treatments-xml/data/03/F4/E5/03F4E51BFFCBFFDFDAEFFE29FBF5FC84.xml

<document id="1F2F899EAE8AA218857F7A9B94B59A0D" ID-DOI="10.5281/zenodo.13154055" ID-ISSN="0253-116X" ID-Zenodo-Dep="13154055" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722537227576" checkinUser="felipe" docAuthor="Zettel, Herbert, Laciny, Alice &amp; T, South" docDate="2021" docId="03F4E51BFFCBFFDFDAEFFE29FBF5FC84" docLanguage="en" docName="LinzerbiolBeitr.2021.53.1.445-449.pdf" docOrigin="Linzer biologische Beiträge 53 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.12:LinzerbiolBeitr.2006-.journal_article.type1" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article.type1" docStyleVersion="12" docTitle="Pleciogonus narumonae Zettel, Laciny &amp; T, 2021, nov.sp." docType="treatment" docVersion="2" lastPageNumber="448" masterDocId="FFCD9D63FFCAFFDCDA5DFF81FFEFFF93" masterDocTitle="A second species of Pleciogonus (Hemiptera, Heteroptera, Gerridae) from Thailand" masterLastPageNumber="449" masterPageNumber="445" pageNumber="446" updateTime="1722540140691" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6AFBBFD97A545C2AA9C99CADF22B8D0B">A second species of Pleciogonus (Hemiptera, Heteroptera, Gerridae) from Thailand</mods:title>
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<mods:namePart id="A7F06AB46F2BD1D024BB1FE00C158490">Zettel, Herbert</mods:namePart>
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<mods:namePart id="E897A23081063EA358F6E52E7D58E8A3">Laciny, Alice</mods:namePart>
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<taxonomicName id="4C5D2F8EFFCBFFDDDAEFFE29FE5EFE2D" authorityName="Zettel &amp; Laciny &amp; T" authorityYear="2021" box="[178,433,424,446]" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="narumonae" status="sp. nov.">
<emphasis id="B929881FFFCBFFDDDAEFFE29FE5EFE2D" bold="true" box="[178,433,424,446]" italics="true" pageId="1" pageNumber="446">Pleciogonus narumonae</emphasis>
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(
<figureCitation id="13664888FFCBFFDDD84CFE28FD8AFE2D" box="[529,613,425,447]" captionStart-0="Fig" captionStart-1="Figs 2-3" captionStartId-0="2.[178,211,1545,1567]" captionStartId-1="3.[178,218,553,575]" captionTargetBox-0="[179,1085,214,1535]" captionTargetBox-1="[185,1083,210,541]" captionTargetId-0="figure-27@2.[179,1085,214,1535]" captionTargetId-1="figure-374@3.[177,1084,210,541]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1: Pleciogonus narumonae nov.sp., habitus of apterous female (holotype). © Alice Laciny / Natural History Museum Vienna, Hemiptera Image Collection." captionText-1="Figs 2-3: Pleciogonus narumonae nov.sp., apterous female (holotype): posterior part of abdomen, (2) in dorsal view (between bases of hind legs), (3) in lateral view; lt6 – laterotergite 6; mfe – metafemur; mt6, mt7 – mediotergites 6-7; mtr – metatrochanter; pr – process of laterotergite 6; st4- st7 – sterna 4-7." figureDoi-0="http://doi.org/10.5281/zenodo.13154059" figureDoi-1="http://doi.org/10.5281/zenodo.13154061" httpUri-0="https://zenodo.org/record/13154059/files/figure.png" httpUri-1="https://zenodo.org/record/13154061/files/figure.png" pageId="1" pageNumber="446">Figs 1-3</figureCitation>
