BA" author="Taylor" box="[648,797,1159,1183]" pageId="8" pageNumber="53" refString="Taylor, E. H. (1965) The serpents of Thailand and adjacent waters. University of Kansas Science Bulletin, 45 (9), 609 - 1096." type="journal article" year="1965">Taylor (1965</bibRefCitation>
|
|||
|
|
; see
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFCC3FB1AFBAB24BA" author="David" box="[852,1050,1159,1184]" pageId="8" pageNumber="53" refString="David, P., Das, I. & Vogel, G. (2011) On some taxonomic and nomenclatural problems in Indian species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae). Zootaxa, 2799, 1 - 14." type="journal article" year="2011">
|
|||
|
|
David
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC08FB14FC602485" box="[927,977,1160,1184]" italics="true" pageId="8" pageNumber="53">et al</emphasis>
|
|||
|
|
. 2011
|
|||
|
|
</bibRefCitation>
|
|||
|
|
), and of
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFBECFB14FA9724BA" author="Wagner" box="[1147,1318,1159,1185]" pageId="8" pageNumber="53" refString="Wagner, F. W. (1975) A revision of the Asian colubrid snakes Oligodon cinereus (Gunther), Oligodon joynsoni (Smith), and Oligodon cyclurus (Cantor). Unpublished MS thesis, Louisiana State University, Baton Rouge (USA), 97 pp." type="book" year="1975">Wagner (1975)</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFAA5FB15FF6E24E1" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFAA5FB15FF6E24E1" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFF70FB31FEF024E1" box="[231,321,1196,1220]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF70FB31FEF024E1" bold="true" box="[231,321,1196,1220]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFE4EFB30FDEA24E1" box="[473,603,1197,1220]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFE4EFB30FDEA24E1" box="[473,603,1197,1220]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFE4EFB30FE4124E1" box="[473,496,1197,1220]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. joynsoni
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by (1) a different pattern,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC1EFB30FBBD24E1" box="[905,1036,1197,1220]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC1EFB30FBBD24E1" box="[905,1036,1197,1220]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFC1EFB30FC1124E1" box="[905,928,1197,1220]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. joynsoni
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
having no dorsal blotches but only more or less visible reticulations on a very dark background, (2) 9–10 maxillary teeth vs.
|
|||
|
|
<quantity id="4CE1A1D91153BB5FFBE9FB4DFB6E24CD" box="[1150,1247,1232,1256]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="8" pageNumber="53" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">11–12 in</quantity>
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFB72FB4DFAD724CD" box="[1253,1382,1232,1256]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFB72FB4DFAD724CD" box="[1253,1382,1232,1256]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFB72FB4DFB4D24C2" box="[1253,1276,1232,1255]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. joynsoni
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and (3) 1 versus usually 2 anterior temporals in
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFD16FB68FCB32529" box="[641,770,1269,1292]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD16FB68FCB32529" box="[641,770,1269,1292]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD16FB68FD292529" box="[641,664,1269,1292]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. joynsoni
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1153BB5FFF52FA85FB0125C1" blockId="8.[153,1436,151,2011]" pageId="8" pageNumber="53">
|
|||
|
|
Currently, the
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFEF0FA85FD892515" box="[359,568,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cyclurus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFEF0FA85FD892515" box="[359,568,1304,1328]" italics="true" pageId="8" pageNumber="53">Oligodon cyclurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group is composed of nine species (see above) which have both forked, non spinose and non papillate hemipenes (although short papillae are present in
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC67FAA0FB242571" box="[1008,1173,1340,1364]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="formosanus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC67FAA0FB242571" box="[1008,1173,1340,1364]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFC67FAA0FBB62571" box="[1008,1031,1341,1364]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. formosanus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
). With 17 (or 15) DSR at midbody,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFEBEFAFDFE53255D" box="[297,482,1376,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFEBEFAFDFE53255D" box="[297,482,1376,1400]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFE7BFAFDFDF9255D" box="[492,584,1376,1400]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFE7BFAFDFDF9255D" bold="true" box="[492,584,1376,1400]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFD73FAFDFCD9255D" box="[740,872,1376,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cyclurus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD73FAFDFCD9255D" box="[740,872,1376,1400]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD73FAFDFD4A2552" box="[740,763,1376,1399]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cyclurus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(19),
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFC26FAFDFBE8255D" box="[945,1113,1376,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="formosanus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC26FAFDFBE8255D" box="[945,1113,1376,1400]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFC26FAFDFC792552" box="[945,968,1376,1399]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. formosanus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(19),
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFB36FAFDFA89255D" box="[1185,1336,1376,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="kheriensis">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFB36FAFDFA89255D" box="[1185,1336,1376,1400]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFB36FAFDFB092552" box="[1185,1208,1376,1399]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. kheriensis
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(19),
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFA16FAFDFEA225B9" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="fasciolatus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFA16FAFDFEA225B9" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFA16FAFDFA292552" box="[1409,1432,1376,1399]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. fasciolatus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(21 or 23),
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFE36FA18FDF025B9" box="[417,577,1412,1436]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="juglandifer">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFE36FA18FDF025B9" box="[417,577,1412,1436]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFE36FA18FE0925B9" box="[417,440,1413,1436]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. juglandifer
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(19), and
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFD2BFA18FCF725B9" box="[700,838,1412,1436]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="ocellatus">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD2BFA18FD6125B9" box="[700,720,1413,1436]" name="Iceland" pageId="8" pageNumber="53">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD2BFA18FD6125B9" box="[700,720,1413,1436]" italics="true" pageId="8" pageNumber="53">O</emphasis>
|
|||
|
|
</collectingCountry>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD76FA19FCF725B9" box="[737,838,1412,1436]" italics="true" pageId="8" pageNumber="53">ocellatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(19). Furthermore,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFBBCFA19FB5525B9" box="[1067,1252,1412,1436]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFBBCFA19FB5525B9" box="[1067,1252,1412,1436]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFB79FA19FAFB25B9" box="[1262,1354,1412,1436]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFB79FA19FAFB25B9" bold="true" box="[1262,1354,1412,1436]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFF4EFA35FEEC25E5" box="[217,349,1448,1472]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cyclurus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF4EFA35FEEC25E5" box="[217,349,1448,1472]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFF4EFA35FF41259A" box="[217,240,1448,1471]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cyclurus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by (1) 17 vs. 19 DSR, (2) 9–10 vs. 12–13 maxillary teeth, (3) 184–193 VEN vs.
