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<document id="7580E6A313BE26ADB23BDE27137AEBA3" ID-DOI="10.5281/zenodo.5306831" ID-ISSN="0374-1036" ID-Zenodo-Dep="5306831" ID-ZooBank="D7B9D474-65C3-4358-8057-DD672465241B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630113600029" checkinUser="marcus" docAuthor="Japoshvili, George, Higashiura, Yoshimitsu &amp; Kamitani, Satoshi" docDate="2016" docId="C16787B1FFE9FFEEFE86F3F5FD142950" docLanguage="en" docName="ActaEntMusNatPra.2016.56.1.345-401.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 56 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Psyllaephagus higashiurai Japoshvili &amp; Higashiura &amp; Kamitani 2016, sp. nov." docType="treatment" docVersion="4" lastPageNumber="389" masterDocId="3D5EFFC9FFC2FFC2FFDEF753FFC52F68" masterDocTitle="A review of Japanese Encyrtidae (Hymenoptera), with descriptions of new species, new records and comments on the types described by Japanese authors" masterLastPageNumber="401" masterPageNumber="345" pageNumber="388" updateTime="1699156291164" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="E04B35F9DDA62166FCE0314D5594DE88" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A7918CAC39A1B4BAB98E67A38CC90575">A review of Japanese Encyrtidae (Hymenoptera), with descriptions of new species, new records and comments on the types described by Japanese authors</mods:title>
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<mods:namePart id="B47FFC7FACEBC3680D6ED33835FAD192">Japoshvili, George</mods:namePart>
<mods:affiliation id="E9FFCA6ADC10C6D44E2BB4C82A9D02F6">Department of Arthropods, Institute of Entomology, Agricultural University of Georgia, 13 km David Agmashenebeli Alley, Tbilisi, Georgia; e-mail: g. japoshvili @ agruni. edu. ge &amp; Invertebrate Research Center, Agladze # 26, Tbilisi, Georgia; e-mail: giorgij 70 @ yahoo. com</mods:affiliation>
</mods:name>
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<mods:namePart id="ED718CBD08EBCEF874408DA608E9968D">Higashiura, Yoshimitsu</mods:namePart>
<mods:affiliation id="F821630FC26EC367DB41D66F0DED1DE5">Citrus Promotion Center, Agricultural Technology Department, Yamaguchi Prefectural Agriculture &amp; Forestry General Technology Center, Higashi-agenosho, Suo-oshima, Yamaguchi, Japan; e-mail: higashiura. yoshimitsu @ pref. yamaguchi. lg. jp</mods:affiliation>
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<mods:namePart id="DCB924562234C4C59776B2F74094FB41">Kamitani, Satoshi</mods:namePart>
<mods:affiliation id="F1A7C728C23B0A6E268DF329E0B56F86">Department of Entomology, Faculty of Agriculture, Kyushu University, Fukuoka, Japan; e-mail: kamitani @ agr. kyushu-u. ac. jp</mods:affiliation>
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<mods:title id="9385DA8D8C427C90A5025D91540742CA">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="FDA7C55F8D04691F35C5EE81113898F1">2016</mods:date>
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<mods:number id="83E6ABFE10DB0F4D535E17F273356F8B">2016-07-15</mods:number>
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<treatment id="C16787B1FFE9FFEEFE86F3F5FD142950" LSID="urn:lsid:plazi:treatment:C16787B1FFE9FFEEFE86F3F5FD142950" httpUri="http://treatment.plazi.org/id/C16787B1FFE9FFEEFE86F3F5FD142950" lastPageId="44" lastPageNumber="389" pageId="43" pageNumber="388">
<subSubSection id="01D4652CFFE9FFE9FE86F3F5FC9F2BD5" box="[344,858,1190,1213]" pageId="43" pageNumber="388" type="nomenclature">
<paragraph id="497136A7FFE9FFE9FE86F3F5FC9F2BD5" blockId="43.[344,858,1190,1213]" box="[344,858,1190,1213]" pageId="43" pageNumber="388">
<heading id="123981CBFFE9FFE9FE86F3F5FC9F2BD5" bold="true" box="[344,858,1190,1213]" centered="true" fontSize="8" level="2" pageId="43" pageNumber="388" reason="3">
<emphasis id="7BBAEAB5FFE9FFE9FE86F3F5FC9F2BD5" bold="true" box="[344,858,1190,1213]" pageId="43" pageNumber="388">
<taxonomicName id="8ECE4D24FFE9FFE9FE86F3F5FD3C2BD5" authority="Japoshvili" authorityName="Japoshvili &amp; Higashiura &amp; Kamitani" authorityYear="2016" box="[344,761,1190,1213]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="388" phylum="Arthropoda" rank="species" species="higashiurai" status="sp. nov.">
