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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1000.59219" ID-GBIF-Dataset="0bdadccc-2790-4c0d-9e04-93845c37e703" ID-PMC="PMC7728727" ID-Pensoft-Pub="1313-2970-1000-31" ID-Pensoft-UUID="D4CBC05E57B055C28B6064FFD38C050E" ID-PubMed="33354133" ID-ZooBank="C7810741557041768F5611BA165889FA" ModsDocID="1313-2970-1000-31" checkinTime="1607107171358" checkinUser="pensoft" docAuthor="Scheffrahn, Rudolf H. &amp; Carrijo, Tiago F." docDate="2020" docId="E821B83226375A5CBC17713977CA1C57" docLanguage="en" docName="ZooKeys 1000: 31-44" docOrigin="ZooKeys 1000" docSource="http://dx.doi.org/10.3897/zookeys.1000.59219" docTitle="Rugitermes rufus Scheffrahn 2020, sp. nov." docType="treatment" docUuid="F4999D2D-8839-43CE-9C91-BDBDC146C59B" docUuidSource="ZooBank" docVersion="5" id="D4CBC05E57B055C28B6064FFD38C050E" lastPageNumber="31" masterDocId="D4CBC05E57B055C28B6064FFD38C050E" masterDocTitle="Three new species of Rugitermes (Isoptera, Kalotermitidae) from Peru and Bolivia" masterLastPageNumber="44" masterPageNumber="31" pageNumber="31" updateTime="1668146359621" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new species of Rugitermes (Isoptera, Kalotermitidae) from Peru and Bolivia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scheffrahn, Rudolf H.</mods:namePart>
<mods:affiliation>University of Florida, Fort Lauderdale Research &amp; Education Center, 3205 College Avenue, Davie, Florida 33314, USA</mods:affiliation>
<mods:nameIdentifier type="email">rhsc@ufl.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Carrijo, Tiago F.</mods:namePart>
<mods:affiliation>Centro de Ciencias Naturais e Humanas, Universidade Federal do ABC, Rua Arcturus 03, Jardim Antares, 09606 - 070 Sao Bernardo do Campo, SP, Brazil</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6308-7252</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>1000</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>31</mods:start>
<mods:end>44</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1000.59219</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1000.59219</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1000-31</mods:identifier>
<mods:identifier type="ZooBank">C7810741557041768F5611BA165889FA</mods:identifier>
<mods:identifier type="Pensoft-UUID">D4CBC05E57B055C28B6064FFD38C050E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="170677953" LSID="urn:lsid:zoobank.org:act:F4999D2D-8839-43CE-9C91-BDBDC146C59B" httpUri="http://treatment.plazi.org/id/E821B83226375A5CBC17713977CA1C57" lastPageNumber="31" pageId="0" pageNumber="31">
<subSubSection pageId="0" pageNumber="31" type="nomenclature">
<paragraph pageId="0" pageNumber="31">
<taxonomicName LSID="E821B832-2637-5A5C-BC17-713977CA1C57" authority="Scheffrahn" authorityName="Scheffrahn" authorityYear="2020" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugitermes rufus" order="Isoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="rufus" status="sp. nov.">Rugitermes rufus Scheffrahn</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="31">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head and thorax of the Rugitermes rufus (A dorsal B lateral) and the R. volcanensis (C dorsal D lateral) imagos." figureDoi="10.3897/zookeys.1000.59219.figure4" httpUri="https://binary.pensoft.net/fig/482870" pageId="0" pageNumber="31">Figures 4A, B</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Head and pronotum of the Rugitermes rufus soldier A dorsal B right (arrow pointing to outer margin of anterolateral ridge) C lateral (arrow pointing to antennal carinae) D ventral." figureDoi="10.3897/zookeys.1000.59219.figure5" httpUri="https://binary.pensoft.net/fig/482871" pageId="0" pageNumber="31">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Field habitus of the imago and soldiers of Rugitermes rufus." figureDoi="10.3897/zookeys.1000.59219.figure6" httpUri="https://binary.pensoft.net/fig/482872" pageId="0" pageNumber="31">, 6</figureCitation>
;
<figureCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (mm) of Rugitermes soldiers." pageId="0" pageNumber="31">Tables 1</figureCitation>
,
<figureCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements (mm) of Rugitermes dealated imagos." pageId="0" pageNumber="31">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="diagnosis">
<paragraph pageId="0" pageNumber="31">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="31">
The coloration and size of the
<taxonomicName lsidName="R. rufus" pageId="0" pageNumber="31" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">R. rufus</emphasis>
</taxonomicName>
imago easily distinguish it from all other species of
<taxonomicName authorityName="Holmgren" authorityYear="1911" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugitermes" order="Blattodea" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Rugitermes</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Scheffrahn" authorityYear="2020" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugitermes rufus" order="Blattodea" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">Rugitermes rufus</emphasis>
</taxonomicName>
is the only
<taxonomicName authorityName="Holmgren" authorityYear="1911" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugitermes" order="Blattodea" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Rugitermes</emphasis>
</taxonomicName>
imago with a reddish-orange head capsule and dark brown coloration of the pronotum and remainder of the body.
