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<document id="7EE6EE25C4CB5A1B53054ED42D19A89B" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629626592536" checkinUser="felipe" docAuthor="Ciplak, Battal, Heller, Klaus-Gerhard &amp; Willemse, Fer" docDate="2009" docId="AD38C73DB902FFDEFF70ED15FD37EFEC" docLanguage="en" docName="zt02156p077.pdf" docOrigin="Zootaxa 2156" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Eupholidoptera werneri Rme, Ramme 1951" docType="treatment" docVersion="3" lastPageNumber="23" masterDocId="5101BF45B917FFC8FFE7E85BFFA8ED23" masterDocTitle="2156" masterLastPageNumber="75" masterPageNumber="1" pageNumber="22" updateTime="1699086057653" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:namePart id="90B7FB4017BFC1DB5529A4932EDE15B5">Ciplak, Battal</mods:namePart>
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<mods:namePart id="D4235B061D9129D943C7B001B56AD68F">Heller, Klaus-Gerhard</mods:namePart>
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<mods:namePart id="C31383DDD4592FF71286272ABAEA85E7">Willemse, Fer</mods:namePart>
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<mods:date id="9426F1EF9323293FA507CCBAC139DD46">2009</mods:date>
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<mods:number id="38FF2A22310B2AF8F8103E18E1D9A66A">2009-07-13</mods:number>
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<treatment id="AD38C73DB902FFDEFF70ED15FD37EFEC" ID-DOI="http://doi.org/10.5281/zenodo.5319580" ID-Zenodo-Dep="5319580" LSID="urn:lsid:plazi:treatment:AD38C73DB902FFDEFF70ED15FD37EFEC" httpUri="http://treatment.plazi.org/id/AD38C73DB902FFDEFF70ED15FD37EFEC" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<subSubSection id="6D8B25A0B902FFDDFF70ED15FDC1E84B" box="[151,617,1358,1384]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="252E762BB902FFDDFF70ED15FDC1E84B" blockId="21.[151,703,1358,1419]" box="[151,617,1358,1384]" pageId="21" pageNumber="22">
<heading id="7E66C147B902FFDDFF70ED15FDC1E84B" bold="true" box="[151,617,1358,1384]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="E2910DA8B902FFDDFF70ED15FDC1E84B" ID-CoL="3CMLK" authority="Ramme, 1951" authorityName="Rme, Ramme" authorityYear="1951" box="[151,617,1358,1384]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B902FFDDFF70ED15FDC1E84B" bold="true" box="[151,617,1358,1384]" pageId="21" pageNumber="22">
<emphasis id="17E5AA39B902FFDDFF70ED15FE10E84B" bold="true" box="[151,440,1358,1384]" italics="true" pageId="21" pageNumber="22">Eupholidoptera werneri</emphasis>
<bibRefCitation id="41000BDAB902FFDDFE58ED15FDC1E84B" author="Ramme, W." box="[447,617,1358,1384]" pageId="21" pageNumber="22" pagination="1 - 431" refId="ref44141" refString="Ramme, W. (1951) Zur Systematik, Faunistik und Biologie der Orthopteren von Sudost-Europa und Vorderasiens. Mitteilungen aus dem Zoologischen Museum in Berlin, 27, 1 - 431." type="journal article" year="1951">Ramme, 1951</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6D8B25A0B902FFDDFF70ED2AFD17E8A8" box="[151,703,1393,1419]" pageId="21" pageNumber="22" type="description">
<paragraph id="252E762BB902FFDDFF70ED2AFD17E8A8" blockId="21.[151,703,1358,1419]" box="[151,703,1393,1419]" pageId="21" pageNumber="22">
(
<figureCitation id="BDAA6AAEB902FFDDFF78ED2AFE88E8A8" box="[159,288,1393,1419]" captionStart="FIGURES 6–19" captionStartId="12.[151,270,1745,1769]" captionTargetBox="[219,1366,223,1641]" captionTargetId="figure-143@12.[188,1398,197,1720]" captionTargetPageId="12" captionText="FIGURES 6–19. Male cerci of Eupholidoptera species (Scale= 2 mm). 6—E. peneri, 7—E. pallipes, 8—E. gemellata, 9—E. akdeniz, 10—E. annulipes, 11—E. marashensis (A, Kahramanmaras; B, Adana), 12—E. uvarovi (A and B—different forms from Karpathos), 13—E. palaestinensis (A–C, different forms from the range of species), 14—E. lyra (A and B, different forms from the range of species), 15—E. ledereri, 16—E. werneri, 17—E. cypria (Cyprus), 18—E. helina, 19—E. smyrnensis (A, Turkey; B and C, Greece) (Of the figures given above: 6–8 from Willemse &amp; Kruseman, 1976; 9 from Ünal, 2002; 13A, 14–15 from Ramme, 1951; 11A, 18B–C, 19A–C, from Willemse 1980)." pageId="21" pageNumber="22">Figures 16</figureCitation>
,
<figureCitation id="BDAA6AAEB902FFDDFECBED2AFEE3E8A8" box="[300,331,1393,1419]" captionStart="FIGURES 52–64" captionStartId="15.[151,272,1529,1553]" captionTargetBox="[229,1354,189,1489]" captionTargetId="figure-156@15.[188,1398,148,1559]" captionTargetPageId="15" captionText="FIGURES 52–64. Male anal tergite of Eupholidoptera species (d—dorsal, c—caudal, L—lateral) (Scale = 2 mm). 52—E. peneri, 53—E. pallipes, 54—E. akdeniz, 55—E. annulipes, 56—E. marashensis (A, Kahramanmaras; B, Adana), 57—E. uvarovi, 58—E. palaestinensis (A, B and C different forms from the range of the species), 59—E. lyra, 60—E. ledereri, 61—E. werneri, 62—E. cypria (A, Turkey; B, Cyprus), 63—E. helina, 64—E. smyrnensis (Of the figures given above: 62, 63 from Ramme, 1951; 54 from Ünal, 2002)." pageId="21" pageNumber="22">61</figureCitation>
