treatments-xml/data/03/8E/8B/038E8B10D85AFFCFBF96E1062D4BFC31.xml

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<mods:title id="F8D8D55BB9FF152CA80860C6AE733240">South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae)</mods:title>
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<mods:namePart id="F075FB6C354EFFC9DCF05EB88BA663EA">Gordon, Robert D.</mods:namePart>
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<mods:namePart id="0B07307BE19ABA3DBE88557B4ED94DC7">Hanley, Guy A.</mods:namePart>
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<subSubSection id="C33D698DD85AFFCEBF96E1062A46FAB1" box="[192,592,1379,1403]" pageId="78" pageNumber="77" type="nomenclature">
<paragraph id="8B983A06D85AFFCEBF96E1062A46FAB1" blockId="78.[192,592,1379,1403]" box="[192,592,1379,1403]" pageId="78" pageNumber="77">
<heading id="D0D08D6AD85AFFCEBF96E1062A46FAB1" bold="true" box="[192,592,1379,1403]" fontSize="10" level="2" pageId="78" pageNumber="77" reason="3">
<emphasis id="B953E614D85AFFCEBF96E1062A46FAB1" bold="true" box="[192,592,1379,1403]" pageId="78" pageNumber="77">
29.
<taxonomicName id="4C274185D85AFFCEBFB8E1062A46FAB1" authority="(Gorham)" baseAuthorityName="Gorham" box="[238,592,1379,1403]" class="Insecta" family="Coccinellidae" genus="Prodilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="77" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B953E614D85AFFCEBFB8E10629D4FAB0" bold="true" box="[238,450,1379,1403]" italics="true" pageId="78" pageNumber="77">Prodilis rugosa</emphasis>
(Gorham)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD85AFFCEBF96E1C12BC4FA11" pageId="78" pageNumber="77" type="reference_group">
<paragraph id="8B983A06D85AFFCEBF96E1C12A6AFA71" blockId="78.[192,978,1443,1499]" box="[192,636,1443,1467]" pageId="78" pageNumber="77">
<treatmentCitationGroup id="AB371D28D85AFFCEBF96E1C12A6AFA71" box="[192,636,1443,1467]" pageId="78" pageNumber="77">
<treatmentCitation id="0A861C17D85AFFCEBF96E1C12A6EFA71" author="Gorham, H. S." box="[192,632,1443,1467]" page="221" pageId="78" pageNumber="77" year="1897">
<taxonomicName id="4C274185D85AFFCEBF96E1C12A6EFA71" authority="Gorham 1897: 221" authorityName="Gorham" authorityPageNumber="221" authorityYear="1897" box="[192,632,1443,1467]" class="Insecta" family="Coccinellidae" genus="Neaporia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="77" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B953E614D85AFFCEBF96E1C1299DFA71" box="[192,395,1444,1467]" italics="true" pageId="78" pageNumber="77">Neaporia rugosa</emphasis>
<bibRefCitation id="EFB647F7D85AFFCEBEC4E1C62A6EFA71" author="Gorham, H. S." box="[402,632,1443,1467]" pageId="78" pageNumber="77" pagination="217 - 240" refId="ref87056" refString="Gorham, H. S. 1897. Biologia Centrali - Americana, Insecta, Coleoptera, Coccinellidae 7: 217 - 240." type="journal article" year="1897">Gorham 1897: 221</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B983A06D85AFFCEBF96E1A62BC4FA11" blockId="78.[192,978,1443,1499]" box="[192,978,1475,1499]" pageId="78" pageNumber="77">
<treatmentCitationGroup id="AB371D28D85AFFCEBF96E1A62BC4FA11" box="[192,978,1475,1499]" pageId="78" pageNumber="77">
<treatmentCitation id="0A861C17D85AFFCEBF96E1A62ABCFA11" author="Korschefsky, R." box="[192,682,1475,1499]" page="110" pageId="78" pageNumber="77" year="1931">
<taxonomicName id="4C274185D85AFFCEBF96E1A62ABCFA11" authority=": Korschefsky 1931: 110" authorityName=": Korschefsky" authorityPageNumber="110" authorityYear="1931" box="[192,682,1475,1499]" class="Insecta" family="Coccinellidae" genus="Prodilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="77" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B953E614D85AFFCEBF96E1A6296BFA11" box="[192,381,1475,1499]" italics="true" pageId="78" pageNumber="77">Prodilis rugosa</emphasis>
:
<bibRefCitation id="EFB647F7D85AFFCEBEC5E1A62ABCFA11" author="Korschefsky, R." box="[403,682,1475,1499]" pageId="78" pageNumber="77" refId="ref87104" refString="Korschefsky, R. 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118, Coleopterorum Catalogus; Berlin. 224 p." type="book" year="1931">Korschefsky 1931: 110</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0A861C17D85AFFCEBDE0E1A62BD8FA11" author="Blackwelder, E." box="[694,974,1475,1499]" page="444" pageId="78" pageNumber="77" year="1945">