<emphasis id="7BBAEAB5FFE9FFE9FE86F3F5FDB92BD5" bold="true" box="[344,636,1190,1213]" italics="true" pageId="43" pageNumber="388">Psyllaephagus higashiurai</emphasis>
Japoshvili
</taxonomicName>
,
<taxonomicNameLabel id="608957CEFFE9FFE9FCDAF3F5FC9F2BD5" box="[772,858,1190,1213]" pageId="43" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="01D4652CFFE9FFE9FDDCF397FD752BBF" box="[514,688,1220,1239]" pageId="43" pageNumber="388" type="description">
<paragraph id="497136A7FFE9FFE9FDDCF397FD752BBF" blockId="43.[113,1090,1220,1414]" box="[514,688,1220,1239]" pageId="43" pageNumber="388">
(
<figureCitation id="D1F52A22FFE9FFE9FDD6F397FDAF2BBF" box="[520,618,1220,1239]" captionStart="Figs 11–15. 11–12" captionStartId="13.[113,150,1059,1078]" captionTargetBox="[116,1085,220,1040]" captionTargetId="figure-155@13.[113,1088,218,1041]" captionTargetPageId="13" captionText="Figs 11–15. 11–12 – Psyllaephagus kamitanii Japoshvili sp. nov., female, head. 13–15 – Psyllaephagus higashiurai Japoshvili sp. nov., female: 13 – head; 14–15 – body." figureDoi="http://doi.org/10.5281/zenodo.5337446" httpUri="https://zenodo.org/record/5337446/files/figure.png" pageId="43" pageNumber="388">Figs 13–15</figureCitation>
,
<figureCitation id="D1F52A22FFE9FFE9FDADF397FD6F2BBF" box="[627,682,1220,1239]" captionStart="Figs" captionStartId="42.[151,190,860,879]" captionTargetBox="[152,1122,217,831]" captionTargetId="figure-10@42.[151,1123,217,831]" captionTargetPageId="42" captionText="Figs. 44–48. 44 – Psyllaephagus kamitanii Japoshvili sp. nov., female, ovipositor.45–48 – Psyllaephagus higashiurai Japoshvili sp. nov., female: 45 – antennae; 46 – midtarsus; 47 – ovipositor; 48 – fore wing." figureDoi="http://doi.org/10.5281/zenodo.5337454" httpUri="https://zenodo.org/record/5337454/files/figure.png" pageId="43" pageNumber="388">45–47</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="01D4652CFFE9FFE9FFAFF3BDFD702AEE" pageId="43" pageNumber="388" type="materials_examined">
<paragraph id="497136A7FFE9FFE9FFAFF3BDFD702AEE" blockId="43.[113,1090,1220,1414]" pageId="43" pageNumber="388">
<emphasis id="7BBAEAB5FFE9FFE9FFAFF3BDFEE62A69" bold="true" box="[113,291,1262,1281]" pageId="43" pageNumber="388">Material examined.</emphasis>
<materialsCitation id="F9A63CFAFFE9FFE9FEF9F3BDFBFB2A69" box="[295,1086,1256,1281]" collectingDate="1994-06-30" collectionCode="ELKU" collectorName="Y. Higashiura" location="Momiki Izumi" pageId="43" pageNumber="388" specimenCount="1" specimenCount-female="1" stateProvince="Kumamoto" typeStatus="holotype">
<typeStatus id="96758805FFE9FFE9FEF9F3BDFEBB2B97" box="[295,382,1262,1281]" pageId="43" pageNumber="388" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="5FC8FD2EFFE9FFE9FE57F3BBFE5C2A69" box="[393,409,1256,1281]" pageId="43" pageNumber="388" type="female">♀</specimenCount>
,
<location id="4C11607CFFE9FFE9FE7DF3BDFDE52A69" LSID="urn:lsid:plazi:treatment:C16787B1FFE9FFEEFE86F3F5FD142950:4C11607CFFE9FFE9FE7DF3BDFDE52A69" box="[419,544,1262,1281]" name="Momiki Izumi" pageId="43" pageNumber="388" stateProvince="Kumamoto">Momiki Izumi</location>
,
<collectingRegion id="8B0AF845FFE9FFE9FDF6F3BDFD7C2A69" box="[552,697,1262,1281]" country="Japan" name="Kumamoto" pageId="43" pageNumber="388">Kumamoto Pref.</collectingRegion>
, swept
<date id="3D701067FFE9FFE9FD28F3BDFC962A69" box="[758,851,1262,1281]" pageId="43" pageNumber="388" value="1994-06-30">
<collectingDate id="2D34E98FFFE9FFE9FD28F3BDFC962A69" box="[758,851,1262,1281]" pageId="43" pageNumber="388" value="1994-06-30">30.vi.1994</collectingDate>
</date>
,
<collectorName id="E43B5371FFE9FFE9FC85F3BDFC0A2A69" box="[859,975,1262,1281]" pageId="43" pageNumber="388">Y. Higashiura</collectorName>
lgt. (
<collectionCode id="2FDFAE62FFE9FFE9FC24F3BDFBFD2A69" box="[1018,1080,1262,1281]" pageId="43" pageNumber="388">ELKU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F9A63CFAFFE9FFE9FFAFF25AFD912A74" box="[113,596,1282,1308]" collectingDate="1994-06-30" collectionCode="ELKU" collectorName="Y. Higashiura" location="Momiki Izumi" pageId="43" pageNumber="388" specimenCount="1" specimenCount-female="1" stateProvince="Kumamoto" typeStatus="paratype">
<typeStatus id="96758805FFE9FFE9FFAFF25AFF0E2A72" box="[113,203,1289,1308]" pageId="43" pageNumber="388" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="5FC8FD2EFFE9FFE9FF0BF25AFF372A73" box="[213,242,1282,1308]" pageId="43" pageNumber="388" type="female">1 ♀</specimenCount>
