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<document id="BC42D3A1F800458BB354955CC18A4D9E" ID-CLB-Dataset="50776" ID-DOI="10.5281/zenodo.171684" ID-GBIF-Dataset="b5adc5a8-a114-4089-bd9f-1f7ddc239f3e" ID-ISSN="1175­5326" ID-Zenodo-Dep="171684" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459756681018" checkinUser="plazi" docAuthor="Bartsch, Ilse" docDate="2006" docId="26686D0AFFCEFF97FEE973C47FE4FA61" docLanguage="en" docName="zt01120p049.pdf" docOrigin="Zootaxa 1120" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Rhombognathus Trouessart 1888" docType="treatment" docVersion="10" lastPageNumber="43" masterDocId="DA511572FFCFFF95FFE170527E7AFFF7" masterDocTitle="A new species and new record of Rhombognathus from Singapore (Acari: Halacaridae)" masterLastPageNumber="49" masterPageNumber="41" pageNumber="42" updateTime="1698218064201" updateUser="ExternalLinkService">
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<mods:title id="74CD62A8911683FFA45B5E29696A9B22">A new species and new record of Rhombognathus from Singapore (Acari: Halacaridae)</mods:title>
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<emphasis id="9CB5000EFFCEFF94FEE973C47C84FC47" bold="true" box="[264,766,918,944]" pageId="1" pageNumber="42">
Genus
<taxonomicName id="69C1A79FFFCEFF94FEBE73C47C84FC47" authority="Trouessart, 1888" authorityName="Trouessart" authorityYear="1888" box="[351,766,918,944]" class="Arachnida" family="Halacaridae" genus="Rhombognathus" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="9CB5000EFFCEFF94FEBE73C47C5DFC47" bold="true" box="[351,551,918,944]" italics="true" pageId="1" pageNumber="42">Rhombognathus</emphasis>
Trouessart, 1888
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</emphasis>
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<paragraph id="AE7EDC1CFFCEFF94FEE973B57CB9FC0A" blockId="1.[264,1324,998,1117]" box="[264,707,998,1022]" pageId="1" pageNumber="42">
<taxonomicName id="69C1A79FFFCEFF94FEE973B57CB9FC0A" authority="Bartsch, 1983" authorityName="Bartsch" authorityYear="1983" box="[264,707,998,1022]" class="Arachnida" family="Halacaridae" genus="Rhombognathus" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="scutulatus">
<emphasis id="9CB5000EFFCEFF94FEE973B57C52FC09" box="[264,552,999,1022]" italics="true" pageId="1" pageNumber="42">Rhombognathus scutulatus</emphasis>
<bibRefCitation id="CA50A1EDFFCEFF94FDCE73B47CB9FC0A" author="Bartsch" box="[559,707,998,1021]" pageId="1" pageNumber="48" refString="Bartsch, I. (1983) Zur Halacaridenfauna der Philippinen. Beschreibung von funf Arten der Gattung Rhombognathus (Acari, Halacaridae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 7, 399 - 416." type="journal article" year="1983">Bartsch, 1983</bibRefCitation>
</taxonomicName>
</paragraph>
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<taxonomicName id="69C1A79FFFCEFF94FEE974557C84FBEA" authority="Bartsch, 1983: 413" authorityName="Bartsch" authorityPageNumber="413" authorityYear="1983" box="[264,766,1030,1054]" class="Arachnida" family="Halacaridae" genus="Rhombognathus" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="scutulatus">
<emphasis id="9CB5000EFFCEFF94FEE974557C53FBE9" box="[264,553,1031,1054]" italics="true" pageId="1" pageNumber="42">Rhombognathus scutulatus</emphasis>
<treatmentCitation id="2F60FA0DFFCEFF94FDD074547C84FBEA" author="Bartsch" box="[561,766,1030,1053]" page="413" pageId="1" pageNumber="42" year="1983">
<bibRefCitation id="CA50A1EDFFCEFF94FDD074547CB0FBEA" author="Bartsch" box="[561,714,1030,1053]" pageId="1" pageNumber="48" refString="Bartsch, I. (1983) Zur Halacaridenfauna der Philippinen. Beschreibung von funf Arten der Gattung Rhombognathus (Acari, Halacaridae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 7, 399 - 416." type="journal article" year="1983">Bartsch, 1983</bibRefCitation>
