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<document id="27240A44AF257F9B99E8743A4EF376C2" ID-CLB-Dataset="300538" ID-DOI="10.17161/jom.i103.15655" ID-GBIF-Dataset="9597d08b-2c12-466b-a7fe-efe79569be3b" ID-ISSN="2325-4467" ID-Zenodo-Dep="13146053" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1722455600613" checkinUser="felipe" docAuthor="Engel, Michael S. & Davis, Steven R." docDate="2021" docId="B82B87C26342F07AF18BFCE958C95240" docLanguage="en" docName="JMelittology.2021.103.1-52.pdf" docOrigin="Journal of Melittology 2021 (103)" docSource="http://dx.doi.org/10.17161/jom.i103.15655" docStyle="DocumentStyle:51747547F162FCF7F4DBBB3D518F67EC.3:JMelittology.2013-.journal_article.2cover" docStyleId="51747547F162FCF7F4DBBB3D518F67EC" docStyleName="JMelittology.2013-.journal_article.2cover" docStyleVersion="3" docTitle="Melikertini" docType="key" docVersion="4" lastPageNumber="11" masterDocId="4412FFBA6349F076F07BFE0B5C595119" masterDocTitle="New genera of melikertine bees with facial modifications in Baltic amber (Hymenoptera: Apidae)" masterLastPageNumber="52" masterPageNumber="1" pageNumber="10" updateTime="1722605887797" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:titleInfo id="5F4DEF09570C5C3850A46FF5153C13DB">
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<mods:title id="C88A4E0D5C6005E13316F61DA009F610">New genera of melikertine bees with facial modifications in Baltic amber (Hymenoptera: Apidae)</mods:title>
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<mods:name id="7698F6465E5D536EC39A5C59B977113E" type="personal">
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<mods:roleTerm id="79BE2E1695163FEC34154563DE947C37">Author</mods:roleTerm>
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<mods:namePart id="1559DCC99DC9C292130322AADBFCA390">Engel, Michael S.</mods:namePart>
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<mods:affiliation id="A93594797ACD8060C64D4B78E08787B8">Division of Entomology, Natural History Museum and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). & Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA (mengel @ amnh. org, sdavis @ amnh. org).</mods:affiliation>
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<mods:roleTerm id="448AACC6965F4BEF63AD1E5AC0938DDB">Author</mods:roleTerm>
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<mods:namePart id="D816EF3079F1169C461F49C4F16097DE">Davis, Steven R.</mods:namePart>
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<mods:affiliation id="C623BD63AD30FCE5CC20E7EF9AA77DBD">Division of Entomology, Natural History Museum and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu).</mods:affiliation>
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<mods:typeOfResource id="9E839A37AC76EDDB6374D45BAFDFF059">text</mods:typeOfResource>
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<mods:title id="637812EEC9C45F474574342DC8244024">Journal of Melittology</mods:title>
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<mods:part id="7D50652978288374B14C35631BCCD710">
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<mods:date id="99AC5EBCB88EBB47E6D2998782DD9B5B">2021</mods:date>
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<mods:number id="A1D979E2398D43625F20996F2AB8FFB7">2021-07-06</mods:number>
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<mods:number id="34401704D93BBE5AF53D53A42300FEB9">2021</mods:number>
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<mods:number id="AE9BE817CA48137C401793EBDB22C1CA">103</mods:number>
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<treatment id="B82B87C26342F07AF18BFCE958C95240" ID-DOI="http://doi.org/10.5281/zenodo.13170213" ID-Zenodo-Dep="13170213" LSID="urn:lsid:plazi:treatment:B82B87C26342F07AF18BFCE958C95240" httpUri="http://treatment.plazi.org/id/B82B87C26342F07AF18BFCE958C95240" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10">
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<subSubSection id="7898655F6342F07DF18BFCE95F1953E0" box="[496,832,737,762]" pageId="11" pageNumber="10" type="nomenclature">