, on slide same data as previous specimens
</materialsCitation>
;
<materialsCitation id="F9A63CFAFFE9FFE9FD84F25AFECC2A5E" collectingDate="1994-06-30" collectorName="Y. Higashiura" location="Momiki Izumi" pageId="43" pageNumber="388" specimenCount="7" specimenCount-female="7" stateProvince="Kumamoto" typeStatus="paratype">
<specimenCount id="5FC8FD2EFFE9FFE9FD84F25AFD432A73" box="[602,646,1282,1308]" pageId="43" pageNumber="388" type="female">7♀♀</specimenCount>
,
<location id="4C11607CFFE9FFE9FD4FF25AFCC92A74" LSID="urn:lsid:plazi:treatment:C16787B1FFE9FFEEFE86F3F5FD142950:4C11607CFFE9FFE9FD4FF25AFCC92A74" box="[657,780,1289,1308]" name="Momiki Izumi" pageId="43" pageNumber="388" stateProvince="Kumamoto">Momiki Izumi</location>
,
<collectingRegion id="8B0AF845FFE9FFE9FCCDF25AFC672A74" box="[787,930,1289,1308]" country="Japan" name="Kumamoto" pageId="43" pageNumber="388">Kumamoto Pref.</collectingRegion>
, swept,
<date id="3D701067FFE9FFE9FC3FF25AFBF82A74" box="[993,1085,1289,1308]" pageId="43" pageNumber="388" value="1994-06-30">
<collectingDate id="2D34E98FFFE9FFE9FC3FF25AFBF82A74" box="[993,1085,1289,1308]" pageId="43" pageNumber="388" value="1994-06-30">30.vi.1994</collectingDate>
</date>
,
<collectorName id="E43B5371FFE9FFE9FFAFF270FF2D2A5E" box="[113,232,1315,1334]" pageId="43" pageNumber="388">Y. Higashiura</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="F9A63CFAFFE9FFE9FEC8F270FD6A2A5E" box="[278,687,1309,1334]" collectingDate="1994-06-30" collectorName="Y. Higashiura" location="Momiki Izumi" pageId="43" pageNumber="388" specimenCount="1" specimenCount-female="1" stateProvince="Kumamoto" typeStatus="paratype">
<specimenCount id="5FC8FD2EFFE9FFE9FEC8F270FEF22A5E" box="[278,311,1309,1334]" pageId="43" pageNumber="388" type="female">1 ♀</specimenCount>
, on slide same data as previous specimens
</materialsCitation>
;
<materialsCitation id="F9A63CFAFFE9FFE9FD66F270FEB72A39" collectingDate="1994-08-07" collectorName="Y. Higashiura" location="Momiki Izumi" pageId="43" pageNumber="388" specimenCount="1" specimenCount-female="1" stateProvince="Kumamoto" typeStatus="paratype">
<specimenCount id="5FC8FD2EFFE9FFE9FD66F270FD1C2A5E" box="[696,729,1309,1334]" pageId="43" pageNumber="388" type="female">1 ♀</specimenCount>
,
<location id="4C11607CFFE9FFE9FD3BF270FCA12A5E" LSID="urn:lsid:plazi:treatment:C16787B1FFE9FFEEFE86F3F5FD142950:4C11607CFFE9FFE9FD3BF270FCA12A5E" box="[741,868,1315,1334]" name="Momiki Izumi" pageId="43" pageNumber="388" stateProvince="Kumamoto">Momiki Izumi</location>
,
<collectingRegion id="8B0AF845FFE9FFE9FCB0F270FBC42A5E" box="[878,1025,1315,1334]" country="Japan" name="Kumamoto" pageId="43" pageNumber="388">Kumamoto Pref.</collectingRegion>
, swept,
<date id="3D701067FFE9FFE9FFAFF26DFF0A2A39" box="[113,207,1342,1361]" pageId="43" pageNumber="388" value="1994-08-07">
<collectingDate id="2D34E98FFFE9FFE9FFAFF26DFF0A2A39" box="[113,207,1342,1361]" pageId="43" pageNumber="388" value="1994-08-07">7.viii.1994</collectingDate>
</date>
,
<collectorName id="E43B5371FFE9FFE9FF07F26DFE952A39" box="[217,336,1342,1361]" pageId="43" pageNumber="388">Y. Higashiura</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="F9A63CFAFFE9FFE9FEA1F26DFBFF2A39" box="[383,1082,1336,1361]" collectingDate="1970-08-22" collectingDateMax="1970-08-23" collectingDateMin="1970-08-22" collectorName="K. Yamagishi" location="Pref" municipality="Kashiwazaki-shi" pageId="43" pageNumber="388" specimenCount="2" specimenCount-female="2" stateProvince="Niigata" typeStatus="paratype">
<specimenCount id="5FC8FD2EFFE9FFE9FEA1F26DFE752A39" box="[383,432,1336,1361]" pageId="43" pageNumber="388" type="female">2 ♀♀</specimenCount>
,
<collectingMunicipality id="A915ACDDFFE9FFE9FE62F26DFD892A39" box="[444,588,1342,1361]" pageId="43" pageNumber="388">Kashiwazaki-shi</collectingMunicipality>
,
<collectingRegion id="8B0AF845FFE9FFE9FD89F26DFD532A39" box="[599,662,1342,1361]" country="Japan" name="Niigata" pageId="43" pageNumber="388">Niigata</collectingRegion>
<location id="4C11607CFFE9FFE9FD42F26DFD072A39" LSID="urn:lsid:plazi:treatment:C16787B1FFE9FFEEFE86F3F5FD142950:4C11607CFFE9FFE9FD42F26DFD072A39" box="[668,706,1342,1361]" municipality="Kashiwazaki-shi" name="Pref" pageId="43" pageNumber="388" stateProvince="Niigata">Pref</location>