: 413
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415, figs 4657;
<treatmentCitation id="2F60FA0DFFCEFF94FC2374547AFEFBEA" author="Bartsch" box="[962,1156,1030,1053]" page="20" pageId="1" pageNumber="42" year="1993">
<bibRefCitation id="CA50A1EDFFCEFF94FC2374547A21FBEA" author="Bartsch" box="[962,1115,1030,1053]" pageId="1" pageNumber="49" refString="Bartsch, I. (1993) Rhombognathine mites (Halacaridae, Acari) from Rottnest Island, Western Australia. In: Wells, F. E., Walker, D. I., Kirkman, H. and Lethbridge, R. (Eds.), The Marine Flora and Fauna of Rottnest Island, Western Australia Western Australian Museum, Perth, pp. 19 - 43." type="book chapter" year="1993">Bartsch, 1993</bibRefCitation>
: 20
</treatmentCitation>
, 21, fig. 1AC;
<treatmentCitation id="2F60FA0DFFCEFF94FED974747C7FFBCA" author="Bartsch" box="[312,517,1062,1085]" page="354" pageId="1" pageNumber="42" year="1999">
<bibRefCitation id="CA50A1EDFFCEFF94FED974747FABFBCA" author="Bartsch" box="[312,465,1062,1085]" pageId="1" pageNumber="49" refString="Bartsch, I. (1999) Halacaridae (Acari) from Rottnest Island, Western Australia. Mites on fronds of the seagrass Amphibolis. In: Walker, D. I. and Wells, F. E. (Eds). The Seagrass Flora and Fauna of Rottnest Island, Western Australian Museum, Perth, pp. 333 - 357." type="book chapter" year="1999">Bartsch, 1999</bibRefCitation>
: 354
</treatmentCitation>
355, fig. 12FH;
<treatmentCitation id="2F60FA0DFFCEFF94FD2D74747DE1FBCA" author="Bartsch" box="[716,923,1062,1085]" page="190" pageId="1" pageNumber="42" year="2000">
<bibRefCitation id="CA50A1EDFFCEFF94FD2D74747D1FFBCA" author="Bartsch" box="[716,869,1062,1085]" pageId="1" pageNumber="49" refString="Bartsch, I. (2000) Rhombognathinae (Acari: Halacaridae) from the Great Barrier Reef, Australia. Memoirs of the Queensland Museum, 45, 165 - 203." type="journal article" year="2000">Bartsch, 2000</bibRefCitation>
: 190
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;
<treatmentCitation id="2F60FA0DFFCEFF94FC4674747A0EFBCA" author="Bartsch" box="[935,1140,1062,1085]" page="273" pageId="1" pageNumber="42" year="2003">
<bibRefCitation id="CA50A1EDFFCEFF94FC4674747A45FBCA" author="Bartsch" box="[935,1087,1062,1085]" pageId="1" pageNumber="49" refString="Bartsch, I. (2003) Rhombognathine mites (Halacaridae, Acari) from Dampier, Western Australia: taxonomy and biogeography. In: Wells, F. E., Walker, D. I. &amp; Jones, D. S. (Eds), The Marine Flora and Fauna of Dampier, Western Australia. Western Australian Museum, Perth, pp. 255 - 280." type="book chapter" year="2003">Bartsch, 2003</bibRefCitation>
: 273
</treatmentCitation>
275, fig. 9AD;
<treatmentCitation id="2F60FA0DFFCEFF94FED974147C5BFBAA" author="Chatterjee" box="[312,545,1094,1117]" page="284" pageId="1" pageNumber="42" year="1995">
<bibRefCitation id="CA50A1EDFFCEFF94FED974147F96FBAA" author="Chatterjee" box="[312,492,1094,1117]" pageId="1" pageNumber="49" refString="Chatterjee, T. (1995) Record of three species of Rhombognathus (Halacaridae: Acari) from Indian Ocean Region. Journal of the Bombay Natural History Society, 92, 282 - 286." type="journal article" year="1995">Chatterjee, 1995</bibRefCitation>
: 284
</treatmentCitation>
, figs 1519;
<treatmentCitation id="2F60FA0DFFCEFF94FD4A74147A62FBAA" author="Chatterjee" box="[683,1048,1094,1117]" page="187" pageId="1" pageNumber="42" year="2000">
<bibRefCitation id="CA50A1EDFFCEFF94FD4A74147D9FFBAA" author="Chatterjee" box="[683,997,1094,1117]" pageId="1" pageNumber="49" refString="Chatterjee, T. &amp; De Troch, M. (2000) Halacaridae (Acari) from Gazi Bay (Kenya): description and biogeography of three new and two known species. Hydrobiologia, 427, 177 - 194." type="journal article" year="2000">Chatterjee &amp; De Troch, 2000</bibRefCitation>
: 187
</treatmentCitation>
189, fig. 6AF.