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<paragraph id="303D36D46342F07DF18BFCE95F1953E0" blockId="11.[496,832,737,762]" box="[496,832,737,762]" pageId="11" pageNumber="10">
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<heading id="6B7581B86342F07DF18BFCE95F1953E0" box="[496,832,737,762]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="10" reason="2">
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Key to Genera of
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<taxonomicName id="F7824D576342F07DF2C6FCEA5F1953E0" box="[701,832,737,761]" class="Insecta" family="Apidae" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Melikertini">Melikertini</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7898655F6342F07DF090FD0A58CE5280" pageId="11" pageNumber="10" type="description">
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<paragraph id="303D36D46342F07DF090FD0A581C5200" blockId="11.[235,1093,769,793]" box="[235,1093,769,793]" pageId="11" pageNumber="10">
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(modified and expanded from that of
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<bibRefCitation id="54134B256342F07DF2D3FD0A5F645200" author="Engel, M. S." box="[680,829,769,793]" pageId="11" pageNumber="10" pagination="1 - 192" refId="ref14691" refString="Engel, M. S. 2001 a. A monograph of the Baltic amber bees and evolution of the Apoidea (Hymenoptera). Bulletin of the American Museum of Natural History 259: 1 - 192." type="journal article" year="2001">
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Engel, 2001
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<bibRefCitation id="54134B256342F07DF34AFD0A5F645200" author="Dewulf, A. & T. De Meulemeester & M. Dehon & M. S. Engel & D. Michez" box="[817,829,769,793]" pageId="11" pageNumber="10" pagination="35 - 48" refId="ref14597" refString="Dewulf, A., T. De Meulemeester, M. Dehon, M. S. Engel, & D. Michez. 2014. A new interpretation of the bee fossil Melita willardi Cockerell (Hymenoptera, Melitidae) based on geometric morphometrics of the wing. ZooKeys 389: 35 - 48." type="journal article" year="2014">a</bibRefCitation>
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</bibRefCitation>
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, and Engel
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<emphasis id="02F6EAC66342F07DF3BDFD095FAA5200" box="[966,1011,769,793]" italics="true" pageId="11" pageNumber="10">et al</emphasis>
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., 2014)
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</paragraph>
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<paragraph id="303D36D46342F07DF0ABFD4958CE5280" blockId="11.[160,1175,833,921]" pageId="11" pageNumber="10">
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As noted by
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<bibRefCitation id="54134B256342F07DF116FD4A5E665240" author="Engel, M. S. & L. C. V. Breitkreuz & M. Ohl" box="[365,575,833,857]" pageId="11" pageNumber="10" pagination="1 - 18" refId="ref15055" refString="Engel, M. S., L. C. V. Breitkreuz, & M. Ohl. 2014. The first male of the extinct bee tribe Melikertini (Hymenoptera: Apidae). Journal of Melitology 30: 1 - 18." type="journal article" year="2014">
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Engel
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<emphasis id="02F6EAC66342F07DF1C2FD495DB15240" box="[441,488,833,857]" italics="true" pageId="11" pageNumber="10">et al</emphasis>
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. (2014)
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</bibRefCitation>
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it remains unknown to what degree some of the characters presented here are gender or caste specific. Accordingly, care should be taken when using the key in relation to new material of hitherto unknown castes or sexes.