, swept,
<date id="3D701067FFE9FFE9FCD4F26DFC5C2A39" box="[778,921,1342,1361]" pageId="43" pageNumber="388" value="1970-08-22" valueMax="1970-08-23" valueMin="1970-08-22">
<collectingDate id="2D34E98FFFE9FFE9FCD4F26DFC5C2A39" box="[778,921,1342,1361]" pageId="43" pageNumber="388" value="1970-08-22" valueMax="1970-08-23" valueMin="1970-08-22">22.–23.viii.1970</collectingDate>
</date>
,
<collectorName id="E43B5371FFE9FFE9FC7DF26DFBDD2A39" box="[931,1048,1342,1361]" pageId="43" pageNumber="388">K. Yamagishi</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="F9A63CFAFFE9FFE9FFAFF20AFD3E2A04" box="[113,763,1362,1388]" collectingDate="1994-08-07" collectorName="Y. Higashiura" location="Momiki Izumi" pageId="43" pageNumber="388" specimenCount="1" specimenCount-female="1" stateProvince="Kumamoto" typeStatus="paratype">
<specimenCount id="5FC8FD2EFFE9FFE9FFAFF20AFF572A03" box="[113,146,1362,1388]" pageId="43" pageNumber="388" type="female">1 ♀</specimenCount>
,
<location id="4C11607CFFE9FFE9FF43F20AFED92A04" LSID="urn:lsid:plazi:treatment:C16787B1FFE9FFEEFE86F3F5FD142950:4C11607CFFE9FFE9FF43F20AFED92A04" box="[157,284,1369,1388]" name="Momiki Izumi" pageId="43" pageNumber="388" stateProvince="Kumamoto">Momiki Izumi</location>
,
<collectingRegion id="8B0AF845FFE9FFE9FEFBF20AFE7D2A04" box="[293,440,1369,1388]" country="Japan" name="Kumamoto" pageId="43" pageNumber="388">Kumamoto Pref.</collectingRegion>
, swept,
<date id="3D701067FFE9FFE9FE23F20AFD9E2A04" box="[509,603,1369,1388]" pageId="43" pageNumber="388" value="1994-08-07">
<collectingDate id="2D34E98FFFE9FFE9FE23F20AFD9E2A04" box="[509,603,1369,1388]" pageId="43" pageNumber="388" value="1994-08-07">7.viii.1994</collectingDate>
</date>
,
<collectorName id="E43B5371FFE9FFE9FDBAF20AFD1F2A04" box="[612,730,1369,1388]" pageId="43" pageNumber="388">Y. Higashiura</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="F9A63CFAFFE9FFE9FCD6F20AFD742AEE" collectingDate="1970-08-22" collectingDateMax="1970-08-23" collectingDateMin="1970-08-22" collectionCode="ELKU, EIAUG" collectorName="K. Yamagishi" location="Kashiwazaki-shi" pageId="43" pageNumber="388" specimenCount="2" specimenCount-female="2" stateProvince="Niigata" typeStatus="paratype">
<specimenCount id="5FC8FD2EFFE9FFE9FCD6F20AFCEC2A03" box="[776,809,1362,1388]" pageId="43" pageNumber="388" type="female">1 ♀</specimenCount>
,
<location id="4C11607CFFE9FFE9FCEAF20AFC012A04" LSID="urn:lsid:plazi:treatment:C16787B1FFE9FFEEFE86F3F5FD142950:4C11607CFFE9FFE9FCEAF20AFC012A04" box="[820,964,1369,1388]" name="Kashiwazaki-shi" pageId="43" pageNumber="388" stateProvince="Niigata">Kashiwazaki-shi</location>
,
<collectingRegion id="8B0AF845FFE9FFE9FC13F20AFBFA2A04" box="[973,1087,1369,1388]" country="Japan" name="Niigata" pageId="43" pageNumber="388">Niigata Pref.</collectingRegion>
, swept,
<date id="3D701067FFE9FFE9FF70F220FEF82AEE" box="[174,317,1395,1414]" pageId="43" pageNumber="388" value="1970-08-22" valueMax="1970-08-23" valueMin="1970-08-22">
<collectingDate id="2D34E98FFFE9FFE9FF70F220FEF82AEE" box="[174,317,1395,1414]" pageId="43" pageNumber="388" value="1970-08-22" valueMax="1970-08-23" valueMin="1970-08-22">22.–23.viii.1970</collectingDate>
</date>
,
<collectorName id="E43B5371FFE9FFE9FE9BF220FE7D2AEE" box="[325,440,1395,1414]" pageId="43" pageNumber="388">K. Yamagishi</collectorName>
lgt. (
<collectionCode id="2FDFAE62FFE9FFE9FE3BF220FDE42AEE" box="[485,545,1395,1414]" pageId="43" pageNumber="388">ELKU</collectionCode>
,
<specimenCount id="5FC8FD2EFFE9FFE9FDF4F220FD8F2AEE" box="[554,586,1389,1414]" pageId="43" pageNumber="388" type="female">1 ♀</specimenCount>
in
<collectionCode id="2FDFAE62FFE9FFE9FDBBF220FD6E2AEE" box="[613,683,1395,1414]" pageId="43" pageNumber="388">EIAUG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="01D4652CFFE9FFEEFFAFF2CCFEA92DF0" lastPageId="44" lastPageNumber="389" pageId="43" pageNumber="388" type="diagnosis">
<paragraph id="497136A7FFE9FFE9FFAFF2CCFC28295E" blockId="43.[113,1089,1439,1590]" pageId="43" pageNumber="388">
<emphasis id="7BBAEAB5FFE9FFE9FFAFF2CCFEFA2ADE" bold="true" box="[113,319,1439,1462]" pageId="43" pageNumber="388">
Diagnosis.
