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<document id="B67BC75F66A54334C170AAD441D5564F" ID-CLB-Dataset="39207" ID-DOI="10.11646/zootaxa.4060.1.4" ID-GBIF-Dataset="20c1a284-64ae-45bf-985c-9d9ba57db703" ID-ISSN="1175-5326" ID-Zenodo-Dep="244553" ID-ZooBank="19B6DC57-023E-415B-936A-09B5DE8E5DBF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461364819441" checkinUser="plazi" docAuthor="Tighe, Kenneth A., Pogonoski, John J., Hibino, Yusuke, Ho, Hsuan-Ching &amp; Nguyen, Quan Van" docDate="2015" docId="03DA87F2FF903845FF22FB24FA5EFDBF" docLanguage="en" docName="zt04060p029.pdf" docOrigin="Zootaxa 4060 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlopsis nanhaiensis Tighe, Ho, Pogonoski &amp; Hibino, sp. nov." docType="treatment" docUuid="EB34A0AD-CE2D-48D0-86E6-6A0BD7A39B7B" docUuidSource="ZooBank" docVersion="10" lastPageNumber="25" masterDocId="FFE3FF8AFF933843FFB5FFD8FFD8FFDF" masterDocTitle="Three new species of the genus Chlopsis (Anguilliformes: Chlopsidae) from the Indo-Pacific" masterLastPageNumber="29" masterPageNumber="19" pageNumber="22" updateTime="1698661925525" updateUser="ExternalLinkService">
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<mods:title id="4A25BCB33F5707AD82916943E2F34F30">Three new species of the genus Chlopsis (Anguilliformes: Chlopsidae) from the Indo-Pacific</mods:title>
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<mods:namePart id="078FCBE4C1CB65033045E7159B359BEA">Tighe, Kenneth A.</mods:namePart>
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<mods:namePart id="DC6ABA0B9CABEFA1CC9CD07012E5A5AD">Pogonoski, John J.</mods:namePart>
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<mods:namePart id="B31F110EF503FADF040CB2A98CFAB531">Hibino, Yusuke</mods:namePart>
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<mods:namePart id="FBDB8615F761AD9B2461B5D83A318FFF">Ho, Hsuan-Ching</mods:namePart>
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<mods:namePart id="06B7196F770BE9F0AF8198D551E3E943">Nguyen, Quan Van</mods:namePart>
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<treatment id="03DA87F2FF903845FF22FB24FA5EFDBF" ID-DOI="http://doi.org/10.5281/zenodo.6094880" ID-GBIF-Taxon="119649111" ID-Zenodo-Dep="6094880" LSID="urn:lsid:zoobank.org:act:EB34A0AD-CE2D-48D0-86E6-6A0BD7A39B7B" httpUri="http://treatment.plazi.org/id/03DA87F2FF903845FF22FB24FA5EFDBF" lastPageId="6" lastPageNumber="25" pageId="3" pageNumber="22">
<subSubSection id="C369656FFF903840FF22FB24FEEFFAE6" pageId="3" pageNumber="22" type="nomenclature">
<paragraph id="8BCC36E4FF903840FF22FB24FCAAFAC9" blockId="3.[151,882,1276,1338]" box="[151,882,1276,1302]" pageId="3" pageNumber="22">
<heading id="D0848188FF903840FF22FB24FCAAFAC9" bold="true" box="[151,882,1276,1302]" fontSize="10" level="2" pageId="3" pageNumber="22" reason="2">
<emphasis id="B907EAF6FF903840FF22FB24FCAAFAC9" bold="true" box="[151,882,1276,1302]" pageId="3" pageNumber="22">
<taxonomicName id="4C734D67FF903840FF22FB24FCD4FAC9" ID-CoL="5XWVC" authority="Tighe, Ho, Pogonoski &amp; Hibino" authorityName="Tighe, Ho, Pogonoski &amp; Hibino" box="[151,780,1276,1302]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="nanhaiensis" status="sp. nov.">
<emphasis id="B907EAF6FF903840FF22FB24FE55FACA" bold="true" box="[151,397,1276,1301]" italics="true" pageId="3" pageNumber="22">Chlopsis nanhaiensis</emphasis>
Tighe, Ho, Pogonoski &amp; Hibino
