treatments-xml/data/03/F0/6D/03F06D13FEAE2167089B1092016AF527.xml

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<document id="703DC123FF21975F030A4A53704B76BE" ID-CLB-Dataset="80832" ID-DOI="10.5281/zenodo.6707142" ID-GBIF-Dataset="ab66b2b7-9544-4411-bf61-5bc3651d7bca" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6707142" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1656002532852" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="03F06D13FEAE2167089B1092016AF527" docLanguage="en" docName="hbmw_7_Cricetidae_0204.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Tylomys panamensis" docType="treatment" docVersion="10" lastPageNumber="535" masterDocId="FFC9156BFFAE20670D37145C0837FFDB" masterDocTitle="Cricetidae" masterLastPageNumber="535" masterPageNumber="204" pageNumber="535" updateTime="1718799336328" updateUser="carolina">
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<mods:title id="68D792A6999D6AF951B91B212D41C6F9">Cricetidae</mods:title>
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<mods:namePart id="6B414563D5CAF4BF667923E3A48FE609">Don E. Wilson</mods:namePart>
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<mods:namePart id="4A7B02D254478FA32FC4ECC11A9DB4B7">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="1C3A8A54608032E63185D424DCCDCDA6">Thomas E. Lacher, Jr</mods:namePart>
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<treatment id="03F06D13FEAE2167089B1092016AF527" ID-DOI="http://doi.org/10.5281/zenodo.6708783" ID-GBIF-Taxon="196221425" ID-Zenodo-Dep="6708783" LSID="urn:lsid:plazi:treatment:03F06D13FEAE2167089B1092016AF527" httpUri="http://treatment.plazi.org/id/03F06D13FEAE2167089B1092016AF527" lastPageNumber="535" pageId="256" pageNumber="535">
<subSubSection id="C3438F8EFEAE2167089B10920DC9FB27" box="[1452,1534,1230,1276]" pageId="256" pageNumber="535" type="multiple">
<paragraph id="8BE6DC05FEAE2167089B10920DC9FB27" blockId="256.[1449,2444,1230,1356]" box="[1452,1534,1230,1276]" pageId="256" pageNumber="535">
<heading id="D0AE6B69FEAE2167089B10920DC9FB27" box="[1452,1534,1230,1276]" pageId="256" pageNumber="535">
<figureCitation id="1362C080FEAE2167089B10920DC9FB27" box="[1452,1534,1230,1276]" captionStart="Plate 31: Cricetidae" captionStartId="245.[107,137,3149,3170]" captionTargetBox="[11,2726,13,3647]" captionTargetPageId="244" captionText="725. Haggards Leaf-eared Mouse (Phyllotis haggardi), 726. Andean Leaf-eared Mouse (Phyllotis andium), 727. Peruvian Leaf-eared Mouse (Phyllotis gerbillus), 728. Friendly Leaf-eared Mouse (Phyllotis amicus), 729. Narrow-nasal Leaf-eared Mouse (Phyllotis stenops), 730. Pearsons Leaf-eared Mouse (Phyllotis pearson), 731. Western Leaf-eared Mouse (Phyllotis occidens), 732. Ancash Leaf-eared Mouse (Phyllotis definitus), 733. Lima Leaf-eared Mouse (Phyllotis limatus), 734. Master Leaf-eared Mouse (Phyllotis magister), 735. Yellow-rumped Leaf-eared Mouse (Phyllotis xanthopygus), 736. Osgoods Leaf-eared Mouse (Phyllotis osgoodi), 737. Bunch Grass Leaf-eared Mouse (Phyllotis osilae), 738. Capricorn Leaf-eared Mouse (Phyllotis caprinus), 739. Tucuman Leaf-eared Mouse (Phyllotis tucumanus), 740. Walnut Leaf-eared Mouse (Phyllotis nogalaris), 741. Darwins Leaf-eared Mouse (Phyllotis darwinii), 742. Los Alisos Leaf-eared Mouse (Phyllotis alisosiensis), 743. Anitas Leaf-eared Mouse (Phyllotis anitae), 744. Bonarian Leaf-eared Mouse (Phyllotis bonariensis), 745. Wolffsohns Leaf-eared Mouse 1 (apecomys wolffsohni), 746. Tapecua Leaf-eared Mouse (Tapecomys primus), 747. Southern Big-eared Mouse (Loxodontomys micropus), 748. Delicate Salt Flat Mouse (Salinomys delicatus), 749. Pearsons Chaco Mouse (Andalgalomys