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<document id="E200F4B73A470B971A8CE1FDE30321C3" ID-DOI="10.11646/zootaxa.4044.2.2" ID-GBIF-Dataset="8b09a6a1-35c3-407c-a4b5-076c4af030db" ID-ISSN="1175-5326" ID-Zenodo-Dep="241241" ID-ZooBank="7000893A-A8BC-4196-B76D-385C97B96FBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461338858631" checkinUser="plazi" docAuthor="Kanao, Taisuke &amp; Maruyama, Munetoshi" docDate="2015" docId="0395F739FFADBF2DFF021944FCE4BB42" docLanguage="en" docName="zt04044p223.pdf" docOrigin="Zootaxa 4044 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Discoxenus assmuthi" docType="treatment" docVersion="11" lastPageNumber="209" masterDocId="FFAC8F41FFA5BF25FF951C5CFFC1BC65" masterDocTitle="Eight new species, a new record, and redescription of the genus Discoxenus Wasmann, 1904: The first record of termitophilous rove beetles in Cambodia (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="223" masterPageNumber="201" pageNumber="209" updateTime="1698619795159" updateUser="plazi">
<mods:mods id="D957792901A8DCA9C9F08F97F9B9E2ED" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C6AED6084A5A662F49A5F5F1E5428857">Eight new species, a new record, and redescription of the genus Discoxenus Wasmann, 1904: The first record of termitophilous rove beetles in Cambodia (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="700BFD80101F192F768B3D7EF308E364">Kanao, Taisuke</mods:namePart>
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<mods:name id="8C5D83C96899FA59F4E73F1967F69ACA" type="personal">
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<mods:namePart id="4D16727A9BDE4C2543DC74E70241E2CE">Maruyama, Munetoshi</mods:namePart>
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<mods:title id="1E89C02FACF285FCBAC47D52FFFB09B5">Zootaxa</mods:title>
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<mods:part id="075BF06C7B077C63F4C37E1FDEB0398A">
<mods:date id="5E4029071B14672C4080384DD89C96A1">2015</mods:date>
<mods:detail id="8E2895E1EE7EB62B36AC32B93E615F99" type="volume">
<mods:number id="65565A1B70F7CE4071E5E433CCB8F29B">4044</mods:number>
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<mods:classification id="657F934EC8638CCDB21BE4989D30BE85">journal article</mods:classification>
<mods:identifier id="8F0732B50A85E9F317194A2D429CB561" type="DOI">10.11646/zootaxa.4044.2.2</mods:identifier>
<mods:identifier id="3A469359C14DBC176777DD94A3D25C43" type="GBIF-Dataset">8b09a6a1-35c3-407c-a4b5-076c4af030db</mods:identifier>
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<treatment id="0395F739FFADBF2DFF021944FCE4BB42" ID-DOI="http://doi.org/10.5281/zenodo.6111779" ID-GBIF-Taxon="119644724" ID-Zenodo-Dep="6111779" LSID="urn:lsid:plazi:treatment:0395F739FFADBF2DFF021944FCE4BB42" httpUri="http://treatment.plazi.org/id/0395F739FFADBF2DFF021944FCE4BB42" lastPageNumber="209" pageId="8" pageNumber="209">
<subSubSection id="C32615A4FFADBF2DFF021944FE2AB957" box="[151,491,1303,1330]" pageId="8" pageNumber="209" type="nomenclature">
<paragraph id="8B83462FFFADBF2DFF021944FE2AB957" blockId="8.[151,491,1303,1330]" box="[151,491,1303,1330]" pageId="8" pageNumber="209">
<heading id="D0CBF143FFADBF2DFF021944FE2AB957" bold="true" box="[151,491,1303,1330]" fontSize="11" level="1" pageId="8" pageNumber="209" reason="1">
<emphasis id="B9489A3DFFADBF2DFF021944FE2AB957" bold="true" box="[151,491,1303,1330]" pageId="8" pageNumber="209">
The
<taxonomicName id="4C3C3DACFFADBF2DFF5A194BFEFDB954" ID-CoL="6CZCF" box="[207,316,1303,1329]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="assmuthi">
<emphasis id="B9489A3DFFADBF2DFF5A194BFEFDB954" bold="true" box="[207,316,1303,1329]" italics="true" pageId="8" pageNumber="209">assmuthi</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFADBF2DFF021902FEB9BA2A" pageId="8" pageNumber="209" type="diagnosis">
<paragraph id="8B83462FFFADBF2DFF021902FEB9BA2A" blockId="8.[151,1437,1374,1831]" pageId="8" pageNumber="209">
<emphasis id="B9489A3DFFADBF2DFF021902FED0B912" bold="true" box="[151,273,1374,1399]" pageId="8" pageNumber="209">Diagnosis.</emphasis>
The
<taxonomicName id="4C3C3DACFFADBF2DFED91902FE71B913" box="[332,432,1374,1398]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="assmuthi">