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</paragraph>
</subSubSection>
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<emphasis id="9CB5000EFFCEFF94FEE974DD7F9CFB5F" box="[264,486,1167,1192]" italics="true" pageId="1" pageNumber="42">Material examined</emphasis>
</paragraph>
<paragraph id="AE7EDC1CFFCEFF94FED974E57CE4FAD6" blockId="1.[264,1323,1167,1313]" pageId="1" pageNumber="42">
One female,
<collectionCode id="C8D044D9FFCEFF94FE3274E57C6AFB26" box="[467,528,1207,1233]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="42">ZRC</collectionCode>
.ARA.489,
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male,
<collectionCode id="C8D044D9FFCEFF94FD1674E57D4EFB26" box="[759,820,1207,1233]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="42">ZRC</collectionCode>
.ARA.490,
<collectingCountry id="D6D69C8CFFCEFF94FC5B74E57A4EFB26" box="[954,1076,1207,1233]" name="Singapore" pageId="1" pageNumber="42">Singapore</collectingCountry>
, Strait of
<collectingCountry id="D6D69C8CFFCEFF94FB4D74E57B5CFB26" box="[1196,1318,1207,1233]" name="Singapore" pageId="1" pageNumber="42">Singapore</collectingCountry>
, Labrador Park (
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,
<geoCoordinate id="CBF5BADBFFCEFF94FDCF748D7CD8FB0E" box="[558,674,1247,1273]" direction="east" orientation="longitude" pageId="1" pageNumber="42" precision="925" value="103.8">103°48E</geoCoordinate>
), patches of small brown and green algae from rocks, mid­tide area,
<date id="DA7FFADCFFCEFF94FE5375557CE1FAD6" box="[434,667,1287,1313]" pageId="1" pageNumber="42" value="2004-09-28">28 September 2004</date>
.
</paragraph>
</subSubSection>
<subSubSection id="E6DB8F97FFCEFF94FEE975057CF0F8DE" pageId="1" pageNumber="42" type="diagnosis">
<paragraph id="AE7EDC1CFFCEFF94FEE975057F05FA87" blockId="1.[264,1323,1367,1833]" box="[264,383,1367,1392]" pageId="1" pageNumber="42">
<emphasis id="9CB5000EFFCEFF94FEE975057F05FA87" box="[264,383,1367,1392]" italics="true" pageId="1" pageNumber="42">Diagnosis</emphasis>
</paragraph>
<paragraph id="AE7EDC1CFFCEFF94FED9752D7CF0F8DE" blockId="1.[264,1323,1367,1833]" pageId="1" pageNumber="42">Idiosomal length of female 360 µm, of male 322 µm. Dorsal plates foveate, fused to a single shield. Ventral plates fused. Area representing PD with single pair of setae. AE and PE generally each with one pair of adjunct setae. Female with five pairs of pgs and two pairs of sgs. Male with 19 plumose pgs, basilar setae included, the latter situated at 0.8. Male genital sclerites with two pairs of sgs. Gnathosoma 1.2 times longer than wide. Length ratio idiosoma:gnathosoma 1:0.23. Length:height ratio of telofemora 2.52.9. Leg chaetotaxy: leg I, 1, 2, 56, 5, 5, 4; leg II, 1, 2, 6, 5, 4; leg III, 1, 2, 3, 3, 5, 4; leg IV, 0, 2, 3, 3, 5, 3. Telofemora I to IV with (3)4/(1)2, 4/2, 3/0, 3/0 dorsal/ventral setae. None of ventral setae of genua enlarged and bipectinate. Tibiae I to IV with 2, 1, 1, 2 bipectinate setae. Paired fossary setae faintly plumose. Two basalmost setae on tarsus III adjacent. Claws rake­like with 1619 tines</paragraph>
</subSubSection>
<subSubSection id="E6DB8F97FFCDFF97FEE9714E7DC9FB51" pageId="2" pageNumber="43" type="discussion">
<paragraph id="AE7EDC1CFFCDFF97FEE9714E7F14FEC2" blockId="2.[264,1324,284,1190]" box="[264,366,284,309]" pageId="2" pageNumber="43">