<bibRefCitation id="EFB647F7D85AFFCEBDE0E1A62BD8FA11" author="Blackwelder, E." box="[694,974,1475,1499]" pageId="78" pageNumber="77" pagination="343 - 550" refId="ref86654" refString="Blackwelder, E. 1945. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies and South America. Part 3, Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 444</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C33D698DD85AFFCFBF96E26629C7FDB1" lastPageId="79" lastPageNumber="78" pageId="78" pageNumber="77" type="description">
<paragraph id="8B983A06D85AFFCFBF96E2662C47FE31" blockId="78.[192,1441,1539,1915]" lastBlockId="79.[192,1440,195,507]" lastPageId="79" lastPageNumber="78" pageId="78" pageNumber="77">
<emphasis id="B953E614D85AFFCEBF96E26629AAF9D1" bold="true" box="[192,444,1539,1563]" pageId="78" pageNumber="77">Description. Male.</emphasis>
Length
<quantity id="4CDF97E3D85AFFCEBD49E2662A6AF9D1" box="[543,636,1539,1563]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="78" pageNumber="77" unit="mm" value="1.7">1.7 mm</quantity>
, width
<quantity id="4CDF97E3D85AFFCEBD84E2662B39F9D1" box="[722,815,1539,1563]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="78" pageNumber="77" unit="mm" value="1.3">1.3 mm</quantity>
; body oval, elytron with side rounded, slightly wider than pronotal base, widest at middle of elytra. Dorsal surface entirely shiny, without trace of microsculpture. Color bluish black; head blue (
<figureCitation id="131C2683D85AFFCEBDACE2262B77F991" box="[762,865,1603,1627]" captionStart="Figures 474490" captionStartId="144.[192,287,1632,1654]" captionTargetBox="[208,1422,215,1599]" captionTargetId="figure-20@144.[194,1440,195,1624]" captionTargetPageId="144" captionText="Figures 474490. Prodilis spp. 474479) Prodilis shelley 474) Habitus. 475) Basal abdominal ventrites. 476) Frons. 477) Phallobase ventral. 478) Phallobase lateral. 479) Sipho. 480484) Prodilis rugosa 480) Habitus. 481) Frons. 482) Phallobase ventral. 483) Phallobase lateral. 484) Sipho. 485490) Prodilis sabrina 485) Habitus. 486) Basal abdominal ventrites. 487) Frons. 488) Phallobase ventral. 489) Phallobase lateral. 490) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170083" httpUri="https://zenodo.org/record/5170083/files/figure.png" pageId="78" pageNumber="77">Fig. 481</figureCitation>
); pronotum bluish black except reflexed lateral margin reddish yellow; elytron with elongate, oval yellow macula at middle of apical ½, reflexed lateral margin reddish yellow (
<figureCitation id="131C2683D85AFFCEBE88E2E62A50F951" box="[478,582,1667,1691]" captionStart="Figures 474490" captionStartId="144.[192,287,1632,1654]" captionTargetBox="[208,1422,215,1599]" captionTargetId="figure-20@144.[194,1440,195,1624]" captionTargetPageId="144" captionText="Figures 474490. Prodilis spp. 474479) Prodilis shelley 474) Habitus. 475) Basal abdominal ventrites. 476) Frons. 477) Phallobase ventral. 478) Phallobase lateral. 479) Sipho. 480484) Prodilis rugosa 480) Habitus. 481) Frons. 482) Phallobase ventral. 483) Phallobase lateral. 484) Sipho. 485490) Prodilis sabrina 485) Habitus. 486) Basal abdominal ventrites. 487) Frons. 488) Phallobase ventral. 489) Phallobase lateral. 490) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170083" httpUri="https://zenodo.org/record/5170083/files/figure.png" pageId="78" pageNumber="77">Fig. 480</figureCitation>