<emphasis id="7BBAEAB5FFE9FFE9FF32F2CCFEFA2ADE" bold="true" box="[236,319,1439,1462]" italics="true" pageId="43" pageNumber="388">Female</emphasis>
</emphasis>
(length about 1.0–
<quantity id="8E369B42FFE9FFE9FDD4F2CCFD992ADE" box="[522,604,1439,1462]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="43" pageNumber="388" unit="mm" value="1.3">1.3 mm</quantity>
) (
<figureCitation id="D1F52A22FFE9FFE9FDAFF2CCFD2B2ADE" box="[625,750,1439,1462]" captionStart="Figs 11–15. 11–12" captionStartId="13.[113,150,1059,1078]" captionTargetBox="[116,1085,220,1040]" captionTargetId="figure-155@13.[113,1088,218,1041]" captionTargetPageId="13" captionText="Figs 11–15. 11–12 – Psyllaephagus kamitanii Japoshvili sp. nov., female, head. 13–15 – Psyllaephagus higashiurai Japoshvili sp. nov., female: 13 – head; 14–15 – body." figureDoi="http://doi.org/10.5281/zenodo.5337446" httpUri="https://zenodo.org/record/5337446/files/figure.png" pageId="43" pageNumber="388">Figs 14–15</figureCitation>
). Head (
<figureCitation id="D1F52A22FFE9FFE9FC99F2CCFC512ADE" box="[839,916,1439,1462]" captionStart="Figs 11–15. 11–12" captionStartId="13.[113,150,1059,1078]" captionTargetBox="[116,1085,220,1040]" captionTargetId="figure-155@13.[113,1088,218,1041]" captionTargetPageId="13" captionText="Figs 11–15. 11–12 – Psyllaephagus kamitanii Japoshvili sp. nov., female, head. 13–15 – Psyllaephagus higashiurai Japoshvili sp. nov., female: 13 – head; 14–15 – body." figureDoi="http://doi.org/10.5281/zenodo.5337446" httpUri="https://zenodo.org/record/5337446/files/figure.png" pageId="43" pageNumber="388">Fig. 13</figureCitation>
) and mesosoma brown with golden green to coppery metallic reflections; antennae generally dirty yellow; not all funicle segments longer than wide; pedicel 1.8× as long as wide and 2.4× as long as F 1; legs generally yellow, only mid and hind coxae, last tarsal segment, hind femur and fourth hind tarsal segment brown; wings hyaline; scape about 3.9× as long as wide; F
</paragraph>
<paragraph id="497136A7FFE9FFE9FC33F17CFC302955" box="[1005,1013,1583,1597]" pageId="43" pageNumber="388">1</paragraph>
<paragraph id="497136A7FFE9FFE9FC27F14CFB85295E" blockId="43.[113,1089,1439,1590]" box="[1017,1088,1567,1590]" pageId="43" pageNumber="388">almost</paragraph>
<caption id="1DB1662FFFEEFFEEFF49F78FFB872F87" box="[151,1090,220,239]" pageId="44" pageNumber="389" startId="44.[151,198,220,239]">
<paragraph id="497136A7FFEEFFEEFF49F78FFB872F87" blockId="44.[151,1090,220,363]" box="[151,1090,220,239]" pageId="44" pageNumber="389">
Table 8. Characters used to separate
<taxonomicName id="8ECE4D24FFEEFFEEFE11F78FFD6B2F87" authority="Japoshvili" authorityName="Japoshvili &amp; Higashiura &amp; Kamitani" authorityYear="2016" box="[463,686,220,239]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="389" phylum="Arthropoda" rank="species" species="higashiurai" status="sp. nov.">
<emphasis id="7BBAEAB5FFEEFFEEFE11F78FFD942F87" box="[463,593,220,239]" italics="true" pageId="44" pageNumber="389">Ps. higashiurai</emphasis>
Japoshvili
</taxonomicName>
<taxonomicNameLabel id="608957CEFFEEFFEEFD6DF78FFD332F87" box="[691,758,220,239]" pageId="44" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="8ECE4D24FFEEFFEEFCC0F78FFBF82F87" authority="Trjapitzin, 1969" authorityName="Trjapitzin" authorityYear="1969" box="[798,1085,220,239]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="389" phylum="Arthropoda" rank="species" species="colposceniae">
<emphasis id="7BBAEAB5FFEEFFEEFCC0F78FFC682F87" box="[798,941,220,239]" italics="true" pageId="44" pageNumber="389">Ps. colposceniae</emphasis>
Trjapitzin, 1969
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="497136A7FFEEFFEEFF41F7ADFCD22E79" blockId="44.[151,1090,220,363]" box="[159,791,254,273]" pageId="44" pageNumber="389">
<emphasis id="7BBAEAB5FFEEFFEEFF41F7ADFCD22E79" bold="true" box="[159,791,254,273]" pageId="44" pageNumber="389">
<taxonomicName id="8ECE4D24FFEEFFEEFF41F7ADFEE02E79" authority="Japoshvili &amp; Higashiura &amp; Kamitani, 2016" authorityName="Japoshvili &amp; Higashiura &amp; Kamitani" authorityYear="2016" box="[159,293,254,273]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="389" phylum="Arthropoda" rank="species" species="higashiurai" status="sp. nov.">