</taxonomicName>
,
<taxonomicNameLabel id="A234578DFF903840FCACFB25FCAAFAC9" box="[793,882,1277,1302]" pageId="3" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC36E4FF903840FF22FAF9FEEFFAE6" blockId="3.[151,882,1276,1338]" box="[151,311,1313,1338]" pageId="3" pageNumber="22">
<figureCitation id="13482A61FF903840FF22FAF9FF3DFAE6" box="[151,229,1313,1338]" captionStart="FIGURE 4" captionStartId="4.[151,244,1099,1121]" captionTargetBox="[151,1435,191,1057]" captionTargetId="figure@4.[151,1435,191,1078]" captionTargetPageId="4" captionText="FIGURE 4. Holotype of Chlopsis nanhaiensis, USNM 408189, 228 mm TL; Dong-gang fish market, Pingtung, Taiwan." httpUri="https://zenodo.org/record/244557/files/figure.png" pageId="3" pageNumber="22">Figs. 4</figureCitation>
,
<figureCitation id="13482A61FF903840FF44FAF9FED9FAE5" box="[241,257,1313,1338]" captionStart="FIGURE 5" captionStartId="4.[151,244,1741,1763]" captionTargetBox="[239,1337,1468,1706]" captionTargetId="figure@4.[233,1354,1457,1720]" captionTargetPageId="4" captionText="FIGURE 5. Head of holotype of Chlopsis nanhaiensis, USNM 408189." httpUri="https://zenodo.org/record/244558/files/figure.png" pageId="3" pageNumber="22">5</figureCitation>
,
<figureCitation id="13482A61FF903840FEB8FAF9FEC5FAE5" box="[269,285,1313,1338]" captionStart="FIGURE 6" captionStartId="5.[151,244,645,667]" captionTargetBox="[298,1286,309,613]" captionTargetId="figure@5.[288,1299,299,624]" captionTargetPageId="5" captionText="FIGURE 6. Upper (A) and lower (B) jaws of holotype of Chlopsis nanhaiensis, USNM 408189." httpUri="https://zenodo.org/record/244559/files/figure.png" pageId="3" pageNumber="22">6</figureCitation>
,
<figureCitation id="13482A61FF903840FE9CFAFAFEEFFAE6" box="[297,311,1314,1337]" captionStart="FIGURE 7" captionStartId="5.[151,244,1990,2012]" captionTargetBox="[151,1436,1111,1966]" captionTargetId="figure@5.[151,1436,1111,1967]" captionTargetPageId="5" captionText="FIGURE 7. Paratype of Chlopsis nanhaiensis, NMMB-P 22369, 270 + mm TL; Dong-gang fish market, Pingtung, Taiwan." httpUri="https://zenodo.org/record/244560/files/figure.png" pageId="3" pageNumber="22">7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C369656FFF903840FF22FAB1FEA6FA31" pageId="3" pageNumber="22" type="materials_examined">
<paragraph id="8BCC36E4FF903840FF22FAB1FE87FA79" blockId="3.[151,1437,1385,2022]" pageId="3" pageNumber="22">
<emphasis id="B907EAF6FF903840FF22FAB1FED5FA5D" bold="true" box="[151,269,1385,1410]" pageId="3" pageNumber="22">
<typeStatus id="54C88846FF903840FF22FAB1FED0FA5D" box="[151,264,1385,1410]" pageId="3" pageNumber="22" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED62AE21FF903840FEA3FAB1FEBFFA5E" LSID="urn:lsid:biocol.org:col:34871" box="[278,359,1385,1409]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="22">USNM</collectionCode>
408189 (
<quantity id="4C8B9B01FF903840FE63FAB1FDEFFA5E" box="[470,567,1385,1409]" metricMagnitude="-1" metricUnit="m" metricValue="2.28" pageId="3" pageNumber="22" unit="mm" value="228.0">228 mm</quantity>
total length); Dong-gang fish market, Pingtung,
<collectingCountry id="F3647674FF903840FBDCFAB1FB66FA5D" box="[1129,1214,1385,1410]" name="Taiwan" pageId="3" pageNumber="22">Taiwan</collectingCountry>
, collected by D. G. Smith,
<date id="FFCD1024FF903840FF5DFA56FE83FA79" box="[232,347,1422,1446]" pageId="3" pageNumber="22" value="2009-11">Nov 2009</date>
.
