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<document id="384E5AFEE1B2EB1D677355C310F88BCF" ID="10.11646/zootaxa.4121.5.3" ID-CLB-Dataset="38882" ID-DOI="10.11646/zootaxa.4121.5.3" ID-GBIF-Dataset="f1dde1c0-56b8-4563-b185-051098ee4a5f" ID-ISSN="1175-5326" ID-Zenodo-Dep="256720" ID-ZooBank="6BD94D27-3820-40E2-A00E-68E32248395A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465799157841" checkinUser="plazi" docAuthor="White, William T. &amp; Last, Peter R." docDate="2016" docId="AD6887FBB97BFFB78EA18CED2C083FE2" docLanguage="en" docName="zootaxa.4121.5.3.pdf" docOrigin="Zootaxa 4121 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Platyrhina psomadakisi White &amp; Last, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="543" masterDocId="5151FF83B97AFFBD8E368C7A29443E16" masterDocTitle="Platyrhina psomadakisi sp. nov., a new species of fanray (Batoidea: Platyrhinidae) from the Andaman Sea, the first record of this family in the Indian Ocean" masterLastPageNumber="544" masterPageNumber="533" pageNumber="534" updateTime="1698662825037" updateUser="ExternalLinkService">
<mods:mods id="0629834FD2D585B1448A7C0D14F43B5C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="74F421DD0ABC820A1A388D70478ECA7E">Platyrhina psomadakisi sp. nov., a new species of fanray (Batoidea: Platyrhinidae) from the Andaman Sea, the first record of this family in the Indian Ocean</mods:title>
</mods:titleInfo>
<mods:name id="4194986F64F90A152AA20266CF1C1D4A" type="personal">
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<mods:namePart id="404482C1A93C8E46C1BA614F96929361">White, William T.</mods:namePart>
</mods:name>
<mods:name id="F87E3EDB8303DA6A9A5B5C731F4F0D7E" type="personal">
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<mods:namePart id="7E2207C391B4A07EC6A7F287DA26F515">Last, Peter R.</mods:namePart>
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<mods:title id="69FCCFC9AF885281E0C7ADB45C40BFD1">Zootaxa</mods:title>
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<mods:part id="F27825833EC137E2143242E1992E2290">
<mods:date id="465BC18A184F0622A6FA514E06A7C180">2016</mods:date>
<mods:detail id="DF87C92F53CA4E5B621B4540B66A7770" type="volume">
<mods:number id="B356969DCA050B828D75A304A86F1D29">4121</mods:number>
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<mods:classification id="299ACF5C9526EFCBD9510B683BBDC667">journal article</mods:classification>
<mods:identifier id="B33CEF3DB64CFB286DAF91EE1A5FB048" type="CLB-Dataset">38882</mods:identifier>
<mods:identifier id="8FF089957B0D5A6C1276E0B82C62F3AD" type="DOI">10.11646/zootaxa.4121.5.3</mods:identifier>
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<treatment id="AD6887FBB97BFFB78EA18CED2C083FE2" ID-DOI="http://doi.org/10.5281/zenodo.5663008" ID-GBIF-Taxon="120827794" ID-Zenodo-Dep="5663008" LSID="urn:lsid:plazi:treatment:AD6887FBB97BFFB78EA18CED2C083FE2" httpUri="http://treatment.plazi.org/id/AD6887FBB97BFFB78EA18CED2C083FE2" lastPageId="10" lastPageNumber="543" pageId="1" pageNumber="534">
<subSubSection id="6DDB6566B97BFFBC8EA18CED28373EE1" pageId="1" pageNumber="534" type="nomenclature">
<paragraph id="257E36EDB97BFFBC8EA18CED2B5E3EA4" blockId="1.[151,538,151,247]" box="[151,538,151,178]" pageId="1" pageNumber="534">
<heading id="7E368181B97BFFBC8EA18CED2B5E3EA4" bold="true" box="[151,538,151,178]" fontSize="11" level="1" pageId="1" pageNumber="534" reason="1">
<emphasis id="17B5EAFFB97BFFBC8EA18CED2B5E3EA4" bold="true" box="[151,538,151,178]" pageId="1" pageNumber="534">
<taxonomicName id="E2C14D6EB97BFFBC8EA18CED28F13EA7" box="[151,437,151,177]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="1" pageNumber="534" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB97BFFBC8EA18CED28F13EA7" bold="true" box="[151,437,151,177]" italics="true" pageId="1" pageNumber="534">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0C865784B97BFFBC8F8A8CE22B5E3EA4" box="[444,538,152,178]" pageId="1" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="257E36EDB97BFFBC8EA18CC0281C3EC5" blockId="1.[151,538,151,247]" box="[151,344,186,211]" pageId="1" pageNumber="534">Andaman Fanray</paragraph>
<paragraph id="257E36EDB97BFFBC8EA18CA428373EE1" blockId="1.[151,538,151,247]" box="[151,371,222,247]" pageId="1" pageNumber="534">
(
<figureCitation id="BDFA2A68B97BFFBC8EA98CA4284F3EE1" box="[159,267,222,247]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="1.[151,250,1967,1989]" captionStartId-1="2.[151,250,1681,1703]" captionStartId-2="3.[151,250,1777,1799]" captionStartId-3="4.[151,250,1539,1561]" captionStartId-4="5.[151,250,1234,1256]" captionStartId-5="5.[151,250,1983,2005]" captionStartId-6="6.[151,250,1746,1768]" captionStartId-7="7.[151,250,1950,1972]" captionTargetBox-0="[157,1428,600,1924]" captionTargetBox-1="[156,1430,209,1620]" captionTargetBox-2="[287,1300,534,1735]" captionTargetBox-3="[341,1249,316,1499]" captionTargetBox-4="[256,1318,440,1190]" captionTargetBox-5="[212,1347,1381,1924]" captionTargetBox-6="[341,1249,204,1694]" captionTargetBox-7="[341,1248,711,1903]" captionTargetId-0="figure@1.[151,1436,563,1945]" captionTargetId-1="figure@2.[151,1436,193,1659]" captionTargetId-2="figure@3.[267,1320,517,1756]" captionTargetId-3="figure@4.[317,1269,301,1518]" captionTargetId-4="figure@5.[235,1352,425,1212]" captionTargetId-5="figure@5.[190,1396,1336,1962]" captionTargetId-6="figure@6.[313,1273,193,1725]" captionTargetId-7="figure@7.[324,1264,697,1929]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="5" captionTargetPageId-6="6" captionTargetPageId-7="7" captionText-0="FIGURE 1. Holotype of Platyrhina psomadakisi sp. nov. (CSIRO H 7861 - 01, adult male 378 mm TL, preserved): (A) dorsal view; (B) ventral view." captionText-1="FIGURE 2. Paratype of Platyrhina psomadakisi sp. nov. (CSIRO H 7859 - 01, female 341 mm TL, fresh): (A) dorsal view; (B) ventral view." captionText-2="FIGURE 3. Dorsal view of juvenile paratype of Platyrhina psomadakisi sp. nov. (CSIRO H 7860 - 01, juvenile male 197 mm TL, preserved)." captionText-3="FIGURE 4. Oronasal region of Platyrhina psomadakisi sp. nov.: (A) holotype (CSIRO H 7861 - 01, adult male 378 mm TL, preserved); (B) paratype (CSIRO H 7859 - 01, female 341 mm TL, preserved)." captionText-4="FIGURE 5. Dorsal view of tail region of Platyrhina psomadakisi sp. nov.: (A) holotype (CSIRO H 7861 - 01, adult male 378 mm TL, preserved); (B) paratype (CSIRO H 7859 - 01, female 341 mm TL, preserved)." captionText-5="FIGURE 6. Lateral view of thorns on midline of head and disc of Platyrhina psomadakisi sp. nov.: (A) holotype (CSIRO H 7861 - 01, adult male 378 mm TL, preserved); (B) paratype (CSIRO H 7859 - 01, female 341 mm TL, preserved)." captionText-6="FIGURE 7. Dorsal view of orbital and scapular region of Platyrhina psomadakisi sp. nov.: (A) holotype (CSIRO H 7861 - 01, adult male 378 mm TL, preserved); (B) paratype (CSIRO H 7859 - 01, female 341 mm TL, preserved)." captionText-7="FIGURE 8. Dorsolateral view of the left orbit and spiracle of Platyrhina psomadakisi sp. nov.: (A) holotype (CSIRO H 7861 - 0 1, adult male 378 mm TL, preserved); (B) paratype (CSIRO H 7859 - 01, female 341 mm TL, preserved)." httpUri-0="https://zenodo.org/record/256721/files/figure.png" httpUri-1="https://zenodo.org/record/256722/files/figure.png" httpUri-2="https://zenodo.org/record/256723/files/figure.png" httpUri-3="https://zenodo.org/record/256724/files/figure.png" httpUri-4="https://zenodo.org/record/256725/files/figure.png" httpUri-5="https://zenodo.org/record/256726/files/figure.png" httpUri-6="https://zenodo.org/record/256727/files/figure.png" httpUri-7="https://zenodo.org/record/256728/files/figure.png" pageId="1" pageNumber="534">Figs. 1–8</figureCitation>
;
<tableCitation id="68430356B97BFFBC8F218CA4282C3EE1" box="[279,360,222,247]" captionStart="TABLE 1" captionStartId="8.[151,239,368,391]" captionText="TABLE 1. Morphometric data for the holotype of Platyrhina psomadakisi sp. nov. and ranges for the three paratypes; expressed as percentages of total length." pageId="1" pageNumber="534">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6DDB6566B97BFFBC8EA18D5C2DF13C01" pageId="1" pageNumber="534" type="materials_examined">
<paragraph id="257E36EDB97BFFBC8EA18D5C2A213F75" blockId="1.[151,1437,294,535]" pageId="1" pageNumber="534">
<emphasis id="17B5EAFFB97BFFBC8EA18D5C284F3F29" bold="true" box="[151,267,294,319]" pageId="1" pageNumber="534">
<typeStatus id="FA7A884FB97BFFBC8EA18D5C28423F29" box="[151,262,294,319]" pageId="1" pageNumber="534" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="43D0AE28B97BFFBC8F2E8D5D282F3F29" box="[280,363,295,319]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="1" pageNumber="534">CSIRO</collectionCode>
H 7861-01, adult male
<quantity id="E2399B08B97BFFBC8CA38D5D2BBC3F28" box="[661,760,295,319]" metricMagnitude="-1" metricUnit="m" metricValue="3.78" pageId="1" pageNumber="534" unit="mm" value="378.0">378 mm</quantity>
TL, west of Lord Loughborough Island, Andaman Sea,
<collectingCountry id="5DD6767DB97BFFBC8EA18D3128423F75" box="[151,262,331,355]" name="Myanmar" pageId="1" pageNumber="534">Myanmar</collectingCountry>
,
<geoCoordinate id="40F5502AB97BFFBC8F278D3028E03F74" box="[273,420,330,355]" direction="north" orientation="latitude" pageId="1" pageNumber="534" precision="15" value="10.349444">10°20’58” N</geoCoordinate>
,
<geoCoordinate id="40F5502AB97BFFBC8F998D302B753F74" box="[431,561,330,355]" direction="east" orientation="longitude" pageId="1" pageNumber="534" precision="15" value="97.76889">97°46’8” E</geoCoordinate>
,
<quantity id="E2399B08B97BFFBC8C0A8D302B313F74" box="[572,629,330,355]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="1" pageNumber="534" unit="m" value="69.0">69 m</quantity>
depth,
<date id="517F102DB97BFFBC8CFE8D312A253F75" box="[712,865,330,355]" pageId="1" pageNumber="534" value="2015-05-29">29 May 2015</date>
.