); antenna, legs yellow; mouthparts yellow except terminal 2/3 of apical maxillary palpomere brown; venter black; abdomen dark brown. Head punctures small, separated by less than a diameter; pronotal punctures larger than on head, separated by a diameter or less; elytral punctures larger than on pronotum, separated by a diameter or less; prosternal punctures large, separated by about a diameter, mesosternal punctures large, separated by less than a diameter, metasternal punctures large, separated by a diameter or less anteriorly and laterally, slightly smaller and separated by a diameter or slightly more medially; abdomen with punctures on ventrites 13 large, separated by a diameter or less, punctures on remaining ventrites smaller, separated by about a diameter. Head with frons narrow, widened from vertex to clypeus, slightly wider than eye measured at vertex; eye canthus short; apical maxillary palpomere long, widened medially. Pronotum widest at middle, reflexed lateral margin narrow, equal in width from base to apex. Epipleuron slightly descending externally, narrow, as wide as pronotal hypomeron, with slight depressions for reception of femoral apex. Prosternum slightly wider than long, slightly longer than mesosternum, base somewhat produced, weakly emarginate medially, lateral carina narrow, short, extended to apex of procoxa. Postcoxal line on ventrite 1 short, rounded, extended slightly more than ½ distance to ventrite apex. Apex of ventrite 5 broadly, shallowly emarginate. Genitalia with basal lobe shorter than paramere, slender, weakly widened from base to emarginate apex, paramere slender, curved in basal 7/8, apical 1/8 recurved to acute apex, with marginal serrations (
<figureCitation id="131C2683D85BFFCFBF9FE586297DFE31" box="[201,363,483,507]" captionStart="Figures 474490" captionStartId="144.[192,287,1632,1654]" captionTargetBox="[208,1422,215,1599]" captionTargetId="figure-20@144.[194,1440,195,1624]" captionTargetPageId="144" captionText="Figures 474490. Prodilis spp. 474479) Prodilis shelley 474) Habitus. 475) Basal abdominal ventrites. 476) Frons. 477) Phallobase ventral. 478) Phallobase lateral. 479) Sipho. 480484) Prodilis rugosa 480) Habitus. 481) Frons. 482) Phallobase ventral. 483) Phallobase lateral. 484) Sipho. 485490) Prodilis sabrina 485) Habitus. 486) Basal abdominal ventrites. 487) Frons. 488) Phallobase ventral. 489) Phallobase lateral. 490) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170083" httpUri="https://zenodo.org/record/5170083/files/figure.png" pageId="79" pageNumber="78">Fig. 482, 483</figureCitation>
); sipho robust, with ventral tumosity in apical 1/8 (
<figureCitation id="131C2683D85BFFCFBC8AE5862C52FE31" box="[988,1092,483,507]" captionStart="Figures 474490" captionStartId="144.[192,287,1632,1654]" captionTargetBox="[208,1422,215,1599]" captionTargetId="figure-20@144.[194,1440,195,1624]" captionTargetPageId="144" captionText="Figures 474490. Prodilis spp. 474479) Prodilis shelley 474) Habitus. 475) Basal abdominal ventrites. 476) Frons. 477) Phallobase ventral. 478) Phallobase lateral. 479) Sipho. 480484) Prodilis rugosa 480) Habitus. 481) Frons. 482) Phallobase ventral. 483) Phallobase lateral. 484) Sipho. 485490) Prodilis sabrina 485) Habitus. 486) Basal abdominal ventrites. 487) Frons. 488) Phallobase ventral. 489) Phallobase lateral. 490) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170083" httpUri="https://zenodo.org/record/5170083/files/figure.png" pageId="79" pageNumber="78">Fig. 484</figureCitation>
).
</paragraph>
<paragraph id="8B983A06D85BFFCFBF96E64629A5FDF1" blockId="79.[192,435,547,571]" box="[192,435,547,571]" pageId="79" pageNumber="78">
<emphasis id="B953E614D85BFFCFBF96E646293AFDF1" bold="true" box="[192,300,547,571]" pageId="79" pageNumber="78">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B983A06D85BFFCFBF96E60629C7FDB1" blockId="79.[192,465,611,635]" box="[192,465,611,635]" pageId="79" pageNumber="78">
<emphasis id="B953E614D85BFFCFBF96E6062954FDB1" bold="true" box="[192,322,611,635]" pageId="79" pageNumber="78">Variation</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD85BFFCFBF96E6C62B46FCF1" pageId="79" pageNumber="78" type="materials_examined">
<paragraph id="8B983A06D85BFFCFBF96E6C62A5CFD71" blockId="79.[192,586,675,699]" box="[192,586,675,699]" pageId="79" pageNumber="78">
<emphasis id="B953E614D85BFFCFBF96E6C62965FD71" bold="true" box="[192,371,675,699]" pageId="79" pageNumber="78">
<typeStatus id="549C84A4D85BFFCFBF96E6C62912FD71" box="[192,260,675,699]" pageId="79" pageNumber="78">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F3307A96D85BFFCFBED7E6C629EBFD71" box="[385,509,675,699]" name="Panama" pageId="79" pageNumber="78">PANAMA</collectingCountry>
: Tolé.
