158 lines
17 KiB
XML
158 lines
17 KiB
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<document id="8948A98C8784B35684797CEBA7F5E486" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE343FFD6FF75F950BC32FDB8" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena bisinuata Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="72" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="69" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:mods id="945D975BB08AB57D66BB9DDDB1088D90" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="13BFBC0A9592198AAB43F089171C722F">
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<mods:title id="8EDBC8511099E0CA0BE25809395491FC">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="586937264B90B8DFE68B0E000A83A7F9" type="personal">
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<mods:roleTerm id="2C4050269444A8C829D520FA62BEF537">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2ACFF9EA355673626204D3BC19811DD6">Perkins, Philip D.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F5864F6DA85D01C415E01795DBD95759">text</mods:typeOfResource>
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<mods:titleInfo id="F61E963AC926A25A7B2014281792123B">
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<mods:title id="D3C86FF29F38BF36AFA6D752CB40B875">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="10BACF7A4ABA6746A154476BEB585AE6">
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<mods:date id="12098F2E98E1A067CF28FCF83EE2F0DD">2017</mods:date>
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<mods:detail id="89FE38B0D6BE0BDAE541CFDF6B0C79CB" type="pubDate">
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<mods:number id="06E4E4F86EF6A13A4320263C9CFF35C2">2017-11-03</mods:number>
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</mods:detail>
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<mods:detail id="86792FEB10A127977D949B9F698B296C" type="volume">
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<mods:number id="C3AE218832CFE03E0A24C8543B48CFB3">4342</mods:number>
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</mods:detail>
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<mods:detail id="468E73D50458FDF777E6897467F6367A" type="issue">
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<mods:number id="98818A105BF052C25815AC038E2E4717">1</mods:number>
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</mods:detail>
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<mods:start id="9ADC8C4AF770BF4D13375C147D366BDF">1</mods:start>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="44998C91C06486CD0BF363A3AC8FE32A">journal article</mods:classification>
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<mods:identifier id="53FA76D2128354665BCB6603A68CDD48" type="CLB-Dataset">31639</mods:identifier>
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<mods:identifier id="0C51BB9E6A515173E034F94094E89FD1" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<mods:identifier id="E0796696754AEF6CA3D2C2D6F352A478" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
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<mods:identifier id="182300CAA69CF44DBB74F3E3828DEAFB" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2E169EDBA05422D8F8FD13A30A72996B" type="Zenodo-Dep">1041066</mods:identifier>
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<mods:identifier id="6BD945E9AE052489645E26B5D4C8E193" type="ZooBank">2ACD54D2-3487-432D-9323-EEC131FE2E64</mods:identifier>
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</mods:mods>
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<treatment id="038587BBE343FFD6FF75F950BC32FDB8" ID-DOI="http://doi.org/10.5281/zenodo.5323534" ID-GBIF-Taxon="137233466" ID-Zenodo-Dep="5323534" LSID="urn:lsid:plazi:treatment:038587BBE343FFD6FF75F950BC32FDB8" httpUri="http://treatment.plazi.org/id/038587BBE343FFD6FF75F950BC32FDB8" lastPageId="71" lastPageNumber="72" pageId="68" pageNumber="69">
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<subSubSection id="C3366526E343FFD5FF75F950BBB4F8B5" pageId="68" pageNumber="69" type="nomenclature">
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<paragraph id="8B9336ADE343FFD5FF75F950BB19F88A" blockId="68.[151,648,1781,1840]" box="[151,549,1781,1807]" pageId="68" pageNumber="69">
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<heading id="D0DB81C1E343FFD5FF75F950BB19F88A" bold="true" box="[151,549,1781,1807]" fontSize="11" level="1" pageId="68" pageNumber="69" reason="1">
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<emphasis id="B958EABFE343FFD5FF75F950BB19F88A" bold="true" box="[151,549,1781,1807]" pageId="68" pageNumber="69">
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<taxonomicName id="4C2C4D2EE343FFD5FF75F950B8BAF88A" ID-CoL="3N3LJ" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,390,1781,1807]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="bisinuata" status="sp. nov.">
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<emphasis id="B958EABFE343FFD5FF75F950B8BAF88A" bold="true" box="[151,390,1781,1807]" italics="true" pageId="68" pageNumber="69">Hydraena bisinuata</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26B57C4E343FFD5FE77F950BB19F88A" box="[405,549,1781,1807]" pageId="68" pageNumber="69" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336ADE343FFD5FF75F8B2BBB4F8B5" blockId="68.[151,648,1781,1840]" box="[151,648,1815,1840]" pageId="68" pageNumber="69">
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<figureCitation id="13172A28E343FFD5FF75F8B2B9CDF8B5" box="[151,241,1815,1840]" captionStart="FIGURE 58" captionStartId="69.[151,250,2009,2031]" captionText="FIGURE 58. Hydraena bisinuata, holotype, habitus dorsal, lateral, and ventral." pageId="68" pageNumber="69">Figs. 58</figureCitation>
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(habitus), 60 (aedeagus), 255 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E343FFD5FF75F8FABAA6F845" pageId="68" pageNumber="69" type="materials_examined">
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<paragraph id="8B9336ADE343FFD5FF75F8FABAA6F845" blockId="68.[151,1436,1887,2020]" pageId="68" pageNumber="69">
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<emphasis id="B958EABFE343FFD5FF75F8FAB8F7F8FD" bold="true" box="[151,459,1887,1912]" pageId="68" pageNumber="69">
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<typeStatus id="5497880FE343FFD5FF75F8FAB9EEF8FD" box="[151,210,1887,1912]" pageId="68" pageNumber="69">Type</typeStatus>
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Material.