<taxonomicName baseAuthorityName="Snyder" baseAuthorityYear="1926" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugitermes costaricensis" order="Blattodea" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis italics="true" pageId="0" pageNumber="31">Rugitermes costaricensis</emphasis>
</taxonomicName>
(Snyder, 1929) and
<taxonomicName lsidName="R. unicolor" pageId="0" pageNumber="31" rank="species" species="unicolor">
<emphasis italics="true" pageId="0" pageNumber="31">R. unicolor</emphasis>
</taxonomicName>
Snyder, 1952 imagos each have a yellow-brown head and yellow pronotum (with less contrast between coloration of head and pronotum in
<taxonomicName lsidName="R. unicolor" pageId="0" pageNumber="31" rank="species" species="unicolor">
<emphasis italics="true" pageId="0" pageNumber="31">R. unicolor</emphasis>
</taxonomicName>
). In all other
<taxonomicName authorityName="Holmgren" authorityYear="1911" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugitermes" order="Blattodea" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Rugitermes</emphasis>
</taxonomicName>
, the imago head is black and the pronotum is yellow [
<taxonomicName lsidName="R. bicolor" pageId="0" pageNumber="31" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="31">R. bicolor</emphasis>
</taxonomicName>
(Emerson, 1926),
<taxonomicName lsidName="R. flavicinctus" pageId="0" pageNumber="31" rank="species" species="flavicinctus">
<emphasis italics="true" pageId="0" pageNumber="31">R. flavicinctus</emphasis>
</taxonomicName>
(Emerson, 1925),
<taxonomicName lsidName="R. magninotus" pageId="0" pageNumber="31" rank="species" species="magninotus">
<emphasis italics="true" pageId="0" pageNumber="31">R. magninotus</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Field habitus of the imagos and soldiers of Rugitermes magninotus from Peru (UFTC no. PU 222)." figureDoi="10.3897/zookeys.1000.59219.figure7" httpUri="https://binary.pensoft.net/fig/482873" pageId="0" pageNumber="31">7</figureCitation>
),
<taxonomicName lsidName="R. kirbyi" pageId="0" pageNumber="31" rank="species" species="kirbyi">
<emphasis italics="true" pageId="0" pageNumber="31">R. kirbyi</emphasis>
</taxonomicName>
(Snyder, 1926),
<taxonomicName lsidName="R. nodulosus" pageId="0" pageNumber="31" rank="species" species="nodulosus">
<emphasis italics="true" pageId="0" pageNumber="31">R. nodulosus</emphasis>
</taxonomicName>
Oliveira, 1979,
<taxonomicName lsidName="R. panamae" pageId="0" pageNumber="31" rank="species" species="panamae">
<emphasis italics="true" pageId="0" pageNumber="31">R. panamae</emphasis>
</taxonomicName>
(Snyder, 1925), and
<taxonomicName lsidName="R. rugosus" pageId="0" pageNumber="31" rank="species" species="rugosus">
<emphasis italics="true" pageId="0" pageNumber="31">R. rugosus</emphasis>
</taxonomicName>
(Hagen, 1858)] or the head and pronotum are both black [(
<taxonomicName lsidName="R. athertoni" pageId="0" pageNumber="31" rank="species" species="athertoni">
<emphasis italics="true" pageId="0" pageNumber="31">R. athertoni</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="R. laticollis" pageId="0" pageNumber="31" rank="species" species="laticollis">
<emphasis italics="true" pageId="0" pageNumber="31">R. laticollis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="R. niger" pageId="0" pageNumber="31" rank="species" species="niger">
<emphasis italics="true" pageId="0" pageNumber="31">R. niger</emphasis>
</taxonomicName>
Oliveira, 1979, and
<taxonomicName lsidName="R. occidentalis" pageId="0" pageNumber="31" rank="species" species="occidentalis">