|
|||
|
|
<quantity id="4CE1A1D91153BB5FFA8DFA35FA2D25E5" box="[1306,1436,1448,1472]" metricMagnitude="0" metricUnit="m" metricValue="4.2291" metricValueMax="4.3688" metricValueMin="4.0893999999999995" pageId="8" pageNumber="53" unit="in" value="166.5" valueMax="172.0" valueMin="161.0">161–172 in</quantity>
|
|||
|
|
males, (4) 1 anterior temporal vs. always 2, and (5) a blotched pattern vs. dorsal reticulations.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1153BB5FFF52FA72FBB327B1" blockId="8.[153,1436,151,2011]" pageId="8" pageNumber="53">
|
|||
|
|
The three remaining species of the
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFDCBFA6DFD51262D" box="[604,736,1520,1544]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cyclurus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFDCBFA6DFD51262D" box="[604,736,1520,1544]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFDCBFA6DFDC22622" box="[604,627,1520,1543]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cyclurus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
-group have 17 DSR at midbody. Besides the morphology of hemipenes,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFEB6F989FE682609" box="[289,473,1556,1580]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFEB6F989FE682609" box="[289,473,1556,1580]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFE74F989FD8F2609" box="[483,574,1556,1580]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFE74F989FD8F2609" bold="true" box="[483,574,1556,1580]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
differs from males of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFCD4F988FC5C2609" box="[835,1005,1557,1580]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="saintgironsi">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFCD4F988FC5C2609" box="[835,1005,1557,1580]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFCD4F988FCEB2609" box="[835,858,1557,1580]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. saintgironsi
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by (1) a shorter tail (0.135–0.146 vs. 0.191–0.203), (2) a higher number of ventrals, 184–193 vs. 166–170, and (3) a distinct pattern; both species have dorsal blotches but they differ in shape (see
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFD15F9C1FCE72651" author="David" box="[642,854,1628,1652]" pageId="8" pageNumber="53" refString="David, P., Vogel, G. & Van Rooijen, J. (2008 b) A revision of the Oligodon taeniatus (Gunther, 1861) (Squamata: Colubridae) group, with the description of three new species from the Indochinese Region. Zootaxa, 1965, 1 - 45." type="journal article" year="2008" yearSuffix="b">
|
|||
|
|
David
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD5AF9C0FD4C2651" box="[717,765,1628,1652]" italics="true" pageId="8" pageNumber="53">et al</emphasis>
|
|||
|
|
. 2008b
|
|||
|
|
</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFCF0F9C1FBAC2651" box="[871,1053,1628,1652]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFCF0F9C1FBAC2651" box="[871,1053,1628,1652]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFBB3F9C1FBCC2651" box="[1060,1149,1628,1652]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFBB3F9C1FBCC2651" bold="true" box="[1060,1149,1628,1652]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFA98F9C0FA2D2651" box="[1295,1436,1629,1652]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="macrurus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFA98F9C0FA2D2651" box="[1295,1436,1629,1652]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFA98F9C0FA972651" box="[1295,1318,1629,1652]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. macrurus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by (1) a shorter tail in males, 0.135–0.146 vs. 0.329–0.373, (2) a lower number of SC in males, 43–47 vs. 73–94, (3) 7–8 vs. 9–10
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFEF1F938FE322699" box="[358,387,1701,1724]" name="Iceland" pageId="8" pageNumber="53">IL</collectingCountry>
|
|||
|
|
, (4) a lower number of maxillary teeth, 9–10 vs. 14–15, and (5) a totally different dorsal pattern,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFF6DF955FE3F26C5" box="[250,398,1736,1760]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="macrurus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF6DF955FE3F26C5" box="[250,398,1736,1760]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFF6DF955FEA026FA" box="[250,273,1736,1759]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. macrurus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
being uniform or showing only faint reticulations on a rather pale background. Lastly,
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFF0EF971FEE12721" box="[153,336,1772,1796]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFF0EF971FEE12721" box="[153,336,1772,1796]" italics="true" pageId="8" pageNumber="53">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551153BB5FFEC0F971FE012721" box="[343,432,1772,1796]" pageId="8" pageNumber="53" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFEC0F971FE012721" bold="true" box="[343,432,1772,1796]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
differs from males of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFD3BF970FC872721" box="[684,822,1772,1796]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="chinensis">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFD3BF970FC872721" box="[684,822,1772,1796]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFD3BF970FD722721" box="[684,707,1773,1796]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. chinensis
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, which occurs in the same region, by (1) a shorter tail, 0.135–0.146 vs. 0.187–0.195, (2) a higher number of ventrals, 184–193 vs. 175–184, (3) a lower number of SC, 43–47 vs. 60–64, (4) 1 anterior temporal vs. usually 2 (in 19 out of 28 occurrences) in
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFB03F8A8FA902769" box="[1172,1313,1844,1868]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="chinensis">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFB03F8A8FA902769" box="[1172,1313,1844,1868]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFB03F8A8FB1A2769" box="[1172,1195,1845,1868]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. chinensis
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and (5) a distinct dorsal pattern, with more dorsal blotches, 27–37 vs. 11–14 blotches of different shape. Additional data on members of the
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFED9F8E0FE7E27B1" box="[334,463,1916,1940]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cyclurus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFED9F8E0FE7E27B1" box="[334,463,1916,1940]" italics="true" pageId="8" pageNumber="53">
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFED9F8E0FED427B1" box="[334,357,1917,1940]" name="Iceland" pageId="8" pageNumber="53">O</collectingCountry>
|
|||
|
|
. cyclurus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group can be found in
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFD44F8E1FC1E27B1" author="David" box="[723,943,1916,1940]" pageId="8" pageNumber="53" refString="David, P., Vogel, G. & Van Rooijen, J. (2008 b) A revision of the Oligodon taeniatus (Gunther, 1861) (Squamata: Colubridae) group, with the description of three new species from the Indochinese Region. Zootaxa, 1965, 1 - 45." type="journal article" year="2008" yearSuffix="b">
|
|||
|
|
David
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFC89F8E0FCFE27B1" box="[798,847,1916,1940]" italics="true" pageId="8" pageNumber="53">et al</emphasis>
|
|||
|
|
. (2008b
|
|||
|
|
</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EF8871CD1153BB5FFC2CF8E1FC4727B1" author="David" box="[955,1014,1916,1940]" pageId="8" pageNumber="53" refString="David, P., Das, I. & Vogel, G. (2011) On some taxonomic and nomenclatural problems in Indian species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae). Zootaxa, 2799, 1 - 14." type="journal article" year="2011">2011</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1153BB5EFF52F83DFCC3211F" blockId="8.[153,1436,151,2011]" lastBlockId="9.[153,1436,151,315]" lastPageId="9" lastPageNumber="54" pageId="8" pageNumber="53">
|
|||
|
|
Nevertheless, because dissected hemipenes are not always available, we further compare our new species with species of
|
|||
|
|
<taxonomicName id="4C1977BF1153BB5FFE98F85EFEC827FE" box="[271,377,1987,2011]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="53" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B96DD02E1153BB5FFE98F85EFEC827FE" box="[271,377,1987,2011]" italics="true" pageId="8" pageNumber="53">Oligodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
inhabiting the Asian mainland and
|
|||
|
|