</paragraph>
<paragraph id="257E36EDB97BFFBC8EF18D142DF13C01" blockId="1.[151,1437,294,535]" pageId="1" pageNumber="534">
<emphasis id="17B5EAFFB97BFFBC8EF18D1428023F91" bold="true" box="[199,326,366,391]" pageId="1" pageNumber="534">
<typeStatus id="FA7A884FB97BFFBC8EF18D1428053F91" box="[199,321,366,391]" pageId="1" pageNumber="534" type="paratype">Paratypes</typeStatus>
.
</emphasis>
(
<specimenCount id="33C7FD64B97BFFBC8F6B8D1528B73F91" box="[349,499,366,391]" pageId="1" pageNumber="534" type="generic">3 specimens</specimenCount>
).
<collectionCode id="43D0AE28B97BFFBC8C3A8D152B1B3F91" box="[524,607,367,391]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="1" pageNumber="534">CSIRO</collectionCode>
H 7859-01, female
<quantity id="E2399B08B97BFFBC8D6D8D152AFB3F90" box="[859,959,367,391]" metricMagnitude="-1" metricUnit="m" metricValue="3.41" pageId="1" pageNumber="534" unit="mm" value="341.0">341 mm</quantity>
TL, northern Andaman Sea,
<collectingCountry id="5DD6767DB97BFFBC8B1F8D152CDC3F91" box="[1321,1432,367,391]" name="Myanmar" pageId="1" pageNumber="534">Myanmar</collectingCountry>
,
<geoCoordinate id="40F5502AB97BFFBC8EA18DE8286F3FBC" box="[151,299,402,427]" direction="north" orientation="latitude" pageId="1" pageNumber="534" precision="15" value="14.932222">14°55’56” N</geoCoordinate>
,
<geoCoordinate id="40F5502AB97BFFBC8F018DE828833FBC" box="[311,455,402,427]" direction="east" orientation="longitude" pageId="1" pageNumber="534" precision="15" value="94.48055">94°28’50” E</geoCoordinate>
,
<quantity id="E2399B08B97BFFBC8FE28DE82B493FBC" box="[468,525,402,427]" metricMagnitude="1" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="534" unit="m" value="58.0">58 m</quantity>
depth,
<date id="517F102DB97BFFBC8C548DE92BBF3FBD" box="[610,763,402,427]" pageId="1" pageNumber="534" value="2015-05-11">11 May 2015</date>
;
<collectionCode id="43D0AE28B97BFFBC8D3E8DE92A1E3FBD" box="[776,858,403,427]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="1" pageNumber="534">CSIRO</collectionCode>
H 7860-01, juvenile male
<quantity id="E2399B08B97BFFBC8ABD8DE92DAD3FBC" box="[1163,1257,403,427]" metricMagnitude="-1" metricUnit="m" metricValue="1.97" pageId="1" pageNumber="534" unit="mm" value="197.0">197 mm</quantity>
TL, west of Sir Charles Metcalfe Island, Andaman Sea,
<collectingCountry id="5DD6767DB97BFFBC8C4E8DCD2BA33FD9" box="[632,743,439,463]" name="Myanmar" pageId="1" pageNumber="534">Myanmar</collectingCountry>
,
<geoCoordinate id="40F5502AB97BFFBC8CC18DCD2AD43FD8" box="[759,912,439,463]" direction="north" orientation="latitude" pageId="1" pageNumber="534" precision="15" value="12.3752775">12°22’31” N</geoCoordinate>
,
<geoCoordinate id="40F5502AB97BFFBC8D978DCD2D723FD8" box="[929,1078,439,463]" direction="east" orientation="longitude" pageId="1" pageNumber="534" precision="15" value="97.027504">97°01’39” E</geoCoordinate>
,
<quantity id="E2399B08B97BFFBC8A7E8DCC2DD13FD8" box="[1096,1173,438,463]" metricMagnitude="2" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="534" unit="m" value="158.0">158 m</quantity>
depth,
<date id="517F102DB97BFFBC8AC28DCD2CD33FD9" box="[1268,1431,438,463]" pageId="1" pageNumber="534" value="2015-05-23">23 May 2015</date>
; Department of Fisheries, Yangon (specimen 604), female
<quantity id="E2399B08B97BFFBC8D718DA02AEE3FE4" box="[839,938,474,499]" metricMagnitude="-1" metricUnit="m" metricValue="2.45" pageId="1" pageNumber="534" unit="mm" value="245.0">245 mm</quantity>
TL, west of Sir Charles Metcalfe Island, Andaman Sea,
<collectingCountry id="5DD6767DB97BFFBC8F778D8528F43C01" box="[321,432,511,535]" name="Myanmar" pageId="1" pageNumber="534">Myanmar</collectingCountry>
, station 137,
<geoCoordinate id="40F5502AB97BFFBC8C738D852B9C3C00" box="[581,728,511,535]" direction="north" orientation="latitude" pageId="1" pageNumber="534" precision="15" value="12.3752775">12°22’31” N</geoCoordinate>
,
<geoCoordinate id="40F5502AB97BFFBC8CD58D852A373C00" box="[739,883,511,535]" direction="east" orientation="longitude" pageId="1" pageNumber="534" precision="15" value="97.027504">97°01’39” E</geoCoordinate>
,
<quantity id="E2399B08B97BFFBC8D488D842A813C00" box="[894,965,510,535]" metricMagnitude="2" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="534" unit="m" value="158.0">158 m</quantity>
depth,
<date id="517F102DB97BFFBC8A2E8D852DF53C01" box="[1048,1201,510,535]" pageId="1" pageNumber="534" value="2015-05-23">23 May 2015</date>
.
</paragraph>
</subSubSection>
<caption id="71BE6665B97BFFBC8EA18BD528C539F5" httpUri="https://zenodo.org/record/256721/files/figure.png" pageId="1" pageNumber="534" targetBox="[157,1428,600,1924]" targetPageId="1">
<paragraph id="257E36EDB97BFFBC8EA18BD528C539F5" blockId="1.[151,1436,1967,2019]" pageId="1" pageNumber="534">
<emphasis id="17B5EAFFB97BFFBC8EA18BD5285739D3" bold="true" box="[151,275,1967,1989]" pageId="1" pageNumber="534">FIGURE 1.</emphasis>
Holotype of
<taxonomicName id="E2C14D6EB97BFFBC8FAA8BD52BC839D2" box="[412,652,1967,1988]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="1" pageNumber="534" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB97BFFBC8FAA8BD52BC839D2" box="[412,652,1967,1988]" italics="true" pageId="1" pageNumber="534">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB97BFFBC8CA28BD52BA139D3" bold="true" box="[660,741,1967,1989]" pageId="1" pageNumber="534">
<taxonomicNameLabel id="0C865784B97BFFBC8CA28BD52BA139D3" box="[660,741,1967,1989]" pageId="1" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CSIRO H 7861-01, adult male 378 mm TL, preserved): (A) dorsal view; (B) ventral view.
</paragraph>
</caption>
<caption id="71BE6665B978FFBF8EA18AEB285E38D3" httpUri="https://zenodo.org/record/256722/files/figure.png" pageId="2" pageNumber="535" targetBox="[156,1430,209,1620]" targetPageId="2">
<paragraph id="257E36EDB978FFBF8EA18AEB285E38D3" blockId="2.[151,1436,1681,1733]" pageId="2" pageNumber="535">
<emphasis id="17B5EAFFB978FFBF8EA18AEB285638B1" bold="true" box="[151,274,1681,1703]" pageId="2" pageNumber="535">FIGURE 2.</emphasis>
Paratype of
<taxonomicName id="E2C14D6EB978FFBF8FA58AEB2BC738B0" box="[403,643,1681,1702]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="535" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB978FFBF8FA58AEB2BC738B0" box="[403,643,1681,1702]" italics="true" pageId="2" pageNumber="535">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB978FFBF8CBC8AEB2B9F38B1" bold="true" box="[650,731,1681,1703]" pageId="2" pageNumber="535">
<taxonomicNameLabel id="0C865784B978FFBF8CBC8AEB2B9F38B1" box="[650,731,1681,1703]" pageId="2" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CSIRO H 7859-01, female 341 mm TL, fresh): (A) dorsal view; (B) ventral view.
</paragraph>
</caption>
<subSubSection id="6DDB6566B978FFBF8EF18A88287B398D" pageId="2" pageNumber="535" type="diagnosis">
<paragraph id="257E36EDB978FFBF8EF18A88287B398D" blockId="2.[151,1436,1778,2019]" pageId="2" pageNumber="535">
<emphasis id="17B5EAFFB978FFBF8EF18A88287E391D" bold="true" box="[199,314,1778,1803]" pageId="2" pageNumber="535">Diagnosis</emphasis>
. A medium-sized fanray with the following combination of characters: a single row of hooked thorns on dorsal midline body and tail (extending from nape to first dorsal fin); thorns on orbital, nape and scapular regions not encircled by pale pigment; dorsal surface of adults with patches of prickly denticles (rough to touch); dorsal fins well separated, interdorsal distance 8.5–9.1% TL; greyish brown dorsally with a series of darker brown saddles on tail.