<emphasis id="7BBAEAB5FFEEFFEEFF41F7ADFEE02E79" bold="true" box="[159,293,254,273]" italics="true" pageId="44" pageNumber="389">Ps. higashiurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="608957CEFFEEFFEEFEF4F7ADFEAB2E79" box="[298,366,254,273]" pageId="44" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="8ECE4D24FFEEFFEEFD58F7ADFCD22E79" authorityName="Trjapitzin" authorityYear="1969" box="[646,791,254,273]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="389" phylum="Arthropoda" rank="species" species="colposceniae">
<emphasis id="7BBAEAB5FFEEFFEEFD58F7ADFCD22E79" bold="true" box="[646,791,254,273]" italics="true" pageId="44" pageNumber="389">Ps. colposceniae</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="497136A7FFEEFFEEFF41F64FFC212E47" blockId="44.[151,1090,220,363]" box="[159,996,284,303]" pageId="44" pageNumber="389">FV almost 0.5× as wide as HW. FV 0.33× or slightly less as wide as HW.</paragraph>
<paragraph id="497136A7FFEEFFEEFF41F669FC502E25" blockId="44.[151,1090,220,363]" box="[159,917,314,333]" pageId="44" pageNumber="389">Ocelli in slightly obtuse triangle. Ocelli in slightly acute triangle.</paragraph>
<paragraph id="497136A7FFEEFFEEFF41F60BFBE62E03" blockId="44.[151,1090,220,363]" box="[159,1059,344,363]" pageId="44" pageNumber="389">Clava almost as long as flagellar segments combined. Clava shorter than flagellar segments combined.</paragraph>
<paragraph id="497136A7FFEEFFEEFF49F552FD502D77" blockId="44.[151,1127,513,1592]" pageId="44" pageNumber="389">quadrate 1× as long as wide; F –F transverse, F 1.1× as long as wide; all funicular segments 2 5 6</paragraph>
<paragraph id="497136A7FFEEFFEEFF49F572FC442D10" blockId="44.[151,1127,513,1592]" pageId="44" pageNumber="389">
with two lines of longitudinal sensilla (
<figureCitation id="D1F52A22FFEEFFEEFDE3F572FD4B2D50" box="[573,654,545,568]" captionStart="Figs" captionStartId="42.[151,190,860,879]" captionTargetBox="[152,1122,217,831]" captionTargetId="figure-10@42.[151,1123,217,831]" captionTargetPageId="42" captionText="Figs. 44–48. 44 – Psyllaephagus kamitanii Japoshvili sp. nov., female, ovipositor.45–48 – Psyllaephagus higashiurai Japoshvili sp. nov., female: 45 – antennae; 46 – midtarsus; 47 – ovipositor; 48 – fore wing." figureDoi="http://doi.org/10.5281/zenodo.5337454" httpUri="https://zenodo.org/record/5337454/files/figure.png" pageId="44" pageNumber="389">Fig. 45</figureCitation>
); frontovertex about 0.33× as wide as head; fore wing (
<figureCitation id="D1F52A22FFEEFFEEFED2F512FE9C2D30" box="[268,345,577,600]" captionStart="Figs" captionStartId="42.[151,190,860,879]" captionTargetBox="[152,1122,217,831]" captionTargetId="figure-10@42.[151,1123,217,831]" captionTargetPageId="42" captionText="Figs. 44–48. 44 – Psyllaephagus kamitanii Japoshvili sp. nov., female, ovipositor.45–48 – Psyllaephagus higashiurai Japoshvili sp. nov., female: 45 – antennae; 46 – midtarsus; 47 – ovipositor; 48 – fore wing." figureDoi="http://doi.org/10.5281/zenodo.5337454" httpUri="https://zenodo.org/record/5337454/files/figure.png" pageId="44" pageNumber="389">Fig. 48</figureCitation>
) about 2.7× as long as wide; middle tibial spur as in
<figureCitation id="D1F52A22FFEEFFEEFC58F512FC112D30" box="[902,980,577,600]" captionStart="Figs" captionStartId="42.[151,190,860,879]" captionTargetBox="[152,1122,217,831]" captionTargetId="figure-10@42.[151,1123,217,831]" captionTargetPageId="42" captionText="Figs. 44–48. 44 – Psyllaephagus kamitanii Japoshvili sp. nov., female, ovipositor.45–48 – Psyllaephagus higashiurai Japoshvili sp. nov., female: 45 – antennae; 46 – midtarsus; 47 – ovipositor; 48 – fore wing." figureDoi="http://doi.org/10.5281/zenodo.5337454" httpUri="https://zenodo.org/record/5337454/files/figure.png" pageId="44" pageNumber="389">Fig. 46</figureCitation>
; exserted part of ovipositor (
<figureCitation id="D1F52A22FFEEFFEEFEF1F532FEB82D10" box="[303,381,609,632]" captionStart="Figs" captionStartId="42.[151,190,860,879]" captionTargetBox="[152,1122,217,831]" captionTargetId="figure-10@42.[151,1123,217,831]" captionTargetPageId="42" captionText="Figs. 44–48. 44 – Psyllaephagus kamitanii Japoshvili sp. nov., female, ovipositor.45–48 – Psyllaephagus higashiurai Japoshvili sp. nov., female: 45 – antennae; 46 – midtarsus; 47 – ovipositor; 48 – fore wing." figureDoi="http://doi.org/10.5281/zenodo.5337454" httpUri="https://zenodo.org/record/5337454/files/figure.png" pageId="44" pageNumber="389">Fig. 47</figureCitation>
) 0.16× as long as gaster including exserted part.