)
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<paragraph id="8BE2540DFFCBFFDDDAEFFE4EFEBEFD87" blockId="1.[178,1086,462,532]" pageId="1" pageNumber="446">
<materialsCitation id="3B355E50FFCBFFDDDAEFFE4EFEA2FD87" collectingDate="2006-11-29" collectorName="H. Zettel" country="Thailand" county="Khuraburi District" location="West of Si" municipality="Baan Tumnang" pageId="1" pageNumber="446" specimenCount="1" stateProvince="Phangnga">
H o l o t y p e (apterous female; Natural History Museum Vienna): 
<collectingCountry id="F34A149DFFCBFFDDD943FE4FFC9EFE77" box="[798,881,462,484]" name="Thailand" pageId="1" pageNumber="446">Thailand</collectingCountry>
, 
<collectingRegion id="49999AEFFFCBFFDDD926FE4FFBD6FE77" box="[891,1081,462,484]" country="Thailand" name="Phangnga" pageId="1" pageNumber="446">Phang Nga Province</collectingRegion>
, 
<collectingCounty id="62832C81FFCBFFDDDA82FE67FE65FE6F" box="[223,394,486,508]" pageId="1" pageNumber="446">Khuraburi District</collectingCounty>
, 
<collectingMunicipality id="6B86CE77FFCBFFDDDBCEFE66FDCDFE6F" box="[403,546,487,508]" pageId="1" pageNumber="446">Baan Tumnang</collectingMunicipality>
, 
<location id="8E8202D6FFCBFFDDD871FE66FD61FE6F" LSID="urn:lsid:plazi:treatment:03F4E51BFFCBFFDFDAEFFE29FBF5FC84:8E8202D6FFCBFFDDD871FE66FD61FE6F" box="[556,654,486,508]" country="Thailand" county="Khuraburi District" municipality="Baan Tumnang" name="West of Si" pageId="1" pageNumber="446" stateProvince="Phangnga">West of Si</location>
<collectingRegion id="49999AEFFFCBFFDDD8C8FE67FD15FE6F" box="[661,762,486,508]" country="Thailand" name="Phangnga" pageId="1" pageNumber="446">Phang Nga</collectingRegion>
<location id="8E8202D6FFCBFFDDD95DFE67FC6EFE6F" LSID="urn:lsid:plazi:treatment:03F4E51BFFCBFFDFDAEFFE29FBF5FC84:8E8202D6FFCBFFDDD95DFE67FC6EFE6F" box="[768,897,486,508]" country="Thailand" county="Khuraburi District" municipality="Baan Tumnang" name="National Park" pageId="1" pageNumber="446" stateProvince="Phangnga">National Park</location>
, 
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<collectingDate id="EFA78B25FFCBFFDDD9D6FE67FC1AFE6F" box="[907,1013,486,508]" pageId="1" pageNumber="446" value="2006-11-29">29.XI.2006</collectingDate>
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, leg. 
<collectorName id="26A831DBFFCBFFDDDE7BFE66FEFBFD87" pageId="1" pageNumber="446">H. Zettel</collectorName>
(#48)
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.