</paragraph>
</subSubSection>
<subSubSection id="6DDB6566B978FFB48EF18BDC2BD238F4" lastPageId="9" lastPageNumber="542" pageId="2" pageNumber="535" type="description">
<paragraph id="257E36EDB978FFBE8EF18BDC28DD3FC6" blockId="2.[151,1436,1778,2019]" lastBlockId="3.[151,1437,151,464]" lastPageId="3" lastPageNumber="536" pageId="2" pageNumber="535">
<emphasis id="17B5EAFFB978FFBF8EF18BDC281D39A9" bold="true" box="[199,345,1958,1983]" pageId="2" pageNumber="535">Description.</emphasis>
Disc broadly rounded to shovel-shaped; anterior margin slightly to moderately convex, apex and posterior margin broadly rounded; slightly longer than wide, much shorter than tail, disc length 0.90 (0.85–0.89) times width. Pelvic fins moderate sized, length 15.5 (15.8–17.3)% TL, 1.40 (1.39–1.45) times base length, 1.66 (1.60–1.96) times width; anterior margin slightly convex to slightly concave, apex broadly rounded, posterior margin nearly straight to moderately convex. Clasper very long, extending well posterior to first dorsal-fin origin in
<typeStatus id="FA7A884FB979FFBE8EA18D7E29B93F0B" box="[151,253,260,285]" pageId="3" pageNumber="536" type="holotype">holotype</typeStatus>
; outer length 12.2% TL in
<typeStatus id="FA7A884FB979FFBE8C038D7E2BDE3F0B" box="[565,666,260,285]" pageId="3" pageNumber="536" type="holotype">holotype</typeStatus>
, 2.1 times pelvic-fin inner margin. Tail elongate, slender anteriorly, tapering gradually; in cross-section nearly flat below, slightly rounded above; tail length from anterior cloaca 1.58 (1.41–1.49) times precloacal length, 1.35 (1.23–1.32) times disc length, 7.57 (7.78–7.95) times its width at pelvicfin insertions. Dermal lateral fold on tail, originating about midway between free rear tip of pelvic fin and first dorsal-fin origin, reaching just behind ventral caudal-fin origin; fold well developed, maximum width below second dorsal-fin base.
</paragraph>
<caption id="71BE6665B979FFBE8EA18A8B28703930" httpUri="https://zenodo.org/record/256723/files/figure.png" pageId="3" pageNumber="536" targetBox="[287,1300,534,1735]" targetPageId="3">
<paragraph id="257E36EDB979FFBE8EA18A8B28703930" blockId="3.[151,1436,1777,1830]" pageId="3" pageNumber="536">
<emphasis id="17B5EAFFB979FFBE8EA18A8B28573911" bold="true" box="[151,275,1777,1799]" pageId="3" pageNumber="536">FIGURE 3.</emphasis>
Dorsal view of juvenile paratype of
<taxonomicName id="E2C14D6EB979FFBE8CB88A882A3B3911" box="[654,895,1778,1799]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="536" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB979FFBE8CB88A882A3B3911" box="[654,895,1778,1799]" italics="true" pageId="3" pageNumber="536">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB979FFBE8DB18A8B2A9C3911" bold="true" box="[903,984,1777,1799]" pageId="3" pageNumber="536">
<taxonomicNameLabel id="0C865784B979FFBE8DB18A8B2A9C3911" box="[903,984,1777,1799]" pageId="3" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CSIRO H 7860-01, juvenile male 197 mm TL, preserved).
</paragraph>
</caption>
<paragraph id="257E36EDB979FFB98EF18B3729A93EEE" blockId="3.[151,1437,1869,2038]" lastBlockId="4.[151,1436,151,248]" lastPageId="4" lastPageNumber="537" pageId="3" pageNumber="536">Head broadly rounded anteriorly with a slightly produced apical lobe, its ventral length 25.6 (25.2–26.6)% TL; snout short, broadly wedge-shaped; preoral length 1.47 (1.28–1.38) times mouth width, 2.95 (2.57–3.09) times internasal distance, 0.68 (0.66–0.75) times dorsal caudal-fin margin, 2.28 (1.92–2.05) times distance from nostril to margin of disc; preorbital snout length (direct) 1.46 (1.27–1.41) times interspiracular length, 2.68 (2.51–2.76) times orbit diameter, 2.25 (1.82–2.24) times interorbital width. Interorbital space slightly concave, relatively broad; orbit small, slightly dorsolateral, not elevated or protruding; diameter 1.60 (1.14–1.36) times spiracle length, 0.84 (0.72– 0.82) times interorbital distance. Spiracles narrowly bean-shaped, relatively large; no skin folds on posterior margin.</paragraph>
<caption id="71BE6665B97EFFB98EA18A792AEE382E" httpUri="https://zenodo.org/record/256724/files/figure.png" pageId="4" pageNumber="537" targetBox="[341,1249,316,1499]" targetPageId="4">
<paragraph id="257E36EDB97EFFB98EA18A792AEE382E" blockId="4.[151,1436,1539,1592]" pageId="4" pageNumber="537">
<emphasis id="17B5EAFFB97EFFB98EA18A792850380F" bold="true" box="[151,276,1539,1561]" pageId="4" pageNumber="537">FIGURE 4.</emphasis>
Oronasal region of
<taxonomicName id="E2C14D6EB97EFFB98FD38A792B9C380E" box="[485,728,1539,1560]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="537" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB97EFFB98FD38A792B9C380E" box="[485,728,1539,1560]" italics="true" pageId="4" pageNumber="537">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB97EFFB98CD68A792A75380F" bold="true" box="[736,817,1539,1561]" pageId="4" pageNumber="537">
<taxonomicNameLabel id="0C865784B97EFFB98CD68A792A75380F" box="[736,817,1539,1561]" pageId="4" pageNumber="537" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) holotype (CSIRO H 7861-01, adult male 378 mm TL, preserved); (B) paratype (CSIRO H 7859-01, female 341 mm TL, preserved).
</paragraph>
</caption>
<paragraph id="257E36EDB97EFFB88EF18A252B553EC3" blockId="4.[151,1436,1631,2016]" lastBlockId="5.[151,1437,151,392]" lastPageId="5" lastPageNumber="538" pageId="4" pageNumber="537">Nostrils relatively narrow, slightly oblique and well separated, nasal flaps well developed; no oronasal grooves; anterior aperture subcircular; nostril length 2.21 (1.94–2.44) times anterior aperture width, 1.21 (1.08– 1.17) times anterior nasal-flap base, 0.82 (0.62–0.76) times distance from nostril to edge of disc, 1.06 (0.83–1.10) times internasal distance. Anterior nasal flap large with long, moderately broad process anteriorly, posterior margin strongly fringed; flap base extending over full length of nostril, length 2.43 (2.08–2.76) times its width at process, 1.82 (1.79–2.12) times anterior aperture width; inserted slightly into internasal space; distance between insertions of flaps 4.67 (3.88–4.18) in greatest distance across nostrils anteriorly, 1.65 (1.45–1.50) in minimum internasal distance; process of flap narrowly rounded distally, overlapping posterolateral nasal flap and largely determining hind margin of anterior aperture. Posterolateral nasal flap lobe-like, broadest anteriorly; originating beside and well away from lateral extremity of anterior nasal aperture, extending posteromedially as a free lobe to beyond middle of posterior aperture. Posterior nasal flap present as a narrow, heavily fringed fold; extending along posterior boarder of nostril from near base of free lobe of posterolateral nasal flap and almost connected to insertion of anterior nasal-flap at its insertion.</paragraph>
<paragraph id="257E36EDB97FFFB88EF18CA528F53F9E" blockId="5.[151,1437,151,392]" pageId="5" pageNumber="538">
Mouth width 1.89 (2.00–2.43) times nostril length, 4.43 (4.10–4.38) in precloacal length; positioned over hind margin of orbit; skin grooves present; upper and lower jaw arched; lower lip pronounced; deep, short lateral grooves around corners of mouth. Teeth very small, rhomboidal, quincuncial; crowns pointed in adult male
<typeStatus id="FA7A884FB97FFFB88EA18D3629B83F73" box="[151,252,332,357]" pageId="5" pageNumber="538" type="holotype">holotype</typeStatus>
, blunt in female and juvenile
<typeStatus id="FA7A884FB97FFFB88C7B8D372BF83F73" box="[589,700,333,357]" pageId="5" pageNumber="538" type="paratype">paratypes</typeStatus>
; ca. 71 rows in upper jaw in
<typeStatus id="FA7A884FB97FFFB88A328D362D2E3F73" box="[1028,1130,332,357]" pageId="5" pageNumber="538" type="holotype">holotype</typeStatus>
; upper and lower jaw teeth similar in shape and size.
</paragraph>
<caption id="71BE6665B97FFFB88EA188A82D473B10" httpUri="https://zenodo.org/record/256725/files/figure.png" pageId="5" pageNumber="538" targetBox="[256,1318,440,1190]" targetPageId="5">
<paragraph id="257E36EDB97FFFB88EA188A82D473B10" blockId="5.[151,1436,1234,1286]" pageId="5" pageNumber="538">
<emphasis id="17B5EAFFB97FFFB88EA188A828503AFE" bold="true" box="[151,276,1234,1256]" pageId="5" pageNumber="538">FIGURE 5.</emphasis>
Dorsal view of tail region of
<taxonomicName id="E2C14D6EB97FFFB88C7E88A82A7E3AF1" box="[584,826,1234,1255]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="538" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB97FFFB88C7E88A82A7E3AF1" box="[584,826,1234,1255]" italics="true" pageId="5" pageNumber="538">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB97FFFB88D7788A82AD73AFE" bold="true" box="[833,915,1234,1256]" pageId="5" pageNumber="538">
<taxonomicNameLabel id="0C865784B97FFFB88D7788A82AD73AFE" box="[833,915,1234,1256]" pageId="5" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) holotype (CSIRO H 7861-01, adult male 378 mm TL, preserved); (B) paratype (CSIRO H 7859-01, female 341 mm TL, preserved).
</paragraph>
</caption>
<caption id="71BE6665B97FFFB88EA18BC52C4539E2" httpUri="https://zenodo.org/record/256726/files/figure.png" pageId="5" pageNumber="538" targetBox="[212,1347,1381,1924]" targetPageId="5">
<paragraph id="257E36EDB97FFFB88EA18BC52C4539E2" blockId="5.[151,1436,1983,2036]" pageId="5" pageNumber="538">
<emphasis id="17B5EAFFB97FFFB88EA18BC5285039C3" bold="true" box="[151,276,1983,2005]" pageId="5" pageNumber="538">FIGURE 6.</emphasis>
Lateral view of thorns on midline of head and disc of
<taxonomicName id="E2C14D6EB97FFFB88D7C8BBA2D7F39C3" box="[842,1083,1984,2005]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="538" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB97FFFB88D7C8BBA2D7F39C3" box="[842,1083,1984,2005]" italics="true" pageId="5" pageNumber="538">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB97FFFB88A728BC52DD139C3" bold="true" box="[1092,1173,1983,2005]" pageId="5" pageNumber="538">
<taxonomicNameLabel id="0C865784B97FFFB88A728BC52DD139C3" box="[1092,1173,1983,2005]" pageId="5" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) holotype (CSIRO H 7861-01, adult male 378 mm TL, preserved); (B) paratype (CSIRO H 7859-01, female 341 mm TL, preserved).