pearsoni), 750. Olrogs Chaco Mouse (Andalgalomys olrogi), 751. Garlepps Mouse (Galenomys garleppii), 752. Painted Big-eared Mouse (Auliscomys pictus), 753. Bolivian Big-eared Mouse (Auliscomys boliviensis), 754. Andean Big-eared Mouse (Auliscomys sublimis), 755. Sumichrasts Vesper Rat (Nyctomys sumichrasti), 756. Yucatan Vesper Rat (Otonyctomys hatti), 757. Big-eared Climbing Rat (Ototylomys phyllotis), 758. La Pera Climbing Rat (Ototylomys chiapensis), 759. Peterss Climbing Rat (Tylomys nudicaudus), 760. Chiapan Climbing Rat (Tylomys bullaris), 761. Tumbala Climbing Rat 1 (ylomys tumbalensis), 762. Watsons Climbing Rat (Tylomys watsoni), 763. Fulvous-bellied Climbing Rat (Tylomys fulviventer), 764. Panama Climbing Rat (Tylomys panamensis), 765. Mira Climbing Rat (Tylomys mirae)" figureDoi="http://doi.org/10.5281/zenodo.6708613" httpUri="https://zenodo.org/record/6708613/files/figure.png" pageId="256" pageNumber="535">764.</figureCitation>
</heading>
</paragraph>
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<subSubSection id="C3438F8EFEAE21670B3810920FEDFB27" box="[1551,2010,1230,1276]" pageId="256" pageNumber="535" type="vernacular_names">
<paragraph id="8BE6DC05FEAE21670B3810920FEDFB27" blockId="256.[1449,2444,1230,1356]" box="[1551,2010,1230,1276]" pageId="256" pageNumber="535">
<heading id="D0AE6B69FEAE21670B3810920FEDFB27" box="[1551,2010,1230,1276]" pageId="256" pageNumber="535">
<vernacularName id="055AAC2BFEAE21670B3810920FEDFB27" ID-CoL="59THK" baseAuthorityName="Gray" baseAuthorityYear="1873" box="[1551,2010,1230,1276]" class="Mammalia" family="Cricetidae" genus="Tylomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="256" pageNumber="535" phylum="Chordata" rank="species" species="panamensis">Panama Climbing Rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE21670514109201BCFB27" box="[2083,2443,1230,1276]" pageId="256" pageNumber="535" type="nomenclature">
<paragraph id="8BE6DC05FEAE21670514109201BCFB27" blockId="256.[1449,2444,1230,1356]" box="[2083,2443,1230,1276]" pageId="256" pageNumber="535">
<heading id="D0AE6B69FEAE21670514109201BCFB27" box="[2083,2443,1230,1276]" pageId="256" pageNumber="535">
<taxonomicName id="4C59A786FEAE21670514109201BCFB27" ID-CoL="59THK" baseAuthorityName="Gray" baseAuthorityYear="1873" box="[2083,2443,1230,1276]" class="Mammalia" family="Cricetidae" genus="Tylomys" kingdom="Animalia" order="Rodentia" pageId="256" pageNumber="535" phylum="Chordata" rank="species" species="panamensis">
<emphasis id="B92D0017FEAE21670514109201BCFB27" box="[2083,2443,1230,1276]" italics="true" pageId="256" pageNumber="535">Tylomys panamensis</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE2167089C11520F9DFA91" pageId="256" pageNumber="535" type="vernacular_names">
<paragraph id="8BE6DC05FEAE2167089C115201B7FAF8" blockId="256.[1449,2444,1230,1356]" box="[1451,2432,1294,1315]" pageId="256" pageNumber="535">
<heading id="D0AE6B69FEAE2167089C115201B7FAF8" box="[1451,2432,1294,1315]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE2167089C11520DC1FAF8" bold="true" box="[1451,1526,1294,1315]" pageId="256" pageNumber="535">French:</emphasis>
<vernacularName id="055AAC2BFEAE216708CC11520EF6FAF8" ID-CoL="59THK" baseAuthorityName="Gray" baseAuthorityYear="1873" box="[1531,1729,1294,1315]" class="Mammalia" family="Cricetidae" genus="Tylomys" kingdom="Animalia" language="fra" order="Rodentia" pageId="256" pageNumber="535" phylum="Chordata" rank="species" species="panamensis">Tylomys du Panama</vernacularName>
/
<emphasis id="B92D0017FEAE21670BE011520F06FAF8" bold="true" box="[1751,1841,1294,1315]" pageId="256" pageNumber="535">German:</emphasis>