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</paragraph>
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</subSubSection>
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<subSubSection id="7898655F6342F07AF0DBFDCA58C95240" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10" type="key">
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<keyStep id="8B762E716342F07DF0DBFDCA58D65543" pageId="11" pageNumber="10">
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<paragraph id="303D36D46342F07DF0DBFDCA58D452E0" blockId="11.[160,1169,961,1785]" pageId="11" pageNumber="10">
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<keyLead id="8B7395E16342F07DF0DBFDCA58D452E0" pageId="11" pageNumber="10">1. Disc of clypeus comparatively flat, without distinct lateral carinae and without bell-like concavity ...................................................................................................... 2</keyLead>
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</paragraph>
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<paragraph id="303D36D46342F07DF0DBFA0958D65543" blockId="11.[160,1169,961,1785]" pageId="11" pageNumber="10">
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<keyLead id="8B7395E16342F07DF0DBFA0958D65543" pageId="11" pageNumber="10">
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—. Clypeus with lateral carinae rising from apex to form margins of bell-like concavity arising from base of clypeus and overhanging disc (
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<figureCitation id="A8B92A516342F07DF3D9FA2A5FA75520" box="[930,1022,1057,1081]" captionStart="Figures 9–10" captionStartId="13.[160,241,1457,1479]" captionTargetBox="[160,1168,224,1451]" captionTargetId="figure-125@13.[160,1168,224,1451]" captionTargetPageId="13" captionText="Figures 9–10. Holotype worker of Aethemelikertes emunctorii, new genus and species, in Eocene (Bartonian) Baltic amber (SMF Be 14262a). 9. Ventral view. 10. Slightly obliquely dorsal facial view emphasizing upper, slightly scalloped margin of clypeal basin." figureDoi="http://doi.org/10.5281/zenodo.13146061" httpUri="https://zenodo.org/record/13146061/files/figure.png" pageId="11" pageNumber="10">Figs. 10</figureCitation>
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,
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<figureCitation id="A8B92A516342F07DF472FA2A587F5520" box="[1033,1062,1057,1081]" captionStart="Figure 12" captionStartId="16.[160,229,1023,1045]" captionTargetBox="[160,1168,224,1016]" captionTargetId="figure-289@16.[160,1168,224,1016]" captionTargetPageId="16" captionText="Figure 12. Ventral facial view of holotype worker of Aethemelikertes emunctorii, new genus and species (SMF Be 14262a). Note carinae demarcating lateral margins of basin extending to clypeal apex where clypeus extends laterally around base of labrum." figureDoi="http://doi.org/10.5281/zenodo.13146067" httpUri="https://zenodo.org/record/13146067/files/figure.png" pageId="11" pageNumber="10">12</figureCitation>
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,
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<figureCitation id="A8B92A516342F07DF44AFA2A582D5520" box="[1073,1140,1057,1081]" captionStart="Figures 13–16" captionStartId="17.[160,241,1192,1214]" captionTargetBox="[160,1168,224,1185]" captionTargetId="figure-247@17.[160,1168,224,1185]" captionTargetPageId="17" captionText="Figures 13–16. X-ray µCT scans of heads of holotype workers of Succinapis micheneri Engel (AMNH B-JH103) and Aethemelikertes emunctorii, new genus and species (SMF (Be 14262a), in Baltic amber; yellow highlights disc and modification of clypeus, orange highlights labrum. 13. Facial view of S. micheneri. 14. Facial view of A. emunctorii. 15. Oblique profile of S. micheneri. 16. Oblique profile of A. emunctorii. Note that scan resolution was insufficient to resolve most setae." figureDoi="http://doi.org/10.5281/zenodo.13146069" httpUri="https://zenodo.org/record/13146069/files/figure.png" pageId="11" pageNumber="10">14, 16</figureCitation>
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) .. .......................................................................................
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<taxonomicName id="F7824D576342F07DF344FA4A5FB05540" authorityName="Engel & Davis" authorityYear="2021" box="[831,1001,1089,1113]" class="Insecta" family="Apidae" genus="Aethemelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