</paragraph>
</caption>
<caption id="71BE6665B97CFFBB8EA18AA82DE43911" httpUri="https://zenodo.org/record/256727/files/figure.png" pageId="6" pageNumber="539" targetBox="[341,1249,204,1694]" targetPageId="6">
<paragraph id="257E36EDB97CFFBB8EA18AA82DE43911" blockId="6.[151,1436,1746,1799]" pageId="6" pageNumber="539">
<emphasis id="17B5EAFFB97CFFBB8EA18AA8285738FE" bold="true" box="[151,275,1746,1768]" pageId="6" pageNumber="539">FIGURE 7.</emphasis>
Dorsal view of orbital and scapular region of
<taxonomicName id="E2C14D6EB97CFFBB8CDA8AA82A9938F1" box="[748,989,1746,1767]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="6" pageNumber="539" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB97CFFBB8CDA8AA82A9938F1" box="[748,989,1746,1767]" italics="true" pageId="6" pageNumber="539">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB97CFFBB8DD38AA82D7238FE" bold="true" box="[997,1078,1746,1768]" pageId="6" pageNumber="539">
<taxonomicNameLabel id="0C865784B97CFFBB8DD38AA82D7238FE" box="[997,1078,1746,1768]" pageId="6" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) holotype (CSIRO H 7861-01, adult male 378 mm TL, preserved); (B) paratype (CSIRO H 7859-01, female 341 mm TL, preserved).
</paragraph>
</caption>
<paragraph id="257E36EDB97CFFBB8EF18B492A7139AE" blockId="6.[151,1436,1843,2012]" pageId="6" pageNumber="539">Gill openings weakly S-shaped, fifth less so; length of third gill slit 2.20 (1.95–2.15) in nostril length, 5.51 (5.36–6.47) in distance between fifth gill slits; distance between first gill slits 1.44 (1.36–1.47) times distance between fifth gill slits; distance between fifth gill slits 2.65 (2.63–3.03) times internasal distance, 1.32 (1.29–1.36) times mouth width, 0.45 (0.48–0.50) of ventral head length.</paragraph>
<paragraph id="257E36EDB97CFFBA8EF18BB92A4A3FC6" blockId="6.[151,1436,1843,2012]" lastBlockId="7.[151,1436,151,645]" lastPageId="7" lastPageNumber="540" pageId="6" pageNumber="539">
Dorsal fins similar in size and shape, well separated, moderately large, subtriangular, relatively upright; anterior margins slightly convex, apices moderately rounded; posterior margins nearly straight to slightly convex; free rear tips relatively short, not produced; first dorsal fin slightly taller than second in all but one
<typeStatus id="FA7A884FB97DFFBA8ADE8CC72C0F3EC3" box="[1256,1355,189,213]" pageId="7" pageNumber="540" type="paratype">paratype</typeStatus>
, length of first 1.07 (1.11–1.52) times its height, base length 2.69 (1.27–2.00) times inner margin length; second dorsal-fin length 1.45 (1.28–1.46) times its height, base length 1.88 (1.66–2.29) times inner margin length. First dorsal fin well behind pelvic-fin rear tip, pelvic-fin insertion to first dorsal-fin origin 1.69 (1.44–0.59) times interdorsal distance; interdorsal space long, 1.89 (1.70–1.90) times second dorsal-fin height, 2.05 (2.10–2.38) times base of first dorsal fin, 1.31 (1.33–1.52) times tail width at origin of first dorsal fin, 1.89 (1.60–2.11) times interspace between second dorsal-fin insertion and upper origin of caudal fin. Caudal fin medium-sized, deep; dorsal caudal margin 1.41 (1.33–1.57) times preventral margin length.
</paragraph>
<paragraph id="257E36EDB97DFFBA8EF18DA62D7A3C93" blockId="7.[151,1436,151,645]" pageId="7" pageNumber="540">
Skin of largest specimens with patches of prickly dermal denticles of varying sizes, greatly enlarged on anterior snout, central disc and in two, well-spaced paired rows along either side of tail. Skin of juvenile
<typeStatus id="FA7A884FB97DFFBA8B078E7A2CD83C0E" box="[1329,1436,512,536]" pageId="7" pageNumber="540" type="paratype">paratypes</typeStatus>
velvety. Thorns well developed, sharp, hooked, not on prominent ridges; a regular median row extending from nape to first dorsal fin, 4–6 main orbital thorns, two symmetrical shoulder patches on each side of disc (outermost patch with 2 thorns, innermost patch one thorn); thornlets on anterior rostral ridges.
</paragraph>
<caption id="71BE6665B97DFFBA8EA18BE42D8239C5" httpUri="https://zenodo.org/record/256728/files/figure.png" pageId="7" pageNumber="540" targetBox="[341,1248,711,1903]" targetPageId="7">
<paragraph id="257E36EDB97DFFBA8EA18BE42D8239C5" blockId="7.[151,1436,1950,2003]" pageId="7" pageNumber="540">
<emphasis id="17B5EAFFB97DFFBA8EA18BE4285639A2" bold="true" box="[151,274,1950,1972]" pageId="7" pageNumber="540">FIGURE 8.</emphasis>
Dorsolateral view of the left orbit and spiracle of
<taxonomicName id="E2C14D6EB97DFFBA8D268BE52D4439A2" box="[784,1024,1951,1972]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="540" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB97DFFBA8D268BE52D4439A2" box="[784,1024,1951,1972]" italics="true" pageId="7" pageNumber="540">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB97DFFBA8A318BE42D1C39A2" bold="true" box="[1031,1112,1950,1972]" pageId="7" pageNumber="540">
<taxonomicNameLabel id="0C865784B97DFFBA8A318BE42D1C39A2" box="[1031,1112,1950,1972]" pageId="7" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) holotype (CSIRO H 7861- 0 1, adult male 378 mm TL, preserved); (B) paratype (CSIRO H 7859-01, female 341 mm TL, preserved).
</paragraph>
</caption>
<paragraph id="257E36EDB972FFB58EF18CED2D193F29" blockId="8.[151,1437,151,320]" pageId="8" pageNumber="541">
Nasal lamellae
<quantity id="E2399B08B972FFB58F498CE228AD3EA6" box="[383,489,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" metricValueMax="8.382" metricValueMin="7.874" pageId="8" pageNumber="541" unit="in" value="32.0" valueMax="33.0" valueMin="31.0">31–33 in</quantity>
<typeStatus id="FA7A884FB972FFB58FC08CED2B1C3EA6" box="[502,600,151,176]" pageId="8" pageNumber="541" type="holotype">holotype</typeStatus>
(
<quantity id="E2399B08B972FFB58C5A8CE22B933EA6" box="[620,727,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="7.493" metricValueMax="7.62" metricValueMin="7.366" pageId="8" pageNumber="541" unit="in" value="29.5" valueMax="30.0" valueMin="29.0">29–30 in</quantity>
larger female
<typeStatus id="FA7A884FB972FFB58DBD8CE22AB53EA6" box="[907,1009,152,176]" pageId="8" pageNumber="541" type="paratype">paratype</typeStatus>
). Pectoral skeleton with 34 (32–33) propterygial, 8–9 (9–11) mesopterygial, 23 (23) metapterygial and 65–66 (64–66) total radials. Total pelvic radials 1 (1) + 19 (~18) excluding clasper in males. Vertebral column with 138 (137–139) total centra; 95 (93–104) total precaudal centra; 25 (23–24) monospondylous precaudal centra; 30 (29–31) diplospondylous predorsal centra; 70 (69–81) diplospondylous precaudal centra; 43 (35–44) diplospondylous caudal centra.
</paragraph>
<caption id="71BE6665B972FFB58EA18D0A2B013FB1" pageId="8" pageNumber="541">
<paragraph id="257E36EDB972FFB58EA18D0A2B013FB1" blockId="8.[151,1436,367,423]" pageId="8" pageNumber="541">
<emphasis id="17B5EAFFB972FFB58EA18D0A284F3F91" bold="true" box="[151,267,368,391]" pageId="8" pageNumber="541">TABLE 1.</emphasis>
Morphometric data for the holotype of
<taxonomicName id="E2C14D6EB972FFB58C888D152AF93F91" box="[702,957,367,391]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="541" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB972FFB58C888D152AF93F91" box="[702,957,367,391]" italics="true" pageId="8" pageNumber="541">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0C865784B972FFB58DF38D0A2D583F91" box="[965,1052,368,391]" pageId="8" pageNumber="541" rank="species">
<emphasis id="17B5EAFFB972FFB58DF38D0A2D583F91" bold="true" box="[965,1052,368,391]" pageId="8" pageNumber="541">sp. nov.</emphasis>
</taxonomicNameLabel>
and ranges for the three paratypes; expressed as percentages of total length.