<emphasis id="B9489A3DFFADBF2DFED91902FE71B913" box="[332,432,1374,1398]" italics="true" pageId="8" pageNumber="209">assmuthi</emphasis>
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group is defined by the following character states: 1) mandible (
<figureCitation id="13075AAAFFADBF2DFB1B1902FAD6B912" box="[1166,1303,1374,1399]" captionStart="FIGURES 54 – 72" captionStartId="9.[151,264,1536,1559]" captionTargetBox="[157,1426,198,1505]" captionTargetId="figure@9.[151,1436,193,1515]" captionTargetPageId="9" captionText="FIGURES 54 – 72. Discoxenus phourini n. sp.: 54. Head, 55. Antenna, 56. Labrum (left side = dorsal view, right side = ventral view), 57. Mandible, left, 58. Mandible, right, 59. Maxilla, 60. Mentum, 61. Labium, 62. Pronotum, 63. Elytron, right, 64. Meso- and metaventrites, 65. Tergite VIII, 66. Sternite VIII, male, 67. Sternite VIII, female, 68. Tergites IX and X, 69. Median lobe of aedeagus, ventral view, 70. Median lobe of aedeagus, lateral view, 71. Paramere, 72. Spermatheca." httpUri="https://zenodo.org/record/241247/files/figure.png" pageId="8" pageNumber="209">Figs. 57, 58</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFAB61902FAADB913" box="[1315,1388,1374,1399]" captionStart="FIGURES 73 – 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 – 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="8" pageNumber="209">76, 77</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFAEC1903FF74B9FE" captionStart="FIGURES 89 – 104" captionStartId="13.[151,264,1283,1306]" captionTargetBox="[162,1429,201,1252]" captionTargetId="figure@13.[151,1436,193,1262]" captionTargetPageId="13" captionText="FIGURES 89 – 104. Discoxenus hirsutus n. sp.: 89. Head, 90. Antenna, 91. Labrum (left side = dorsal view, right side = ventral view), 92. Mandible, left, 93. Mandible, right, 94. Mentum, 95. Pronotum, 96. Elytron, right, 97. Tergite VIII, 98. Sternite VIII, male, 99. Sternite VIII, female, 100. Tergites IX and X. 101. Median lobe of aedeagus, ventral view, 102. Median lobe of aedeagus, lateral view, 103. Paramere, 104. Spermatheca." httpUri="https://zenodo.org/record/241249/files/figure.png" pageId="8" pageNumber="209">92, 93</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFF5119DFFEEBB9FE" box="[196,298,1411,1435]" captionStart="FIGURES 105 – 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 – 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="8" pageNumber="209">108, 109</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFEAC19DFFE61B9FE" box="[313,416,1410,1435]" captionStart="FIGURES 121 – 136" captionStartId="16.[151,264,1630,1653]" captionTargetBox="[168,1426,489,1599]" captionTargetId="figure@16.[151,1436,481,1609]" captionTargetPageId="16" captionText="FIGURES 121 – 136. Discoxenus lucidus n. sp.: 121. Head, 122. Antenna, 123. Labrum (left side = dorsal view, right side = ventral view), 124. Mandible, left, 125. Mandible, right, 126. Mentum, 127. Pronotum, 128. Elytron, right, 129. Tergite VIII, 130. Sternite VIII, male, 131. Sternite VIII, female, 132. Tergites IX and X, 133. Median lobe of aedeagus, ventral view, 134. Median lobe of aedeagus, lateral view, 135. Paramere, 136. Spermatheca." httpUri="https://zenodo.org/record/241251/files/figure.png" pageId="8" pageNumber="209">124, 125</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFE3B19DFFDD4B9FE" box="[430,533,1411,1435]" captionStart="FIGURES 137 – 148" captionStartId="18.[151,264,1794,1817]" captionTargetBox="[176,1420,892,1760]" captionTargetId="figure@18.[151,1436,879,1772]" captionTargetPageId="18" captionText="FIGURES 137 – 148. Discoxenus kakizoei n. sp.: 137. Head, 138. Antenna, 139. Labrum (left side = dorsal view, right side = ventral view), 140. Mandible, left, 141. Mandible, right, 142. Mentum, 143. Pronotum, 144. Elytron, right, 145. Tergite VIII, 146. Sternite VIII, 147. Tergites IX and X, 148. Spermatheca." httpUri="https://zenodo.org/record/241252/files/figure.png" pageId="8" pageNumber="209">140, 141</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFDB119DEFD48B9FE" box="[548,649,1410,1435]" captionStart="FIGURES 149 – 164" captionStartId="20.