</paragraph>
<paragraph id="8BCC36E4FF903840FF72FA69FEA6FA31" blockId="3.[151,1437,1385,2022]" pageId="3" pageNumber="22">
<emphasis id="B907EAF6FF903840FF72FA69FEE5FA15" bold="true" box="[199,317,1457,1482]" pageId="3" pageNumber="22">
<typeStatus id="54C88846FF903840FF72FA69FEE0FA15" box="[199,312,1457,1482]" pageId="3" pageNumber="22" type="paratype">Paratype</typeStatus>
:
</emphasis>
NMMB-P 22369 (270+ mm total length); Dong-gang fish market, Pingtung,
<collectingCountry id="F3647674FF903840FB04FA69FADEFA15" box="[1201,1286,1457,1482]" name="Taiwan" pageId="3" pageNumber="22">Taiwan</collectingCountry>
, collected by H. Ho,
<date id="FFCD1024FF903840FF5CFA0DFEA3FA31" box="[233,379,1493,1518]" pageId="3" pageNumber="22" value="2015-04-26">26 Apr 2015</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C369656FFF903840FF72FA21FCEEF985" pageId="3" pageNumber="22" type="diagnosis">
<paragraph id="8BCC36E4FF903840FF72FA21FCEEF985" blockId="3.[151,1437,1385,2022]" pageId="3" pageNumber="22">
<emphasis id="B907EAF6FF903840FF72FA21FE99F9CD" bold="true" box="[199,321,1529,1554]" pageId="3" pageNumber="22">Diagnosis.</emphasis>
Distinguished from all other members of the genus
<taxonomicName id="4C734D67FF903840FC23FA21FC20F9CE" box="[918,1016,1529,1553]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B907EAF6FF903840FC23FA21FC20F9CE" box="[918,1016,1529,1553]" italics="true" pageId="3" pageNumber="22">Chlopsis</emphasis>
</taxonomicName>
by the combination of the following characters: pigmentation bicolored, dorsal origin approximately one eye-diameter behind gill opening, and low vertebral number (120 and 118+ in only known specimens).
</paragraph>
</subSubSection>
<subSubSection id="C369656FFF903846FF72F9BDFB87FC67" lastPageId="5" lastPageNumber="24" pageId="3" pageNumber="22" type="description">
<paragraph id="8BCC36E4FF903840FF72F9BDFCF8F8EE" blockId="3.[151,1437,1385,2022]" pageId="3" pageNumber="22">
<emphasis id="B907EAF6FF903840FF72F9BDFE81F9A1" bold="true" box="[199,345,1637,1662]" pageId="3" pageNumber="22">Description.</emphasis>
Total vertebrae 120 (118+, incomplete, tail regenerated), predorsal vertebrae 12 (11), preanal vertebrae 35 (33), precaudal vertebrae 54 (52), dorsal rays 353 (incomplete), anal rays 316 (incomplete), dorsal rays anterior to anal-fin origin 67 (78). Proportions as percent of total length: predorsal length 14.5 (incomplete), preanal length 33.8 (incomplete), head length 12.3 (incomplete), depth at anus 3.3 (incomplete). Proportions as percent of head length: eye diameter 10.7 (9.1), interorbital width 16.1 (15.2), snout length 16.8 (20.6), tip of snout to rictus 32.1 (30.6), tip of lower jaw to rictus 28.6 (27.9).
</paragraph>
<paragraph id="8BCC36E4FF903840FF72F8E5FBBFF839" blockId="3.[151,1437,1385,2022]" pageId="3" pageNumber="22">
Body moderately elongate, slightly compressed, body depth nearly constant, gradually tapering over last 20% of total length to tip of tail (
<figureCitation id="13482A61FF903840FE64F8B9FDCFF8A6" box="[465,535,1889,1914]" captionStart="FIGURE 4" captionStartId="4.[151,244,1099,1121]" captionTargetBox="[151,1435,191,1057]" captionTargetId="figure@4.[151,1435,191,1078]" captionTargetPageId="4" captionText="FIGURE 4. Holotype of Chlopsis nanhaiensis, USNM 408189, 228 mm TL; Dong-gang fish market, Pingtung, Taiwan." httpUri="https://zenodo.org/record/244557/files/figure.png" pageId="3" pageNumber="22">Fig. 4</figureCitation>
). Dorsal-fin origin slightly more than one eye diameter posterior to gill opening (
<figureCitation id="13482A61FF903840FF2AF85DFF3CF841" box="[159,228,1925,1950]" captionStart="FIGURE 5" captionStartId="4.[151,244,1741,1763]" captionTargetBox="[239,1337,1468,1706]" captionTargetId="figure@4.[233,1354,1457,1720]" captionTargetPageId="4" captionText="FIGURE 5. Head of holotype of Chlopsis nanhaiensis, USNM 408189." httpUri="https://zenodo.org/record/244558/files/figure.png" pageId="3" pageNumber="22">Fig. 5</figureCitation>
). Head moderate in length, relatively deep. Snout relatively broad. Gape short, rictus at posterior margin of eye. Anterior nostril tubular, slightly behind tip of snout, directed anterolaterally. Posterior nostril a posteroventrally directed low tubular opening (covered by a flap) on lip in front of eye.