</paragraph>
<paragraph id="8B983A06D85BFFCFBF96E6862A1CFD31" blockId="79.[192,522,739,763]" box="[192,522,739,763]" pageId="79" pageNumber="78">
<emphasis id="B953E614D85BFFCFBF96E686298BFD31" bold="true" box="[192,413,739,763]" pageId="79" pageNumber="78">
<typeStatus id="549C84A4D85BFFCFBF96E6862912FD31" box="[192,260,739,763]" pageId="79" pageNumber="78">Type</typeStatus>
depository
</emphasis>
. BMNH.
</paragraph>
<paragraph id="8B983A06D85BFFCFBF96E7462B46FCF1" blockId="79.[192,848,803,827]" box="[192,848,803,827]" pageId="79" pageNumber="78">
<emphasis id="B953E614D85BFFCFBF96E7462A3CFCF1" bold="true" box="[192,554,803,827]" pageId="79" pageNumber="78">Geographical distribution</emphasis>
. 1.
<collectingCountry id="F3307A96D85BFFCFBD13E7462ADDFCF1" box="[581,715,803,827]" name="Panama" pageId="79" pageNumber="78">PANAMA</collectingCountry>
: (
<collectionCode id="ED36A2C3D85BFFCFBDB0E7462B52FCF1" box="[742,836,803,827]" pageId="79" pageNumber="78">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD85BFFCFBF96E7062D4BFC31" pageId="79" pageNumber="78" type="discussion">
<paragraph id="8B983A06D85BFFCFBF96E7062D4BFC31" blockId="79.[192,1441,867,1019]" pageId="79" pageNumber="78">
<emphasis id="B953E614D85BFFCFBF96E7062952FCB1" bold="true" box="[192,324,867,891]" pageId="79" pageNumber="78">Remarks.</emphasis>
This distinctive species is characterized by a mostly blue dorsal surface, dense, deep punctures on both dorsal and ventral surfaces, and the elytral color pattern.
<bibRefCitation id="EFB647F7D85BFFCFBB7AE7E62CF0FC51" author="Gorham, H. S." box="[1068,1254,899,923]" pageId="79" pageNumber="78" pagination="217 - 240" refId="ref87056" refString="Gorham, H. S. 1897. Biologia Centrali - Americana, Insecta, Coleoptera, Coccinellidae 7: 217 - 240." type="journal article" year="1897">Gorham (1897)</bibRefCitation>
named it “rugosa” because of the densely punctured dorsal surface resulting in a somewhat rough aspect. The
<typeStatus id="549C84A4D85BFFCFBA35E7C628E2FC16" pageId="79" pageNumber="78" type="holotype">holotype</typeStatus>
is labeled “Tolé,
<collectingCountry id="F3307A96D85BFFCFBEEBE7A12A34FC11" box="[445,546,964,987]" name="Panama" pageId="79" pageNumber="78">Panama</collectingCountry>
, Champion./
<taxonomicName id="4C274185D85BFFCFBDEEE7A12BD5FC11" authority="Gorh" authorityName="Gorham" authorityYear="1897" box="[696,963,963,987]" class="Insecta" family="Coccinellidae" genus="Neaporia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="78" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B953E614D85BFFCFBDEEE7A12B69FC11" box="[696,895,964,987]" italics="true" pageId="79" pageNumber="78">Neaporia rugosa</emphasis>
Gorh
</taxonomicName>
(handwritten)/Sp. figured/
<typeStatus id="549C84A4D85BFFCFBA53E7A62DB7FC11" box="[1285,1441,963,987]" pageId="79" pageNumber="78" type="holotype">HOLOTYPE</typeStatus>
(red bordered disc)/B.C.A.,Col.,VII./
<typeStatus id="549C84A4D85BFFCFBD31E7862B19FC31" box="[615,783,995,1019]" pageId="79" pageNumber="78" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C274185D85BFFCFBC40E7812C38FC31" authority="Gorh." authorityName="Gorham" authorityYear="1897" box="[790,1070,995,1019]" class="Insecta" family="Coccinellidae" genus="Neaporia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="78" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B953E614D85BFFCFBC40E7812BF6FC31" box="[790,992,996,1019]" italics="true" pageId="79" pageNumber="78">Neaporia rugosa</emphasis>
Gorh.
</taxonomicName>
det. R.G. Booth 2015.”Ó
</paragraph>
</subSubSection>
</treatment>
</document>