<vernacularName id="055AAC2BFEAE21670A0C11520FC9FAF8" ID-CoL="59THK" baseAuthorityName="Gray" baseAuthorityYear="1873" box="[1851,2046,1294,1315]" class="Mammalia" family="Cricetidae" genus="Tylomys" kingdom="Animalia" language="deu" order="Rodentia" pageId="256" pageNumber="535" phylum="Chordata" rank="species" species="panamensis">Panama-Kletterratte</vernacularName>
/
<emphasis id="B92D0017FEAE2167052411520059FAF8" bold="true" box="[2067,2158,1294,1315]" pageId="256" pageNumber="535">Spanish:</emphasis>
<vernacularName id="055AAC2BFEAE2167054E115201B7FAF8" ID-CoL="59THK" baseAuthorityName="Gray" baseAuthorityYear="1873" box="[2169,2432,1294,1315]" class="Mammalia" family="Cricetidae" genus="Tylomys" kingdom="Animalia" language="esp" order="Rodentia" pageId="256" pageNumber="535" phylum="Chordata" rank="species" species="panamensis">Rata trepadora de Panama</vernacularName>
</heading>
</paragraph>
<paragraph id="8BE6DC05FEAE2167089D11690F9DFA91" blockId="256.[1449,2444,1230,1356]" box="[1450,1962,1333,1354]" pageId="256" pageNumber="535">
<heading id="D0AE6B69FEAE2167089D11690F9DFA91" box="[1450,1962,1333,1354]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE2167089D11690E96FA91" bold="true" box="[1450,1697,1333,1354]" pageId="256" pageNumber="535">Other common names:</emphasis>
<vernacularName id="055AAC2BFEAE21670B9C11690F9DFA91" ID-CoL="59THK" baseAuthorityName="Gray" baseAuthorityYear="1873" box="[1707,1962,1333,1354]" class="Mammalia" family="Cricetidae" genus="Tylomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="256" pageNumber="535" phylum="Chordata" rank="species" species="panamensis">Panamanian Climbing Rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE21670539112400A8FA1B" pageId="256" pageNumber="535" type="reference_group">
<paragraph id="8BE6DC05FEAE21670539112400A8FA1B" blockId="256.[2062,2656,1400,1827]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE216705391124009EFA42" bold="true" box="[2062,2217,1400,1433]" pageId="256" pageNumber="535">Taxonomy.</emphasis>
<taxonomicName id="4C59A786FEAE2167058F11240260FA42" authority="Gray, 1873" authorityName="Gray" authorityYear="1873" box="[2232,2647,1400,1433]" class="Mammalia" family="Cricetidae" genus="Neomys" kingdom="Animalia" order="Rodentia" pageId="256" pageNumber="535" phylum="Chordata" rank="species" species="panamensis">Neomys panamensis Gray, 1873</taxonomicName>
,
<materialsCitation id="3B31D658FEAE2167053811C300A8FA1B" ID-GBIF-Occurrence="3818814843" box="[2063,2207,1439,1472]" pageId="256" pageNumber="535">
<collectingCountry id="F34E9C95FEAE2167052A11C300BAFA1B" box="[2077,2189,1439,1472]" name="Panama" pageId="256" pageNumber="535">
<collectingRegion id="499D12E7FEAE2167052A11C300BAFA1B" box="[2077,2189,1439,1472]" country="Panama" name="Panama" pageId="256" pageNumber="535">Panama</collectingRegion>
</collectingCountry>
.”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE2167052511970061F9ED" pageId="256" pageNumber="535" type="discussion">
<paragraph id="8BE6DC05FEAE2167052511970061F9ED" blockId="256.[2062,2656,1400,1827]" pageId="256" pageNumber="535">
<taxonomicName id="4C59A786FEAE2167052511970126FA33" authorityName="Gray" authorityYear="1873" box="[2066,2321,1483,1512]" class="Mammalia" family="Cricetidae" genus="Tylomys" kingdom="Animalia" order="Rodentia" pageId="256" pageNumber="535" phylum="Chordata" rank="species" species="panamensis">Tylomys panamensis</taxonomicName>
is known from only two specimens collected in the 1850s. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE21670539126100E1F977" pageId="256" pageNumber="535" type="distribution">