</paragraph>
<paragraph id="497136A7FFEEFFEEFF6BF5D2FEA92DF0" blockId="44.[151,1127,513,1592]" box="[181,364,641,664]" pageId="44" pageNumber="389">
<emphasis id="7BBAEAB5FFEEFFEEFF6BF5D2FF302DF0" bold="true" box="[181,245,641,664]" pageId="44" pageNumber="389">
<emphasis id="7BBAEAB5FFEEFFEEFF6BF5D2FF282DF0" bold="true" box="[181,237,641,664]" italics="true" pageId="44" pageNumber="389">Male</emphasis>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="01D4652CFFEEFFEEFF49F5F2FE7A2AF0" pageId="44" pageNumber="389" type="description">
<paragraph id="497136A7FFEEFFEEFF49F5F2FC192DD0" blockId="44.[151,1127,513,1592]" box="[151,988,673,696]" pageId="44" pageNumber="389">
<emphasis id="7BBAEAB5FFEEFFEEFF49F5F2FE472DD0" bold="true" box="[151,386,673,696]" pageId="44" pageNumber="389">
Description.
<emphasis id="7BBAEAB5FFEEFFEEFEF6F5F2FE472DD0" bold="true" box="[296,386,673,696]" italics="true" pageId="44" pageNumber="389">Female.</emphasis>
</emphasis>
<typeStatus id="96758805FFEEFFEEFE57F5F2FE292DD0" box="[393,492,673,696]" pageId="44" pageNumber="389" type="holotype">Holotype</typeStatus>
<quantity id="8E369B42FFEEFFEEFE2DF5F2FD822DD0" box="[499,583,673,696]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="44" pageNumber="389" unit="mm" value="1.2">1.2 mm</quantity>
, including exserted part of ovipositor.
</paragraph>
<paragraph id="497136A7FFEEFFEEFF6BF592FB902C90" blockId="44.[151,1127,513,1592]" pageId="44" pageNumber="389">Head metallic green with golden reflections and translucent white setae; front above torulli and in front of anterior ocellus with brown band without metallic reflection; genae with goldengreenish reflection. Malar sulcus prominent. Antenna with scape almost yellowish, funicular segments yellow; clava with C 2 –C 3 dirty yellow. Palpi light yellow. Pronotum with at least some kind of coppery with golden green reflections; mesoscutum and axillae golden-green with very reduced coppery reflections; scutellum with golden-green reflections; mesopleuron with coppery golden reflections, with coppery reflection more intensive; tegula brown, with golden metallic reflections; metanotum and propodeum with copperygolden reflections; gaster dorsal side with first tergite coppery golden, remaining tergites with coppery silver reflections; ventral surface with golden-green to coppery reflections; visible parts of gonostyli brown.</paragraph>
<paragraph id="497136A7FFEEFFEEFF6BF352FE7A2AF0" blockId="44.[151,1127,513,1592]" pageId="44" pageNumber="389">
Head in frontal view 1.3× as wide as high; almost 3× as wide as FV; head on frontovertex with fine, raised, regular, coriaceous-granular sculpture, cells smaller than diameter of eye facet; face and temple with similar, but more elongate and very small reticulate-imbricate sculpture; gena with similar sculpture; ocellar angle about 100°; occipital margin sharp, carinate; eye reaching occipital margin; mesoscutum and scutellum with similar, small, regular polygonal coriaceous structure, however mesoscutum with sculpture deeper, which makes it different from that of scutellum; distance between toruli almost 1.56× as long as distance between toruli and clypeal margin and 1.08× as long as distance from torulus to eye; sensilla visible only on F 6; mandible with one tooth and truncation; head and mesosoma almost 1.7× as long as gaster; middle tibial spur as long as basitarsus (
<figureCitation id="D1F52A22FFEEFFEEFCD4F272FC9F2A50" box="[778,858,1313,1336]" captionStart="Figs" captionStartId="42.[151,190,860,879]" captionTargetBox="[152,1122,217,831]" captionTargetId="figure-10@42.[151,1123,217,831]" captionTargetPageId="42" captionText="Figs. 44–48. 44 – Psyllaephagus kamitanii Japoshvili sp. nov., female, ovipositor.45–48 – Psyllaephagus higashiurai Japoshvili sp. nov., female: 45 – antennae; 46 – midtarsus; 47 – ovipositor; 48 – fore wing." figureDoi="http://doi.org/10.5281/zenodo.5337454" httpUri="https://zenodo.org/record/5337454/files/figure.png" pageId="44" pageNumber="389">Fig. 46</figureCitation>
); hypopygium extending to almost basal half; ovipositor exserted; ovipositor as in
<figureCitation id="D1F52A22FFEEFFEEFCD7F212FC9F2A30" box="[777,858,1345,1368]" captionStart="Figs" captionStartId="42.[151,190,860,879]" captionTargetBox="[152,1122,217,831]" captionTargetId="figure-10@42.[151,1123,217,831]" captionTargetPageId="42" captionText="Figs. 44–48. 44 – Psyllaephagus kamitanii Japoshvili sp. nov., female, ovipositor.45–48 – Psyllaephagus higashiurai Japoshvili sp. nov., female: 45 – antennae; 46 – midtarsus; 47 – ovipositor; 48 – fore wing." figureDoi="http://doi.org/10.5281/zenodo.5337454" httpUri="https://zenodo.org/record/5337454/files/figure.png" pageId="44" pageNumber="389">Fig. 48</figureCitation>
. Relative measurements: HW 44, FV 20, FVL 22, OD 3, OOL 2, OCL 4, AOL 7, POL 10, SL 20, SW 5, MS 14, EL 13, EW 19, ScL 24, WS 12.