</paragraph>
<caption id="DF0C666CFF973847FF22FB93FAC4FBBE" box="[151,1308,1099,1121]" httpUri="https://zenodo.org/record/244557/files/figure.png" pageId="4" pageNumber="23" targetBox="[151,1435,191,1057]" targetPageId="4">
<paragraph id="8BCC36E4FF973847FF22FB93FAC4FBBE" blockId="4.[151,1308,1099,1121]" box="[151,1308,1099,1121]" pageId="4" pageNumber="23">
<emphasis id="B907EAF6FF973847FF22FB93FED4FBBE" bold="true" box="[151,268,1099,1121]" pageId="4" pageNumber="23">FIGURE 4.</emphasis>
Holotype of
<taxonomicName id="4C734D67FF973847FE3EFB94FD8FFBBE" box="[395,599,1100,1121]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="nanhaiensis">
<emphasis id="B907EAF6FF973847FE3EFB94FD8FFBBE" box="[395,599,1100,1121]" italics="true" pageId="4" pageNumber="23">Chlopsis nanhaiensis</emphasis>
</taxonomicName>
, USNM 408189, 228 mm TL; Dong-gang fish market, Pingtung, Taiwan.
</paragraph>
</caption>
<paragraph id="8BCC36E4FF973847FF72FB69FE73FAA1" blockId="4.[151,1436,1201,1406]" pageId="4" pageNumber="23">
Lateral line on body absent except for one pore in branchial region, dorsal and anterior to gill opening (
<figureCitation id="13482A61FF973847FAFFFB69FA56FB15" box="[1354,1422,1201,1226]" captionStart="FIGURE 5" captionStartId="4.[151,244,1741,1763]" captionTargetBox="[239,1337,1468,1706]" captionTargetId="figure@4.[233,1354,1457,1720]" captionTargetPageId="4" captionText="FIGURE 5. Head of holotype of Chlopsis nanhaiensis, USNM 408189." httpUri="https://zenodo.org/record/244558/files/figure.png" pageId="4" pageNumber="23">Fig. 5</figureCitation>
). Supraorbital pores three: first (ethmoidal) at anteroventral tip of snout, second anteromedial to base of anterior nostril, and last above and slightly behind anterior nostril. Infraorbital pores four: first just behind anterior nostril, second midway between anterior and posterior nostrils, third just behind and slightly below posterior nostril, and last below middle of eye. Preoperculomandibular pores five, first near tip of lower jaw and last just behind middle of eye, anterior to rictus.
</paragraph>
<caption id="DF0C666CFF973847FF22F915FC93F93D" box="[151,843,1741,1763]" httpUri="https://zenodo.org/record/244558/files/figure.png" pageId="4" pageNumber="23" targetBox="[239,1337,1468,1706]" targetPageId="4">
<paragraph id="8BCC36E4FF973847FF22F915FC93F93D" blockId="4.[151,843,1741,1763]" box="[151,843,1741,1763]" pageId="4" pageNumber="23">
<emphasis id="B907EAF6FF973847FF22F915FED4F93C" bold="true" box="[151,268,1741,1763]" pageId="4" pageNumber="23">FIGURE 5.</emphasis>
Head of holotype of
<taxonomicName id="4C734D67FF973847FE6DF915FD7CF93D" box="[472,676,1741,1762]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="nanhaiensis">
<emphasis id="B907EAF6FF973847FE6DF915FD7CF93D" box="[472,676,1741,1762]" italics="true" pageId="4" pageNumber="23">Chlopsis nanhaiensis</emphasis>
</taxonomicName>
, USNM 408189.
</paragraph>
</caption>
<paragraph id="8BCC36E4FF973846FF72F8EBFDCCFF27" blockId="4.[151,1437,1843,2012]" lastBlockId="5.[151,1436,151,248]" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23">
Maxillary teeth (
<figureCitation id="13482A61FF973847FE36F8EBFE1FF893" box="[387,455,1843,1868]" captionStart="FIGURE 6" captionStartId="5.[151,244,645,667]" captionTargetBox="[298,1286,309,613]" captionTargetId="figure@5.[288,1299,299,624]" captionTargetPageId="5" captionText="FIGURE 6. Upper (A) and lower (B) jaws of holotype of Chlopsis nanhaiensis, USNM 408189." httpUri="https://zenodo.org/record/244559/files/figure.png" pageId="4" pageNumber="23">Fig. 6</figureCitation>
) conical, slightly recurved, in 2 irregular rows, increasing in size from outer to inner, a total of 22 teeth in inner row. Mandibular teeth like those of maxilla, in 2 rows with 2122 teeth in the slightly enlarged, inner row; 89 of the anterior teeth enlarged compared to the rest of the mandibular teeth posteriorly. Intermaxillary teeth conical, slightly recurved, with approximately 18 teeth in patch; 9 small teeth around outside of patch and 9 enlarged teeth arranged in 3 rows of 3 teeth arranged longitudinally in center of intermaxillary tooth patch. Vomerine dentition shorter and stouter, slightly compressed, in two longitudinal series of 1214 teeth anteriorly and reduced to a single median row of 5 teeth posteriorly. Description of teeth based on
<typeStatus id="54C88846FF963846FAB3FF64FAB3FF0A" box="[1286,1387,188,213]" pageId="5" pageNumber="24" type="holotype">holotype</typeStatus>
, but dentition of
<typeStatus id="54C88846FF963846FE95FF38FE58FF27" box="[288,384,224,248]" pageId="5" pageNumber="24" type="paratype">paratype</typeStatus>
very similar.