<caption id="DF268C8DFEAE21670539126100E1F977" ID-DOI="http://doi.org/10.5281/zenodo.6708345" ID-Zenodo-Dep="6708345" httpUri="https://zenodo.org/record/6708345/files/figure.png" inLine="true" pageId="256" pageNumber="535" targetBox="[1447,2038,1407,1821]" targetPageId="256">
<paragraph id="8BE6DC05FEAE21670539126100E1F977" blockId="256.[2062,2656,1400,1827]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE2167053912610089F985" bold="true" box="[2062,2238,1597,1630]" pageId="256" pageNumber="535">Distribution.</emphasis>
E Panama at an elevation of 600 m; known localities are now in Darién National Park.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C3438F8EFEAE2167053912EF0E4DF786" pageId="256" pageNumber="535" type="description">
<paragraph id="8BE6DC05FEAE2167053912EF0E4DF786" blockId="256.[2062,2656,1400,1827]" lastBlockId="256.[1443,2655,1837,2812]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE2167053912EF012EF90F" bold="true" box="[2062,2329,1715,1748]" pageId="256" pageNumber="535">Descriptive notes.</emphasis>
Head—body 226 mm, tail 199 mm. No specific data are available for body weight. The Panama Climbing Rat is the size of the Brown Rat (
<taxonomicName id="4C59A786FEAE2167050913710109F891" baseAuthorityName="Berkenhout" baseAuthorityYear="1769" box="[2110,2366,1837,1866]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="256" pageNumber="535" phylum="Chordata" rank="species" species="norvegicus">Rattus norvegicus</taxonomicName>
). Back is dark gray (blackish washed), with long hairs; sides of head and body are quite pale; throat, chest, lower body, and interior of legs are white; and legs are brown, with white hairs on nails. Tail is bicolored, naked, shiny, and flaky; two-thirds from base are dark and remaining one-third is white. Karyotype is 2n = 52; long pair is submetacentric, two pairs are small and submetacentric, and 22 pairs are medium and small acrocentric. X-chromosome is submetacentric, and Y-chromosome is subteocentric.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE2167089E1C380ECDF720" pageId="256" pageNumber="535" type="biology_ecology">
<paragraph id="8BE6DC05FEAE2167089E1C380ECDF720" blockId="256.[1443,2655,1837,2812]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE2167089E1C380E2FF75E" bold="true" box="[1449,1560,2148,2181]" pageId="256" pageNumber="535">Habitat.</emphasis>
Tropical wet forest and tropical rainforest, composed of
<taxonomicName id="4C59A786FEAE2167046C1C380E2EF777" class="Magnoliopsida" family="Fabaceae" genus="Bauhinia" kingdom="Plantae" order="Fabales" pageId="256" pageNumber="535" phylum="Tracheophyta" rank="species" species="undetermined">Bauhinia spp. (Fabaceae)</taxonomicName>
,
<taxonomicName id="4C59A786FEAE21670B181CD30F36F777" box="[1583,1793,2191,2220]" class="Liliopsida" family="Arecaceae" genus="Oenocarpus" kingdom="Plantae" order="Arecales" pageId="256" pageNumber="535" phylum="Tracheophyta" rank="species" species="undetermined">Oenocarpus spp.</taxonomicName>
and
<taxonomicName id="4C59A786FEAE21670A661CD30FDBF777" box="[1873,2028,2191,2220]" class="Liliopsida" family="Arecaceae" genus="Attalea" kingdom="Plantae" order="Arecales" pageId="256" pageNumber="535" phylum="Tracheophyta" rank="species" species="undetermined">Attalea spp.</taxonomicName>
(both
<taxonomicName id="4C59A786FEAE216705621CD300DBF777" box="[2133,2284,2191,2220]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="256" pageNumber="535" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
), Orchids (
<taxonomicName id="4C59A786FEAE216704A61CD3027BF777" box="[2449,2636,2191,2220]" class="Liliopsida" family="Orchidaceae" kingdom="Plantae" order="Asparagales" pageId="256" pageNumber="535" phylum="Tracheophyta" rank="family">Orchidaceae</taxonomicName>