[151,264,1496,1519]" captionTargetBox="[157,1431,367,1462]" captionTargetId="figure@20.[151,1436,357,1491]" captionTargetPageId="20" captionText="FIGURES 149 – 164. Discoxenus katayamai: 149. Head, 150. Antenna, 151. Labrum (left side = dorsal view, right side = ventral view), 152. Mandible, left, 153. Mandible, right, 154. Mentum, 155. Pronotum, 156. Elytron, right, 157. Tergite VIII, 158. Sternite VIII, male, 159. Sternite VIII, female, 160. Tergites IX and X, 161. Median lobe of aedeagus, ventral view, 162. Median lobe of aedeagus, lateral view, 163. Paramere, 164. Spermatheca." httpUri="https://zenodo.org/record/241253/files/figure.png" pageId="8" pageNumber="209">152, 153</figureCitation>
) covered with 15–40 pores, 2) maxillary basistipes (
<figureCitation id="13075AAAFFADBF2DFB7E19DEFA81B9FE" box="[1259,1344,1410,1435]" captionStart="FIGURES 54 – 72" captionStartId="9.[151,264,1536,1559]" captionTargetBox="[157,1426,198,1505]" captionTargetId="figure@9.[151,1436,193,1515]" captionTargetPageId="9" captionText="FIGURES 54 – 72. Discoxenus phourini n. sp.: 54. Head, 55. Antenna, 56. Labrum (left side = dorsal view, right side = ventral view), 57. Mandible, left, 58. Mandible, right, 59. Maxilla, 60. Mentum, 61. Labium, 62. Pronotum, 63. Elytron, right, 64. Meso- and metaventrites, 65. Tergite VIII, 66. Sternite VIII, male, 67. Sternite VIII, female, 68. Tergites IX and X, 69. Median lobe of aedeagus, ventral view, 70. Median lobe of aedeagus, lateral view, 71. Paramere, 72. Spermatheca." httpUri="https://zenodo.org/record/241247/files/figure.png" pageId="8" pageNumber="209">Fig. 59</figureCitation>
) almost fused with palpifer, 3) sternite VIII (
<figureCitation id="13075AAAFFADBF2DFDD319FAFD14B9DA" box="[582,725,1446,1471]" captionStart="FIGURES 54 – 72" captionStartId="9.[151,264,1536,1559]" captionTargetBox="[157,1426,198,1505]" captionTargetId="figure@9.[151,1436,193,1515]" captionTargetPageId="9" captionText="FIGURES 54 – 72. Discoxenus phourini n. sp.: 54. Head, 55. Antenna, 56. Labrum (left side = dorsal view, right side = ventral view), 57. Mandible, left, 58. Mandible, right, 59. Maxilla, 60. Mentum, 61. Labium, 62. Pronotum, 63. Elytron, right, 64. Meso- and metaventrites, 65. Tergite VIII, 66. Sternite VIII, male, 67. Sternite VIII, female, 68. Tergites IX and X, 69. Median lobe of aedeagus, ventral view, 70. Median lobe of aedeagus, lateral view, 71. Paramere, 72. Spermatheca." httpUri="https://zenodo.org/record/241247/files/figure.png" pageId="8" pageNumber="209">Figs. 66, 67</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFD7019FBFCF0B9DA" box="[741,817,1447,1471]" captionStart="FIGURES 73 – 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 – 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="8" pageNumber="209">82, 83</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFCD419FBFC4CB9DA" box="[833,909,1447,1471]" captionStart="FIGURES 89 – 104" captionStartId="13.[151,264,1283,1306]" captionTargetBox="[162,1429,201,1252]" captionTargetId="figure@13.[151,1436,193,1262]" captionTargetPageId="13" captionText="FIGURES 89 – 104. Discoxenus hirsutus n. sp.: 89. Head, 90. Antenna, 91. Labrum (left side = dorsal view, right side = ventral view), 92. Mandible, left, 93. Mandible, right, 94. Mentum, 95. Pronotum, 96. Elytron, right, 97. Tergite VIII, 98. Sternite VIII, male, 99. Sternite VIII, female, 100. Tergites IX and X. 101. Median lobe of aedeagus, ventral view, 102. Median lobe of aedeagus, lateral view, 103. Paramere, 104. Spermatheca." httpUri="https://zenodo.org/record/241249/files/figure.png" pageId="8" pageNumber="209">98, 99</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFC0819FBFBC3B9DA" box="[925,1026,1446,1471]" captionStart="FIGURES 105 – 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 – 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="8" pageNumber="209">114, 115</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFB8719FBFBBBB9DA" box="[1042,1146,1447,1471]" captionStart="FIGURES 121 – 136" captionStartId="16.[151,264,1630,1653]" captionTargetBox="[168,1426,489,1599]" captionTargetId="figure@16.