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<emphasis id="02F6EAC66342F07DF344FA4A5FB05540" box="[831,1001,1089,1113]" italics="true" pageId="11" pageNumber="10">Aethemelikertes</emphasis>
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</taxonomicName>
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Engel,
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<taxonomicNameLabel id="19C557BD6342F07DF43BFA4958D65543" box="[1088,1167,1090,1114]" pageId="11" pageNumber="10" rank="genus">n. gen.</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="8B762E716342F07DF0DBFA6A58C955E0" pageId="11" pageNumber="10">
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<paragraph id="303D36D46342F07DF0DBFA6A58C955C0" blockId="11.[160,1169,961,1785]" pageId="11" pageNumber="10">
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<keyLead id="8B7395E16342F07DF0DBFA6A58C955C0" pageId="11" pageNumber="10">
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2(1). Clypeal protrusion present,
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<emphasis id="02F6EAC66342F07DF244FA695E0E5560" box="[575,599,1122,1145]" italics="true" pageId="11" pageNumber="10">i.e</emphasis>
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., base of clypeus produced into variously modified facial prominences, prominence bending upward over fronto-clypeal portion of epistomal sulcus and obscuring supraclypeal area or even lowermost frons in facial view (
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<figureCitation id="A8B92A516342F07DF19AFACA5E1155C0" box="[481,584,1217,1241]" captionStart="Figures 3–5" captionStartId="7.[160,241,1424,1446]" captionTargetBox="[160,1168,215,1417]" captionTargetId="figure-138@7.[160,1168,215,1417]" captionTargetPageId="7" captionText="Figures 3–5. Worker of Succinapis micheneri Engel (SEMC B-018). 3. Facial view. 4. Dorsal view of head and mesoscutum. 5. Right lateral view." figureDoi="http://doi.org/10.5281/zenodo.13146057" httpUri="https://zenodo.org/record/13146057/files/figure.png" pageId="11" pageNumber="10">Figs. 3, 4</figureCitation>
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,
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<figureCitation id="A8B92A516342F07DF228FACA5E3B55C0" box="[595,610,1217,1241]" captionStart="Figures 6–8" captionStartId="8.[160,241,1281,1303]" captionTargetBox="[160,1168,224,1275]" captionTargetId="figure-207@8.[160,1168,224,1275]" captionTargetPageId="8" captionText="Figures 6–8. Worker of Succinapis micheneri Engel (SEMC B-017). 6. Oblique facial view. 7. Outer (prolateral) view of metatibia and metabasitarsus. 8. Detail of fringe setae of metatibial posterior/upper margin; white arrow indicates a deceptive internal microfracture of resin trail with branched fractal patern (such fracture planes and trails are a common taphonomic structure among or even running alongside many fine setae in amber inclusions, and can give such setae an artificially plumose appearance; accordingly, care must be taken before concluding whether or not amber-included setae are branched); black arrow highlights one of many truly plumose setae, separate from the internal microfracture planes." figureDoi="http://doi.org/10.5281/zenodo.13146059" httpUri="https://zenodo.org/record/13146059/files/figure.png" pageId="11" pageNumber="10">6</figureCitation>
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,
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<figureCitation id="A8B92A516342F07DF216FACA5EE955C0" box="[621,688,1217,1241]" captionStart="Figures 23–26" captionStartId="24.[160,241,1266,1288]" captionTargetBox="[160,1168,224,1258]" captionTargetId="figure-175@24.[160,1168,224,1258]" captionTargetPageId="24" captionText="Figures 23–26. X-ray µCT scans of heads of holotype workers of Haidomelikertes proboscidea (Engel), new combination (AMNH B-JH96), and H. uraeus, new genus and species (SMF (Be 14263a), in Baltic amber; yellow highlights disc and modification of clypeus, orange highlights labrum. 23. Facial view of H. proboscidea. 24. Facial view of H. uraeus. 25. Oblique profile of H. proboscidea. 26. Oblique profile of H. uraeus. Note that scan resolution was insufficient to resolve most setae." figureDoi="http://doi.org/10.5281/zenodo.13146081" httpUri="https://zenodo.org/record/13146081/files/figure.png" pageId="11" pageNumber="10">23–26</figureCitation>
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) ................................................................... 3
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</keyLead>
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</paragraph>