<collectingCountry id="F30E4CAC1153BB5FFC9FF85FFCEB27FF" box="[776,858,1986,2010]" name="Taiwan" pageId="8" pageNumber="53">Taiwan</collectingCountry>
|
|||
|
|
with either 17 or 15 DSR at midbody, regardless of their current group; species from
|
|||
|
|
<collectingCountry id="F30E4CAC1152BB5EFD85FF0AFD35208A" box="[530,644,151,175]" name="Sri Lanka" pageId="9" pageNumber="54">Sri Lanka</collectingCountry>
|
|||
|
|
, the Indo-Malayan Archipelago and the
|
|||
|
|
<collectingCountry id="F30E4CAC1152BB5EFBDBFF0AFB7C208A" box="[1100,1229,151,175]" name="Philippines" pageId="9" pageNumber="54">Philippines</collectingCountry>
|
|||
|
|
, not present in the region of
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFE90FF26FE0F20F6" box="[263,446,187,211]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFE90FF26FE0F20F6" box="[263,446,187,211]" italics="true" pageId="9" pageNumber="54">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551152BB5EFE51FF26FD9020F6" box="[454,545,187,211]" pageId="9" pageNumber="54" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFE51FF26FD9020F6" bold="true" box="[454,545,187,211]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
, are excluded. The number of DSR at midbody is a major diagnostic character in the genus
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFEBEFF40FE2220D0" box="[297,403,221,245]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFEBEFF40FE2220D0" box="[297,403,221,245]" italics="true" pageId="9" pageNumber="54">Oligodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(see David
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFD8BFF43FDFC20D0" box="[540,589,221,245]" italics="true" pageId="9" pageNumber="54">et al</emphasis>
|
|||
|
|
. 2008a-b). This number is usually constant within a given species but may sometimes vary due to an anomalous position of the dorsal scale row reductions, as it is likely the case with
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0EFEBEFEE1211E" box="[153,336,291,315]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0EFEBEFEE1211E" box="[153,336,291,315]" italics="true" pageId="9" pageNumber="54">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551152BB5EFEC1FEBEFE01211E" box="[342,432,291,315]" pageId="9" pageNumber="54" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFEC1FEBEFE01211E" bold="true" box="[342,432,291,315]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
Data are summarized in
|
|||
|
|
<tableCitation id="C69B39871152BB5EFD5EFEBFFC96211F" box="[713,807,290,314]" captionStart="TABLE 3" captionStartId="9.[151,235,356,377]" captionTargetBox="[151,1436,464,1022]" captionTargetPageId="9" captionText="TABLE 3. Main characters of mainland Oligodon species with 15 dorsal scale rows at midbody. Exceptional values are placed in brackets. See Materials and Methods for abbreviations, with the addition of: Cloacal plate: D—divided; E—entire;? — data unavailable." httpUri="http://table.plazi.org/id/DF665CB41152BB5EFF00FEF9FEA22192" pageId="9" pageNumber="54" tableUuid="DF665CB41152BB5EFF00FEF9FEA22192">Tables 3</tableCitation>
|
|||
|
|
and 4.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF665CB41152BB5EFF00FEF9FEA22192" ID-Table-UUID="DF665CB41152BB5EFF00FEF9FEA22192" httpUri="http://table.plazi.org/id/DF665CB41152BB5EFF00FEF9FEA22192" pageId="9" pageNumber="54" targetBox="[151,1436,464,1022]" targetIsTable="true" targetPageId="9">
|
|||
|
|
<paragraph id="8BA60C3C1152BB5EFF00FEF9FEA22192" blockId="9.[151,1436,355,439]" pageId="9" pageNumber="54">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF00FEF9FEB5215C" bold="true" box="[151,260,356,377]" pageId="9" pageNumber="54">TABLE 3.</emphasis>
|
|||
|
|
Main characters of mainland
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFDA4FEF9FD23215F" box="[563,658,356,378]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFDA4FEF9FD23215F" box="[563,658,356,378]" italics="true" pageId="9" pageNumber="54">Oligodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species with 15 dorsal scale rows at midbody. Exceptional values are placed in brackets. See Materials and Methods for abbreviations, with the addition of: Cloacal plate: D—divided; E—entire;?—data unavailable.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA60C3C1152BB5EFF0BFE4DFAE923DB" pageId="9" pageNumber="54">
|
|||
|
|
<table id="F919FE9C115244A8FF00FE4DFA2D23DB" box="[151,1436,464,1022]" gridcols="6" gridrows="9" pageId="9" pageNumber="54">
|
|||
|
|
<tr id="35290E7E115244A8FF00FE4DFA2D220C" box="[151,1436,464,553]" gridrow="0" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FE4DFE10220C" box="[151,417,464,553]" gridcol="0" gridrow="0" pageId="9" pageNumber="54">
|
|||
|
|
Species
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFD8FFEFB220C" box="[156,330,530,553]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0BFD8FFEFB220C" box="[156,330,530,553]" italics="true" pageId="9" pageNumber="54">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551152BB5EFEC6FD89FE10220C" box="[337,417,532,553]" pageId="9" pageNumber="54" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFEC6FD89FE10220C" bold="true" box="[337,417,532,553]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
</th>
|
|||
|
|
<th id="76F86702115244A8FE7AFE4DFDEE220C" box="[493,607,464,553]" gridcol="1" gridrow="0" pageId="9" pageNumber="54">Hemipenes Not forked</th>
|
|||
|
|
<th id="76F86702115244A8FD27FE4DFC03220C" box="[688,946,464,553]" gridcol="2" gridrow="0" pageId="9" pageNumber="54">Hemipenes length DEN (SC) 16 9–10</th>
|
|||
|
|
<th id="76F86702115244A8FC73FE4DFB80220C" box="[996,1073,464,553]" gridcol="3" gridrow="0" pageId="9" pageNumber="54">Cloacal plate E</th>
|
|||
|
|
<th id="76F86702115244A8FBF8FE4DFB78220C" box="[1135,1225,464,553]" gridcol="4" gridrow="0" pageId="9" pageNumber="54">VEN 184–193</th>
|
|||
|
|
<th id="76F86702115244A8FA8EFE4DFA2D220C" box="[1305,1436,464,553]" gridcol="5" gridrow="0" pageId="9" pageNumber="54">SC 43–47</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FDA5FA2D2257" box="[151,1436,568,626]" gridrow="1" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FDA5FE102257" box="[151,417,568,626]" gridcol="0" gridrow="1" pageId="9" pageNumber="54">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0BFDA4FEBC2257" italics="true" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFDA4FE9D226B" box="[156,300,568,590]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="brevicauda">O. brevicauda</taxonomicName>
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFDC1FEBC2257" box="[156,269,604,626]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="dorsalis">O. dorsalis</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FE7AFDA5FDEE2257" box="[493,607,568,626]" gridcol="1" gridrow="1" pageId="9" pageNumber="54">? Forked</td>
|
|||
|
|
<td id="76F86702115244A8FD27FDA5FC032257" box="[688,946,568,626]" gridcol="2" gridrow="1" pageId="9" pageNumber="54">? 7–8 11 6–7</td>
|
|||
|
|
<td id="76F86702115244A8FC73FDA5FB802257" box="[996,1073,568,626]" gridcol="3" gridrow="1" pageId="9" pageNumber="54">D D</td>
|
|||
|
|
<td id="76F86702115244A8FBF8FDA5FB782257" box="[1135,1225,568,626]" gridcol="4" gridrow="1" pageId="9" pageNumber="54">158–176 162–188</td>
|
|||
|
|
<td id="76F86702115244A8FA8EFDA5FA2D2257" box="[1305,1436,568,626]" gridcol="5" gridrow="1" pageId="9" pageNumber="54">25–29 27–51</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FD1DFA2D229F" box="[151,1436,640,698]" gridrow="2" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FD1DFE10229F" box="[151,417,640,698]" gridcol="0" gridrow="2" pageId="9" pageNumber="54">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0BFD1CFEAB229F" italics="true" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFD1CFEF222B3" box="[156,323,640,662]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="erythrorachis">O. erythrorachis</taxonomicName>
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFD39FEAB229F" box="[156,282,676,698]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="hamptoni">O. hamptoni</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FE7AFD1DFDEE229F" box="[493,607,640,698]" gridcol="1" gridrow="2" pageId="9" pageNumber="54">? Not forked</td>
|
|||
|
|
<td id="76F86702115244A8FD27FD1DFC03229F" box="[688,946,640,698]" gridcol="2" gridrow="2" pageId="9" pageNumber="54">? 7–8 11 7–8</td>
|
|||
|
|
<td id="76F86702115244A8FC73FD1DFB80229F" box="[996,1073,640,698]" gridcol="3" gridrow="2" pageId="9" pageNumber="54">D D</td>
|
|||
|
|
<td id="76F86702115244A8FBF8FD1DFB78229F" box="[1135,1225,640,698]" gridcol="4" gridrow="2" pageId="9" pageNumber="54">154 160–175</td>
|
|||
|
|