<emphasis id="9CB5000EFFCDFF97FEE9714E7F14FEC2" box="[264,366,284,309]" italics="true" pageId="2" pageNumber="43">Remarks</emphasis>
</paragraph>
<paragraph id="AE7EDC1CFFCDFF97FED971167D7EFE21" blockId="2.[264,1324,284,1190]" pageId="2" pageNumber="43">
<taxonomicName id="69C1A79FFFCDFF97FED971167C0FFEAA" box="[312,629,324,349]" class="Arachnida" family="Halacaridae" genus="Rhombognathus" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="scutulatus">
<emphasis id="9CB5000EFFCDFF97FED971167C0FFEAA" box="[312,629,324,349]" italics="true" pageId="2" pageNumber="43">Rhombognathus scutulatus</emphasis>
</taxonomicName>
is one of the most wide­spread
<taxonomicName id="69C1A79FFFCDFF97FC1171167AD5FEAA" box="[1008,1199,324,349]" class="Arachnida" family="Halacaridae" genus="Rhombognathus" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="9CB5000EFFCDFF97FC1171167AD5FEAA" box="[1008,1199,324,349]" italics="true" pageId="2" pageNumber="43">Rhombognathus</emphasis>
</taxonomicName>
species in the Indo­West Pacific area (
<bibRefCitation id="CA50A1EDFFCDFF97FDB1713E7D2FFE71" author="Bartsch" box="[592,853,364,390]" pageId="2" pageNumber="49" refString="Bartsch, I. (2003) Rhombognathine mites (Halacaridae, Acari) from Dampier, Western Australia: taxonomy and biogeography. In: Wells, F. E., Walker, D. I. &amp; Jones, D. S. (Eds), The Marine Flora and Fauna of Dampier, Western Australia. Western Australian Museum, Perth, pp. 255 - 280." type="book chapter" year="2003">Bartsch, 2003: fig. 11</bibRefCitation>
). There seem to be differences between populations of various localities, namely in the size of the claw pectines and their number of tines and in the length of the telofemora.
</paragraph>
<paragraph id="AE7EDC1CFFCDFF97FED971B67DF5FC41" blockId="2.[264,1324,284,1190]" pageId="2" pageNumber="43">
The largest number of tines, 2428, was found in specimens from Rottnest Island, off Perth, Western
<collectingCountry id="D6D69C8CFFCDFF97FE5D725E7C52FDD1" box="[444,552,524,550]" name="Australia" pageId="2" pageNumber="43">Australia</collectingCountry>
(
<bibRefCitation id="CA50A1EDFFCDFF97FDD6725E7C9AFDD1" author="Bartsch" box="[567,736,524,550]" pageId="2" pageNumber="49" refString="Bartsch, I. (1993) Rhombognathine mites (Halacaridae, Acari) from Rottnest Island, Western Australia. In: Wells, F. E., Walker, D. I., Kirkman, H. and Lethbridge, R. (Eds.), The Marine Flora and Fauna of Rottnest Island, Western Australia Western Australian Museum, Perth, pp. 19 - 43." type="book chapter" year="1993">Bartsch, 1993</bibRefCitation>
). Specimens from the Great Barrier Reef, tropical Queensland, had 2224 tines (
<bibRefCitation id="CA50A1EDFFCDFF97FD9572667D65FDB9" author="Bartsch" box="[628,799,564,590]" pageId="2" pageNumber="49" refString="Bartsch, I. (2000) Rhombognathinae (Acari: Halacaridae) from the Great Barrier Reef, Australia. Memoirs of the Queensland Museum, 45, 165 - 203." type="journal article" year="2000">Bartsch, 2000</bibRefCitation>
), and those from the Dampier Archipelago, tropical Western
<collectingCountry id="D6D69C8CFFCDFF97FE3E720E7C37FD81" box="[479,589,604,630]" name="Australia" pageId="2" pageNumber="43">Australia</collectingCountry>
, 1820 tines (