[151,1436,481,1609]" captionTargetPageId="16" captionText="FIGURES 121 – 136. Discoxenus lucidus n. sp.: 121. Head, 122. Antenna, 123. Labrum (left side = dorsal view, right side = ventral view), 124. Mandible, left, 125. Mandible, right, 126. Mentum, 127. Pronotum, 128. Elytron, right, 129. Tergite VIII, 130. Sternite VIII, male, 131. Sternite VIII, female, 132. Tergites IX and X, 133. Median lobe of aedeagus, ventral view, 134. Median lobe of aedeagus, lateral view, 135. Paramere, 136. Spermatheca." httpUri="https://zenodo.org/record/241251/files/figure.png" pageId="8" pageNumber="209">130, 131</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFB1F19FAFB77B9DA" box="[1162,1206,1446,1471]" captionStart="FIGURES 137 – 148" captionStartId="18.[151,264,1794,1817]" captionTargetBox="[176,1420,892,1760]" captionTargetId="figure@18.[151,1436,879,1772]" captionTargetPageId="18" captionText="FIGURES 137 – 148. Discoxenus kakizoei n. sp.: 137. Head, 138. Antenna, 139. Labrum (left side = dorsal view, right side = ventral view), 140. Mandible, left, 141. Mandible, right, 142. Mentum, 143. Pronotum, 144. Elytron, right, 145. Tergite VIII, 146. Sternite VIII, 147. Tergites IX and X, 148. Spermatheca." httpUri="https://zenodo.org/record/241252/files/figure.png" pageId="8" pageNumber="209">146</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFB5319FAFAECB9DA" box="[1222,1325,1446,1471]" captionStart="FIGURES 149 – 164" captionStartId="20.[151,264,1496,1519]" captionTargetBox="[157,1431,367,1462]" captionTargetId="figure@20.[151,1436,357,1491]" captionTargetPageId="20" captionText="FIGURES 149 – 164. Discoxenus katayamai: 149. Head, 150. Antenna, 151. Labrum (left side = dorsal view, right side = ventral view), 152. Mandible, left, 153. Mandible, right, 154. Mentum, 155. Pronotum, 156. Elytron, right, 157. Tergite VIII, 158. Sternite VIII, male, 159. Sternite VIII, female, 160. Tergites IX and X, 161. Median lobe of aedeagus, ventral view, 162. Median lobe of aedeagus, lateral view, 163. Paramere, 164. Spermatheca." httpUri="https://zenodo.org/record/241253/files/figure.png" pageId="8" pageNumber="209">158, 159</figureCitation>
) sparsely covered with minute pores anteriorly, 4) tergite X (
<figureCitation id="13075AAAFFADBF2DFD611996FC96B986" box="[756,855,1482,1507]" captionStart="FIGURES 54 – 72" captionStartId="9.[151,264,1536,1559]" captionTargetBox="[157,1426,198,1505]" captionTargetId="figure@9.[151,1436,193,1515]" captionTargetPageId="9" captionText="FIGURES 54 – 72. Discoxenus phourini n. sp.: 54. Head, 55. Antenna, 56. Labrum (left side = dorsal view, right side = ventral view), 57. Mandible, left, 58. Mandible, right, 59. Maxilla, 60. Mentum, 61. Labium, 62. Pronotum, 63. Elytron, right, 64. Meso- and metaventrites, 65. Tergite VIII, 66. Sternite VIII, male, 67. Sternite VIII, female, 68. Tergites IX and X, 69. Median lobe of aedeagus, ventral view, 70. Median lobe of aedeagus, lateral view, 71. Paramere, 72. Spermatheca." httpUri="https://zenodo.org/record/241247/files/figure.png" pageId="8" pageNumber="209">Figs. 68</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFCF31997FC44B986" box="[870,901,1483,1507]" captionStart="FIGURES 73 – 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 – 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="8" pageNumber="209">84</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFC001997FC00B986" box="[917,961,1483,1507]" captionStart="FIGURES 89 – 104" captionStartId="13.[151,264,1283,1306]" captionTargetBox="[162,1429,201,1252]" captionTargetId="figure@13.[151,1436,193,1262]" captionTargetPageId="13" captionText="FIGURES 89 – 104. Discoxenus hirsutus n. sp.: 89. Head, 90. Antenna, 91. Labrum (left side = dorsal view, right side = ventral view), 92. Mandible, left, 93. Mandible, right, 94. Mentum, 95. Pronotum, 96. Elytron, right, 97. Tergite VIII, 98. Sternite VIII, male, 99. Sternite VIII, female, 100. Tergites IX and X. 101. Median lobe of aedeagus, ventral view, 102. Median lobe of aedeagus, lateral view, 103. Paramere, 104. Spermatheca." httpUri="https://zenodo.org/record/241249/files/figure.png" pageId="8" pageNumber="209">100</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFC441996FC3CB986" box="[977,1021,1482,1507]" captionStart="FIGURES 105 – 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 – 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="8" pageNumber="209">116</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFB981997FBF8B986" box="[1037,1081,1483,1507]" captionStart="FIGURES 121 – 136" captionStartId="16.