</paragraph>
</caption>
<paragraph id="257E36EDB972FFB58C8A8DB82D5D3FCE" blockId="8.[159,1432,450,2001]" box="[700,1049,450,472]" pageId="8" pageNumber="541">
<typeStatus id="FA7A884FB972FFB58C8A8DB82A5E3FCE" box="[700,794,450,472]" pageId="8" pageNumber="541" type="holotype">Holotype</typeStatus>
<typeStatus id="FA7A884FB972FFB58D818DB92D5D3FCE" box="[951,1049,451,472]" pageId="8" pageNumber="541" type="paratype">Paratypes</typeStatus>
</paragraph>
<paragraph id="257E36EDB972FFB58D818D962D9439BE" blockId="8.[159,1432,450,2001]" pageId="8" pageNumber="541">Min. Max. Total length (mm) 378.0 197.0 341.0 Disc width—maximum 50.5 53.1 54.3 Disc length 45.2 45.3 48.5 Head length—dorsal 17.4 16.8 17.9 Head length—ventral 25.6 25.2 26.6 Snout length—presocket 10.9 10.4 10.7 Orbit diameter 4.1 3.8 4.2 Spiracle length 2.5 3.1 3.3 Orbit and spiracle length 5.4 5.9 6.3 Interorbital width 4.9 4.7 5.9 Interspiracular width 7.5 7.4 8.4 Preoral length 12.9 11.9 13.3 Mouth width 8.7 9.3 9.8 Prenarial distance 9.8 8.8 9.3 Nostril length 4.6 3.8 4.9 Anterior aperture—width 2.1 1.9 2.1 Anterior nasal flap—base length 3.8 3.6 4.4 Anterior nasal flap—width 1.6 1.3 2.1 Distance across anterior nasal apertures 12.3 12.0 12.6 Internarial distance—minimum width 4.4 4.2 4.8 Distance between anterior nasal flaps 2.6 2.9 3.2 Distance from nostril to disc margin 5.6 6.2 6.5 Third gill opening—width 2.1 1.9 2.4 Distance between first gill openings 16.6 17.5 18.6 Distance between fifth gill openings 11.5 12.4 12.8 Pelvic fin—length 15.5 15.8 17.3 Pelvic fin—anterior margin length 9.1 9.8 10.3 Pelvic fin—width 9.3 8.8 9.9 Pelvic fin—base length 11.1 11.4 11.9 Pelvic fin—inner margin length 5.7 6.1 7.0 First dorsal fin—length 6.2 5.9 7.1 First dorsal fin—anterior margin length 8.4 7.8 8.2 First dorsal fin—height 5.8 4.6 5.7 First dorsal fin—base length 4.4 3.8 4.3 First dorsal fin—inner margin length 1.6 2.1 3.0</paragraph>
<paragraph id="257E36EDB972FFB48A548BC62D5D3EC9" blockId="8.[159,1432,450,2001]" lastBlockId="9.[159,1254,160,264]" lastPageId="9" lastPageNumber="542" pageId="8" pageNumber="541">
<emphasis id="17B5EAFFB972FFB58A548BC62CDC39C7" box="[1122,1432,1980,2001]" italics="true" pageId="8" pageNumber="541">......continued on the next page</emphasis>
<typeStatus id="FA7A884FB973FFB48C8A8CB32A5E3EC9" box="[700,794,201,223]" pageId="9" pageNumber="542" type="holotype">Holotype</typeStatus>
<typeStatus id="FA7A884FB973FFB48D818CB02D5D3EC9" box="[951,1049,202,223]" pageId="9" pageNumber="542" type="paratype">Paratypes</typeStatus>
</paragraph>
<caption id="71BE6665B973FFB48EA98CDA28C93EA0" ID-Table-UUID="71BE6665B973FFB48EA98CDA28C93EA0" box="[159,397,160,182]" httpUri="http://table.plazi.org/id/71BE6665B973FFB48EA98CDA28C93EA0" pageId="9" pageNumber="542" targetBox="[159,1254,284,1422]" targetIsTable="true" targetPageId="9">
<paragraph id="257E36EDB973FFB48EA98CDA28C93EA0" blockId="9.[159,1254,160,264]" box="[159,397,160,182]" pageId="9" pageNumber="542">
<emphasis id="17B5EAFFB973FFB48EA98CDA28483EA0" bold="true" box="[159,268,160,182]" pageId="9" pageNumber="542">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="257E36EDB973FFB48D818C882D3C3844" blockId="9.[159,1254,160,264]" lastBlockId="9.[151,1436,1485,2014]" pageId="9" pageNumber="542">
Min. Max.
<emphasis id="17B5EAFFB973FFB48EF189B728643BF0" bold="true" box="[199,288,1485,1510]" pageId="9" pageNumber="542">Colour.</emphasis>
(Based on
<typeStatus id="FA7A884FB973FFB48F9B89B42B493BF0" box="[429,525,1486,1510]" pageId="9" pageNumber="542" type="paratype">paratype</typeStatus>
CSIRO H 7859-01). Dorsal surfaces medium brown with a series of mostly indistinct cross bars on disc and five broad dark brown saddles on tail; dorsal fins similar colour to disc, slightly darker at anterior of base adjacent to dark saddles on tail dorsal-fin bases. No pale areas around the large thorns on head, body or tail. Ventral surface whitish, slightly darker near posterior margins of disc.
</paragraph>
<paragraph id="257E36EDB973FFB48EA98D662D943B98" pageId="9" pageNumber="542">
<table id="57C1C44DB97300428EA98D662DA23B98" box="[159,1254,284,1422]" gridcols="4" gridrows="28" pageId="9" pageNumber="542">
<tr id="9BF134AFB97300428EA98D662DA23F24" box="[159,1254,284,306]" gridrow="0" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98D662B0F3F24" box="[159,587,284,306]" gridcol="0" gridrow="0" pageId="9" pageNumber="542">Second dorsal fin—length</th>
<th id="D8205DD3B97300428C8D8D662BAC3F24" box="[699,744,284,306]" gridcol="1" gridrow="0" pageId="9" pageNumber="542">6.9</th>
<th id="D8205DD3B97300428D818D662AA73F24" box="[951,995,284,306]" gridcol="2" gridrow="0" pageId="9" pageNumber="542">6.5</th>
<th id="D8205DD3B97300428A868D662DA23F24" box="[1200,1254,284,306]" gridcol="3" gridrow="0" pageId="9" pageNumber="542">6.8</th>
</tr>
<tr id="9BF134AFB97300428EA98D3F2DA23F4D" box="[159,1254,325,347]" gridrow="1" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98D3F2B0F3F4D" box="[159,587,325,347]" gridcol="0" gridrow="1" pageId="9" pageNumber="542">Second dorsal fin—anterior margin length</th>
<td id="D8205DD3B97300428C8D8D3F2BAC3F4D" box="[699,744,325,347]" gridcol="1" gridrow="1" pageId="9" pageNumber="542">7.7</td>
<td id="D8205DD3B97300428D818D3F2AA73F4D" box="[951,995,325,347]" gridcol="2" gridrow="1" pageId="9" pageNumber="542">7.8</td>
<td id="D8205DD3B97300428A868D3F2DA23F4D" box="[1200,1254,325,347]" gridcol="3" gridrow="1" pageId="9" pageNumber="542">8.4</td>
</tr>
<tr id="9BF134AFB97300428EA98D142DA23F92" box="[159,1254,366,388]" gridrow="2" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98D142B0F3F92" box="[159,587,366,388]" gridcol="0" gridrow="2" pageId="9" pageNumber="542">Second dorsal fin—height</th>
<td id="D8205DD3B97300428C8D8D142BAC3F92" box="[699,744,366,388]" gridcol="1" gridrow="2" pageId="9" pageNumber="542">4.8</td>
<td id="D8205DD3B97300428D818D142AA73F92" box="[951,995,366,388]" gridcol="2" gridrow="2" pageId="9" pageNumber="542">4.4</td>
<td id="D8205DD3B97300428A868D142DA23F92" box="[1200,1254,366,388]" gridcol="3" gridrow="2" pageId="9" pageNumber="542">5.4</td>
</tr>
<tr id="9BF134AFB97300428EA98DE22DA23FB8" box="[159,1254,408,430]" gridrow="3" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98DE22B0F3FB8" box="[159,587,408,430]" gridcol="0" gridrow="3" pageId="9" pageNumber="542">Second dorsal fin—base length</th>
<td id="D8205DD3B97300428C8D8DE22BAC3FB8" box="[699,744,408,430]" gridcol="1" gridrow="3" pageId="9" pageNumber="542">5.0</td>
<td id="D8205DD3B97300428D818DE22AA73FB8" box="[951,995,408,430]" gridcol="2" gridrow="3" pageId="9" pageNumber="542">4.6</td>
<td id="D8205DD3B97300428A868DE22DA23FB8" box="[1200,1254,408,430]" gridcol="3" gridrow="3" pageId="9" pageNumber="542">5.0</td>
</tr>
<tr id="9BF134AFB97300428EA98DBB2DA23FC1" box="[159,1254,449,471]" gridrow="4" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98DBB2B0F3FC1" box="[159,587,449,471]" gridcol="0" gridrow="4" pageId="9" pageNumber="542">Second dorsal fin—inner margin length</th>
<td id="D8205DD3B97300428C8D8DBB2BAC3FC1" box="[699,744,449,471]" gridcol="1" gridrow="4" pageId="9" pageNumber="542">2.7</td>
<td id="D8205DD3B97300428D818DBB2AA73FC1" box="[951,995,449,471]" gridcol="2" gridrow="4" pageId="9" pageNumber="542">2.2</td>
<td id="D8205DD3B97300428A868DBB2DA23FC1" box="[1200,1254,449,471]" gridcol="3" gridrow="4" pageId="9" pageNumber="542">2.8</td>
</tr>
<tr id="9BF134AFB97300428EA98D902DA23C16" box="[159,1254,490,512]" gridrow="5" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98D902B0F3C16" box="[159,587,490,512]" gridcol="0" gridrow="5" pageId="9" pageNumber="542">Caudal fin—dorsal margin</th>
<td id="D8205DD3B97300428C8D8D902BAC3C16" box="[699,744,490,512]" gridcol="1" gridrow="5" pageId="9" pageNumber="542">18.9</td>
<td id="D8205DD3B97300428D818D902AA73C16" box="[951,995,490,512]" gridcol="2" gridrow="5" pageId="9" pageNumber="542">17.6</td>
<td id="D8205DD3B97300428A868D902DA23C16" box="[1200,1254,490,512]" gridcol="3" gridrow="5" pageId="9" pageNumber="542">19.1</td>
</tr>
<tr id="9BF134AFB97300428EA98E6E2DA23C3C" box="[159,1254,532,554]" gridrow="6" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98E6E2B0F3C3C" box="[159,587,532,554]" gridcol="0" gridrow="6" pageId="9" pageNumber="542">Caudal fin—preventral margin</th>
<td id="D8205DD3B97300428C8D8E6E2BAC3C3C" box="[699,744,532,554]" gridcol="1" gridrow="6" pageId="9" pageNumber="542">13.4</td>
<td id="D8205DD3B97300428D818E6E2AA73C3C" box="[951,995,532,554]" gridcol="2" gridrow="6" pageId="9" pageNumber="542">11.2</td>