</paragraph>
<caption id="DF0C666CFF963846FF22FD5DFBE1FD44" box="[151,1081,645,667]" httpUri="https://zenodo.org/record/244559/files/figure.png" pageId="5" pageNumber="24" targetBox="[298,1286,309,613]" targetPageId="5">
<paragraph id="8BCC36E4FF963846FF22FD5DFBE1FD44" blockId="5.[151,1081,645,667]" box="[151,1081,645,667]" pageId="5" pageNumber="24">
<emphasis id="B907EAF6FF963846FF22FD5DFED4FD44" bold="true" box="[151,268,645,667]" pageId="5" pageNumber="24">FIGURE 6.</emphasis>
Upper (A) and lower (B) jaws of holotype of
<taxonomicName id="4C734D67FF963846FD73FD5EFC4BFD44" box="[710,915,646,667]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="nanhaiensis">
<emphasis id="B907EAF6FF963846FD73FD5EFC4BFD44" box="[710,915,646,667]" italics="true" pageId="5" pageNumber="24">Chlopsis nanhaiensis</emphasis>
</taxonomicName>
, USNM 408189.
</paragraph>
</caption>
<paragraph id="8BCC36E4FF963846FF72FD33FB87FC67" blockId="5.[151,1436,747,1060]" pageId="5" pageNumber="24">
Color of body dark grayish brown above and distinctly lighter ventrally; ventral light area starts on snout, in front of unpigmented anterior nostril and continuing back above posterior nostril to anterior margin of eye; tip of lower jaw also pigmented (
<figureCitation id="13482A61FF963846FE78FCEBFDCAFC93" box="[461,530,819,844]" captionStart="FIGURE 5" captionStartId="4.[151,244,1741,1763]" captionTargetBox="[239,1337,1468,1706]" captionTargetId="figure@4.[233,1354,1457,1720]" captionTargetPageId="4" captionText="FIGURE 5. Head of holotype of Chlopsis nanhaiensis, USNM 408189." httpUri="https://zenodo.org/record/244558/files/figure.png" pageId="5" pageNumber="24">Fig. 5</figureCitation>
); posterior to eye, dorsal edge of ventral light area becomes more irregular, but extends dorsally to gill opening; ventral light area then tapers to base of anal fin slightly behind anus, but continues along base of anal fin for approximately 2/3 of body length; anal fin remains unpigmented to near tip of tail; ventral base of caudal fin and posterior portion of anal fin base much darker than rest of body.
</paragraph>
</subSubSection>
<subSubSection id="C369656FFF963846FF72FC1BFF19FBFB" pageId="5" pageNumber="24" type="etymology">
<paragraph id="8BCC36E4FF963846FF72FC1BFF19FBFB" blockId="5.[151,1436,747,1060]" pageId="5" pageNumber="24">
<emphasis id="B907EAF6FF963846FF72FC1BFE96FC03" bold="true" box="[199,334,963,988]" pageId="5" pageNumber="24">Etymology.</emphasis>
The name
<taxonomicName id="4C734D67FF963846FE61FC1CFD82FC03" box="[468,602,964,988]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="nanhaiensis">
<emphasis id="B907EAF6FF963846FE61FC1CFD82FC03" box="[468,602,964,988]" italics="true" pageId="5" pageNumber="24">nanhaiensis</emphasis>
</taxonomicName>
is from Nan Hai, the Chinese name for the South
<collectingCountry id="F3647674FF963846FB19FC1BFB37FC03" box="[1196,1263,963,988]" name="China" pageId="5" pageNumber="24">China</collectingCountry>
Sea where the
<typeStatus id="54C88846FF963846FF22FC3FFF21FBDF" box="[151,249,999,1024]" pageId="5" pageNumber="24" type="holotype">holotype</typeStatus>
apparently was captured by commercial fishermen, and the Latin adjective
<emphasis id="B907EAF6FF963846FBD6FC30FB43FC20" box="[1123,1179,1000,1023]" italics="true" pageId="5" pageNumber="24">ensis</emphasis>
, meaning “originating in.”