[151,264,1630,1653]" captionTargetBox="[168,1426,489,1599]" captionTargetId="figure@16.[151,1436,481,1609]" captionTargetPageId="16" captionText="FIGURES 121 – 136. Discoxenus lucidus n. sp.: 121. Head, 122. Antenna, 123. Labrum (left side = dorsal view, right side = ventral view), 124. Mandible, left, 125. Mandible, right, 126. Mentum, 127. Pronotum, 128. Elytron, right, 129. Tergite VIII, 130. Sternite VIII, male, 131. Sternite VIII, female, 132. Tergites IX and X, 133. Median lobe of aedeagus, ventral view, 134. Median lobe of aedeagus, lateral view, 135. Paramere, 136. Spermatheca." httpUri="https://zenodo.org/record/241251/files/figure.png" pageId="8" pageNumber="209">132</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFBDC1997FBB4B986" box="[1097,1141,1483,1507]" captionStart="FIGURES 137 – 148" captionStartId="18.[151,264,1794,1817]" captionTargetBox="[176,1420,892,1760]" captionTargetId="figure@18.[151,1436,879,1772]" captionTargetPageId="18" captionText="FIGURES 137 – 148. Discoxenus kakizoei n. sp.: 137. Head, 138. Antenna, 139. Labrum (left side = dorsal view, right side = ventral view), 140. Mandible, left, 141. Mandible, right, 142. Mentum, 143. Pronotum, 144. Elytron, right, 145. Tergite VIII, 146. Sternite VIII, 147. Tergites IX and X, 148. Spermatheca." httpUri="https://zenodo.org/record/241252/files/figure.png" pageId="8" pageNumber="209">147</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFB131996FB70B986" box="[1158,1201,1482,1507]" captionStart="FIGURES 149 – 164" captionStartId="20.[151,264,1496,1519]" captionTargetBox="[157,1431,367,1462]" captionTargetId="figure@20.[151,1436,357,1491]" captionTargetPageId="20" captionText="FIGURES 149 – 164. Discoxenus katayamai: 149. Head, 150. Antenna, 151. Labrum (left side = dorsal view, right side = ventral view), 152. Mandible, left, 153. Mandible, right, 154. Mentum, 155. Pronotum, 156. Elytron, right, 157. Tergite VIII, 158. Sternite VIII, male, 159. Sternite VIII, female, 160. Tergites IX and X, 161. Median lobe of aedeagus, ventral view, 162. Median lobe of aedeagus, lateral view, 163. Paramere, 164. Spermatheca." httpUri="https://zenodo.org/record/241253/files/figure.png" pageId="8" pageNumber="209">160</figureCitation>
) with posterior half covered with a batch of setae, 5) median lobe of male aedeagus with basal capsule small (Figs, 69, 85, 101, 117, 133, 161), and 6) basal capsule of male aedeagal median lobe with distal crest strongly developed (
<figureCitation id="13075AAAFFADBF2DFA991A4EFAADBA4E" box="[1292,1388,1554,1579]" captionStart="FIGURES 54 – 72" captionStartId="9.[151,264,1536,1559]" captionTargetBox="[157,1426,198,1505]" captionTargetId="figure@9.[151,1436,193,1515]" captionTargetPageId="9" captionText="FIGURES 54 – 72. Discoxenus phourini n. sp.: 54. Head, 55. Antenna, 56. Labrum (left side = dorsal view, right side = ventral view), 57. Mandible, left, 58. Mandible, right, 59. Maxilla, 60. Mentum, 61. Labium, 62. Pronotum, 63. Elytron, right, 64. Meso- and metaventrites, 65. Tergite VIII, 66. Sternite VIII, male, 67. Sternite VIII, female, 68. Tergites IX and X, 69. Median lobe of aedeagus, ventral view, 70. Median lobe of aedeagus, lateral view, 71. Paramere, 72. Spermatheca." httpUri="https://zenodo.org/record/241247/files/figure.png" pageId="8" pageNumber="209">Figs. 70</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFAEC1A4EFA56BA4E" box="[1401,1431,1554,1579]" captionStart="FIGURES 73 – 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 – 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="8" pageNumber="209">86</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFF021A6BFF02BA2A" box="[151,195,1591,1615]" captionStart="FIGURES 89 – 104" captionStartId="13.[151,264,1283,1306]" captionTargetBox="[162,1429,201,1252]" captionTargetId="figure@13.