<td id="76F86702115244A8FA8EFD1DFA2D229F" box="[1305,1436,640,698]" gridcol="5" gridrow="2" pageId="9" pageNumber="54">46 30–32</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FD55FA2D2327" box="[151,1436,712,770]" gridrow="3" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FD55FE102327" box="[151,417,712,770]" gridcol="0" gridrow="3" pageId="9" pageNumber="54">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0BFD54FE912327" italics="true" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFD54FEAD22FB" box="[156,284,713,734]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="inornatus">O. inornatus</taxonomicName>
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFD71FE912327" box="[156,288,748,770]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="jintakunei">O. jintakunei</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FE7AFD55FDEE2327" box="[493,607,712,770]" gridcol="1" gridrow="3" pageId="9" pageNumber="54">Not forked?</td>
|
|||
|
|
<td id="76F86702115244A8FD27FD55FC032327" box="[688,946,712,770]" gridcol="2" gridrow="3" pageId="9" pageNumber="54">10–12? 11–12? 6</td>
|
|||
|
|
<td id="76F86702115244A8FC73FD55FB802327" box="[996,1073,712,770]" gridcol="3" gridrow="3" pageId="9" pageNumber="54">E D</td>
|
|||
|
|
<td id="76F86702115244A8FBF8FD55FB782327" box="[1135,1225,712,770]" gridcol="4" gridrow="3" pageId="9" pageNumber="54">169–174 189</td>
|
|||
|
|
<td id="76F86702115244A8FA8EFD55FA2D2327" box="[1305,1436,712,770]" gridcol="5" gridrow="3" pageId="9" pageNumber="54">31–42 46</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FC8DFA2D236F" box="[151,1436,784,842]" gridrow="4" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FC8DFE10236F" box="[151,417,784,842]" gridcol="0" gridrow="4" pageId="9" pageNumber="54">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0BFC8CFEF8236F" italics="true" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFC8CFEB82303" box="[156,265,784,806]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="lacroixi">O. lacroixi</taxonomicName>
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFCA9FEF8236F" box="[156,329,820,842]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="lungshenensis">O. lungshenensis</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FE7AFC8DFDEE236F" box="[493,607,784,842]" gridcol="1" gridrow="4" pageId="9" pageNumber="54">Not forked Not forked</td>
|
|||
|
|
<td id="76F86702115244A8FD27FC8DFC03236F" box="[688,946,784,842]" gridcol="2" gridrow="4" pageId="9" pageNumber="54">6–8 8–12 10?</td>
|
|||
|
|
<td id="76F86702115244A8FC73FC8DFB80236F" box="[996,1073,784,842]" gridcol="3" gridrow="4" pageId="9" pageNumber="54">D D</td>
|
|||
|
|
<td id="76F86702115244A8FBF8FC8DFB78236F" box="[1135,1225,784,842]" gridcol="4" gridrow="4" pageId="9" pageNumber="54">162–178 166–180</td>
|
|||
|
|
<td id="76F86702115244A8FA8EFC8DFA2D236F" box="[1305,1436,784,842]" gridcol="5" gridrow="4" pageId="9" pageNumber="54">29–? 31–38</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FCC5FA2D234B" box="[151,1436,856,878]" gridrow="5" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FCC5FE10234B" box="[151,417,856,878]" gridcol="0" gridrow="5" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFCC4FEAA234B" box="[156,283,856,878]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="melaneus">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0BFCC4FEAA234B" box="[156,283,856,878]" italics="true" pageId="9" pageNumber="54">O. melaneus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FE7AFCC5FDEE234B" box="[493,607,856,878]" gridcol="1" gridrow="5" pageId="9" pageNumber="54">Not forked</td>
|
|||
|
|
<td id="76F86702115244A8FD27FCC5FC03234B" box="[688,946,856,878]" gridcol="2" gridrow="5" pageId="9" pageNumber="54">15 7</td>
|
|||
|
|
<td id="76F86702115244A8FC73FCC5FB80234B" box="[996,1073,856,878]" gridcol="3" gridrow="5" pageId="9" pageNumber="54">D</td>
|
|||
|
|
<td id="76F86702115244A8FBF8FCC5FB78234B" box="[1135,1225,856,878]" gridcol="4" gridrow="5" pageId="9" pageNumber="54">152–160</td>
|
|||
|
|
<td id="76F86702115244A8FA8EFCC5FA2D234B" box="[1305,1436,856,878]" gridcol="5" gridrow="5" pageId="9" pageNumber="54">39–40</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FCE1FA2D23B7" box="[151,1436,892,914]" gridrow="6" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FCE1FE1023B7" box="[151,417,892,914]" gridcol="0" gridrow="6" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFCE1FF4D23B7" box="[156,252,892,914]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="nikhili">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0BFCE1FF4D23B7" box="[156,252,892,914]" italics="true" pageId="9" pageNumber="54">O. nikhili</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FE7AFCE1FDEE23B7" box="[493,607,892,914]" gridcol="1" gridrow="6" pageId="9" pageNumber="54">?</td>
|
|||
|
|
<td id="76F86702115244A8FD27FCE1FC0323B7" box="[688,946,892,914]" gridcol="2" gridrow="6" pageId="9" pageNumber="54">??</td>
|
|||
|
|
<td id="76F86702115244A8FC73FCE1FB8023B7" box="[996,1073,892,914]" gridcol="3" gridrow="6" pageId="9" pageNumber="54">D</td>
|
|||
|
|
<td id="76F86702115244A8FBF8FCE1FB7823B7" box="[1135,1225,892,914]" gridcol="4" gridrow="6" pageId="9" pageNumber="54">144</td>
|
|||
|
|
<td id="76F86702115244A8FA8EFCE1FA2D23B7" box="[1305,1436,892,914]" gridcol="5" gridrow="6" pageId="9" pageNumber="54">33</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FC3DFA2D23FF" box="[151,1436,928,986]" gridrow="7" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FC3DFE1023FF" box="[151,417,928,986]" gridcol="0" gridrow="7" pageId="9" pageNumber="54">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0BFC3CFE9A23FF" italics="true" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFC3CFEB92393" box="[156,264,929,950]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="ornatus">O. ornatus</taxonomicName>
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFC59FE9A23FF" box="[156,299,964,986]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="taeniolatus">O. taeniolatus</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FE7AFC3DFDEE23FF" box="[493,607,928,986]" gridcol="1" gridrow="7" pageId="9" pageNumber="54">Not forked Forked</td>
|
|||
|
|
<td id="76F86702115244A8FD27FC3DFC0323FF" box="[688,946,928,986]" gridcol="2" gridrow="7" pageId="9" pageNumber="54">9 6–8 9–11 6–7</td>
|
|||
|
|
<td id="76F86702115244A8FC73FC3DFB8023FF" box="[996,1073,928,986]" gridcol="3" gridrow="7" pageId="9" pageNumber="54">D D</td>
|
|||
|
|
<td id="76F86702115244A8FBF8FC3DFB7823FF" box="[1135,1225,928,986]" gridcol="4" gridrow="7" pageId="9" pageNumber="54">156–182 158–218</td>
|
|||
|
|
<td id="76F86702115244A8FA8EFC3DFA2D23FF" box="[1305,1436,928,986]" gridcol="5" gridrow="7" pageId="9" pageNumber="54">27–44 29–59</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FC75FA2D23DB" box="[151,1436,1000,1022]" gridrow="8" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FC75FE1023DB" box="[151,417,1000,1022]" gridcol="0" gridrow="8" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF0BFC74FEAA23DB" box="[156,283,1001,1022]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="torquatus">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0BFC74FEAA23DB" box="[156,283,1001,1022]" italics="true" pageId="9" pageNumber="54">O. torquatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FE7AFC75FDEE23DB" box="[493,607,1000,1022]" gridcol="1" gridrow="8" pageId="9" pageNumber="54">Not forked</td>
|
|||
|
|
<td id="76F86702115244A8FD27FC75FC0323DB" box="[688,946,1000,1022]" gridcol="2" gridrow="8" pageId="9" pageNumber="54">8 15–16</td>
|
|||
|
|
<td id="76F86702115244A8FC73FC75FB8023DB" box="[996,1073,1000,1022]" gridcol="3" gridrow="8" pageId="9" pageNumber="54">D</td>
|
|||
|
|
<td id="76F86702115244A8FBF8FC75FB7823DB" box="[1135,1225,1000,1022]" gridcol="4" gridrow="8" pageId="9" pageNumber="54">114–169</td>
|
|||
|
|
<td id="76F86702115244A8FA8EFC75FA2D23DB" box="[1305,1436,1000,1022]" gridcol="5" gridrow="8" pageId="9" pageNumber="54">25–34</td>
|
|||
|
|
</tr>
|
|||
|
|
</table>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1152BB5EFF00FBBFFABB265A" pageId="9" pageNumber="54">
|
|||
|
|
<table id="F919FE9C115244A8FF00FBBFFACF265A" box="[151,1406,1058,1663]" gridcols="6" gridrows="15" pageId="9" pageNumber="54">
|
|||
|
|
<tr id="35290E7E115244A8FF00FBBFFACF241D" box="[151,1406,1058,1080]" gridrow="0" pageId="9" pageNumber="54" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
|||
|
|
<th id="76F86702115244A8FF00FBBFFE46241D" box="[151,503,1058,1080]" gridcol="0" gridrow="0" pageId="9" pageNumber="54">continued.</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FBCCFACF248E" box="[151,1406,1105,1195]" gridrow="1" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FBCCFE46248E" box="[151,503,1105,1195]" gridcol="0" gridrow="1" pageId="9" pageNumber="54">
|
|||
|
|
Species SL