<emphasis italics="true" pageId="0" pageNumber="31">R. occidentalis</emphasis>
</taxonomicName>
(Silvestri, 1901)]. The head width of the
<taxonomicName lsidName="R. rufus" pageId="0" pageNumber="31" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">R. rufus</emphasis>
</taxonomicName>
imago is among the smallest: in range of
<taxonomicName lsidName="R. flavicinctus" pageId="0" pageNumber="31" rank="species" species="flavicinctus">
<emphasis italics="true" pageId="0" pageNumber="31">R. flavicinctus</emphasis>
</taxonomicName>
(1.2 mm) and
<taxonomicName lsidName="R. magninotus" pageId="0" pageNumber="31" rank="species" species="magninotus">
<emphasis italics="true" pageId="0" pageNumber="31">R. magninotus</emphasis>
</taxonomicName>
(1.25 mm). The soldier of
<taxonomicName lsidName="R. rufus" pageId="0" pageNumber="31" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">R. rufus</emphasis>
</taxonomicName>
is smaller in all measurements compared with
<taxonomicName lsidName="R. volcanensis" pageId="0" pageNumber="31" rank="species" species="volcanensis">
<emphasis italics="true" pageId="0" pageNumber="31">R. volcanensis</emphasis>
</taxonomicName>
. The antennal carina in
<taxonomicName lsidName="R. rufus" pageId="0" pageNumber="31" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">R. rufus</emphasis>
</taxonomicName>
forms a smaller ovoid shelf of the gena and, in lateral oblique view, projects acutely from the frontal margin. In
<taxonomicName lsidName="R. volcanensis" pageId="0" pageNumber="31" rank="species" species="volcanensis">
<emphasis italics="true" pageId="0" pageNumber="31">R. volcanensis</emphasis>
</taxonomicName>
, the shelf formed by the antennal carina is larger and, in the lateral oblique view, weakly projects above the frontal margin. The third antennal article of
<taxonomicName lsidName="R. rufus" pageId="0" pageNumber="31" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">R. rufus</emphasis>
</taxonomicName>
is proportionally narrower and longer than that of
<taxonomicName lsidName="R. volcanensis" pageId="0" pageNumber="31" rank="species" species="volcanensis">
<emphasis italics="true" pageId="0" pageNumber="31">R. volcanensis</emphasis>
</taxonomicName>
. See also diagnosis of
<taxonomicName lsidName="R. aridus" pageId="0" pageNumber="31" rank="species" species="aridus">
<emphasis italics="true" pageId="0" pageNumber="31">R. aridus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="type locality">
<paragraph pageId="0" pageNumber="31">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="31">
Bolivia,
<normalizedToken originalValue="Amboró">Amboro</normalizedToken>
National Park, Refugio Los Volcanes, Santa Cruz District (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Biomes for new Rugitermes A R. aridus B R. rufus and R. volcanensis." figureDoi="10.3897/zookeys.1000.59219.figure9" httpUri="https://binary.pensoft.net/fig/482875" pageId="0" pageNumber="31">9B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="materials_examined">
<paragraph pageId="0" pageNumber="31">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="31">
<materialsCitation ID-GBIF-Occurrence="2989740311" collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectionCode="UFTC" collectorName="R. Scheffrahn" elevation="0" latitude="-18.11968" location="Amboro National Park" longLatPrecision="1" longitude="-63.60881" municipality="Bolivia" specimenCode="B01021" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">
<typeStatus>Holotype</typeStatus>
.