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<subSubSection id="C3470786FFCBFFDFDAEFFDA2FBF5FC84" lastPageId="3" lastPageNumber="448" pageId="1" pageNumber="446" type="description">
<paragraph id="8BE2540DFFCBFFDDDAEFFDA2FDC0FD7B" blockId="1.[178,1086,547,745]" pageId="1" pageNumber="446">D e s c r i p t i o n: Measurements: Body length 5.60, maximum width (at mesopleura) 2.11. Head length 0.71, width (including eyes) 1.06; minimum distance of eyes 0.43. Pronotum length 0.63, width 1.21. Mesonotum length (at midline) 1.87. Metanotum length (at midline) 0.70. Combined length of abdominal tergites 1-7, 1.70. Length of antennomeres 1-4: 2.38, 0.64, 0.96, 0.64. Lengths of leg joints: Profemur 2.52, protibia 1.91, protarsus 0.90 + 0.71, mesofemur 7.58, mesotibia 1.82, mesotarsus 0.13 + 0.14, metafemur 7.42, metatibia 1.02, metatarsus 0.33.</paragraph>
<paragraph id="8BE2540DFFCBFFDDDAEFFD79FBC3FCDA" blockId="1.[178,1086,760,841]" pageId="1" pageNumber="446">
Colour: Dorsal pattern as in 
<figureCitation id="13664888FFCBFFDDDB81FD79FDDAFC9E" box="[476,565,760,782]" captionStart="Fig" captionStartId="2.[178,211,1545,1567]" captionTargetBox="[179,1085,214,1535]" captionTargetId="figure-27@2.[179,1085,214,1535]" captionTargetPageId="2" captionText="Fig. 1: Pleciogonus narumonae nov.sp., habitus of apterous female (holotype). © Alice Laciny / Natural History Museum Vienna, Hemiptera Image Collection." figureDoi="http://doi.org/10.5281/zenodo.13154059" httpUri="https://zenodo.org/record/13154059/files/figure.png" pageId="1" pageNumber="446">Figure 1</figureCitation>
. Sides yellow to pale orange. Venter pale yellow. Antennae black. Legs: coxae and trochanters chiefly yellow; profemur chiefly black; mesofemur black except extreme base yellow; metafemur, and all tibiae and tarsi black.
</paragraph>
<paragraph id="8BE2540DFFCBFFDDDAEFFCD9FC8DFA9B" blockId="1.[178,1085,856,1289]" pageId="1" pageNumber="446">
Structures: Head anteriorly only slightly protruded, antennal tubercles rounded. Antennomere 1 hardly longer than antennomeres 2-4 combined; antennomere 3 longer than each, antennomere 2 and 4. Pronotum transverse; sides rounded. Meso- and metanotum without special structures; hind margin of metanotum deeply sinuated. Profemur hardly incrassate, flexor side in distal half with numerous short black oblique spines; protarsomere 1 much longer than 2. Meso- and metafemur subequal in length, each longer than body. Mesotarsomeres subequal in length; metatarsomeres fused. Claws of middle and hindlegs reduced. Abdomen short. Mediotergites 1-6 transverse; 1 about 1.5 times longer than 2; 6 slightly shorter than 4 and 5 combined. Mediotergite 7 (
<figureCitation id="13664888FFCBFFDDD95DFBC2FCAFFBCB" box="[768,832,1091,1113]" captionStart="Figs 2-3" captionStartId="3.[178,218,553,575]" captionTargetBox="[185,1083,210,541]" captionTargetId="figure-374@3.[177,1084,210,541]" captionTargetPageId="3" captionText="Figs 2-3: Pleciogonus narumonae nov.sp., apterous female (holotype): posterior part of abdomen, (2) in dorsal view (between bases of hind legs), (3) in lateral view; lt6 – laterotergite 6; mfe – metafemur; mt6, mt7 – mediotergites 6-7; mtr – metatrochanter; pr – process of laterotergite 6; st4- st7 – sterna 4-7." figureDoi="http://doi.org/10.5281/zenodo.13154061" httpUri="https://zenodo.org/record/13154061/files/figure.png" pageId="1" pageNumber="446">Fig. 2</figureCitation>
) about as long as wide; anterior two-fifths flat, posterior third folded in a roof-like shape; hind margin with two roundish convexities separated by a deep incision. Laterotergites steep; connexival margin strongly sinuate in dorsal view. Laterotergite 6 (
<figureCitation id="13664888FFCBFFDDD8F5FB1AFCE3FB23" box="[680,780,1179,1201]" captionStart="Figs 2-3" captionStartId="3.[178,218,553,575]" captionTargetBox="[185,1083,210,541]" captionTargetId="figure-374@3.[177,1084,210,541]" captionTargetPageId="3" captionText="Figs 2-3: Pleciogonus narumonae nov.sp., apterous female (holotype): posterior part of abdomen, (2) in dorsal view (between bases of hind legs), (3) in lateral view; lt6 – laterotergite 6; mfe – metafemur; mt6, mt7 – mediotergites 6-7; mtr – metatrochanter; pr – process of laterotergite 6; st4- st7 – sterna 4-7." figureDoi="http://doi.org/10.5281/zenodo.13154061" httpUri="https://zenodo.org/record/13154061/files/figure.png" pageId="1" pageNumber="446">Figs. 2, 3</figureCitation>
) with a short, triangular protrusion. Sternum 7 as characteristic for the genus: sides posteriorly curved towards middle, posterodorsally broadly rounded; middle with a large lobe that is wider than hind margin of mediotergite 7. Segment 8 and genitalia retracted into segment 7.