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FB09FEFC248E" box="[159,333,1172,1195]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FB09FEFC248E" box="[159,333,1172,1195]" italics="true" pageId="9" pageNumber="54">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551152BB5EFEC3FB08FE15248F" box="[340,420,1173,1194]" pageId="9" pageNumber="54" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFEC3FB08FE15248F" bold="true" box="[340,420,1173,1194]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
7 or 8
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FBCCFDFB248E" box="[547,586,1105,1195]" gridcol="1" gridrow="1" pageId="9" pageNumber="54">InN 1</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FBCCFD1D248E" box="[624,684,1105,1195]" gridcol="2" gridrow="1" pageId="9" pageNumber="54">Lor 1</td>
|
|||
|
|
<td id="76F86702115244A8FD49FBCCFCB3248E" box="[734,770,1105,1195]" gridcol="3" gridrow="1" pageId="9" pageNumber="54">Ate 1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FBCCFC60248E" box="[837,977,1105,1195]" gridcol="4" gridrow="1" pageId="9" pageNumber="54">Ratio TaL/TL (males) 0.135–0.146</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFBCCFACF248E" box="[1069,1406,1105,1195]" gridcol="5" gridrow="1" pageId="9" pageNumber="54">Dorsal pattern Dark blotches</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FB24FACF24EA" box="[151,1406,1209,1231]" gridrow="2" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FB24FE4624EA" box="[151,503,1209,1231]" gridcol="0" gridrow="2" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FB27FE9E24EA" box="[159,303,1209,1231]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="brevicauda">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FB27FE9E24EA" box="[159,303,1209,1231]" italics="true" pageId="9" pageNumber="54">O. brevicauda</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
7
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FB24FDFB24EA" box="[547,586,1209,1231]" gridcol="1" gridrow="2" pageId="9" pageNumber="54">0</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FB24FD1D24EA" box="[624,684,1209,1231]" gridcol="2" gridrow="2" pageId="9" pageNumber="54">0</td>
|
|||
|
|
<td id="76F86702115244A8FD49FB24FCB324EA" box="[734,770,1209,1231]" gridcol="3" gridrow="2" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FB24FC6024EA" box="[837,977,1209,1231]" gridcol="4" gridrow="2" pageId="9" pageNumber="54">?</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFB24FACF24EA" box="[1069,1406,1209,1231]" gridcol="5" gridrow="2" pageId="9" pageNumber="54">Stripes + spots</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FB40FACF24D6" box="[151,1406,1245,1267]" gridrow="3" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FB40FE4624D6" box="[151,503,1245,1267]" gridcol="0" gridrow="3" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FB43FEA124D6" box="[159,272,1245,1267]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="dorsalis">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FB43FEA124D6" box="[159,272,1245,1267]" italics="true" pageId="9" pageNumber="54">O. dorsalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
7
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FB40FDFB24D6" box="[547,586,1245,1267]" gridcol="1" gridrow="3" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FB40FD1D24D6" box="[624,684,1245,1267]" gridcol="2" gridrow="3" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FD49FB40FCB324D6" box="[734,770,1245,1267]" gridcol="3" gridrow="3" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FB40FC6024D6" box="[837,977,1245,1267]" gridcol="4" gridrow="3" pageId="9" pageNumber="54">?</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFB40FACF24D6" box="[1069,1406,1245,1267]" gridcol="5" gridrow="3" pageId="9" pageNumber="54">Stripes</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FA9CFACF2532" box="[151,1406,1281,1303]" gridrow="4" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FA9CFE462532" box="[151,503,1281,1303]" gridcol="0" gridrow="4" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FA9FFEF62532" box="[159,327,1281,1303]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="erythrorachis">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FA9FFEF62532" box="[159,327,1281,1303]" italics="true" pageId="9" pageNumber="54">O. erythrorachis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
7
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FA9CFDFB2532" box="[547,586,1281,1303]" gridcol="1" gridrow="4" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FA9CFD1D2532" box="[624,684,1281,1303]" gridcol="2" gridrow="4" pageId="9" pageNumber="54">0</td>
|
|||
|
|
<td id="76F86702115244A8FD49FA9CFCB32532" box="[734,770,1281,1303]" gridcol="3" gridrow="4" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FA9CFC602532" box="[837,977,1281,1303]" gridcol="4" gridrow="4" pageId="9" pageNumber="54">?</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFA9CFACF2532" box="[1069,1406,1281,1303]" gridcol="5" gridrow="4" pageId="9" pageNumber="54">Vertebral stripe + black crossbars</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FAB8FACF251E" box="[151,1406,1317,1339]" gridrow="5" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FAB8FE46251E" box="[151,503,1317,1339]" gridcol="0" gridrow="5" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FABBFEAF251E" box="[159,286,1317,1339]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="hamptoni">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FABBFEAF251E" box="[159,286,1317,1339]" italics="true" pageId="9" pageNumber="54">O. hamptoni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
5
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FAB8FDFB251E" box="[547,586,1317,1339]" gridcol="1" gridrow="5" pageId="9" pageNumber="54">0</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FAB8FD1D251E" box="[624,684,1317,1339]" gridcol="2" gridrow="5" pageId="9" pageNumber="54">0 or 1</td>
|
|||
|
|
<td id="76F86702115244A8FD49FAB8FCB3251E" box="[734,770,1317,1339]" gridcol="3" gridrow="5" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FAB8FC60251E" box="[837,977,1317,1339]" gridcol="4" gridrow="5" pageId="9" pageNumber="54">?</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFAB8FACF251E" box="[1069,1406,1317,1339]" gridcol="5" gridrow="5" pageId="9" pageNumber="54">Stripes</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FAD4FACF257A" box="[151,1406,1353,1375]" gridrow="6" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FAD4FE46257A" box="[151,503,1353,1375]" gridcol="0" gridrow="6" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FAD7FEAE257A" box="[159,287,1354,1375]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="inornatus">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FAD7FEAE257A" box="[159,287,1354,1375]" italics="true" pageId="9" pageNumber="54">O. inornatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
8
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FAD4FDFB257A" box="[547,586,1353,1375]" gridcol="1" gridrow="6" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FAD4FD1D257A" box="[624,684,1353,1375]" gridcol="2" gridrow="6" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FD49FAD4FCB3257A" box="[734,770,1353,1375]" gridcol="3" gridrow="6" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FAD4FC60257A" box="[837,977,1353,1375]" gridcol="4" gridrow="6" pageId="9" pageNumber="54">0.141–0.144</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFAD4FACF257A" box="[1069,1406,1353,1375]" gridcol="5" gridrow="6" pageId="9" pageNumber="54">Uniform or reticulations</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FAF0FACF25A6" box="[151,1406,1389,1411]" gridrow="7" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FAF0FE4625A6" box="[151,503,1389,1411]" gridcol="0" gridrow="7" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FAF3FE9225A6" box="[159,291,1389,1411]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="jintakunei">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FAF3FE9225A6" box="[159,291,1389,1411]" italics="true" pageId="9" pageNumber="54">O. jintakunei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
7
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FAF0FDFB25A6" box="[547,586,1389,1411]" gridcol="1" gridrow="7" pageId="9" pageNumber="54">0</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FAF0FD1D25A6" box="[624,684,1389,1411]" gridcol="2" gridrow="7" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FD49FAF0FCB325A6" box="[734,770,1389,1411]" gridcol="3" gridrow="7" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FAF0FC6025A6" box="[837,977,1389,1411]" gridcol="4" gridrow="7" pageId="9" pageNumber="54">----</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFAF0FACF25A6" box="[1069,1406,1389,1411]" gridcol="5" gridrow="7" pageId="9" pageNumber="54">Pale rings</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FA0CFACF2582" box="[151,1406,1425,1447]" gridrow="8" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FA0CFE462582" box="[151,503,1425,1447]" gridcol="0" gridrow="8" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FA0FFEBD2582" box="[159,268,1425,1447]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="lacroixi">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FA0FFEBD2582" box="[159,268,1425,1447]" italics="true" pageId="9" pageNumber="54">O. lacroixi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