<td id="D8205DD3B97300428A868E6E2DA23C3C" box="[1200,1254,532,554]" gridcol="3" gridrow="6" pageId="9" pageNumber="542">13.6</td>
</tr>
<tr id="9BF134AFB97300428EA98E472DA23C45" box="[159,1254,573,595]" gridrow="7" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98E472B0F3C45" box="[159,587,573,595]" gridcol="0" gridrow="7" pageId="9" pageNumber="542">Caudal fin—postventral margin</th>
<td id="D8205DD3B97300428C8D8E472BAC3C45" box="[699,744,573,595]" gridcol="1" gridrow="7" pageId="9" pageNumber="542">7.0</td>
<td id="D8205DD3B97300428D818E472AA73C45" box="[951,995,573,595]" gridcol="2" gridrow="7" pageId="9" pageNumber="542">6.5</td>
<td id="D8205DD3B97300428A868E472DA23C45" box="[1200,1254,573,595]" gridcol="3" gridrow="7" pageId="9" pageNumber="542">6.9</td>
</tr>
<tr id="9BF134AFB97300428EA98E1C2DA23C6A" box="[159,1254,614,636]" gridrow="8" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98E1C2B0F3C6A" box="[159,587,614,636]" gridcol="0" gridrow="8" pageId="9" pageNumber="542">Snout to first dorsal-fin origin</th>
<td id="D8205DD3B97300428C8D8E1C2BAC3C6A" box="[699,744,614,636]" gridcol="1" gridrow="8" pageId="9" pageNumber="542">59.4</td>
<td id="D8205DD3B97300428D818E1C2AA73C6A" box="[951,995,614,636]" gridcol="2" gridrow="8" pageId="9" pageNumber="542">59.0</td>
<td id="D8205DD3B97300428A868E1C2DA23C6A" box="[1200,1254,614,636]" gridcol="3" gridrow="8" pageId="9" pageNumber="542">59.9</td>
</tr>
<tr id="9BF134AFB97300428EA98EEA2DA23CB0" box="[159,1254,656,678]" gridrow="9" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98EEA2B0F3CB0" box="[159,587,656,678]" gridcol="0" gridrow="9" pageId="9" pageNumber="542">Snout to second dorsal-fin origin</th>
<td id="D8205DD3B97300428C8D8EEA2BAC3CB0" box="[699,744,656,678]" gridcol="1" gridrow="9" pageId="9" pageNumber="542">71.7</td>
<td id="D8205DD3B97300428D818EEA2AA73CB0" box="[951,995,656,678]" gridcol="2" gridrow="9" pageId="9" pageNumber="542">71.2</td>
<td id="D8205DD3B97300428A868EEA2DA23CB0" box="[1200,1254,656,678]" gridcol="3" gridrow="9" pageId="9" pageNumber="542">72.1</td>
</tr>
<tr id="9BF134AFB97300428EA98EC32DA23CD9" box="[159,1254,697,719]" gridrow="10" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98EC32B0F3CD9" box="[159,587,697,719]" gridcol="0" gridrow="10" pageId="9" pageNumber="542">Snout to upper caudal-fin origin</th>
<td id="D8205DD3B97300428C8D8EC32BAC3CD9" box="[699,744,697,719]" gridcol="1" gridrow="10" pageId="9" pageNumber="542">81.5</td>
<td id="D8205DD3B97300428D818EC32AA73CD9" box="[951,995,697,719]" gridcol="2" gridrow="10" pageId="9" pageNumber="542">80.0</td>
<td id="D8205DD3B97300428A868EC32DA23CD9" box="[1200,1254,697,719]" gridcol="3" gridrow="10" pageId="9" pageNumber="542">81.5</td>
</tr>
<tr id="9BF134AFB97300428EA98E982DA23CEE" box="[159,1254,738,760]" gridrow="11" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98E982B0F3CEE" box="[159,587,738,760]" gridcol="0" gridrow="11" pageId="9" pageNumber="542">Snout to lower caudal-fin origin</th>
<td id="D8205DD3B97300428C8D8E982BAC3CEE" box="[699,744,738,760]" gridcol="1" gridrow="11" pageId="9" pageNumber="542">82.0</td>
<td id="D8205DD3B97300428D818E982AA73CEE" box="[951,995,738,760]" gridcol="2" gridrow="11" pageId="9" pageNumber="542">81.7</td>
<td id="D8205DD3B97300428A868E982DA23CEE" box="[1200,1254,738,760]" gridcol="3" gridrow="11" pageId="9" pageNumber="542">83.7</td>
</tr>
<tr id="9BF134AFB97300428EA98F762DA23D34" box="[159,1254,780,802]" gridrow="12" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98F762B0F3D34" box="[159,587,780,802]" gridcol="0" gridrow="12" pageId="9" pageNumber="542">Snout to pelvic-fin origin</th>
<td id="D8205DD3B97300428C8D8F762BAC3D34" box="[699,744,780,802]" gridcol="1" gridrow="12" pageId="9" pageNumber="542">36.9</td>
<td id="D8205DD3B97300428D818F762AA73D34" box="[951,995,780,802]" gridcol="2" gridrow="12" pageId="9" pageNumber="542">37.5</td>
<td id="D8205DD3B97300428A868F762DA23D34" box="[1200,1254,780,802]" gridcol="3" gridrow="12" pageId="9" pageNumber="542">39.7</td>
</tr>
<tr id="9BF134AFB97300428EA98F4F2DA23D5D" box="[159,1254,821,843]" gridrow="13" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98F4F2B0F3D5D" box="[159,587,821,843]" gridcol="0" gridrow="13" pageId="9" pageNumber="542">Snout to anterior vent</th>
<td id="D8205DD3B97300428C8D8F4F2BAC3D5D" box="[699,744,821,843]" gridcol="1" gridrow="13" pageId="9" pageNumber="542">38.7</td>
<td id="D8205DD3B97300428D818F4F2AA73D5D" box="[951,995,821,843]" gridcol="2" gridrow="13" pageId="9" pageNumber="542">40.1</td>
<td id="D8205DD3B97300428A868F4F2DA23D5D" box="[1200,1254,821,843]" gridcol="3" gridrow="13" pageId="9" pageNumber="542">41.4</td>
</tr>
<tr id="9BF134AFB97300428EA98F242DA23D62" box="[159,1254,862,884]" gridrow="14" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98F242B0F3D62" box="[159,587,862,884]" gridcol="0" gridrow="14" pageId="9" pageNumber="542">Pelvic-fin insertion to dorsal-fin origin</th>
<td id="D8205DD3B97300428C8D8F242BAC3D62" box="[699,744,862,884]" gridcol="1" gridrow="14" pageId="9" pageNumber="542">15.3</td>
<td id="D8205DD3B97300428D818F242AA73D62" box="[951,995,862,884]" gridcol="2" gridrow="14" pageId="9" pageNumber="542">12.2</td>
<td id="D8205DD3B97300428A868F242DA23D62" box="[1200,1254,862,884]" gridcol="3" gridrow="14" pageId="9" pageNumber="542">14.5</td>
</tr>
<tr id="9BF134AFB97300428EA98FF22DA23D88" box="[159,1254,904,926]" gridrow="15" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98FF22B0F3D88" box="[159,587,904,926]" gridcol="0" gridrow="15" pageId="9" pageNumber="542">Interdorsal distance</th>
<td id="D8205DD3B97300428C8D8FF22BAC3D88" box="[699,744,904,926]" gridcol="1" gridrow="15" pageId="9" pageNumber="542">9.0</td>
<td id="D8205DD3B97300428D818FF22AA73D88" box="[951,995,904,926]" gridcol="2" gridrow="15" pageId="9" pageNumber="542">8.5</td>
<td id="D8205DD3B97300428A868FF22DA23D88" box="[1200,1254,904,926]" gridcol="3" gridrow="15" pageId="9" pageNumber="542">9.1</td>
</tr>
<tr id="9BF134AFB97300428EA98FCB2DA23DD1" box="[159,1254,945,967]" gridrow="16" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98FCB2B0F3DD1" box="[159,587,945,967]" gridcol="0" gridrow="16" pageId="9" pageNumber="542">Caudal peduncle length (dorsal)</th>
<td id="D8205DD3B97300428C8D8FCB2BAC3DD1" box="[699,744,945,967]" gridcol="1" gridrow="16" pageId="9" pageNumber="542">4.8</td>
<td id="D8205DD3B97300428D818FCB2AA73DD1" box="[951,995,945,967]" gridcol="2" gridrow="16" pageId="9" pageNumber="542">4.3</td>
<td id="D8205DD3B97300428A868FCB2DA23DD1" box="[1200,1254,945,967]" gridcol="3" gridrow="16" pageId="9" pageNumber="542">5.3</td>
</tr>
<tr id="9BF134AFB97300428EA98FA02DA23DE6" box="[159,1254,986,1008]" gridrow="17" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA98FA02B0F3DE6" box="[159,587,986,1008]" gridcol="0" gridrow="17" pageId="9" pageNumber="542">Body width—pelvic insertion</th>
<td id="D8205DD3B97300428C8D8FA02BAC3DE6" box="[699,744,986,1008]" gridcol="1" gridrow="17" pageId="9" pageNumber="542">8.1</td>
<td id="D8205DD3B97300428D818FA02AA73DE6" box="[951,995,986,1008]" gridcol="2" gridrow="17" pageId="9" pageNumber="542">7.5</td>
<td id="D8205DD3B97300428A868FA02DA23DE6" box="[1200,1254,986,1008]" gridcol="3" gridrow="17" pageId="9" pageNumber="542">7.6</td>
</tr>
<tr id="9BF134AFB97300428EA9887E2DA23A0C" box="[159,1254,1028,1050]" gridrow="18" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA9887E2B0F3A0C" box="[159,587,1028,1050]" gridcol="0" gridrow="18" pageId="9" pageNumber="542">Disc width—anterior orbit</th>
<td id="D8205DD3B97300428C8D887E2BAC3A0C" box="[699,744,1028,1050]" gridcol="1" gridrow="18" pageId="9" pageNumber="542">30.7</td>
<td id="D8205DD3B97300428D81887E2AA73A0C" box="[951,995,1028,1050]" gridcol="2" gridrow="18" pageId="9" pageNumber="542">29.6</td>
<td id="D8205DD3B97300428A86887E2DA23A0C" box="[1200,1254,1028,1050]" gridcol="3" gridrow="18" pageId="9" pageNumber="542">37.6</td>
</tr>
<tr id="9BF134AFB97300428EA988572DA23A55" box="[159,1254,1069,1091]" gridrow="19" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA988572B0F3A55" box="[159,587,1069,1091]" gridcol="0" gridrow="19" pageId="9" pageNumber="542">Body width—first dorsal–fin origin</th>
<td id="D8205DD3B97300428C8D88572BAC3A55" box="[699,744,1069,1091]" gridcol="1" gridrow="19" pageId="9" pageNumber="542">6.9</td>
<td id="D8205DD3B97300428D8188572AA73A55" box="[951,995,1069,1091]" gridcol="2" gridrow="19" pageId="9" pageNumber="542">6.0</td>