</paragraph>
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C o m p a r a t i v e n o t e s: 
<taxonomicName id="4C5D2F8EFFCBFFDDD84BFA99FCE0FABD" authorityName="Zettel &amp; Laciny &amp; T" authorityYear="2021" box="[534,783,1304,1326]" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="narumonae" status="sp. nov.">
<emphasis id="B929881FFFCBFFDDD84BFA99FCE0FABD" box="[534,783,1304,1326]" italics="true" pageId="1" pageNumber="446">Pleciogonus narumonae</emphasis>
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can be easily distinguished from 
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<emphasis id="B929881FFFCBFFDDDB63FAB7FE56FAD8" box="[318,441,1334,1355]" italics="true" pageId="1" pageNumber="446">P. wongsirii</emphasis>
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by the structure of connexivum 6 of the female. In 
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<emphasis id="B929881FFFCBFFDDD99CFAB7FBD3FAD8" box="[961,1084,1334,1355]" italics="true" pageId="1" pageNumber="446">P. wongsirii</emphasis>
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it is posteriorly protracted in a long, slender, hirsute process that almost reaches or even surpasses the hind margin of tergite 7 (cf. 
<bibRefCitation id="EFCC29FCFFCBFFDDD835FAF1FCF5FA16" author="CHEN P. - P. &amp; NIESER N. &amp; W. WATTANACHAIYINGCHAROEN" box="[616,794,1392,1414]" pageId="1" pageNumber="446" pagination="193 - 212" refId="ref1760" refString="CHEN P. - P., NIESER N. &amp; W. WATTANACHAIYINGCHAROEN (2002): A new genus, Pleciogonus and four new species of semiaquatic and aquatic bugs from Thailand (Heteroptera: Gerromorpha, Nepomorpha). - Tijdschrift voor Entomologie 145 / 2: 193 - 212." type="journal article" year="2002">CHEN et al. 2002</bibRefCitation>
: figs. 28, 30), whereas in 
<taxonomicName id="4C5D2F8EFFCBFFDDDE7AFAF0FECAFA30" authorityName="Zettel &amp; Laciny &amp; T" authorityYear="2021" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="narumonae" status="sp. nov.">
<emphasis id="B929881FFFCBFFDDDE7AFAF0FECAFA30" italics="true" pageId="1" pageNumber="446">P. narumonae</emphasis>
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it forms a short, inconspicuous, slightly acute angle, that has the length of one fourth of tergite 7 and does not bear any longer setae (
<figureCitation id="13664888FFCBFFDDD969FA2AFC9DFA53" box="[820,882,1451,1473]" captionStart="Figs 2-3" captionStartId="3.[178,218,553,575]" captionTargetBox="[185,1083,210,541]" captionTargetId="figure-374@3.[177,1084,210,541]" captionTargetPageId="3" captionText="Figs 2-3: Pleciogonus narumonae nov.sp., apterous female (holotype): posterior part of abdomen, (2) in dorsal view (between bases of hind legs), (3) in lateral view; lt6 – laterotergite 6; mfe – metafemur; mt6, mt7 – mediotergites 6-7; mtr – metatrochanter; pr – process of laterotergite 6; st4- st7 – sterna 4-7." figureDoi="http://doi.org/10.5281/zenodo.13154061" httpUri="https://zenodo.org/record/13154061/files/figure.png" pageId="1" pageNumber="446">Fig. 3</figureCitation>