</emphasis>
One
<specimenCount type="female">female</specimenCount>
delate imago:
<collectingMunicipality>Bolivia</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E821B83226375A5CBC17713977CA1C57:1DEF03C7FA911F42E8861E59BA2949B1" latitude="-18.11968" longLatPrecision="1" longitude="-63.60881" municipality="Bolivia" name="Amboro National Park">
<normalizedToken originalValue="Amboró">Amboro</normalizedToken>
National Park
</location>
,
<location LSID="urn:lsid:plazi:treatment:E821B83226375A5CBC17713977CA1C57:53DDA05EAE650411E4DC6FA581A4CB23" latitude="-18.11968" longLatPrecision="1" longitude="-63.60881" municipality="Bolivia" name="hillside above Refugio Los Volcanes">hillside above Refugio Los Volcanes</location>
,
<location LSID="urn:lsid:plazi:treatment:E821B83226375A5CBC17713977CA1C57:60347A76EF09347D089ED03665C0D5DB" latitude="-18.11968" longLatPrecision="1" longitude="-63.60881" municipality="Bolivia" name="Santa Cruz District">Santa Cruz District</location>
(
<geoCoordinate degrees="18.119680" direction="south" orientation="latitude" precision="1" value="-18.11968">-18.119680</geoCoordinate>
,
<geoCoordinate degrees="63.608810" direction="west" orientation="longitude" precision="1" value="-63.60881">-63.608810</geoCoordinate>
) elev.
<elevation>1402 m</elevation>
,
<collectingDate>2 Jun. 2014</collectingDate>
,
<collectorName>R. Scheffrahn</collectorName>
et al.,
<collectionCode>UFTC</collectionCode>
no.
<specimenCode>B01021</specimenCode>
<emphasis bold="true" pageId="0" pageNumber="31">
.
<materialsCitation ID-GBIF-Occurrence="2989740305" collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectionCode="UFTC" collectorName="R. Scheffrahn" elevation="0" latitude="-18.11968" location="Amboro National Park" longLatPrecision="1" longitude="-63.60881" municipality="Bolivia" specimenCount="2" specimenCount-soldier="1" typeStatus="paratype">
<emphasis italics="true" pageId="0" pageNumber="31">
<typeStatus>Paratypes</typeStatus>
</emphasis>
. Three
<specimenCount type="soldier">soldiers</specimenCount>
,
<specimenCount>pseudergates</specimenCount>
, same colony sample as holotype.
</materialsCitation>
</emphasis>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2989740307" collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectionCode="UFTC" collectorName="R. Scheffrahn" elevation="0" latitude="-18.1004" location="Amboro National Park" longLatPrecision="7" longitude="-63.5934" municipality="Bolivia" specimenCode="BO1055" specimenCount="2">
<specimenCount>Five soldiers</specimenCount>
,
<specimenCount>pseudergates</specimenCount>
,
<collectingMunicipality>Bolivia</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E821B83226375A5CBC17713977CA1C57:C8DBEA7AC486228F0070D6FDC7347BE4" latitude="-18.1004" longLatPrecision="7" longitude="-63.5934" municipality="Bolivia" name="Amboro National Park">
<normalizedToken originalValue="Amboró">Amboro</normalizedToken>
National Park
</location>
,
<location LSID="urn:lsid:plazi:treatment:E821B83226375A5CBC17713977CA1C57:383F9999BD199E3F628B6A25BCDF5515" latitude="-18.1004" longLatPrecision="7" longitude="-63.5934" municipality="Bolivia" name="forest surrounding Refugio Los Volcanes">forest surrounding Refugio Los Volcanes</location>
,
<location LSID="urn:lsid:plazi:treatment:E821B83226375A5CBC17713977CA1C57:89F8830DA5D75C29FE21FF7451C1E721" latitude="-18.1004" longLatPrecision="7" longitude="-63.5934" municipality="Bolivia" name="Santa Cruz District">Santa Cruz District</location>
(
<geoCoordinate degrees="18.1004" direction="south" orientation="latitude" precision="5" value="-18.1004">-18.1004</geoCoordinate>
,
<geoCoordinate degrees="63.5934" direction="west" orientation="longitude" precision="5" value="-63.5934">-63.5934</geoCoordinate>
) elev.