5
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FA0CFDFB2582" box="[547,586,1425,1447]" gridcol="1" gridrow="8" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FA0CFD1D2582" box="[624,684,1425,1447]" gridcol="2" gridrow="8" pageId="9" pageNumber="54">0</td>
|
|||
|
|
<td id="76F86702115244A8FD49FA0CFCB32582" box="[734,770,1425,1447]" gridcol="3" gridrow="8" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FA0CFC602582" box="[837,977,1425,1447]" gridcol="4" gridrow="8" pageId="9" pageNumber="54">?</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFA0CFACF2582" box="[1069,1406,1425,1447]" gridcol="5" gridrow="8" pageId="9" pageNumber="54">Spots + stripes</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FA28FACF25EE" box="[151,1406,1461,1483]" gridrow="9" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FA28FE4625EE" box="[151,503,1461,1483]" gridcol="0" gridrow="9" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FA2BFEFD25EE" box="[159,332,1461,1483]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="lungshenensis">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FA2BFEFD25EE" box="[159,332,1461,1483]" italics="true" pageId="9" pageNumber="54">O. lungshenensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
6
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FA28FDFB25EE" box="[547,586,1461,1483]" gridcol="1" gridrow="9" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FA28FD1D25EE" box="[624,684,1461,1483]" gridcol="2" gridrow="9" pageId="9" pageNumber="54">0</td>
|
|||
|
|
<td id="76F86702115244A8FD49FA28FCB325EE" box="[734,770,1461,1483]" gridcol="3" gridrow="9" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FA28FC6025EE" box="[837,977,1461,1483]" gridcol="4" gridrow="9" pageId="9" pageNumber="54">?</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFA28FACF25EE" box="[1069,1406,1461,1483]" gridcol="5" gridrow="9" pageId="9" pageNumber="54">Crossbars</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FA44FACF25CA" box="[151,1406,1497,1519]" gridrow="10" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FA44FE4625CA" box="[151,503,1497,1519]" gridcol="0" gridrow="10" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FA47FEAF25CA" box="[159,286,1497,1519]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="melaneus">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FA47FEAF25CA" box="[159,286,1497,1519]" italics="true" pageId="9" pageNumber="54">O. melaneus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
7
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FA44FDFB25CA" box="[547,586,1497,1519]" gridcol="1" gridrow="10" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FA44FD1D25CA" box="[624,684,1497,1519]" gridcol="2" gridrow="10" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FD49FA44FCB325CA" box="[734,770,1497,1519]" gridcol="3" gridrow="10" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FA44FC6025CA" box="[837,977,1497,1519]" gridcol="4" gridrow="10" pageId="9" pageNumber="54">?</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFA44FACF25CA" box="[1069,1406,1497,1519]" gridcol="5" gridrow="10" pageId="9" pageNumber="54">Dark with speckling</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00FA60FACF2636" box="[151,1406,1533,1555]" gridrow="11" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00FA60FE462636" box="[151,503,1533,1555]" gridcol="0" gridrow="11" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08FA63FF4E2636" box="[159,255,1533,1555]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="nikhili">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08FA63FF4E2636" box="[159,255,1533,1555]" italics="true" pageId="9" pageNumber="54">O. nikhili</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
7
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4FA60FDFB2636" box="[547,586,1533,1555]" gridcol="1" gridrow="11" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FDE7FA60FD1D2636" box="[624,684,1533,1555]" gridcol="2" gridrow="11" pageId="9" pageNumber="54">0</td>
|
|||
|
|
<td id="76F86702115244A8FD49FA60FCB32636" box="[734,770,1533,1555]" gridcol="3" gridrow="11" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2FA60FC602636" box="[837,977,1533,1555]" gridcol="4" gridrow="11" pageId="9" pageNumber="54">0.173</td>
|
|||
|
|
<td id="76F86702115244A8FBBAFA60FACF2636" box="[1069,1406,1533,1555]" gridcol="5" gridrow="11" pageId="9" pageNumber="54">Stripes</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00F9BCFACF2612" box="[151,1406,1569,1591]" gridrow="12" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00F9BCFE462612" box="[151,503,1569,1591]" gridcol="0" gridrow="12" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08F9BFFEBA2612" box="[159,267,1570,1591]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="ornatus">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08F9BFFEBA2612" box="[159,267,1570,1591]" italics="true" pageId="9" pageNumber="54">O. ornatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
6–7
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4F9BCFDFB2612" box="[547,586,1569,1591]" gridcol="1" gridrow="12" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FDE7F9BCFD1D2612" box="[624,684,1569,1591]" gridcol="2" gridrow="12" pageId="9" pageNumber="54">0</td>
|
|||
|
|
<td id="76F86702115244A8FD49F9BCFCB32612" box="[734,770,1569,1591]" gridcol="3" gridrow="12" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2F9BCFC602612" box="[837,977,1569,1591]" gridcol="4" gridrow="12" pageId="9" pageNumber="54">0.146–0.182</td>
|
|||
|
|
<td id="76F86702115244A8FBBAF9BCFACF2612" box="[1069,1406,1569,1591]" gridcol="5" gridrow="12" pageId="9" pageNumber="54">Widely separated blotches</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00F9D8FACF267E" box="[151,1406,1605,1627]" gridrow="13" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00F9D8FE46267E" box="[151,503,1605,1627]" gridcol="0" gridrow="13" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08F9DBFE9E267E" box="[159,303,1605,1627]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="taeniolatus">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08F9DBFE9E267E" box="[159,303,1605,1627]" italics="true" pageId="9" pageNumber="54">O. taeniolatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
7
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4F9D8FDFB267E" box="[547,586,1605,1627]" gridcol="1" gridrow="13" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FDE7F9D8FD1D267E" box="[624,684,1605,1627]" gridcol="2" gridrow="13" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FD49F9D8FCB3267E" box="[734,770,1605,1627]" gridcol="3" gridrow="13" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2F9D8FC60267E" box="[837,977,1605,1627]" gridcol="4" gridrow="13" pageId="9" pageNumber="54">?</td>
|
|||
|
|
<td id="76F86702115244A8FBBAF9D8FACF267E" box="[1069,1406,1605,1627]" gridcol="5" gridrow="13" pageId="9" pageNumber="54">Blotched or spotted</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="35290E7E115244A8FF00F9F4FACF265A" box="[151,1406,1641,1663]" gridrow="14" pageId="9" pageNumber="54">
|
|||
|
|
<th id="76F86702115244A8FF00F9F4FE46265A" box="[151,503,1641,1663]" gridcol="0" gridrow="14" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF08F9F7FEAF265A" box="[159,286,1642,1663]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="torquatus">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF08F9F7FEAF265A" box="[159,286,1642,1663]" italics="true" pageId="9" pageNumber="54">O. torquatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
7
|
|||
|
|
</th>
|
|||
|
|
<td id="76F86702115244A8FDB4F9F4FDFB265A" box="[547,586,1641,1663]" gridcol="1" gridrow="14" pageId="9" pageNumber="54">2</td>
|
|||
|
|
<td id="76F86702115244A8FDE7F9F4FD1D265A" box="[624,684,1641,1663]" gridcol="2" gridrow="14" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FD49F9F4FCB3265A" box="[734,770,1641,1663]" gridcol="3" gridrow="14" pageId="9" pageNumber="54">1</td>
|
|||
|
|
<td id="76F86702115244A8FCD2F9F4FC60265A" box="[837,977,1641,1663]" gridcol="4" gridrow="14" pageId="9" pageNumber="54">?</td>
|
|||
|
|
<td id="76F86702115244A8FBBAF9F4FACF265A" box="[1069,1406,1641,1663]" gridcol="5" gridrow="14" pageId="9" pageNumber="54">Stripes + reticulations</td>
|
|||
|
|
</tr>
|
|||
|
|
</table>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1152BB5EFF52F959FC542743" blockId="9.[153,1436,1732,1895]" pageId="9" pageNumber="54">