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<typeStatus id="5497880FE343FFD5FEBCF8FAB8F7F8FD" box="[350,459,1887,1912]" pageId="68" pageNumber="69" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):
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<collectingRegion id="49E8F84FE343FFD5FDDBF8FABB87F8FD" box="[569,699,1887,1912]" country="Madagascar" name="Toamasina" pageId="68" pageNumber="69">
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<emphasis id="B958EABFE343FFD5FDDBF8FABB87F8FD" bold="true" box="[569,699,1887,1912]" pageId="68" pageNumber="69">Toamasina</emphasis>
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</collectingRegion>
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, Tamatave
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<quantity id="4CD49B48E343FFD5FCAEF8FABA9DF8FD" box="[844,929,1887,1912]" metricMagnitude="3" metricUnit="m" metricValue="6.3" pageId="68" pageNumber="69" unit="km" value="6.3">6.3 km</quantity>
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S Ambanizona, Andranobe Field Station, rainforest, sifted litter from flat sandy areas, some from slope, elev.
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<quantity id="4CD49B48E343FFD5FC7FF826BAE6F81E" box="[925,986,1923,1948]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="68" pageNumber="69" unit="m" value="25.0">25 m</quantity>
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,
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<geoCoordinate id="EE18506AE343FFD5FC05F826BD6DF819" box="[999,1105,1923,1948]" direction="south" orientation="latitude" pageId="68" pageNumber="69" precision="925" value="-15.683333">15° 41' S</geoCoordinate>
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,
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<geoCoordinate id="EE18506AE343FFD5FBBCF821BDF7F81E" box="[1118,1227,1923,1948]" direction="east" orientation="longitude" pageId="68" pageNumber="69" precision="925" value="49.95">49° 57' E</geoCoordinate>
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,
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<date id="FF92106DE343FFD5FB3AF821BC69F819" box="[1240,1365,1923,1948]" pageId="68" pageNumber="69" value="1993-11-14">14 xi 1993</date>
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, B. L. Fisher (#886) (
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<collectionCode id="ED3DAE68E343FFD5FEDFF80DB8A8F845" box="[317,404,1960,1984]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="68" pageNumber="69">FMNH</collectionCode>
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).