<elevation>1374 m</elevation>
,
<collectingDate>3 Jun. 2014</collectingDate>
,
<collectorName>R. Scheffrahn</collectorName>
et al.,
<collectionCode>UFTC</collectionCode>
no.
<specimenCode>BO1055</specimenCode>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2989740302" collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectionCode="BO" collectorName="R. Scheffrahn" elevation="0" latitude="-18.1004" location="Four" longLatPrecision="7" longitude="-63.5934" municipality="Bolivia" specimenCode="BO1061, BO1062, BO1070" specimenCount="7">
<location LSID="urn:lsid:plazi:treatment:E821B83226375A5CBC17713977CA1C57:44E1B5D8154FEF885CB5D3C4453E0907" name="Four">Four</location>
colonies, same locality as BO1055:
<specimenCount>three soldiers</specimenCount>
,
<specimenCount>pseudergates</specimenCount>
(
<specimenCode>BO1061</specimenCode>
),
<specimenCount>four soldiers</specimenCount>
,
<specimenCount>pseudergates</specimenCount>
(
<specimenCode>BO1062</specimenCode>
),
<specimenCount>two male</specimenCount>
<specimenCount>dealate imagos</specimenCount>
,
<specimenCount>pseudergates</specimenCount>
(
<specimenCode>BO1070</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="description">
<paragraph pageId="0" pageNumber="31">Description.</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Dealated imago</emphasis>
(Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head and thorax of the Rugitermes rufus (A dorsal B lateral) and the R. volcanensis (C dorsal D lateral) imagos." figureDoi="10.3897/zookeys.1000.59219.figure4" httpUri="https://binary.pensoft.net/fig/482870" pageId="0" pageNumber="31">4A, B</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Field habitus of the imago and soldiers of Rugitermes rufus." figureDoi="10.3897/zookeys.1000.59219.figure6" httpUri="https://binary.pensoft.net/fig/482872" pageId="0" pageNumber="31">6</figureCitation>
). Head capsule reddish orange; pronotum, thorax, abdomen, and wing scales concolorous dark brown. Compound eye small; trapezoidal; with shorter straight margin below ocellus and more curved margin near gena. Ocellus ovoid, concolorous with interior of antennal socket; one-half diameter removed from eye margin. Head vertex and frons covered with about a dozen erect setae ca 0.15-mm-long; frons darker, with faint rugosity at frontal flange. Pronotum slightly wider than head capsule; anterior margin slightly incised; anterolateral corners nearly rectate, posterior margin narrowly concave; margins with few longer and many shorter setae. Pronotum interior pilosity congruent with vertex. Antennae with basal article relative lengths 1&gt;2=3&lt;4. Wing scale covered with about 20 setae of similar length and density as those on head and pronotum. Legs with femora light orange-brown; tibia much lighter. Arolium present. Measurements of three imagoes from two colonies are shown in Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements (mm) of Rugitermes dealated imagos." httpUri="http://table.plazi.org/id/4618DD7D7C048B84F497B0C28CBE3C44" pageId="0" pageNumber="31" tableUuid="4618DD7D7C048B84F497B0C28CBE3C44">2</tableCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1000.59219.figure4" httpUri="https://binary.pensoft.net/fig/482870" pageId="0" pageNumber="31" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figure 4.</emphasis>
Head and thorax of the
<taxonomicName authorityName="Scheffrahn" authorityYear="2020" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugitermes rufus" order="Blattodea" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">Rugitermes rufus</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="31">A</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="31">B</emphasis>
lateral) and the
<taxonomicName lsidName="R. volcanensis" pageId="0" pageNumber="31" rank="species" species="volcanensis">
<emphasis italics="true" pageId="0" pageNumber="31">R. volcanensis</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="31">C</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="31">D</emphasis>
lateral) imagos.