[151,1436,193,1262]" captionTargetPageId="13" captionText="FIGURES 89 – 104. Discoxenus hirsutus n. sp.: 89. Head, 90. Antenna, 91. Labrum (left side = dorsal view, right side = ventral view), 92. Mandible, left, 93. Mandible, right, 94. Mentum, 95. Pronotum, 96. Elytron, right, 97. Tergite VIII, 98. Sternite VIII, male, 99. Sternite VIII, female, 100. Tergites IX and X. 101. Median lobe of aedeagus, ventral view, 102. Median lobe of aedeagus, lateral view, 103. Paramere, 104. Spermatheca." httpUri="https://zenodo.org/record/241249/files/figure.png" pageId="8" pageNumber="209">102</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFF5A1A6BFF3BBA2A" box="[207,250,1591,1615]" captionStart="FIGURES 105 – 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 – 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="8" pageNumber="209">118</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFE931A6BFEF3BA2A" box="[262,306,1591,1615]" captionStart="FIGURES 121 – 136" captionStartId="16.[151,264,1630,1653]" captionTargetBox="[168,1426,489,1599]" captionTargetId="figure@16.[151,1436,481,1609]" captionTargetPageId="16" captionText="FIGURES 121 – 136. Discoxenus lucidus n. sp.: 121. Head, 122. Antenna, 123. Labrum (left side = dorsal view, right side = ventral view), 124. Mandible, left, 125. Mandible, right, 126. Mentum, 127. Pronotum, 128. Elytron, right, 129. Tergite VIII, 130. Sternite VIII, male, 131. Sternite VIII, female, 132. Tergites IX and X, 133. Median lobe of aedeagus, ventral view, 134. Median lobe of aedeagus, lateral view, 135. Paramere, 136. Spermatheca." httpUri="https://zenodo.org/record/241251/files/figure.png" pageId="8" pageNumber="209">134</figureCitation>
,
<figureCitation id="13075AAAFFADBF2DFEAB1A6AFEADBA2A" box="[318,364,1590,1615]" captionStart="FIGURES 149 – 164" captionStartId="20.[151,264,1496,1519]" captionTargetBox="[157,1431,367,1462]" captionTargetId="figure@20.[151,1436,357,1491]" captionTargetPageId="20" captionText="FIGURES 149 – 164. Discoxenus katayamai: 149. Head, 150. Antenna, 151. Labrum (left side = dorsal view, right side = ventral view), 152. Mandible, left, 153. Mandible, right, 154. Mentum, 155. Pronotum, 156. Elytron, right, 157. Tergite VIII, 158. Sternite VIII, male, 159. Sternite VIII, female, 160. Tergites IX and X, 161. Median lobe of aedeagus, ventral view, 162. Median lobe of aedeagus, lateral view, 163. Paramere, 164. Spermatheca." httpUri="https://zenodo.org/record/241253/files/figure.png" pageId="8" pageNumber="209">162</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFADBF2DFF521A06FCE4BB42" pageId="8" pageNumber="209" type="discussion">
<paragraph id="8B83462FFFADBF2DFF521A06FB2BBADE" blockId="8.[151,1437,1374,1831]" pageId="8" pageNumber="209">
<emphasis id="B9489A3DFFADBF2DFF521A06FE8EBA16" bold="true" box="[199,335,1626,1651]" pageId="8" pageNumber="209">Comments.</emphasis>
The
<taxonomicName id="4C3C3DACFFADBF2DFE031A07FE3BBA16" box="[406,506,1627,1651]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="assmuthi">
<emphasis id="B9489A3DFFADBF2DFE031A07FE3BBA16" box="[406,506,1627,1651]" italics="true" pageId="8" pageNumber="209">assmuthi</emphasis>
</taxonomicName>
species group includes 11 species:
<taxonomicName id="4C3C3DACFFADBF2DFC241A07FAC8BA16" authority="Wasmann, 1904" authorityName="Wasmann" authorityYear="1904" box="[945,1289,1627,1651]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="assmuthi">
<emphasis id="B9489A3DFFADBF2DFC241A07FBFFBA16" box="[945,1086,1627,1651]" italics="true" pageId="8" pageNumber="209">D. assmuthi</emphasis>
<bibRefCitation id="EFAD3BDEFFADBF2DFBD91A07FAC8BA16" author="Wasmann" box="[1100,1289,1627,1651]" pageId="8" pageNumber="209" refString="Wasmann, E. (1904) Zur Kenntnis der Gaste der Treiberameisen und ihrer Wirthe am obern Congo nach den Sammlungen und Beobachyungen. Zoologische Jahrbucher, 7, 611 - 682." type="journal article" year="1904">Wasmann, 1904</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C3C3DACFFADBF2DFA891A07FE8FBAF2" authority="Wasmann, 1904" authorityName="Wasmann" authorityYear="1904" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="lepisma">
<emphasis id="B9489A3DFFADBF2DFA891A07FA5DBA16" box="[1308,1436,1627,1651]" italics="true" pageId="8" pageNumber="209">D. lepisma</emphasis>