<bibRefCitation id="CA50A1EDFFCDFF97FCE6720E7DC3FD81" author="Bartsch" box="[775,953,604,630]" pageId="2" pageNumber="49" refString="Bartsch, I. (2003) Rhombognathine mites (Halacaridae, Acari) from Dampier, Western Australia: taxonomy and biogeography. In: Wells, F. E., Walker, D. I. &amp; Jones, D. S. (Eds), The Marine Flora and Fauna of Dampier, Western Australia. Western Australian Museum, Perth, pp. 255 - 280." type="book chapter" year="2003">Bartsch, 2003</bibRefCitation>
). In the two specimens from
<collectingCountry id="D6D69C8CFFCDFF97FEE972D67F05FD69" box="[264,383,644,670]" name="Singapore" pageId="2" pageNumber="43">Singapore</collectingCountry>
claws I and II had 1617 tines, claws
<date id="DA7FFADCFFCDFF97FCA472D67A76FD69" box="[837,1036,644,670]" pageId="2" pageNumber="43" value="1917-03-19" valueMax="1917-04-19" valueMin="1917-03-19">III and IV 1719</date>
tines. The
<typeStatus id="717A62BEFFCDFF97FB7172D67AB9FD69" box="[1168,1219,644,670]" pageId="2" pageNumber="43">type</typeStatus>
material from the
<collectingCountry id="D6D69C8CFFCDFF97FE9272FE7F8DFD31" box="[371,503,684,710]" name="Philippines" pageId="2" pageNumber="43">Philippines</collectingCountry>
had 1719 tines. A smaller number of tines was found in two females and males from the southern coast of
<collectingCountry id="D6D69C8CFFCDFF97FD1272867D16FD19" box="[755,876,724,750]" name="Sri Lanka" pageId="2" pageNumber="43">Sri Lanka</collectingCountry>
(unpublished record, material from Ahangama, from
<taxonomicName id="69C1A79FFFCDFF97FE3B72AE7C86FCE1" authority="(Chlorophyta)" baseAuthorityName="Chlorophyta" box="[474,764,764,790]" class="Bryopsidophyceae" family="Halimedaceae" genus="Halimeda" higherTaxonomySource="GBIF" kingdom="Plantae" order="Bryopsidales" pageId="2" pageNumber="43" phylum="Chlorophyta" rank="genus">
<emphasis id="9CB5000EFFCDFF97FE3B72AE7C37FCE2" box="[474,589,764,789]" italics="true" pageId="2" pageNumber="43">Halimeda</emphasis>
(Chlorophyta)
</taxonomicName>
on a moderately exposed fringing reef flat,
<date id="DA7FFADCFFCDFF97FAEC72AE7FC3FCC9" pageId="2" pageNumber="43" value="1993-02-28">28 February 1993</date>
, coll. H.­G. Müller); claws I and II of these specimens bear 1314 tines, claws
<date id="DA7FFADCFFCDFF97FEB4731E7F82FC91" box="[341,504,844,870]" pageId="2" pageNumber="43" value="1915-03" valueMax="1915-04-31" valueMin="1915-03-15">III and IV 15</date>
tines. Specimens from
<collectingCountry id="D6D69C8CFFCDFF97FCF8731E7D2FFC91" box="[793,853,844,870]" name="India" pageId="2" pageNumber="43">India</collectingCountry>
and eastern Africa (
<collectingCountry id="D6D69C8CFFCDFF97FBAE731E7AE5FC91" box="[1103,1183,844,870]" name="Kenya" pageId="2" pageNumber="43">Kenya</collectingCountry>
) bear more than 17 tines (
<bibRefCitation id="CA50A1EDFFCDFF97FE5473267C05FC79" author="Chatterjee" box="[437,639,884,910]" pageId="2" pageNumber="49" refString="Chatterjee, T. (1995) Record of three species of Rhombognathus (Halacaridae: Acari) from Indian Ocean Region. Journal of the Bombay Natural History Society, 92, 282 - 286." type="journal article" year="1995">Chatterjee, 1995</bibRefCitation>
;
<bibRefCitation id="CA50A1EDFFCDFF97FD6C73267D89FC79" author="Chatterjee" box="[653,1011,884,910]" pageId="2" pageNumber="49" refString="Chatterjee, T. &amp; De Troch, M. (2000) Halacaridae (Acari) from Gazi Bay (Kenya): description and biogeography of three new and two known species. Hydrobiologia, 427, 177 - 194." type="journal article" year="2000">Chatterjee &amp; De Troch, 2000</bibRefCitation>
). An increased number of tines may be correlated with life in very agitated water.