</paragraph>
</caption>
<caption ID-Table-UUID="4618DD7D7C048B84F497B0C28CBE3C44" httpUri="http://table.plazi.org/id/4618DD7D7C048B84F497B0C28CBE3C44" pageId="0" pageNumber="31" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Table 2.</emphasis>
Measurements (mm) of
<taxonomicName authorityName="Holmgren" authorityYear="1911" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugitermes" order="Blattodea" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Rugitermes</emphasis>
</taxonomicName>
dealated imagos.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="31">
<table pageId="0" pageNumber="31">
<tr pageId="0" pageNumber="31">
<th colspan="1" pageId="0" pageNumber="31" rowspan="2">Character</th>
<th colspan="1" pageId="0" pageNumber="31" rowspan="1">
<taxonomicName lsidName="R. rufus" pageId="0" pageNumber="31" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">R. rufus</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="31" rowspan="1">
<taxonomicName lsidName="R. rufus" pageId="0" pageNumber="31" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">R. rufus</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="31" rowspan="1">
<taxonomicName lsidName="R. volcanensis" pageId="0" pageNumber="31" rank="species" species="volcanensis">
<emphasis italics="true" pageId="0" pageNumber="31">R. volcanensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="31">
<th colspan="1" pageId="0" pageNumber="31" rowspan="1">Range</th>
<th colspan="1" pageId="0" pageNumber="31" rowspan="1">Mean</th>
<th colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</th>
</tr>
<tr pageId="0" pageNumber="31">
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">Head width, maximum (w/eyes)</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">1.19-1.26</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">1.23</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">1.21</td>
</tr>
<tr pageId="0" pageNumber="31">
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">Pronotum width, max.</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">1.21-1.29</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">1.24</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">1.27</td>
</tr>
<tr pageId="0" pageNumber="31">
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">Eye diam. ocellus, max.</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">0.08-0.10</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">0.09</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">0.07</td>
</tr>
<tr pageId="0" pageNumber="31">
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">Eye diam. compound, max.</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">0.27-0.30</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">0.28</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">0.27</td>
</tr>
<tr pageId="0" pageNumber="31">
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">Body length</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">6.00-6.90</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">6.33</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">7.2</td>
</tr>
<tr pageId="0" pageNumber="31">
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">Right forewing length</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="31">
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">Body length with wings</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="31">
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">No. antennal articles, max.</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">9 (broken)</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">10 (broken)</td>
</tr>
<tr pageId="0" pageNumber="31">
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="31">n</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">3</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">1</td>
</tr>
<tr pageId="0" pageNumber="31">
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">No. colonies</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">2</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="31" rowspan="1">1</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Soldier</emphasis>
(Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Head and pronotum of the Rugitermes rufus soldier A dorsal B right (arrow pointing to outer margin of anterolateral ridge) C lateral (arrow pointing to antennal carinae) D ventral." figureDoi="10.3897/zookeys.1000.59219.figure5" httpUri="https://binary.pensoft.net/fig/482871" pageId="0" pageNumber="31">5</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Field habitus of the imago and soldiers of Rugitermes rufus." figureDoi="10.3897/zookeys.1000.59219.figure6" httpUri="https://binary.pensoft.net/fig/482872" pageId="0" pageNumber="31">6</figureCitation>