), Bromeliads (Bromelaceae), ferns, other epiphytes of family
<taxonomicName id="4C59A786FEAE216704391CEF01EDF70F" box="[2318,2522,2227,2260]" class="Liliopsida" family="Cyclanthaceae" kingdom="Plantae" order="Pandanales" pageId="256" pageNumber="535" phylum="Tracheophyta" rank="family">Cyclanthaceae</taxonomicName>
, and Anthuriums spp. (
<taxonomicName id="4C59A786FEAE21670B421C820EDAF720" box="[1653,1773,2270,2299]" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" pageId="256" pageNumber="535" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE2167089E1D5E0E35F692" pageId="256" pageNumber="535" type="food_feeding">
<paragraph id="8BE6DC05FEAE2167089E1D5E0E35F692" blockId="256.[1443,2655,1837,2812]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE2167089E1D5E0E86F6F8" bold="true" box="[1449,1713,2306,2339]" pageId="256" pageNumber="535">Food and Feeding.</emphasis>
The Panama Climbing Rat probably feeds on fruits and shoots of plants.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE2167089E1D080F29F6AA" box="[1449,1822,2388,2417]" pageId="256" pageNumber="535" type="breeding">
<paragraph id="8BE6DC05FEAE2167089E1D080F29F6AA" blockId="256.[1443,2655,1837,2812]" box="[1449,1822,2388,2417]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE2167089E1D080E18F6AA" bold="true" box="[1449,1583,2388,2417]" pageId="256" pageNumber="535">Breeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE216708901D240EA0F63C" pageId="256" pageNumber="535" type="activity">
<paragraph id="8BE6DC05FEAE216708901D240EA0F63C" blockId="256.[1443,2655,1837,2812]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE216708901D240EA5F642" bold="true" box="[1447,1682,2424,2457]" pageId="256" pageNumber="535">Activity patterns.</emphasis>
The Panama Climbing Rat is strictly nocturnal, with activity between 21:00 h and 06:00 h. It is most active around midnight and during rainy season in June-September.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE2167089E1DB20234F5ED" pageId="256" pageNumber="535" type="biology_ecology">
<paragraph id="8BE6DC05FEAE2167089E1DB20234F5ED" blockId="256.[1443,2655,1837,2812]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE2167089E1DB20050F5D4" bold="true" box="[1449,2151,2542,2575]" pageId="256" pageNumber="535">Movements, Home range and Social organization.</emphasis>
The Panama Climbing Rat is probably solitary. In Darien,it was reported moving at 17 cm/s in the jungle canopy.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE2167089E1E1C001FF577" pageId="256" pageNumber="535" type="conservation">
<paragraph id="8BE6DC05FEAE2167089E1E1C001FF577" blockId="256.[1443,2655,1837,2812]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE2167089E1E1C0F35F586" bold="true" box="[1449,1794,2624,2653]" pageId="256" pageNumber="535">Status and Conservation.</emphasis>
Classified as Data Deficient on The IUCN Red List. Data on distribution, threats,status, ecological requirements, and taxonomic validity are needed to make an accurate conservation assessment.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFEAE2167089E1E9C016AF527" pageId="256" pageNumber="535" type="bibRefCitation_list">
<paragraph id="8BE6DC05FEAE2167089E1E9C016AF527" blockId="256.[1443,2655,1837,2812]" pageId="256" pageNumber="535">
<emphasis id="B92D0017FEAE2167089E1E9C0E76F50E" bold="true" box="[1449,1601,2752,2773]" pageId="256" pageNumber="535">Bibliography.</emphasis>
Espinoza (2002), Goodwin (1955b), Handley (1966), Méndez-Carvajal et al. (2015), Musser &amp; Carleton (2005), Nowak (1999), Pathak et al. (1973), Reid (2009), Wilson &amp; Cole (2000).
</paragraph>
</subSubSection>
</treatment>
</document>