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<typeStatus id="5497880FE343FFD5FE45F80DBB31F845" box="[423,525,1960,1984]" pageId="68" pageNumber="69" type="paratype">Paratype</typeStatus>
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: Same data as
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<typeStatus id="5497880FE343FFD5FD53F802BA2FF845" box="[689,787,1959,1984]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus>
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(1
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<collectionCode id="ED3DAE68E343FFD5FCDAF80DBAB2F845" box="[824,910,1960,1984]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="68" pageNumber="69">FMNH</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E343FFD4FF25F86EBB4FFF51" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69" type="diagnosis">
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<paragraph id="8B9336ADE343FFD4FF25F86EBB4FFF51" blockId="68.[151,1436,1887,2020]" lastBlockId="69.[151,1436,151,789]" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
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<emphasis id="B958EABFE343FFD5FF25F86EB8EAF861" bold="true" box="[199,470,1995,2020]" pageId="68" pageNumber="69">Differential Diagnosis.</emphasis>
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Similar to
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<taxonomicName id="4C2C4D2EE343FFD5FDBFF868BA31F861" authorityName="Perkins" authorityYear="2017" box="[605,781,1995,2020]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="rubrifurcata">
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<emphasis id="B958EABFE343FFD5FDBFF868BA31F861" box="[605,781,1995,2020]" italics="true" pageId="68" pageNumber="69">H. rubrifurcata</emphasis>
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</taxonomicName>
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in size, habitus, and reddish brown color; differentiated therefrom by the flat, non-carinate metaventral plaques and the less strongly punctate dorsum. The aedeagi of the two species markedly differ (
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<figureCitation id="13172A28E342FFD4FE3FFF19BB00FF50" box="[477,572,188,213]" captionStart="FIGURES 59 – 60" captionStartId="70.[151,264,1876,1899]" captionText="FIGURES 59 – 60. Hydraena, aedeagi of holotypes." pageId="69" pageNumber="70">Figs. 60</figureCitation>
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,
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<figureCitation id="13172A28E342FFD4FDA5FF19BB5AFF51" box="[583,614,188,212]" captionStart="FIGURES 71 – 72" captionStartId="82.[151,264,1998,2021]" captionText="FIGURES 71 – 72. Hydraena, aedeagi of holotypes." pageId="69" pageNumber="70">71</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E342FFD6FF25FF7ABAAFFE55" lastPageId="71" lastPageNumber="72" pageId="69" pageNumber="70" type="description">
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<paragraph id="8B9336ADE342FFD4FF25FF7ABA37FE99" blockId="69.[151,1436,151,789]" pageId="69" pageNumber="70">
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<emphasis id="B958EABFE342FFD4FF25FF7AB86DFF7D" bold="true" box="[199,337,223,248]" pageId="69" pageNumber="70">Description</emphasis>
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. Size:
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<typeStatus id="5497880FE342FFD4FE41FF7ABB39FF7D" box="[419,517,223,248]" pageId="69" pageNumber="70" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.79/0.77; head width 0.44; pronotum 0.40/0.57, PA 0.47, PB 0.48; elytra 1.10/0.77.
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</paragraph>
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<paragraph id="8B9336ADE342FFD4FF75FE8DBBDBFE0D" blockId="69.[151,1436,151,789]" pageId="69" pageNumber="70">Dorsum general coloration dark reddish brown; head and pronotal disc darker, almost piceous; anterior border of pronotum very slightly lighter (when teneral pronotal fascia is more apparent); legs dark brown; maxillary palpi reddish brown, distal ½ of last palpomere not darker.</paragraph>
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<paragraph id="8B9336ADE342FFD4FF25FE31BAD6FDB8" blockId="69.[151,1436,151,789]" pageId="69" pageNumber="70">Head with clypeus finely moderately densely punctate; frons slightly more coarsely punctate laterally than medially where interstices ca. 1–2xpd, each puncture with short distinctive seta. Pronotum in anterior and posterior 1/3 slightly more densely punctate than on disc, interstices ca. 0.5–1xpd; punctures on disc smaller, interstices ca. 1–1.5xpd, shining. Mentum moderately coarsely, sparsely punctate, interstices shining; postmentum finely densely punctate. Genae raised, shining, with posterior ridge, interrupted in midline.</paragraph>
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<paragraph id="8B9336ADE342FFD4FF25FDEDBA71FD00" blockId="69.[151,1436,151,789]" pageId="69" pageNumber="70">Pronotum bisinuate; anterior margin straight behind eyes, emarginate behind frons; PF1 and PF2 absent; PF3 shallow; PF4 very shallow; PF3 and PF4 shallowly confluent.</paragraph>
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<paragraph id="8B9336ADE342FFD4FF25FD2ABDA6FC90" blockId="69.[151,1436,151,789]" pageId="69" pageNumber="70">Elytra weakly arcuate laterally; summit of moderately steep posterior declivity at ca. midlength; lateral explanate margins wide; serial punctures small, round, separated by ca. 1xpd or less, slightly smaller than largest pronotal punctures. Intervals ca. 1xpd; discal intervals weakly raised, shining. Apices in dorsal aspect rather sharply conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
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<caption id="DF536625E342FFD4FF75F87CBAFCF86A" box="[151,960,2009,2031]" pageId="69" pageNumber="70">
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<paragraph id="8B9336ADE342FFD4FF75F87CBAFCF86A" blockId="69.[151,960,2009,2031]" box="[151,960,2009,2031]" pageId="69" pageNumber="70">
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<emphasis id="B958EABFE342FFD4FF75F87CB81CF86B" bold="true" box="[151,288,2009,2031]" pageId="69" pageNumber="70">FIGURE 58.</emphasis>
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<taxonomicName id="4C2C4D2EE342FFD4FEC7F87CB8D3F86A" authorityName="Perkins" authorityYear="2017" box="[293,495,2009,2031]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="bisinuata">
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<emphasis id="B958EABFE342FFD4FEC7F87CB8D3F86A" box="[293,495,2009,2031]" italics="true" pageId="69" pageNumber="70">Hydraena bisinuata</emphasis>
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</taxonomicName>
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, holotype, habitus dorsal, lateral, and ventral.