). Head capsule, in dorsal view, dark brown along margin of anteclypeus, anterolateral ridges, and antennal carinae grading to orange brown at frons, orange at mid-capsule and yellow at occiput; mandibles nearly black throughout. In ventral view, postmentum light orange-brown contrasting with light yellowish head capsule. Head capsule long, rectangular; lateral margins parallel, covered with a few setae except at occiput. Pronotum much wider than long; with a few long setae at margins and middle. Frontal flange and frons forming shallow furrow; angled 40° with plane of vertex. In dorsal view, antennal carina forms small ovoid shelf over the gena. In lateral oblique view, antennal carinae project acutely from the lateral margin of frons (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Head and pronotum of the Rugitermes rufus soldier A dorsal B right (arrow pointing to outer margin of anterolateral ridge) C lateral (arrow pointing to antennal carinae) D ventral." figureDoi="10.3897/zookeys.1000.59219.figure5" httpUri="https://binary.pensoft.net/fig/482871" pageId="0" pageNumber="31">5C</figureCitation>
). Antennal carinae and anterolateral ridges weakly rugose. The outer margin of each ridge form angle of ca 110° (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Head and pronotum of the Rugitermes rufus soldier A dorsal B right (arrow pointing to outer margin of anterolateral ridge) C lateral (arrow pointing to antennal carinae) D ventral." figureDoi="10.3897/zookeys.1000.59219.figure5" httpUri="https://binary.pensoft.net/fig/482871" pageId="0" pageNumber="31">5B</figureCitation>
). Eye spots large, narrowly oblique, barely lighter than head capsule. Third antennal article narrow at the base and longer than second and fourth article combined. Mandibles about one-half length of head capsule; tip curves about 60°; outer margin of mid-blade slightly curved, shallow hump at base. Measurements of 14 soldiers from four colonies are shown in Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (mm) of Rugitermes soldiers." httpUri="http://table.plazi.org/id/8DABB5E3DDCEACD2D17CDFFD32E2FB91" pageId="0" pageNumber="31" tableUuid="8DABB5E3DDCEACD2D17CDFFD32E2FB91">1</tableCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1000.59219.figure5" httpUri="https://binary.pensoft.net/fig/482871" pageId="0" pageNumber="31" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figure 5.</emphasis>
Head and pronotum of the
<taxonomicName authorityName="Scheffrahn" authorityYear="2020" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugitermes rufus" order="Blattodea" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">Rugitermes rufus</emphasis>
</taxonomicName>
soldier
<emphasis bold="true" pageId="0" pageNumber="31">A</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="31">B</emphasis>
right (arrow pointing to outer margin of anterolateral ridge)
<emphasis bold="true" pageId="0" pageNumber="31">C</emphasis>
lateral (arrow pointing to antennal carinae)
<emphasis bold="true" pageId="0" pageNumber="31">D</emphasis>
ventral.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1000.59219.figure6" httpUri="https://binary.pensoft.net/fig/482872" pageId="0" pageNumber="31" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figure 6.</emphasis>
Field habitus of the imago and soldiers of
<taxonomicName authorityName="Scheffrahn" authorityYear="2020" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugitermes rufus" order="Blattodea" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="rufus">
<emphasis italics="true" pageId="0" pageNumber="31">Rugitermes rufus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="distribution">
<paragraph pageId="0" pageNumber="31">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="31">
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Localities of the new Rugitermes species described herein." figureDoi="10.3897/zookeys.1000.59219.figure1" httpUri="https://binary.pensoft.net/fig/482867" pageId="0" pageNumber="31">1</figureCitation>
).
<normalizedToken originalValue="Amboró">Amboro</normalizedToken>
National Park, a transition of the Tucuman-Bolivian forest and the montane Chaco.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="etymology">
<paragraph pageId="0" pageNumber="31">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="31">
The species name &quot;
<taxonomicName authorityName="Scheffrahn" authorityYear="2020" class="Insecta" family="Kalotermitidae" genus="Rugitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="rufus" order="Blattodea" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="rufus">rufus</taxonomicName>
&quot; describes the unique reddish-orange coloration of the imago head.
</paragraph>
</subSubSection>
</treatment>
</document>
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