</paragraph>
</subSubSection>
<caption id="DF0C666CFF963846FF22F81EFAECF803" box="[151,1332,1990,2012]" httpUri="https://zenodo.org/record/244560/files/figure.png" pageId="5" pageNumber="24" targetBox="[151,1436,1111,1966]" targetPageId="5">
<paragraph id="8BCC36E4FF963846FF22F81EFAECF803" blockId="5.[151,1332,1990,2012]" box="[151,1332,1990,2012]" pageId="5" pageNumber="24">
<emphasis id="B907EAF6FF963846FF22F81EFED4F803" bold="true" box="[151,268,1990,2012]" pageId="5" pageNumber="24">FIGURE 7.</emphasis>
Paratype of
<taxonomicName id="4C734D67FF963846FE33F81FFD8AF803" box="[390,594,1991,2012]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="nanhaiensis">
<emphasis id="B907EAF6FF963846FE33F81FFD8AF803" box="[390,594,1991,2012]" italics="true" pageId="5" pageNumber="24">Chlopsis nanhaiensis</emphasis>
</taxonomicName>
, NMMB-P 22369, 270+ mm TL; Dong-gang fish market, Pingtung, Taiwan.
</paragraph>
</caption>
<subSubSection id="C369656FFF953845FF72FF4FFA5EFDBF" pageId="6" pageNumber="25" type="distribution">
<paragraph id="8BCC36E4FF953845FF72FF4FFC89FF0B" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="25">
<emphasis id="B907EAF6FF953845FF72FF4FFEB8FF6F" bold="true" box="[199,352,151,176]" pageId="6" pageNumber="25">Distribution.</emphasis>
Known from the
<typeStatus id="54C88846FF953845FD91FF4FFD5EFF6F" box="[548,646,151,176]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
and
<typeStatus id="54C88846FF953845FD0EFF40FCC3FF6F" box="[699,795,152,176]" pageId="6" pageNumber="25" type="paratype">paratype</typeStatus>
(
<figureCitation id="13482A61FF953845FC9FFF4FFCB4FF6F" box="[810,876,151,176]" captionStart="FIGURE 7" captionStartId="5.[151,244,1990,2012]" captionTargetBox="[151,1436,1111,1966]" captionTargetId="figure@5.[151,1436,1111,1967]" captionTargetPageId="5" captionText="FIGURE 7. Paratype of Chlopsis nanhaiensis, NMMB-P 22369, 270 + mm TL; Dong-gang fish market, Pingtung, Taiwan." httpUri="https://zenodo.org/record/244560/files/figure.png" pageId="6" pageNumber="25">Fig. 7</figureCitation>
) collected by fishermen off the southwest coast of
<collectingCountry id="F3647674FF953845FF22FF64FF35FF0A" box="[151,237,188,213]" name="Taiwan" pageId="6" pageNumber="25">Taiwan</collectingCountry>
; presumably more widespread in the South
<collectingCountry id="F3647674FF953845FD62FF64FCC2FF0A" box="[727,794,188,213]" name="China" pageId="6" pageNumber="25">China</collectingCountry>
Sea.
</paragraph>
<paragraph id="8BCC36E4FF953845FF72FF38FA5EFDBF" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="25">
<taxonomicName id="4C734D67FF953845FF72FF38FE6DFF27" box="[199,437,224,248]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="nanhaiensis">
<emphasis id="B907EAF6FF953845FF72FF38FE6DFF27" box="[199,437,224,248]" italics="true" pageId="6" pageNumber="25">Chlopsis nanhaiensis</emphasis>
</taxonomicName>
is similar in overall appearance to several other members of the genus
<taxonomicName id="4C734D67FF953845FB7FFF38FAF4FF27" box="[1226,1324,224,248]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B907EAF6FF953845FB7FFF38FAF4FF27" box="[1226,1324,224,248]" italics="true" pageId="6" pageNumber="25">Chlopsis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C734D67FF953845FA8FFF38FF33FEC3" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="apterus">
<emphasis id="B907EAF6FF953845FA8FFF38FF33FEC3" italics="true" pageId="6" pageNumber="25">Chlopsis apterus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C734D67FF953845FF4DFEDCFE5CFEC3" box="[248,388,260,284]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="bicollaris">
<emphasis id="B907EAF6FF953845FF4DFEDCFE5CFEC3" box="[248,388,260,284]" italics="true" pageId="6" pageNumber="25">C. bicollaris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D67FF953845FE0FFEDCFDF1FEC3" box="[442,553,260,284]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="kazuko">
<emphasis id="B907EAF6FF953845FE0FFEDCFDF1FEC3" box="[442,553,260,284]" italics="true" pageId="6" pageNumber="25">C. kazuko</emphasis>
</taxonomicName>