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</paragraph>
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</caption>
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<caption id="DF536625E341FFD7FF75F8F1BB98F8EE" box="[151,676,1876,1899]" pageId="70" pageNumber="71">
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<paragraph id="8B9336ADE341FFD7FF75F8F1BB98F8EE" blockId="70.[151,676,1876,1899]" box="[151,676,1876,1899]" pageId="70" pageNumber="71">
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<emphasis id="B958EABFE341FFD7FF75F8F1B868F8EF" bold="true" box="[151,340,1876,1899]" pageId="70" pageNumber="71">FIGURES 59–60.</emphasis>
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<taxonomicName id="4C2C4D2EE341FFD7FEB8F8F0B883F8EE" box="[346,447,1877,1899]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE341FFD7FEB8F8F0B883F8EE" box="[346,447,1877,1899]" italics="true" pageId="70" pageNumber="71">Hydraena</emphasis>
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</taxonomicName>
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, aedeagi of holotypes.
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</paragraph>
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</caption>
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<paragraph id="8B9336ADE340FFD6FF25FF32BAAFFE55" blockId="71.[151,1437,151,573]" pageId="71" pageNumber="72">Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/3/0.75. P1 almost as wide as P2; median carina very weakly sinuate in profile. P2 l/w ca.2/1, sides parallel, apex blunt and on same level as anteromedian carina of metaventrite. Plaques roundly raised, converging but not contiguous anteriorly, separated basally by slightly less than plaque width, located at sides of median depression. Metaventrite anteromedian carina shorter than plaques, lengths as 3 to 5, joining anterior level of plaques, together forming inverted Y-shape; long and strong, slightly arcuate longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at arcuate posterior margin ca. 1.5x P2. Protibia straight, slender. Mesotibia very slightly arcuate, medial margin with 3 or 4 short sharp spines, surface of tibia notched between spines. Metatibia straight, slender. Metafemur moderately slender. Abdominal apex with small apicomedian notch.</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E340FFD6FF25FE79BA63FE71" box="[199,863,476,501]" pageId="71" pageNumber="72" type="etymology">
|
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|
|
<paragraph id="8B9336ADE340FFD6FF25FE79BA63FE71" blockId="71.[151,1437,151,573]" box="[199,863,476,501]" pageId="71" pageNumber="72">
|
|||
|
|
<emphasis id="B958EABFE340FFD6FF25FE79B872FE70" bold="true" box="[199,334,476,501]" pageId="71" pageNumber="72">Etymology.</emphasis>
|
|||
|
|
Named in reference to the bisinuate pronotum.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3366526E340FFD6FF25FE5ABC32FDB8" pageId="71" pageNumber="72" type="discussion">
|
|||
|
|
<paragraph id="8B9336ADE340FFD6FF25FE5ABC32FDB8" blockId="71.[151,1437,151,573]" pageId="71" pageNumber="72">
|
|||
|
|
<emphasis id="B958EABFE340FFD6FF25FE5AB807FD9D" bold="true" box="[199,315,511,536]" pageId="71" pageNumber="72">Remarks.</emphasis>
|
|||
|
|
The
|
|||
|
|
<typeStatus id="5497880FE340FFD6FE98FE5AB8E0FD9D" box="[378,476,511,536]" pageId="71" pageNumber="72" type="holotype">holotype</typeStatus>
|
|||
|
|
is slightly teneral; the pronotal fascia is not as apparent in mature specimens. The aedeagal flagellum illustrated (
|
|||
|
|
<figureCitation id="13172A28E340FFD6FE13FD81BB7FFDB8" box="[497,579,548,573]" captionStart="FIGURES 59 – 60" captionStartId="70.[151,264,1876,1899]" captionText="FIGURES 59 – 60. Hydraena, aedeagi of holotypes." pageId="71" pageNumber="72">Fig. 60</figureCitation>
|
|||
|
|
) is from a
|
|||
|
|
<typeStatus id="5497880FE340FFD6FD5FFD80BA1DFDB8" box="[701,801,549,573]" pageId="71" pageNumber="72" type="paratype">paratype</typeStatus>
|
|||
|
|
; the flagellum of the
|
|||
|
|
<typeStatus id="5497880FE340FFD6FBEDFD81BD4DFDB8" box="[1039,1137,548,573]" pageId="71" pageNumber="72" type="holotype">holotype</typeStatus>
|
|||
|
|
is broken off.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|