<td id="D8205DD3B97300428A8688572DA23A55" box="[1200,1254,1069,1091]" gridcol="3" gridrow="19" pageId="9" pageNumber="542">6.4</td>
</tr>
<tr id="9BF134AFB97300428EA9882C2DA23A7A" box="[159,1254,1110,1132]" gridrow="20" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA9882C2B0F3A7A" box="[159,587,1110,1132]" gridcol="0" gridrow="20" pageId="9" pageNumber="542">Body width—second dorsal–fin origin</th>
<td id="D8205DD3B97300428C8D882C2BAC3A7A" box="[699,744,1110,1132]" gridcol="1" gridrow="20" pageId="9" pageNumber="542">4.6</td>
<td id="D8205DD3B97300428D81882C2AA73A7A" box="[951,995,1110,1132]" gridcol="2" gridrow="20" pageId="9" pageNumber="542">4.2</td>
<td id="D8205DD3B97300428A86882C2DA23A7A" box="[1200,1254,1110,1132]" gridcol="3" gridrow="20" pageId="9" pageNumber="542">4.5</td>
</tr>
<tr id="9BF134AFB97300428EA988FA2DA23A80" box="[159,1254,1152,1174]" gridrow="21" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA988FA2B0F3A80" box="[159,587,1152,1174]" gridcol="0" gridrow="21" pageId="9" pageNumber="542">Body depth—maximum</th>
<td id="D8205DD3B97300428C8D88FA2BAC3A80" box="[699,744,1152,1174]" gridcol="1" gridrow="21" pageId="9" pageNumber="542">6.0</td>
<td id="D8205DD3B97300428D8188FA2AA73A80" box="[951,995,1152,1174]" gridcol="2" gridrow="21" pageId="9" pageNumber="542">5.4</td>
<td id="D8205DD3B97300428A8688FA2DA23A80" box="[1200,1254,1152,1174]" gridcol="3" gridrow="21" pageId="9" pageNumber="542">5.6</td>
</tr>
<tr id="9BF134AFB97300428EA988D32DA23AA9" box="[159,1254,1193,1215]" gridrow="22" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA988D32B0F3AA9" box="[159,587,1193,1215]" gridcol="0" gridrow="22" pageId="9" pageNumber="542">Body depth—pelvic–fin insertion</th>
<td id="D8205DD3B97300428C8D88D32BAC3AA9" box="[699,744,1193,1215]" gridcol="1" gridrow="22" pageId="9" pageNumber="542">4.8</td>
<td id="D8205DD3B97300428D8188D32AA73AA9" box="[951,995,1193,1215]" gridcol="2" gridrow="22" pageId="9" pageNumber="542">4.2</td>
<td id="D8205DD3B97300428A8688D32DA23AA9" box="[1200,1254,1193,1215]" gridcol="3" gridrow="22" pageId="9" pageNumber="542">4.8</td>
</tr>
<tr id="9BF134AFB97300428EA988A82DA23AFE" box="[159,1254,1234,1256]" gridrow="23" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA988A82B0F3AFE" box="[159,587,1234,1256]" gridcol="0" gridrow="23" pageId="9" pageNumber="542">Body depth—first dorsal–fin origin</th>
<td id="D8205DD3B97300428C8D88A82BAC3AFE" box="[699,744,1234,1256]" gridcol="1" gridrow="23" pageId="9" pageNumber="542">3.5</td>
<td id="D8205DD3B97300428D8188A82AA73AFE" box="[951,995,1234,1256]" gridcol="2" gridrow="23" pageId="9" pageNumber="542">3.4</td>
<td id="D8205DD3B97300428A8688A82DA23AFE" box="[1200,1254,1234,1256]" gridcol="3" gridrow="23" pageId="9" pageNumber="542">3.8</td>
</tr>
<tr id="9BF134AFB97300428EA988862DA23B04" box="[159,1254,1276,1298]" gridrow="24" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA988862B0F3B04" box="[159,587,1276,1298]" gridcol="0" gridrow="24" pageId="9" pageNumber="542">Body depth—second dorsal–fin origin</th>
<td id="D8205DD3B97300428C8D88862BAC3B04" box="[699,744,1276,1298]" gridcol="1" gridrow="24" pageId="9" pageNumber="542">2.4</td>
<td id="D8205DD3B97300428D8188862AA73B04" box="[951,995,1276,1298]" gridcol="2" gridrow="24" pageId="9" pageNumber="542">2.4</td>
<td id="D8205DD3B97300428A8688862DA23B04" box="[1200,1254,1276,1298]" gridcol="3" gridrow="24" pageId="9" pageNumber="542">3.0</td>
</tr>
<tr id="9BF134AFB97300428EA9895F2DA23B2D" box="[159,1254,1317,1339]" gridrow="25" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA9895F2B0F3B2D" box="[159,587,1317,1339]" gridcol="0" gridrow="25" pageId="9" pageNumber="542">Clasper outer length</th>
<td id="D8205DD3B97300428C8D895F2BAC3B2D" box="[699,744,1317,1339]" gridcol="1" gridrow="25" pageId="9" pageNumber="542">12.2</td>
<td id="D8205DD3B97300428D81895F2AA73B2D" box="[951,995,1317,1339]" gridcol="2" gridrow="25" pageId="9" pageNumber="542">0.0</td>
<td id="D8205DD3B97300428A86895F2DA23B2D" box="[1200,1254,1317,1339]" gridcol="3" gridrow="25" pageId="9" pageNumber="542">0.0</td>
</tr>
<tr id="9BF134AFB97300428EA989342DA23B72" box="[159,1254,1358,1380]" gridrow="26" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA989342B0F3B72" box="[159,587,1358,1380]" gridcol="0" gridrow="26" pageId="9" pageNumber="542">Clasper inner length</th>
<td id="D8205DD3B97300428C8D89342BAC3B72" box="[699,744,1358,1380]" gridcol="1" gridrow="26" pageId="9" pageNumber="542">17.3</td>
<td id="D8205DD3B97300428D8189342AA73B72" box="[951,995,1358,1380]" gridcol="2" gridrow="26" pageId="9" pageNumber="542">0.0</td>
<td id="D8205DD3B97300428A8689342DA23B72" box="[1200,1254,1358,1380]" gridcol="3" gridrow="26" pageId="9" pageNumber="542">0.0</td>
</tr>
<tr id="9BF134AFB97300428EA989022DA23B98" box="[159,1254,1400,1422]" gridrow="27" pageId="9" pageNumber="542">
<th id="D8205DD3B97300428EA989022B0F3B98" box="[159,587,1400,1422]" gridcol="0" gridrow="27" pageId="9" pageNumber="542">Clasper base width</th>
<td id="D8205DD3B97300428C8D89022BAC3B98" box="[699,744,1400,1422]" gridcol="1" gridrow="27" pageId="9" pageNumber="542">2.2</td>
<td id="D8205DD3B97300428D8189022AA73B98" box="[951,995,1400,1422]" gridcol="2" gridrow="27" pageId="9" pageNumber="542">0.0</td>
<td id="D8205DD3B97300428A8689022DA23B98" box="[1200,1254,1400,1422]" gridcol="3" gridrow="27" pageId="9" pageNumber="542">0.0</td>
</tr>
</table>
</paragraph>
<paragraph id="257E36EDB973FFB48EF18A2728B1388C" blockId="9.[151,1436,1485,2014]" pageId="9" pageNumber="542">
<emphasis id="17B5EAFFB973FFB48EF18A2729BA3860" bold="true" box="[199,254,1629,1654]" pageId="9" pageNumber="542">Size.</emphasis>
Known from the four
<typeStatus id="FA7A884FB973FFB48FCF8A242B6D3860" box="[505,553,1630,1654]" pageId="9" pageNumber="542">type</typeStatus>
specimens; an adult male (
<quantity id="E2399B08B973FFB48D618A242AF13863" box="[855,949,1630,1654]" metricMagnitude="-1" metricUnit="m" metricValue="3.78" pageId="9" pageNumber="542" unit="mm" value="378.0">378 mm</quantity>
TL), a juvenile male (
<quantity id="E2399B08B973FFB48A9B8A242C4F3863" box="[1197,1291,1630,1654]" metricMagnitude="-1" metricUnit="m" metricValue="1.97" pageId="9" pageNumber="542" unit="mm" value="197.0">197 mm</quantity>
TL) and two females (341 and
<quantity id="E2399B08B973FFB48F698AFB28F8388F" box="[351,444,1665,1690]" metricMagnitude="-1" metricUnit="m" metricValue="2.45" pageId="9" pageNumber="542" unit="mm" value="245.0">245 mm</quantity>
TL).
</paragraph>
<paragraph id="257E36EDB973FFB48EF18ADF2BD238F4" blockId="9.[151,1436,1485,2014]" pageId="9" pageNumber="542">
<emphasis id="17B5EAFFB973FFB48EF18ADF2B4F38A8" bold="true" box="[199,523,1701,1726]" pageId="9" pageNumber="542">Geographical distribution.</emphasis>
The
<typeStatus id="FA7A884FB973FFB48C678ADC2BCB38A8" box="[593,655,1702,1726]" pageId="9" pageNumber="542">types</typeStatus>
, and only known specimens, were collected off
<collectingCountry id="5DD6767DB973FFB48AEA8ADC2C0D38A8" box="[1244,1353,1702,1726]" name="Myanmar" pageId="9" pageNumber="542">Myanmar</collectingCountry>
in the Andaman Sea at depths of
<quantity id="E2399B08B973FFB48FF28AB32B7138F7" box="[452,565,1737,1762]" metricMagnitude="2" metricUnit="m" metricValue="1.08" metricValueMax="1.58" metricValueMin="0.58" pageId="9" pageNumber="542" unit="m" value="108.0" valueMax="158.0" valueMin="58.0">58–158 m</quantity>
(
<figureCitation id="BDFA2A68B973FFB48C728AB32BCC38F4" box="[580,648,1737,1762]" captionStart="FIGURE 9" captionStartId="10.[151,250,1269,1291]" captionTargetBox="[340,1246,555,1247]" captionTargetId="figure@10.[340,1247,555,1247]" captionTargetPageId="10" captionText="FIGURE 9. Map showing capture locations for the holotype (red star) and paratypes (red circles) of Platyrhina psomadakisi sp. nov. (Map data © 2016 Terrametrics, Google Earth)." httpUri="https://zenodo.org/record/256729/files/figure.png" pageId="9" pageNumber="542">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6DDB6566B973FFB48EF18A972C5739CB" pageId="9" pageNumber="542" type="etymology">
<paragraph id="257E36EDB973FFB48EF18A972D593958" blockId="9.[151,1436,1485,2014]" pageId="9" pageNumber="542">
<emphasis id="17B5EAFFB973FFB48EF18A97280A3910" bold="true" box="[199,334,1773,1798]" pageId="9" pageNumber="542">Etymology.</emphasis>
Named after Dr Peter N. Psomadakis whose efforts to obtain valuable specimens of sharks and rays, as well as many teleosts, from a number of poorly sampled locations in the northern Indian Ocean have greatly improved our understanding of the biogeography of this complex region.