). A distinct spot of golden brownish pilosity at the middle of the mesonotal hind margin of 
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<emphasis id="B929881FFFCBFFDDD9F8FA48FBCCFA4D" box="[933,1059,1481,1502]" italics="true" pageId="1" pageNumber="446">P. wongsirii</emphasis>
</taxonomicName>
is lacking in 
<taxonomicName id="4C5D2F8EFFCBFFDDDB7FFA67FE5BFA68" authorityName="Zettel &amp; Laciny &amp; T" authorityYear="2021" box="[290,436,1510,1531]" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="narumonae" status="sp. nov.">
<emphasis id="B929881FFFCBFFDDDB7FFA67FE5BFA68" box="[290,436,1510,1531]" italics="true" pageId="1" pageNumber="446">P. narumonae</emphasis>
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Antennomere 1 is relatively shorter (compared to antennomeres 2-4) in 
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<emphasis id="B929881FFFCBFFDDDBDEF982FDF9F98B" box="[387,534,1539,1560]" italics="true" pageId="1" pageNumber="446">P. narumonae</emphasis>
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than in 
<taxonomicName id="4C5D2F8EFFCBFFDDD89AF982FCA8F98B" authorityName="CHEN, NIESER &amp; WATTANACHAIYINGCHAROEN" authorityYear="2002" box="[711,839,1539,1560]" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="wongsirii">
<emphasis id="B929881FFFCBFFDDD89AF982FCA8F98B" box="[711,839,1539,1560]" italics="true" pageId="1" pageNumber="446">P. wongsirii</emphasis>
</taxonomicName>
. The black areas of 
<taxonomicName id="4C5D2F8EFFCBFFDDDE7AF982FECAF9A5" authorityName="Zettel &amp; Laciny &amp; T" authorityYear="2021" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="narumonae" status="sp. nov.">
<emphasis id="B929881FFFCBFFDDDE7AF982FECAF9A5" italics="true" pageId="1" pageNumber="446">P. narumonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A3564FFCBFFDDDB76F9A1FE9AF9A6" box="[299,373,1568,1589]" pageId="1" pageNumber="446" rank="species">nov.sp.</taxonomicNameLabel>
are comparatively more widely extended, especially on the profemur, but the pattern of 
<taxonomicName id="4C5D2F8EFFC9FFDFDB34FD23FE09FD24" authorityName="CHEN, NIESER &amp; WATTANACHAIYINGCHAROEN" authorityYear="2002" box="[361,486,674,695]" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="wongsirii">
<emphasis id="B929881FFFC9FFDFDB34FD23FE09FD24" box="[361,486,674,695]" italics="true" pageId="3" pageNumber="448">P. wongsirii</emphasis>
</taxonomicName>
shows some variation in colour. We could not find differences in the genus-specific structures of abdominal segment 7.