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<paragraph id="303D36D46342F07DF0DBFAE958C955E0" blockId="11.[160,1169,961,1785]" box="[160,1168,1249,1273]" pageId="11" pageNumber="10">
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<keyLead id="8B7395E16342F07DF0DBFAE958C955E0" box="[160,1168,1249,1273]" pageId="11" pageNumber="10">—. Clypeal protrusion absent ......................................................................................... 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="8B762E716342F07DF0DBFB0A58C95463" pageId="11" pageNumber="10">
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<paragraph id="303D36D46342F07DF0DBFB0A58C95420" blockId="11.[160,1169,961,1785]" pageId="11" pageNumber="10">
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<keyLead id="8B7395E16342F07DF0DBFB0A58C95420" pageId="11" pageNumber="10">
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3(2). Apex of clypeal protrusion narrow, narrower than intertorular distance (
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<figureCitation id="A8B92A516342F07DF423FB0A5D445420" captionStart="Figures 3–5" captionStartId="7.[160,241,1424,1446]" captionTargetBox="[160,1168,215,1417]" captionTargetId="figure-138@7.[160,1168,215,1417]" captionTargetPageId="7" captionText="Figures 3–5. Worker of Succinapis micheneri Engel (SEMC B-018). 3. Facial view. 4. Dorsal view of head and mesoscutum. 5. Right lateral view." figureDoi="http://doi.org/10.5281/zenodo.13146057" httpUri="https://zenodo.org/record/13146057/files/figure.png" pageId="11" pageNumber="10">Figs. 3, 4</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="A8B92A516342F07DF152FB2A5D6E5420" box="[297,311,1313,1337]" captionStart="Figures 6–8" captionStartId="8.[160,241,1281,1303]" captionTargetBox="[160,1168,224,1275]" captionTargetId="figure-207@8.[160,1168,224,1275]" captionTargetPageId="8" captionText="Figures 6–8. Worker of Succinapis micheneri Engel (SEMC B-017). 6. Oblique facial view. 7. Outer (prolateral) view of metatibia and metabasitarsus. 8. Detail of fringe setae of metatibial posterior/upper margin; white arrow indicates a deceptive internal microfracture of resin trail with branched fractal patern (such fracture planes and trails are a common taphonomic structure among or even running alongside many fine setae in amber inclusions, and can give such setae an artificially plumose appearance; accordingly, care must be taken before concluding whether or not amber-included setae are branched); black arrow highlights one of many truly plumose setae, separate from the internal microfracture planes." figureDoi="http://doi.org/10.5281/zenodo.13146059" httpUri="https://zenodo.org/record/13146059/files/figure.png" pageId="11" pageNumber="10">6</figureCitation>
|
|||
|
|
) .................................................................................................
|
|||
|
|
<taxonomicName id="F7824D576342F07DF3A9FB29581E5420" authorityName="Engel" authorityYear="2001" box="[978,1095,1314,1337]" class="Insecta" family="Apidae" genus="Succinapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="02F6EAC66342F07DF3A9FB29581E5420" box="[978,1095,1314,1337]" italics="true" pageId="11" pageNumber="10">Succinapis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Engel
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="303D36D46342F07DF0DBFB4958C95463" blockId="11.[160,1169,961,1785]" pageId="11" pageNumber="10">
|
|||
|
|
<keyLead id="8B7395E16342F07DF0DBFB4958C95463" pageId="11" pageNumber="10">
|
|||
|
|
—. Apex of clypeal protrusion broad, as wide as or slightly wider than intertorular distance (
|
|||
|
|
<figureCitation id="A8B92A516342F07DF11FFB6A5DBC5460" box="[356,485,1377,1401]" captionStart="Figures 23–26" captionStartId="24.[160,241,1266,1288]" captionTargetBox="[160,1168,224,1258]" captionTargetId="figure-175@24.[160,1168,224,1258]" captionTargetPageId="24" captionText="Figures 23–26. X-ray µCT scans of heads of holotype workers of Haidomelikertes proboscidea (Engel), new combination (AMNH B-JH96), and H. uraeus, new genus and species (SMF (Be 14263a), in Baltic amber; yellow highlights disc and modification of clypeus, orange highlights labrum. 23. Facial view of H. proboscidea. 24. Facial view of H. uraeus. 25. Oblique profile of H. proboscidea. 26. Oblique profile of H. uraeus. Note that scan resolution was insufficient to resolve most setae." figureDoi="http://doi.org/10.5281/zenodo.13146081" httpUri="https://zenodo.org/record/13146081/files/figure.png" pageId="11" pageNumber="10">Figs. 23–26</figureCitation>
|
|||
|
|
) .................................................