</paragraph>
<paragraph id="257E36EDB973FFB48EF18B232C5739CB" blockId="9.[151,1436,1485,2014]" pageId="9" pageNumber="542">
<emphasis id="17B5EAFFB973FFB48EF18B232B5F3964" bold="true" box="[199,539,1881,1906]" pageId="9" pageNumber="542">Comparison with congeners.</emphasis>
<taxonomicName id="E2C14D6EB973FFB48C178B232A6F3967" box="[545,811,1881,1905]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="542" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB973FFB48C178B232A6F3967" box="[545,811,1881,1905]" italics="true" pageId="9" pageNumber="542">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB973FFB48D078B232ACE3964" bold="true" box="[817,906,1881,1906]" pageId="9" pageNumber="542">
<taxonomicNameLabel id="0C865784B973FFB48D078B232ACE3964" box="[817,906,1881,1906]" pageId="9" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be readily distinguished from
<taxonomicName id="E2C14D6EB973FFB48B3A8B202CC43967" box="[1292,1408,1882,1905]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="542" phylum="Chordata" rank="species" species="sinensis">
<emphasis id="17B5EAFFB973FFB48B3A8B202CC43967" box="[1292,1408,1882,1905]" italics="true" pageId="9" pageNumber="542">P. sinensis</emphasis>
</taxonomicName>
in having a single row of hooked thorns on the dorsal midline of tail (
<emphasis id="17B5EAFFB973FFB48DBC8B042AE53983" box="[906,929,1918,1941]" italics="true" pageId="9" pageNumber="542">vs</emphasis>
. 2 rows in
<emphasis id="17B5EAFFB973FFB48A168B042DD53983" box="[1056,1169,1918,1941]" italics="true" pageId="9" pageNumber="542">P. s i ne n s i s</emphasis>
). It also differs from
<taxonomicName id="E2C14D6EB973FFB48BB18B0429B439AF" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="542" phylum="Chordata" rank="species" species="sinensis">
<emphasis id="17B5EAFFB973FFB48BB18B0429B439AF" italics="true" pageId="9" pageNumber="542">P. sinensis</emphasis>
</taxonomicName>
in: having a much shorter snout (preoral length 11.9–13.3
<emphasis id="17B5EAFFB973FFB48DBE8BD82ADB39AF" box="[904,927,1954,1977]" italics="true" pageId="9" pageNumber="542">vs</emphasis>
. 15–17% TL; preorbital length 10.4–10.9
<emphasis id="17B5EAFFB973FFB48B4B8BD82CD039AF" box="[1405,1428,1954,1977]" italics="true" pageId="9" pageNumber="542">vs</emphasis>
. 15–16% TL) and distance from pelvic-fin insertion to first dorsal-fin origin 12.2–15.3
<emphasis id="17B5EAFFB973FFB48A568BBC2D3339CB" box="[1120,1143,1990,2013]" italics="true" pageId="9" pageNumber="542">vs</emphasis>
. 17–25% TL.
</paragraph>
</subSubSection>
<subSubSection id="6DDB6566B970FFB78EF18CE22C083FE2" pageId="10" pageNumber="543" type="reference_group">
<paragraph id="257E36EDB970FFB78EF18CE22DA13FBB" blockId="10.[151,1437,151,501]" pageId="10" pageNumber="543">
<taxonomicName id="E2C14D6EB970FFB78EF18CE228943EA6" box="[199,464,152,176]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="543" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB970FFB78EF18CE228943EA6" box="[199,464,152,176]" italics="true" pageId="10" pageNumber="543">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB970FFB78FE18CED2B743EA6" bold="true" box="[471,560,151,176]" pageId="10" pageNumber="543">
<taxonomicNameLabel id="0C865784B970FFB78FE18CED2B743EA6" box="[471,560,151,176]" pageId="10" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="E2C14D6EB970FFB78CF48CE32A1D3EA6" box="[706,857,152,176]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="543" phylum="Chordata" rank="species" species="hyugaensis">
<emphasis id="17B5EAFFB970FFB78CF48CE32A1D3EA6" box="[706,857,152,176]" italics="true" pageId="10" pageNumber="543">P. hyugaensis</emphasis>
</taxonomicName>
,
<emphasis id="17B5EAFFB970FFB78D518CE32A923EA6" box="[871,982,152,176]" italics="true" pageId="10" pageNumber="543">P. s i ne n s i s</emphasis>
and
<taxonomicName id="E2C14D6EB970FFB78A268CE32D213EB9" box="[1040,1125,152,176]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="543" phylum="Chordata" rank="species" species="tangi">
<emphasis id="17B5EAFFB970FFB78A268CE32D213EB9" box="[1040,1125,152,176]" italics="true" pageId="10" pageNumber="543">P. tangi</emphasis>
</taxonomicName>
in the following characters: dorsal fins much more widely separated (interdorsal space 8.5–9.1
<emphasis id="17B5EAFFB970FFB78DBF8CC72AE43EC2" box="[905,928,189,212]" italics="true" pageId="10" pageNumber="543">vs</emphasis>
. 4–6% TL), more pectoral radials (64–66
<emphasis id="17B5EAFFB970FFB78B4B8CC72CD03EC2" box="[1405,1428,189,212]" italics="true" pageId="10" pageNumber="543">vs</emphasis>
. 56–61), and a series of darker brown bands on body and tail (
<emphasis id="17B5EAFFB970FFB78D708C9B2A193EEE" box="[838,861,225,248]" italics="true" pageId="10" pageNumber="543">vs</emphasis>
. no banding evident). The
<typeStatus id="FA7A884FB970FFB78ABA8C9A2D833EEE" box="[1164,1223,224,248]" pageId="10" pageNumber="543">types</typeStatus>
of the new species also have more vertebrae than the three Pacific species based on data presented in
<bibRefCitation id="41504B1CB970FFB78A088D7E2C773F0A" box="[1086,1331,260,285]" pageId="10" pageNumber="543" refString="Iwatsuki, Y., Miyamoto, K., Nakaya, K. &amp; Zhang, J. (2011) A review of the genus Platyrhina (Chondrichthys: Platyrhinidae) from the northwestern Pacific, with descriptions of two new species. Zootaxa, 2738, 26 - 40." type="journal article">
Iwatsuki
<emphasis id="17B5EAFFB970FFB78A908D7F2DA43F0A" box="[1190,1248,260,284]" italics="true" pageId="10" pageNumber="543">et al.</emphasis>
(2011)
</bibRefCitation>
, i.e. total centra 137–139
<emphasis id="17B5EAFFB970FFB78F788D5328213F56" box="[334,357,297,320]" italics="true" pageId="10" pageNumber="543">vs</emphasis>
. 125–134. However, two
<taxonomicName id="E2C14D6EB970FFB78CBB8D532BA73F29" box="[653,739,296,320]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="543" phylum="Chordata" rank="species" species="tangi">
<emphasis id="17B5EAFFB970FFB78CBB8D532BA73F29" box="[653,739,296,320]" italics="true" pageId="10" pageNumber="543">P. tangi</emphasis>
</taxonomicName>
specimens (CSIRO H 7395-13 and -14) from
<collectingCountry id="5DD6767DB970FFB78AC08D5D2C0F3F56" box="[1270,1355,295,320]" name="Taiwan" pageId="10" pageNumber="543">Taiwan</collectingCountry>
, which were also radiographed in this study were both found to have 140 total centra. This discrepancy needs to be further investigated to determine whether this extends the range of total centra for
<taxonomicName id="E2C14D6EB970FFB78A358D0B2D183F91" box="[1027,1116,368,392]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="543" phylum="Chordata" rank="species" species="tangi">
<emphasis id="17B5EAFFB970FFB78A358D0B2D183F91" box="[1027,1116,368,392]" italics="true" pageId="10" pageNumber="543">P. tangi</emphasis>
</taxonomicName>
(reported to be
<quantity id="E2399B08B970FFB78B2C8D0A2CD83F9E" box="[1306,1436,367,392]" metricMagnitude="0" metricUnit="m" metricValue="3.3909" metricValueMax="3.4036" metricValueMin="3.3781999999999996" pageId="10" pageNumber="543" unit="in" value="133.5" valueMax="134.0" valueMin="133.0">133–134 in</quantity>
<bibRefCitation id="41504B1CB970FFB78EA18DEE28393FBA" author="Iwatsuki" box="[151,381,404,429]" pageId="10" pageNumber="543" refString="Iwatsuki, Y., Miyamoto, K., Nakaya, K. &amp; Zhang, J. (2011) A review of the genus Platyrhina (Chondrichthys: Platyrhinidae) from the northwestern Pacific, with descriptions of two new species. Zootaxa, 2738, 26 - 40." type="journal article" year="2011">
Iwatsuki
<emphasis id="17B5EAFFB970FFB78EC88DEF28723FBA" box="[254,310,404,428]" italics="true" pageId="10" pageNumber="543">et al.</emphasis>
, 2011
</bibRefCitation>
) or whether there are differences in count methodology or radiograph quality.
</paragraph>
<paragraph id="257E36EDB970FFB78EF18DC22C083FE2" blockId="10.[151,1437,151,501]" pageId="10" pageNumber="543">
<taxonomicName id="E2C14D6EB970FFB78EF18DC228963FC6" box="[199,466,440,464]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="543" phylum="Chordata" rank="species" species="psomadakisi" status="sp. nov.">
<emphasis id="17B5EAFFB970FFB78EF18DC228963FC6" box="[199,466,440,464]" italics="true" pageId="10" pageNumber="543">Platyrhina psomadakisi</emphasis>
</taxonomicName>
<emphasis id="17B5EAFFB970FFB78FEC8DCD2B713FC6" bold="true" box="[474,565,439,464]" pageId="10" pageNumber="543">
<taxonomicNameLabel id="0C865784B970FFB78FEC8DCD2B713FC6" box="[474,565,439,464]" pageId="10" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also easily distinguished from
<taxonomicName id="E2C14D6EB970FFB78D848DC32D4D3FD9" box="[946,1033,440,464]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="543" phylum="Chordata" rank="species" species="tangi">
<emphasis id="17B5EAFFB970FFB78D848DC32D4D3FD9" box="[946,1033,440,464]" italics="true" pageId="10" pageNumber="543">P. tangi</emphasis>
</taxonomicName>
in lacking pale pigment around the hooked thorns on the orbital, nape and scapular regions (
<emphasis id="17B5EAFFB970FFB78D258DA72A6E3FE2" box="[787,810,477,500]" italics="true" pageId="10" pageNumber="543">vs</emphasis>
. distinctly encircled in pale pigment in
<taxonomicName id="E2C14D6EB970FFB78AD18DA72C7F3FE2" box="[1255,1339,477,500]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="543" phylum="Chordata" rank="species" species="tangi">
<emphasis id="17B5EAFFB970FFB78AD18DA72C7F3FE2" box="[1255,1339,477,500]" italics="true" pageId="10" pageNumber="543">P. tangi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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