<bibRefCitation id="EFAD3BDEFFADBF2DFF021A23FE8FBAF2" author="Wasmann" box="[151,334,1663,1687]" pageId="8" pageNumber="209" refString="Wasmann, E. (1904) Zur Kenntnis der Gaste der Treiberameisen und ihrer Wirthe am obern Congo nach den Sammlungen und Beobachyungen. Zoologische Jahrbucher, 7, 611 - 682." type="journal article" year="1904">Wasmann, 1904</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C3C3DACFFADBF2DFECC1A23FDAEBAF2" authority="Kistner, 1982" authorityName="Kistner" authorityYear="1982" box="[345,623,1662,1687]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9489A3DFFADBF2DFECC1A23FE0CBAF3" box="[345,461,1662,1686]" italics="true" pageId="8" pageNumber="209">D. indicus</emphasis>
<bibRefCitation id="EFAD3BDEFFADBF2DFE411A22FDAEBAF2" author="Kistner" box="[468,623,1662,1687]" pageId="8" pageNumber="209" refString="Kistner, D. H. (1982) A revision of the termitophilous genus Discoxenus with a study of the relationships of the genus and notes on its behavior (Coleoptera, Staphylinidae). Sociobiology, 7 (2), 165 - 186." type="journal article" year="1982">Kistner, 1982</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C3C3DACFFADBF2DFD3E1A23FC3EBAF2" authority="Kistner, 1982" authorityName="Kistner" authorityYear="1982" box="[683,1023,1662,1687]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="malaysiensis">
<emphasis id="B9489A3DFFADBF2DFD3E1A23FC9DBAF3" box="[683,860,1662,1686]" italics="true" pageId="8" pageNumber="209">D. malaysiensis</emphasis>
<bibRefCitation id="EFAD3BDEFFADBF2DFCF11A22FC3EBAF2" author="Kistner" box="[868,1023,1662,1687]" pageId="8" pageNumber="209" refString="Kistner, D. H. (1982) A revision of the termitophilous genus Discoxenus with a study of the relationships of the genus and notes on its behavior (Coleoptera, Staphylinidae). Sociobiology, 7 (2), 165 - 186." type="journal article" year="1982">Kistner, 1982</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C3C3DACFFADBF2DFB9F1A23FB4DBAF3" box="[1034,1164,1662,1686]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="phourini" status="sp. nov.">
<emphasis id="B9489A3DFFADBF2DFB9F1A23FBDFBAF3" box="[1034,1054,1663,1686]" italics="true" pageId="8" pageNumber="209">D</emphasis>
.
<emphasis id="B9489A3DFFADBF2DFBB91A22FB4DBAF3" box="[1068,1164,1662,1686]" italics="true" pageId="8" pageNumber="209">phourini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFADBF2DFB011A22FB15BAF2" box="[1172,1236,1662,1687]" pageId="8" pageNumber="209" rank="species">
<emphasis id="B9489A3DFFADBF2DFB011A22FB15BAF2" bold="true" box="[1172,1236,1662,1687]" pageId="8" pageNumber="209">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C3C3DACFFADBF2DFB771A23FA5DBAF3" box="[1250,1436,1662,1686]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="kohkongensis" status="sp. nov.">
<emphasis id="B9489A3DFFADBF2DFB771A23FB37BAF3" box="[1250,1270,1663,1686]" italics="true" pageId="8" pageNumber="209">D</emphasis>
.
<emphasis id="B9489A3DFFADBF2DFA901A22FA5DBAF3" box="[1285,1436,1662,1686]" italics="true" pageId="8" pageNumber="209">kohkongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFADBF2DFF021AFEFF14BADE" box="[151,213,1698,1723]" pageId="8" pageNumber="209" rank="species">
<emphasis id="B9489A3DFFADBF2DFF021AFEFF14BADE" bold="true" box="[151,213,1698,1723]" pageId="8" pageNumber="209">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C3C3DACFFADBF2DFF711AFFFEA1BADE" box="[228,352,1699,1723]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B9489A3DFFADBF2DFF711AFFFF39BADF" box="[228,248,1699,1722]" italics="true" pageId="8" pageNumber="209">D</emphasis>
.
<emphasis id="B9489A3DFFADBF2DFE931AFFFEA1BADE" box="[262,352,1699,1723]" italics="true" pageId="8" pageNumber="209">hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFADBF2DFEF21AFEFE67BADE" box="[359,422,1698,1723]" pageId="8" pageNumber="209" rank="species">
<emphasis id="B9489A3DFFADBF2DFEF21AFEFE67BADE" bold="true" box="[359,422,1698,1723]" pageId="8" pageNumber="209">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C3C3DACFFADBF2DFE211AFFFDEEBADF" box="[436,559,1699,1722]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B9489A3DFFADBF2DFE211AFFFE09BADF" box="[436,456,1699,1722]" italics="true" pageId="8" pageNumber="209">D</emphasis>
.