|
|||
|
|
<taxonomicName id="F7824D576342F07DF344FB6A5FB55460" authorityName="Engel & Davis" authorityYear="2021" box="[831,1004,1377,1401]" class="Insecta" family="Apidae" genus="Haidomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="02F6EAC66342F07DF344FB6A5FB55460" box="[831,1004,1377,1401]" italics="true" pageId="11" pageNumber="10">Haidomelikertes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Engel,
|
|||
|
|
<taxonomicNameLabel id="19C557BD6342F07DF439FB6958C95463" box="[1090,1168,1378,1402]" pageId="11" pageNumber="10" rank="genus">n. gen.</taxonomicNameLabel>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="8B762E716342F07DF0DBFB8A58C95700" pageId="11" pageNumber="10">
|
|||
|
|
<paragraph id="303D36D46342F07DF0DBFB8A58D554A0" blockId="11.[160,1169,961,1785]" pageId="11" pageNumber="10">
|
|||
|
|
<keyLead id="8B7395E16342F07DF0DBFB8A58D554A0" pageId="11" pageNumber="10">4(2). Mesoscutellum without tongue-like medioapical extension [females: workers] ....................................................................................................................................... 5</keyLead>
|
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|
|
</paragraph>
|
|||
|
|
<paragraph id="303D36D46342F07DF0DBFBC958C95700" blockId="11.[160,1169,961,1785]" pageId="11" pageNumber="10">
|
|||
|
|
<keyLead id="8B7395E16342F07DF0DBFBC958C95700" pageId="11" pageNumber="10">
|
|||
|
|
—. Mesoscutellum with prominent, tongue-like medioapical extension projecting over metanotum, propodeum, and portions of metasoma, dorsal surface of extension densely setose [males: drones] ......................
|
|||
|
|
<taxonomicName id="F7824D576342F07DF32AF80A58575700" authorityName="Engel, Breitkreuz & Ohl" authorityYear="2014" box="[849,1038,1537,1561]" class="Insecta" family="Apidae" genus="Mochlomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="02F6EAC66342F07DF32AF80A58575700" box="[849,1038,1537,1561]" italics="true" pageId="11" pageNumber="10">Mochlomelikertes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Engel
|
|||
|
|
<emphasis id="02F6EAC66342F07DF426F80958D05700" box="[1117,1161,1537,1561]" italics="true" pageId="11" pageNumber="10">et al</emphasis>
|
|||
|
|
.
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="8B762E716342F07DF0DBF82A58C957E0" pageId="11" pageNumber="10">
|
|||
|
|
<paragraph id="303D36D46342F07DF0DBF82A58C95780" blockId="11.[160,1169,961,1785]" pageId="11" pageNumber="10">
|
|||
|
|
<keyLead id="8B7395E16342F07DF0DBF82A58C95780" pageId="11" pageNumber="10">5(4). Mesoscutellum bulging, overhanging metanotum and propodeum; apical margins of metasomal terga distinctly lighter than remainder of metasoma, thus metasoma appears banded; anterior and posterior margins of metabasitarus distinctly converging toward apex .......................................................................... 6</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="303D36D46342F07DF0DBF8A958C957E0" blockId="11.[160,1169,961,1785]" pageId="11" pageNumber="10">
|
|||
|
|
<keyLead id="8B7395E16342F07DF0DBF8A958C957E0" pageId="11" pageNumber="10">—. Mesoscutellum not bulging, not overhanging metanotum or propodeum; metasomal terga uniformly colored; anterior and posterior margins of metabasitarsus approximately parallel ........................................................................................ 7</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="8B762E716345F07AF0DBFEEA58D75340" pageId="12" pageNumber="11">
|
|||
|
|
<paragraph id="303D36D46345F07AF0DBFEEA58D75080" blockId="12.[160,1169,225,857]" pageId="12" pageNumber="11">
|
|||
|
|
<keyLead id="8B7395E16345F07AF0DBFEEA58D75080" pageId="12" pageNumber="11">
|
|||
|
|
6(5). Forewing with anterior margin of first submarginal cell approximately equal to length of anterior margin of second submarginal (
|
|||
|
|
<emphasis id="02F6EAC66345F07AF31CFF095FD95000" box="[871,896,258,281]" italics="true" pageId="12" pageNumber="11">i.e</emphasis>
|
|||
|
|
., r-rs as long as immediately succeeding abscissa of Rs); compound eyes converging below; second flagellomere distinctly shorter than third flagellomere, first flagellomere distinctly shorter than combined lengths of second and third flagellomeres ............. ..............................................................................................................