, all from the eastern Pacific Ocean, are all also bicolored, as is
<taxonomicName id="4C734D67FF953845FB58FEDCFA87FEC3" box="[1261,1375,260,284]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B907EAF6FF953845FB58FEDCFA87FEC3" box="[1261,1375,260,284]" italics="true" pageId="6" pageNumber="25">C. bicolor</emphasis>
</taxonomicName>
from the Atlantic Ocean,
<taxonomicName id="4C734D67FF953845FECEFEF0FDCAFE9F" box="[379,530,296,320]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="bidentatus">
<emphasis id="B907EAF6FF953845FECEFEF0FDCAFE9F" box="[379,530,296,320]" italics="true" pageId="6" pageNumber="25">C. bidentatus</emphasis>
</taxonomicName>
from the southwestern Pacific Ocean, and
<taxonomicName id="4C734D67FF953845FC4BFEF0FB54FE9F" box="[1022,1164,296,320]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B907EAF6FF953845FC4BFEF0FB54FE9F" box="[1022,1164,296,320]" italics="true" pageId="6" pageNumber="25">C. orientalis</emphasis>
</taxonomicName>
(described herein). The posterior origin of the dorsal fin (at least one eye diameter behind the gill opening) separates
<taxonomicName id="4C734D67FF953845FB03FE94FA86FEBB" box="[1206,1374,332,356]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="nanhaiensis">
<emphasis id="B907EAF6FF953845FB03FE94FA86FEBB" box="[1206,1374,332,356]" italics="true" pageId="6" pageNumber="25">C. nanhaiensis</emphasis>
</taxonomicName>
from
<taxonomicName id="4C734D67FF953845FF22FEA8FEFFFE57" box="[151,295,368,392]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="bicollaris">
<emphasis id="B907EAF6FF953845FF22FEA8FEFFFE57" box="[151,295,368,392]" italics="true" pageId="6" pageNumber="25">C. bicollaris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D67FF953845FED1FEA8FE0DFE57" box="[356,469,368,392]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="kazuko">
<emphasis id="B907EAF6FF953845FED1FEA8FE0DFE57" box="[356,469,368,392]" italics="true" pageId="6" pageNumber="25">C. kazuko</emphasis>
</taxonomicName>
.
<taxonomicName id="4C734D67FF953845FE52FEA8FD70FE57" box="[487,680,368,392]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="apterus">
<emphasis id="B907EAF6FF953845FE52FEA8FD70FE57" box="[487,680,368,392]" italics="true" pageId="6" pageNumber="25">Chlopsis apterus</emphasis>
</taxonomicName>
,
<emphasis id="B907EAF6FF953845FD0CFEA8FC0DFE57" box="[697,981,368,392]" italics="true" pageId="6" pageNumber="25">
<taxonomicName id="4C734D67FF953845FD0CFEA8FCF5FE57" box="[697,813,368,392]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="bicolor">C. bicolor</taxonomicName>
,
<taxonomicName id="4C734D67FF953845FC89FEA8FC0DFE57" box="[828,981,368,392]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="bidentatus">C. bidentatus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C734D67FF953845FBA6FEA8FB7AFE57" box="[1043,1186,368,392]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B907EAF6FF953845FBA6FEA8FB7AFE57" box="[1043,1186,368,392]" italics="true" pageId="6" pageNumber="25">C. orientalis</emphasis>
</taxonomicName>
, which also have the dorsal origin behind the gill opening, have more vertebrae (
<quantity id="4C8B9B01FF953845FCF4FE4CFC53FE72" box="[833,907,404,429]" metricMagnitude="0" metricUnit="m" metricValue="3.048" pageId="6" pageNumber="25" unit="in" value="120.0">120 in</quantity>
<taxonomicName id="4C734D67FF953845FC21FE4CFBE4FE73" box="[916,1084,404,428]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="nanhaiensis">
<emphasis id="B907EAF6FF953845FC21FE4CFBE4FE73" box="[916,1084,404,428]" italics="true" pageId="6" pageNumber="25">C. nanhaiensis</emphasis>
</taxonomicName>
versus
<quantity id="4C8B9B01FF953845FB20FE4CFACDFE72" box="[1173,1301,404,429]" metricMagnitude="0" metricUnit="m" metricValue="3.4798" metricValueMax="3.556" metricValueMin="3.4036" pageId="6" pageNumber="25" unit="in" value="137.0" valueMax="140.0" valueMin="134.0">134140 in</quantity>
<taxonomicName id="4C734D67FF953845FAABFE4CFA4DFE73" box="[1310,1429,404,428]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="apterus">