<emphasis id="B9489A3DFFADBF2DFE431AFFFDEEBADF" box="[470,559,1699,1722]" italics="true" pageId="8" pageNumber="209">minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFADBF2DFDA31AFEFDB4BADE" box="[566,629,1698,1723]" pageId="8" pageNumber="209" rank="species">
<emphasis id="B9489A3DFFADBF2DFDA31AFEFDB4BADE" bold="true" box="[566,629,1698,1723]" pageId="8" pageNumber="209">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C3C3DACFFADBF2DFD161AFFFD37BADE" box="[643,758,1699,1723]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="lucidus" status="sp. nov.">
<emphasis id="B9489A3DFFADBF2DFD161AFFFD56BADF" box="[643,663,1699,1722]" italics="true" pageId="8" pageNumber="209">D</emphasis>
.
<emphasis id="B9489A3DFFADBF2DFD301AFFFD37BADE" box="[677,758,1699,1723]" italics="true" pageId="8" pageNumber="209">lucidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFADBF2DFD681AFEFCFDBADE" box="[765,828,1698,1723]" pageId="8" pageNumber="209" rank="species">
<emphasis id="B9489A3DFFADBF2DFD681AFEFCFDBADE" bold="true" box="[765,828,1698,1723]" pageId="8" pageNumber="209">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C3C3DACFFADBF2DFCDF1AFFFC09BADE" box="[842,968,1699,1723]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="kakizoei" status="sp. nov.">
<emphasis id="B9489A3DFFADBF2DFCDF1AFFFC9FBADF" box="[842,862,1699,1722]" italics="true" pageId="8" pageNumber="209">D</emphasis>
.
<emphasis id="B9489A3DFFADBF2DFCF91AFFFC09BADE" box="[876,968,1699,1723]" italics="true" pageId="8" pageNumber="209">kakizoei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFADBF2DFC5A1AFEFBCFBADE" box="[975,1038,1698,1723]" pageId="8" pageNumber="209" rank="species">
<emphasis id="B9489A3DFFADBF2DFC5A1AFEFBCFBADE" bold="true" box="[975,1038,1698,1723]" pageId="8" pageNumber="209">n. sp.</emphasis>
</taxonomicNameLabel>
, and
<taxonomicName id="4C3C3DACFFADBF2DFBD91AFFFB26BADE" box="[1100,1255,1699,1723]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="katayamai">
<emphasis id="B9489A3DFFADBF2DFBD91AFFFBA1BADF" box="[1100,1120,1699,1722]" italics="true" pageId="8" pageNumber="209">D</emphasis>
.
<emphasis id="B9489A3DFFADBF2DFBFB1AFFFB26BADE" box="[1134,1255,1699,1723]" italics="true" pageId="8" pageNumber="209">katayamai</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B83462FFFADBF2DFF521A9AFCE4BB42" blockId="8.[151,1437,1374,1831]" pageId="8" pageNumber="209">
The affiliation of the four previously known species,
<taxonomicName id="4C3C3DACFFADBF2DFC8C1A9BFC5EBABB" box="[793,927,1734,1758]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="assmuthi">
<emphasis id="B9489A3DFFADBF2DFC8C1A9BFC5EBABB" box="[793,927,1734,1758]" italics="true" pageId="8" pageNumber="209">D. assmuthi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3C3DACFFADBF2DFC381A9BFBE7BABB" box="[941,1062,1734,1758]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="lepisma">
<emphasis id="B9489A3DFFADBF2DFC381A9BFBE7BABB" box="[941,1062,1734,1758]" italics="true" pageId="8" pageNumber="209">D. lepisma</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3C3DACFFADBF2DFBA11A9BFB67BABB" box="[1076,1190,1734,1758]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9489A3DFFADBF2DFBA11A9BFB67BABB" box="[1076,1190,1734,1758]" italics="true" pageId="8" pageNumber="209">D. indicus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C3C3DACFFADBF2DFB711A9BFA54BABB" box="[1252,1429,1734,1758]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="malaysiensis">
<emphasis id="B9489A3DFFADBF2DFB711A9BFA54BABB" box="[1252,1429,1734,1758]" italics="true" pageId="8" pageNumber="209">D. malaysiensis</emphasis>
</taxonomicName>
, are determined based on the character states of their mouthparts and/or male aedeagus (
<bibRefCitation id="EFAD3BDEFFADBF2DFBF41AB6FB36BB66" author="Kistner" box="[1121,1271,1770,1795]" pageId="8" pageNumber="209" refString="Kistner, D. H. (1982) A revision of the termitophilous genus Discoxenus with a study of the relationships of the genus and notes on its behavior (Coleoptera, Staphylinidae). Sociobiology, 7 (2), 165 - 186." type="journal article" year="1982">Kistner 1982</bibRefCitation>
), and expected to share other character states typical of the species group.
</paragraph>
</subSubSection>
</treatment>
</document>
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