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF52F959FECC26F9" box="[197,381,1732,1756]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF52F959FECC26F9" box="[197,381,1732,1756]" italics="true" pageId="9" pageNumber="54">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551152BB5EFE12F959FE5026F9" box="[389,481,1732,1756]" pageId="9" pageNumber="54" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFE12F959FE5026F9" bold="true" box="[389,481,1732,1756]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
might be confused only with
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFCAFF958FC0A26F9" box="[824,955,1733,1756]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFCAFF958FC0A26F9" box="[824,955,1733,1756]" italics="true" pageId="9" pageNumber="54">
|
|||
|
|
<collectingCountry id="F30E4CAC1152BB5EFCAFF958FCFE26F9" box="[824,847,1733,1756]" name="Iceland" pageId="9" pageNumber="54">O</collectingCountry>
|
|||
|
|
. joynsoni
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, of which we examined five specimens to which we add the four specimens identified by
|
|||
|
|
<bibRefCitation id="EF8871CD1152BB5EFD7AF97BFC3E26DB" author="Taylor" box="[749,911,1766,1790]" pageId="9" pageNumber="54" refString="Taylor, E. H. (1965) The serpents of Thailand and adjacent waters. University of Kansas Science Bulletin, 45 (9), 609 - 1096." type="journal article" year="1965">Taylor (1965)</bibRefCitation>
|
|||
|
|
as
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFC52F97AFB7C26DB" box="[965,1229,1766,1790]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="subSpecies" species="cinereus" subSpecies="swinhonis">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFC52F97AFB7C26DB" box="[965,1229,1766,1790]" italics="true" pageId="9" pageNumber="54">
|
|||
|
|
<collectingCountry id="F30E4CAC1152BB5EFC52F97AFC6D26DB" box="[965,988,1767,1790]" name="Iceland" pageId="9" pageNumber="54">O</collectingCountry>
|
|||
|
|
. cinereus swinhonis
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFA83F97AFE872704" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="subSpecies" species="cinereus" subSpecies="multifasciatus">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFA83F97AFE872704" italics="true" pageId="9" pageNumber="54">
|
|||
|
|
<collectingCountry id="F30E4CAC1152BB5EFA83F97AFA9A26DB" box="[1300,1323,1767,1790]" name="Iceland" pageId="9" pageNumber="54">O</collectingCountry>
|
|||
|
|
. cinereus multifasciatus
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(see
|
|||
|
|
<bibRefCitation id="EF8871CD1152BB5EFEE7F894FD842704" author="David" box="[368,565,1801,1825]" pageId="9" pageNumber="54" refString="David, P., Das, I. & Vogel, G. (2011) On some taxonomic and nomenclatural problems in Indian species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae). Zootaxa, 2799, 1 - 14." type="journal article" year="2011">
|
|||
|
|
David
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFE2BF897FE452704" box="[444,500,1801,1825]" italics="true" pageId="9" pageNumber="54">et al.</emphasis>
|
|||
|
|
2011
|
|||
|
|
</bibRefCitation>
|
|||
|
|
). However none of these specimens has a blotched pattern as
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFB72F894FA2D2704" box="[1253,1436,1801,1825]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFB72F894FA2D2704" box="[1253,1436,1801,1825]" italics="true" pageId="9" pageNumber="54">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551152BB5EFF0EF8B1FF462761" box="[153,247,1836,1860]" pageId="9" pageNumber="54" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF0EF8B1FF462761" bold="true" box="[153,247,1836,1860]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
On the basis of this constant character, together with differences in dentition and head scalation, we consider
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFF68F8D3FE042742" box="[255,437,1870,1895]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFF68F8D3FE042742" box="[255,437,1870,1895]" italics="true" pageId="9" pageNumber="54">Oligodon nagao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25E6D551152BB5EFE2BF8D2FDA72742" box="[444,534,1871,1895]" pageId="9" pageNumber="54" rank="species">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFE2BF8D2FDA72742" bold="true" box="[444,534,1871,1895]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
to be a species distinct from
|
|||
|
|
<taxonomicName id="4C1977BF1152BB5EFCCAF8D2FC6F2743" box="[861,990,1871,1894]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="joynsoni">
|
|||
|
|
<emphasis id="B96DD02E1152BB5EFCCAF8D2FC6F2743" box="[861,990,1871,1894]" italics="true" pageId="9" pageNumber="54">
|
|||
|
|
<collectingCountry id="F30E4CAC1152BB5EFCCAF8D2FCC52743" box="[861,884,1871,1894]" name="Iceland" pageId="9" pageNumber="54">O</collectingCountry>
|
|||
|
|
. joynsoni
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3035FB71150BB5CFF52FF05FAFE2139" pageId="11" pageNumber="56" type="etymology">
|
|||
|
|
<paragraph id="8BA60C3C1150BB5CFF52FF05FB3220F1" blockId="11.[151,1437,151,824]" pageId="11" pageNumber="56">
|
|||
|
|
<emphasis id="B96DD02E1150BB5CFF52FF05FEF52095" bold="true" box="[197,324,152,176]" pageId="11" pageNumber="56">Etymology</emphasis>
|
|||
|
|
. This new species is named in honour of the
|
|||
|
|
<taxonomicName id="4C1977BF1150BB5CFCC6FF05FC2B2095" box="[849,922,152,176]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="59" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">Nagao</taxonomicName>
|
|||
|
|
Natural Environment Foundation,
|
|||
|
|
<collectingCountry id="F30E4CAC1150BB5CFABCFF05FADF208A" box="[1323,1390,152,175]" name="Japan" pageId="11" pageNumber="56">Japan</collectingCountry>
|
|||
|
|
, for the support of natural sciences research and conservation in developing countries in Asia.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA60C3C1150BB5CFF00FF7DFAFE2139" blockId="11.[151,1437,151,824]" pageId="11" pageNumber="56">
|
|||
|
|
<collectingCountry id="F30E4CAC1150BB5CFF00FF7DFF0720DD" box="[151,182,224,248]" name="American Samoa" pageId="11" pageNumber="56">As</collectingCountry>
|
|||
|
|
common names, we suggest
|
|||
|
|
<taxonomicName id="4C1977BF1150BB5CFD9FFF7DFDE020DD" box="[520,593,224,248]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="59" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">Nagao</taxonomicName>
|
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Kukri Snake (English name),
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<emphasis id="B96DD02E1150BB5CFC38FF7CFC7120DD" box="[943,960,225,248]" italics="true" pageId="11" pageNumber="56">R</emphasis>
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ắ
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<emphasis id="B96DD02E1150BB5CFC59FF7CFBB620DD" box="[974,1031,224,248]" italics="true" pageId="11" pageNumber="56">n khi</emphasis>
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ế
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<emphasis id="B96DD02E1150BB5CFB83FF7CFB3120DD" box="[1044,1152,225,248]" italics="true" pageId="11" pageNumber="56">m na-gao</emphasis>
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(Vietnamese name), and
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<emphasis id="B96DD02E1150BB5CFF00FE98FE102139" box="[151,417,260,284]" italics="true" pageId="11" pageNumber="56">Fang Ban Xiao Tou She</emphasis>
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(Chinese name, meaning the “Kukri Snake with rectangular blotches on the back”).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3035FB71150BB5CFF52FEBAFAA72312" pageId="11" pageNumber="56" type="distribution">
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<paragraph id="8BA60C3C1150BB5CFF52FEBAFD292141" blockId="11.[151,1437,151,824]" pageId="11" pageNumber="56">
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<emphasis id="B96DD02E1150BB5CFF52FEBAFE052165" bold="true" box="[197,436,295,320]" pageId="11" pageNumber="56">
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Distribution (
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<figureCitation id="132210B91150BB5CFEF0FEBAFE1C2165" box="[359,429,295,320]" captionStart="FIGURE 6" captionStartId="11.[151,249,1932,1955]" captionTargetBox="[169,1414,874,1905]" captionTargetId="figure@11.[163,1424,863,1914]" captionTargetPageId="11" captionText="FIGURE 6. Distribution of Oligodon nagao sp. nov.: (1) Huu Lien Nature Reserve in Lang Son Province and (2) Ha Lang forest in Cao Bang Province, Vietnam; (3) Nonggang National Nature Reserve in Guangxi Autonomous Region, China; and (4) Ban Nathan in Khammouane Province, Laos." httpUri="https://zenodo.org/record/214044/files/figure.png" pageId="11" pageNumber="56">Fig. 6</figureCitation>
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)
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</emphasis>
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.