|
|||
|
|
<taxonomicName id="F7824D576345F07AF3A3FF8A581D5080" authorityName="Engel" authorityYear="2001" box="[984,1092,385,409]" class="Insecta" family="Apidae" genus="Melissites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="02F6EAC66345F07AF3A3FF8A581D5080" box="[984,1092,385,409]" italics="true" pageId="12" pageNumber="11">Melissites</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Engel
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="303D36D46345F07AF0DBFFA958D75340" blockId="12.[160,1169,225,857]" pageId="12" pageNumber="11">
|
|||
|
|
<keyLead id="8B7395E16345F07AF0DBFFA958D75340" pageId="12" pageNumber="11">
|
|||
|
|
—. Forewing with anterior margin of first submarginal cell many times longer than length of anterior margin of second submarginal cell (
|
|||
|
|
<emphasis id="02F6EAC66345F07AF3D1FFC95F9B50C0" box="[938,962,450,473]" italics="true" pageId="12" pageNumber="11">i.e</emphasis>
|
|||
|
|
., r-rs many times longer than immediately succeeding abscissa of Rs); compound eyes approximately parallel; second and third flagellomeres approximately equal in length, first flagellomere approximately equal to combined lengths of second and third flagellomeres ..............................................................................
|
|||
|
|
<taxonomicName id="F7824D576345F07AF3DCFC4958D75340" authority="Manning" authorityName="Manning" authorityYear="1960" box="[935,1166,577,601]" class="Insecta" family="Apidae" genus="Roussyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="02F6EAC66345F07AF3DCFC4958425340" box="[935,1051,578,601]" italics="true" pageId="12" pageNumber="11">Roussyana</emphasis>
|
|||
|
|
Manning
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="8B762E716345F07AF0DBFC6A58C953E3" pageId="12" pageNumber="11">
|
|||
|
|
<paragraph id="303D36D46345F07AF0DBFC6A58C95380" blockId="12.[160,1169,225,857]" pageId="12" pageNumber="11">
|
|||
|
|
<keyLead id="8B7395E16345F07AF0DBFC6A58C95380" pageId="12" pageNumber="11">
|
|||
|
|
7(5). Apical margin of clypeus straight, flat, not flared anteriorly and not projecting over plane of labrum (genus
|
|||
|
|
<taxonomicName id="F7824D576345F07AF238FC8A5EED5380" authorityName="Engel" authorityYear="1998" box="[579,692,641,665]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="02F6EAC66345F07AF238FC8A5EED5380" box="[579,692,641,665]" italics="true" pageId="12" pageNumber="11">Melikertes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Engel,
|
|||
|
|
<taxonomicNameLabel id="19C557BD6345F07AF372FC8A5F7F5380" box="[777,806,641,665]" pageId="12" pageNumber="11" sensu="lato">
|
|||
|
|
<emphasis id="02F6EAC66345F07AF372FC8A5F785380" box="[777,801,641,665]" italics="true" pageId="12" pageNumber="11">s.l</emphasis>
|
|||
|
|
.