</paragraph>
</subSubSection>
<subSubSection id="01D4652CFFEEFFEEFF49F2F2FBA32AB0" pageId="44" pageNumber="389" type="diagnosis">
<paragraph id="497136A7FFEEFFEEFF49F2F2FBA32AB0" blockId="44.[151,1127,513,1592]" pageId="44" pageNumber="389">
<emphasis id="7BBAEAB5FFEEFFEEFF49F2F2FE4A2AD0" bold="true" box="[151,399,1441,1464]" pageId="44" pageNumber="389">Differential diagnosis.</emphasis>
The overall morphology of this new species is very close to
<taxonomicName id="8ECE4D24FFEEFFEEFC21F2F2FF1E2AB0" baseAuthorityName="Tachikawa" baseAuthorityYear="1955" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="389" phylum="Arthropoda" rank="species" species="syntomozae">
<emphasis id="7BBAEAB5FFEEFFEEFC21F2F2FF1E2AB0" italics="true" pageId="44" pageNumber="389">Ps. syntomozae</emphasis>
</taxonomicName>
and
<taxonomicName id="8ECE4D24FFEEFFEEFED3F292FDB52AB0" authority="Trjapitzin, 1969" authorityName="Trjapitzin" authorityYear="1969" box="[269,624,1473,1496]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="389" phylum="Arthropoda" rank="species" species="colposceniae">
<emphasis id="7BBAEAB5FFEEFFEEFED3F292FE7B2AB0" box="[269,446,1473,1496]" italics="true" pageId="44" pageNumber="389">Ps. colposceniae</emphasis>
Trjapitzin, 1969
</taxonomicName>
, but differs in the character given in the
<tableCitation id="044C031CFFEEFFEEFBCCF292FBA62AB0" box="[1042,1123,1473,1496]" captionStart="Table 8" captionStartId="44.[151,198,220,239]" captionText="Table 8. Characters used to separate Ps. higashiurai Japoshvili sp. nov. and Ps. colposceniae Trjapitzin, 1969." pageId="44" pageNumber="389">Table 8</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="01D4652CFFEEFFEEFF49F2B2FE562970" pageId="44" pageNumber="389" type="etymology">
<paragraph id="497136A7FFEEFFEEFF49F2B2FE562970" blockId="44.[151,1127,513,1592]" pageId="44" pageNumber="389">
<emphasis id="7BBAEAB5FFEEFFEEFF49F2B2FEDD2A90" bold="true" box="[151,280,1505,1528]" pageId="44" pageNumber="389">Etymology.</emphasis>
Named after Japanese biologist and biocontrol specialist working on
<taxonomicName id="8ECE4D24FFEEFFEEFC33F2B2FE562970" authority=", Yoshimitsu Higashiura." authorityName="Yoshimitsu Higashiura." class="Insecta" family="Encyrtidae" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="389" phylum="Arthropoda" rank="family">Encyrtidae, Yoshimitsu Higashiura.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="01D4652CFFEEFFEEFF49F172FD142950" box="[151,721,1569,1592]" pageId="44" pageNumber="389" type="distribution">
<paragraph id="497136A7FFEEFFEEFF49F172FD142950" blockId="44.[151,1127,513,1592]" box="[151,721,1569,1592]" pageId="44" pageNumber="389">
<emphasis id="7BBAEAB5FFEEFFEEFF49F172FEEF2950" bold="true" box="[151,298,1569,1592]" pageId="44" pageNumber="389">Distribution.</emphasis>
<collectingCountry id="31D97637FFEEFFEEFEEEF172FEB42950" box="[304,369,1569,1592]" name="Japan" pageId="44" pageNumber="389">Japan</collectingCountry>
: Honshu (new record). Endemic.
</paragraph>
</subSubSection>
</treatment>
</document>
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