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<taxonomicName id="4C1977BF1150BB5CFE54FEB5FDCB2165" box="[451,634,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="nagao" status="sp. nov.">
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<emphasis id="B96DD02E1150BB5CFE54FEB5FDCB2165" box="[451,634,296,320]" italics="true" pageId="11" pageNumber="56">Oligodon nagao</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25E6D551150BB5CFD15FEB5FD6D2165" box="[642,732,296,320]" pageId="11" pageNumber="56" rank="species">
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<emphasis id="B96DD02E1150BB5CFD15FEB5FD6D2165" bold="true" box="[642,732,296,320]" pageId="11" pageNumber="56">sp. nov.</emphasis>
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</taxonomicNameLabel>
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is currently known from a small area straddling over
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<collectingCountry id="F30E4CAC1150BB5CFAAAFEBAFA2D211A" box="[1341,1436,295,319]" name="Vietnam" pageId="11" pageNumber="56">Vietnam</collectingCountry>
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and
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<collectingCountry id="F30E4CAC1150BB5CFF5FFED1FEBF2141" box="[200,270,332,356]" name="China" pageId="11" pageNumber="56">China</collectingCountry>
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, and from central
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<collectingCountry id="F30E4CAC1150BB5CFE4EFED0FDA32141" box="[473,530,333,356]" name="Laos" pageId="11" pageNumber="56">Laos</collectingCountry>
|
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, as follows:
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</paragraph>
|
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<paragraph id="8BA60C3C1150BB5CFF52FEF2FD3B21EA" blockId="11.[151,1437,151,824]" pageId="11" pageNumber="56">
|
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<collectingCountry id="F30E4CAC1150BB5CFF52FEF2FE9321A2" box="[197,290,367,391]" name="Vietnam" pageId="11" pageNumber="56">Vietnam</collectingCountry>
|
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. Known from the provinces of Lang Son (Huu Lung District) and Cao Bang (Ha Lang District).
|
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<collectingCountry id="F30E4CAC1150BB5CFF0EFE09FE602189" box="[153,465,404,428]" name="China" pageId="11" pageNumber="56">People’s Republic of China</collectingCountry>
|
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. Known from Guangxi Autonomous Region (Longzhou County).
|
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<collectingCountry id="F30E4CAC1150BB5CFB52FE08FB4A2189" box="[1221,1275,405,428]" name="Laos" pageId="11" pageNumber="56">Laos</collectingCountry>
|
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. Known from Khammouane Province (Hin Boun District).
|
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</paragraph>
|
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<paragraph id="8BA60C3C1150BB5CFF52FE41FAA72312" blockId="11.[151,1437,151,824]" pageId="11" pageNumber="56">
|
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<emphasis id="B96DD02E1150BB5CFF52FE41FE3321D1" bold="true" box="[197,386,475,500]" pageId="11" pageNumber="56">Natural history</emphasis>
|
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. This species has been found only in karst environment. The Vietnamese and Chinese specimens were all collected at night in karst forests. The specimen from Cao Bang was found at night (21:00) near the limestone cliff surrounded by secondary forest made of short hardwood, shrubs and vines. No water was observed in the vicinity. The Laotian specimen was collected in a large cave of a karst massif located in the corridor connecting Phou Hin Boun National Park to Nakai Nam Theu National Park. In this totally mineral biotope, this adult male was active around noon in nearly complete darkness on the ground beneath a large boulder. A few amphibian species were found there, such as
|
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<taxonomicName id="4C1977BF1150BB5CFD4CFD29FC7E22E9" box="[731,975,692,716]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="nigrovittata">
|
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<emphasis id="B96DD02E1150BB5CFD4CFD29FC7E22E9" box="[731,975,692,716]" italics="true" pageId="11" pageNumber="56">Hylarana nigrovittata</emphasis>
|
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</taxonomicName>
|
|||
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and
|
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<taxonomicName id="4C1977BF1150BB5CFB92FD29FB5122E9" box="[1029,1248,692,716]" class="Amphibia" family="Microhylidae" genus="Micryletta" kingdom="Animalia" order="Anura" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="inornata">
|
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<emphasis id="B96DD02E1150BB5CFB92FD29FB5122E9" box="[1029,1248,692,716]" italics="true" pageId="11" pageNumber="56">Micryletta inornata</emphasis>
|
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</taxonomicName>
|
|||
|
|
, as well as some other reptile species:
|
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<taxonomicName id="4C1977BF1150BB5CFE00FD45FD8522D5" box="[407,564,728,752]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="56" phylum="Chordata" rank="genus">
|
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|
|
<emphasis id="B96DD02E1150BB5CFE00FD45FD8522D5" box="[407,564,728,752]" italics="true" pageId="11" pageNumber="56">Cyrtodactylus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. and
|
|||
|
|
<taxonomicName id="4C1977BF1150BB5CFD37FD45FBA322D5" box="[672,1042,728,752]" class="Squamata" family="Viperidae" genus="Triceratolepidophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="sieversorum">
|
|||
|
|
<emphasis id="B96DD02E1150BB5CFD37FD45FBA322D5" box="[672,1042,728,752]" italics="true" pageId="11" pageNumber="56">Triceratolepidophis sieversorum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The
|
|||
|
|
<taxonomicName id="4C1977BF1150BB5CFBCBFD45FB7722D5" box="[1116,1222,728,752]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="56" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B96DD02E1150BB5CFBCBFD45FB7722D5" box="[1116,1222,728,752]" italics="true" pageId="11" pageNumber="56">Oligodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
specimen did not attempt to bite when it was collected but it showed the usual behaviour of many species of the genus
|
|||
|
|
<taxonomicName id="4C1977BF1150BB5CFAA5FD61FA2D2331" box="[1330,1436,764,788]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="56" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B96DD02E1150BB5CFAA5FD61FA2D2331" box="[1330,1436,764,788]" italics="true" pageId="11" pageNumber="56">Oligodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
when they feel threatened, i.e. showing the bright colour of the ventral side of its tail curled in a spiral.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|