</paragraph>
<caption id="DF220485FFC8FFDEDAEFF988FD1FF9A5" ID-DOI="http://doi.org/10.5281/zenodo.13154059" ID-Zenodo-Dep="13154059" httpUri="https://zenodo.org/record/13154059/files/figure.png" pageId="2" pageNumber="447" startId="2.[178,211,1545,1567]" targetBox="[179,1085,214,1535]" targetPageId="2" targetType="figure">
<paragraph id="8BE2540DFFC8FFDEDAEFF988FD1FF9A5" blockId="2.[178,1083,1544,1590]" pageId="2" pageNumber="447">
<emphasis id="B929881FFFC8FFDEDAEFF988FF05F98D" bold="true" box="[178,234,1545,1567]" pageId="2" pageNumber="447">Fig. 1</emphasis>
: 
<taxonomicName id="4C5D2F8EFFC8FFDEDAA4F988FE37F98C" authorityName="Zettel &amp; Laciny &amp; T" authorityYear="2021" box="[249,472,1545,1567]" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="narumonae" status="sp. nov.">
<emphasis id="B929881FFFC8FFDEDAA4F988FE37F98C" box="[249,472,1545,1567]" italics="true" pageId="2" pageNumber="447">Pleciogonus narumonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A3564FFC8FFDEDBBCF98BFDCAF98C" box="[481,549,1546,1567]" pageId="2" pageNumber="447" rank="species">nov.sp.</taxonomicNameLabel>
, habitus of apterous female (holotype). © Alice Laciny / Natural History Museum Vienna, 
<taxonomicName id="4C5D2F8EFFC8FFDEDBBAF9A1FD6AF9A6" authority="Image" authorityName="Image" box="[487,645,1568,1590]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="order">Hemiptera Image</taxonomicName>
Collection.
</paragraph>
</caption>
<caption id="DF220485FFC9FFDFDAEFFDA8FEACFD11" ID-DOI="http://doi.org/10.5281/zenodo.13154061" ID-Zenodo-Dep="13154061" httpUri="https://zenodo.org/record/13154061/files/figure.png" pageId="3" pageNumber="448" startId="3.[178,218,553,575]" targetBox="[185,1083,210,541]" targetPageId="3" targetType="figure">
<paragraph id="8BE2540DFFC9FFDFDAEFFDA8FEACFD11" blockId="3.[178,1085,553,643]" pageId="3" pageNumber="448">
<emphasis id="B929881FFFC9FFDFDAEFFDA8FF10FDAD" bold="true" box="[178,255,553,575]" pageId="3" pageNumber="448">Figs 2-3</emphasis>
: 
<taxonomicName id="4C5D2F8EFFC9FFDFDB50FDA8FE03FDAC" authorityName="Zettel &amp; Laciny &amp; T" authorityYear="2021" box="[269,492,553,575]" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="narumonae" status="sp. nov.">
<emphasis id="B929881FFFC9FFDFDB50FDA8FE03FDAC" box="[269,492,553,575]" italics="true" pageId="3" pageNumber="448">Pleciogonus narumonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A3564FFC9FFDFDBAEFDABFDD7FDAC" box="[499,568,554,575]" pageId="3" pageNumber="448" rank="species">nov.sp.</taxonomicNameLabel>
, apterous female (holotype): posterior part of abdomen, (
<emphasis id="B929881FFFC9FFDFDAE4FDC1FF2BFDC6" bold="true" box="[185,196,576,597]" pageId="3" pageNumber="448">2</emphasis>
) in dorsal view (between bases of hind legs), (
<emphasis id="B929881FFFC9FFDFD8D6FDC1FD79FDC6" bold="true" box="[651,662,576,597]" pageId="3" pageNumber="448">3</emphasis>
) in lateral view; lt6 – laterotergite 6; mfe – metafemur; mt6, mt7 – mediotergites 6-7; mtr – metatrochanter; pr – process of laterotergite 6; st4- st7 – sterna 4-7.
</paragraph>
</caption>
<paragraph id="8BE2540DFFC9FFDFDAEFFD65FBF5FC84" blockId="3.[178,1085,740,791]" pageId="3" pageNumber="448">
E t y m o l o g y: This new species is dedicated to Associate Professor Dr. Narumon Sangpradub of the 
<collectingRegion id="49999AEFFFC9FFDFDB2FFC80FE0AFC84" box="[370,485,769,791]" country="Thailand" name="Khon Kaen" pageId="3" pageNumber="448">Khon Kaen</collectingRegion>
University for her work on aquatic insects of 
<collectingCountry id="F34A149DFFC9FFDFD9E5FC80FBFAFC84" box="[952,1045,769,791]" name="Thailand" pageId="3" pageNumber="448">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>