<emphasis id="B907EAF6FF953845FAABFE4CFA4DFE73" box="[1310,1429,404,428]" italics="true" pageId="6" pageNumber="25">C. apterus</emphasis>
</taxonomicName>
,
<quantity id="4C8B9B01FF953845FF22FE60FECEFE0F" box="[151,278,439,464]" metricMagnitude="0" metricUnit="m" metricValue="3.3147" metricValueMax="3.4036" metricValueMin="3.2257999999999996" pageId="6" pageNumber="25" unit="in" value="130.5" valueMax="134.0" valueMin="127.0">127134 in</quantity>
<taxonomicName id="4C734D67FF953845FEABFE60FE57FE0F" box="[286,399,440,464]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B907EAF6FF953845FEABFE60FE57FE0F" box="[286,399,440,464]" italics="true" pageId="6" pageNumber="25">C. bicolor</emphasis>
</taxonomicName>
,
<quantity id="4C8B9B01FF953845FE28FE6FFDC5FE0F" box="[413,541,439,464]" metricMagnitude="0" metricUnit="m" metricValue="3.2131" metricValueMax="3.2512" metricValueMin="3.175" pageId="6" pageNumber="25" unit="in" value="126.5" valueMax="128.0" valueMin="125.0">125128 in</quantity>
<taxonomicName id="4C734D67FF953845FD91FE60FD62FE0F" box="[548,698,440,464]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="bidentatus">
<emphasis id="B907EAF6FF953845FD91FE60FD62FE0F" box="[548,698,440,464]" italics="true" pageId="6" pageNumber="25">C. bidentatus</emphasis>
</taxonomicName>
and
<quantity id="4C8B9B01FF953845FD47FE6FFCE1FE0F" box="[754,825,439,464]" metricMagnitude="0" metricUnit="m" metricValue="3.429" pageId="6" pageNumber="25" unit="in" value="135.0">135 in</quantity>
<taxonomicName id="4C734D67FF953845FCF5FE60FC15FE0F" box="[832,973,440,464]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B907EAF6FF953845FCF5FE60FC15FE0F" box="[832,973,440,464]" italics="true" pageId="6" pageNumber="25">C. orientalis</emphasis>
</taxonomicName>
). All other species in the genus
<taxonomicName id="4C734D67FF953845FA8FFE60FA44FE0F" box="[1338,1436,440,464]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B907EAF6FF953845FA8FFE60FA44FE0F" box="[1338,1436,440,464]" italics="true" pageId="6" pageNumber="25">Chlopsis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C734D67FF953845FE75FE04FDBFFE2B" box="[448,615,476,500]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="nanhaiensis">
<emphasis id="B907EAF6FF953845FE75FE04FDBFFE2B" box="[448,615,476,500]" italics="true" pageId="6" pageNumber="25">C. nanhaiensis</emphasis>
</taxonomicName>
by coloration.
<taxonomicName id="4C734D67FF953845FCA5FE04FBC1FE2B" box="[784,1049,476,500]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="sagmacollaris">
<emphasis id="B907EAF6FF953845FCA5FE04FBC1FE2B" box="[784,1049,476,500]" italics="true" pageId="6" pageNumber="25">Chlopsis sagmacollaris</emphasis>
</taxonomicName>
also has
<taxonomicName id="4C734D67FF953845FB35FE04FB17FE2A" box="[1152,1231,476,501]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="bicolor">bicolor</taxonomicName>
pigmentation, but is much lighter in overall coloration and has the distinctive, darker pigment patch on the nape of the neck.
<taxonomicName id="4C734D67FF953845FA8FFDD8FF3BFDE3" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="olokun">
<emphasis id="B907EAF6FF953845FA8FFDD8FF3BFDE3" italics="true" pageId="6" pageNumber="25">Chlopsis olokun</emphasis>
</taxonomicName>
, from the eastern Atlantic, is a fairly uniform tan or gray color, while
<taxonomicName id="4C734D67FF953845FBA4FDFCFB4BFDE3" box="[1041,1171,548,572]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="B907EAF6FF953845FBA4FDFCFB4BFDE3" box="[1041,1171,548,572]" italics="true" pageId="6" pageNumber="25">C. dentatus</emphasis>
</taxonomicName>
(from the Atlantic and Indian Oceans) and
<taxonomicName id="4C734D67FF953845FEC2FD90FDF8FDBF" box="[375,544,584,608]" class="Actinopterygii" family="Chlopsidae" genus="Chlopsis" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="slusserorum">
<emphasis id="B907EAF6FF953845FEC2FD90FDF8FDBF" box="[375,544,584,608]" italics="true" pageId="6" pageNumber="25">C. slusserorum</emphasis>
</taxonomicName>
(from the southwestern Pacific) have banded, blotched or mottled coloration.
</paragraph>
</subSubSection>
</treatment>
</document>