</paragraph>
<paragraph id="AE7EDC1CFFCDFF97FED973967DC9FB51" blockId="2.[264,1324,284,1190]" pageId="2" pageNumber="43">
The telofemora in the specimens from
<collectingCountry id="D6D69C8CFFCDFF97FCCC73967DDDFC29" box="[813,935,964,990]" name="Singapore" pageId="2" pageNumber="43">Singapore</collectingCountry>
, as well as in those from the
<collectingCountry id="D6D69C8CFFCDFF97FEE973BE7FF5FBF1" box="[264,399,1004,1030]" name="Philippines" pageId="2" pageNumber="43">Philippines</collectingCountry>
, are slender, 2.52.9 times longer than high, whereas in specimens from western and eastern
<collectingCountry id="D6D69C8CFFCDFF97FDFB74467CFCFBD9" box="[538,646,1044,1070]" name="Australia" pageId="2" pageNumber="43">Australia</collectingCountry>
(Rottnest Island, Dampier and Great Barrier Reef, respectively) the length:height ratio ranged from 2.1 to 2.5. Specimens from
<collectingCountry id="D6D69C8CFFCDFF97FB66746E7A83FBA1" box="[1159,1273,1084,1110]" name="Sri Lanka" pageId="2" pageNumber="43">Sri Lanka</collectingCountry>
had short telofemora, about twice as long as high. A similar wide range in the length:height ratio is else not documented in the genus
<taxonomicName id="69C1A79FFFCDFF97FD0C74DE7DD1FB52" box="[749,939,1164,1189]" class="Arachnida" family="Halacaridae" genus="Rhombognathus" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="9CB5000EFFCDFF97FD0C74DE7DD1FB52" box="[749,939,1164,1189]" italics="true" pageId="2" pageNumber="43">Rhombognathus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E6DB8F97FFCDFF97FEE9748E7FE4FA61" pageId="2" pageNumber="43" type="distribution">
<paragraph id="AE7EDC1CFFCDFF97FEE9748E7C56FB02" blockId="2.[264,1323,1244,1430]" box="[264,556,1244,1269]" pageId="2" pageNumber="43">
<emphasis id="9CB5000EFFCDFF97FEE9748E7C56FB02" box="[264,556,1244,1269]" italics="true" pageId="2" pageNumber="43">Distribution and Biology</emphasis>
</paragraph>
<paragraph id="AE7EDC1CFFCDFF97FED975567FE4FA61" blockId="2.[264,1323,1244,1430]" pageId="2" pageNumber="43">
<taxonomicName id="69C1A79FFFCDFF97FED975567C0CFAEA" box="[312,630,1284,1309]" class="Arachnida" family="Halacaridae" genus="Rhombognathus" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="scutulatus">
<emphasis id="9CB5000EFFCDFF97FED975567C0CFAEA" box="[312,630,1284,1309]" italics="true" pageId="2" pageNumber="43">Rhombognathus scutulatus</emphasis>
</taxonomicName>
is spread throughout the Indo­West Pacific area, from the
<collectingCountry id="D6D69C8CFFCDFF97FEE9757E7FF1FAB1" box="[264,395,1324,1350]" name="Philippines" pageId="2" pageNumber="43">Philippines</collectingCountry>
in the east to
<collectingCountry id="D6D69C8CFFCDFF97FDCD757E7C02FAB1" box="[556,632,1324,1350]" name="Kenya" pageId="2" pageNumber="43">Kenya</collectingCountry>
in the west. The species lives in a variety of mid­tide, lowtide and shallow sublittoral habitats, on shores exposed to moderate as well as rather strong swell.
</paragraph>
</subSubSection>
</treatment>
</document>