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
) ................................................. 8
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="303D36D46345F07AF0DBFCA958C953E3" blockId="12.[160,1169,225,857]" pageId="12" pageNumber="11">
|
|||
|
|
<keyLead id="8B7395E16345F07AF0DBFCA958C953E3" pageId="12" pageNumber="11">
|
|||
|
|
—. Apical margin of clypeus flared and projecting anteriorly over plane with labrum, medioapically with shallow U-shaped emargination (
|
|||
|
|
<figureCitation id="A8B92A516345F07AF3C3FCCA58D553C0" box="[952,1164,705,729]" captionStart="Figures 34–39" captionStartId="28.[160,241,1410,1432]" captionTargetBox="[160,1168,224,1404]" captionTargetId="figure-161@28.[160,1168,224,1404]" captionTargetPageId="28" captionText="Figures 34–39. PPC-SRµCT scan of worker head of Amelikertotes clypeata (Engel), new combination; upper right worker from figure 29. 34. Dorsal view. 35. Ventral view. 36. Facial view. 37. Posterior view. 37. Right lateral view. 38. Left lateral view. Arrows point to flared and cleft clypeus. Note that scan resolution was insufficient to resolve most fine setae." figureDoi="http://doi.org/10.5281/zenodo.13146089" httpUri="https://zenodo.org/record/13146089/files/figure.png" pageId="12" pageNumber="11">Figs. 34–36, 38, 39</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="A8B92A516345F07AF08FFCEA5DDE53E0" box="[244,391,737,761]" captionStart="Figures 57–62" captionStartId="32.[160,241,1387,1409]" captionTargetBox="[160,1168,211,1380]" captionTargetId="figure-163@32.[160,1168,211,1380]" captionTargetPageId="32" captionText="Figures 57–62. PPC-SRµCT scan of worker head of Amelikertotes clypeata (Engel), new combination; lower left worker from figure 29. 57. Facial view. 58. Right lateral view. 59. Posterior view. 60. Left lateral view. 61. Ventral view. 62. Ventral view with mandibles removed to reveal labrum. Scale bar in center for 57–60, scale bar at botom left for 61 and 62. Note that scan resolution was insufficient to resolve most fine setae." figureDoi="http://doi.org/10.5281/zenodo.13146097" httpUri="https://zenodo.org/record/13146097/files/figure.png" pageId="12" pageNumber="11">57, 58, 60–62</figureCitation>
|
|||
|
|
) ...................................................................
|
|||
|
|
<taxonomicName id="F7824D576345F07AF323FCEA5FB553E0" authorityName="Engel & Davis" authorityYear="2021" box="[856,1004,737,761]" class="Insecta" family="Apidae" genus="Amelikertotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="02F6EAC66345F07AF323FCEA5FB553E0" box="[856,1004,737,761]" italics="true" pageId="12" pageNumber="11">Amelikertotes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Engel,
|
|||
|
|
<taxonomicNameLabel id="19C557BD6345F07AF439FCE958C953E3" box="[1090,1168,738,762]" pageId="12" pageNumber="11" rank="genus">n. gen.</taxonomicNameLabel>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="8B762E716345F07AF0DBFD0A58C95240" pageId="12" pageNumber="11">
|
|||
|
|
<paragraph id="303D36D46345F07AF0DBFD0A58C95220" blockId="12.[160,1169,225,857]" pageId="12" pageNumber="11">
|
|||
|
|
<keyLead id="8B7395E16345F07AF0DBFD0A58C95220" pageId="12" pageNumber="11">
|
|||
|
|
8(7). Forewing with two submarginal cells (1rs-m absent) .............................................. .........................................................................
|
|||
|
|
<taxonomicName id="F7824D576345F07AF29BFD2A5F275220" authorityName="Engel & Ortega-Blanco" authorityYear="2013" box="[736,894,801,825]" class="Insecta" family="Apidae" genus="Paramelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="02F6EAC66345F07AF29BFD2A5F275220" box="[736,894,801,825]" italics="true" pageId="12" pageNumber="11">Paramelikertes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Engel & Ortega-Blanco
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="303D36D46345F07AF0DBFD4958C95240" blockId="12.[160,1169,225,857]" box="[160,1168,833,857]" pageId="12" pageNumber="11">
|
|||
|
|
<keyLead id="8B7395E16345F07AF0DBFD4958C95240" box="[160,1168,833,857]" pageId="12" pageNumber="11">
|
|||
|
|
—. Forewing with three submarginal cells (1rs-m present) ....
|
|||
|
|
<taxonomicName id="F7824D576345F07AF3EEFD4A585F5240" authorityName="Engel" authorityYear="1998" box="[917,1030,833,857]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="02F6EAC66345F07AF3EEFD4A585F5240" box="[917,1030,833,857]" italics="true" pageId="12" pageNumber="11">Melikertes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Engel,
|
|||
|
|
<taxonomicNameLabel id="19C557BD6345F07AF420FD4958C95240" box="[1115,1168,834,857]" pageId="12" pageNumber="11" sensu="stricto">
|
|||
|
|
<emphasis id="02F6EAC66345F07AF420FD4958D35240" box="[1115,1162,834,857]" italics="true" pageId="12" pageNumber="11">s.str</emphasis>
|
|||
|
|
.
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|