,
<figureCitation id="BDAA6AAEB902FFDDFEBFED2AFE2EE8A8" box="[344,390,1393,1419]" captionStart="FIGURES 97–110" captionStartId="23.[151,269,1827,1851]" captionTargetBox="[193,1386,190,1787]" captionTargetId="figure-103@23.[188,1398,159,1842]" captionTargetPageId="23" captionText="FIGURES 97–110. Male subgenital plate (and spines under the styli) of Eupholidoptera species (L— lateral view, V— ventral view) (Scale = 2 mm). 97— E. peneri, 98— E. pallipes, 99—E. gemellata, 100— E. akdeniz, 101—E. annulipes, 102—E. marashensis (A, Kahramanmaras; B, Adana), 103—E. uvarovi, 104—E. palaestinensis, 105—E. lyra, 106—E. ledereri, 107—E. werneri, 108—E. cypria, 109— E. helina, 110—E. smyrnensis (Of the figures given above: 98–99 are from Willemse &amp; Heller, 2001; 100 from Ünal, 2002)." pageId="21" pageNumber="22">107</figureCitation>
,
<figureCitation id="BDAA6AAEB902FFDDFE75ED2AFE68E8A8" box="[402,448,1393,1419]" captionStart="FIGURES 149–153" captionStartId="33.[151,269,1857,1881]" captionTargetBox="[208,1376,199,1825]" captionTargetId="figure-59@33.[188,1398,116,1953]" captionTargetPageId="33" captionText="FIGURES 149–153. Titillators of Eupholidoptera species (d— Dorsal view, L— lateral view, V— ventral view) (Scale= 2 mm). 149—E. uvarovi, 150—E. palaestinensis, 151—E. lyra, 152—E. ledereri, 153—E. werneri (Of the figures given above 152, 153 from Ramme, 1951)." pageId="21" pageNumber="22">153</figureCitation>
,
<figureCitation id="BDAA6AAEB902FFDDFE2AED2AFE53E8A8" box="[461,507,1393,1419]" captionStart="FIGURES 189–200" captionStartId="45.[151,270,1867,1891]" captionTargetBox="[264,1348,186,1844]" captionTargetId="figure-96@45.[254,1350,176,1861]" captionTargetPageId="45" captionText="FIGURES 189–200. Female subgenital plate of Eupholidoptera species (L— lateral view, V— ventral view) (Scale=2 mm). 189— E. peneri, 190— E. pallipes, 191—E. annulipes (A and B, different forms from the range), 192—E. marashensis, 193—E. uvarovi, 194—E. palaestinensis, 195—E. lyra, 196—E. ledereri, 197—E. werneri, 198—E. cypria (A, Turkey; B, Cyprus), 199— E. helina, 200— E. smyrnensis (Of the figures given above: 190 from Willemse &amp; Heller, 2001; 196 from Ramme, 1951)." pageId="21" pageNumber="22">197</figureCitation>
,
<figureCitation id="BDAA6AAEB902FFDDFDE0ED2AFD9DE8A8" box="[519,565,1393,1419]" captionStart="FIGURES 237–238" captionStartId="55.[151,271,1931,1955]" captionTargetBox="[151,1433,201,1907]" captionTargetPageId="55" captionText="FIGURES 237–238. Distribution maps of the species belonging to the E. chabrieri group. 237—Distribution of the species six European species from E. chabrieri group, 238— Distribution of the Aegean, Anatolian and Levatine species of E. chabrieri group." pageId="21" pageNumber="22">237</figureCitation>
, Appendix)
</paragraph>
</subSubSection>
<subSubSection id="6D8B25A0B902FFDDFF70EDE5FD0EE8F6" box="[151,678,1470,1493]" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="252E762BB902FFDDFF70EDE5FD0EE8F6" blockId="21.[151,678,1470,1493]" box="[151,678,1470,1493]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="05815105B902FFDDFF70EDE5FD0EE8F6" box="[151,678,1470,1493]" pageId="21" pageNumber="22">
<treatmentCitation id="A430503AB902FFDDFF70EDE5FD0AE8F6" author="Ramme, W." box="[151,674,1470,1493]" page="202" pageId="21" pageNumber="22" year="1951">
<taxonomicName id="E2910DA8B902FFDDFF70EDE5FD0AE8F6" ID-CoL="3CMLK" authority="Ramme, 1951: 201 - 202" authorityName="Rme, Ramme" authorityPageNumber="201 - 202" authorityYear="1951" box="[151,674,1470,1493]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B902FFDDFF70EDE5FE30E8F6" box="[151,408,1470,1493]" italics="true" pageId="21" pageNumber="22">Eupholidoptera werneri</emphasis>
<bibRefCitation id="41000BDAB902FFDDFE79EDE5FD0AE8F6" author="Ramme, W." box="[414,674,1470,1493]" pageId="21" pageNumber="22" pagination="1 - 431" refId="ref44141" refString="Ramme, W. (1951) Zur Systematik, Faunistik und Biologie der Orthopteren von Sudost-Europa und Vorderasiens. Mitteilungen aus dem Zoologischen Museum in Berlin, 27, 1 - 431." type="journal article" year="1951">Ramme, 1951: 201–202</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6D8B25A0B902FFDDFF70EE5EFD13EB4F" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="252E762BB902FFDDFF70EE5EFC6FEB65" blockId="21.[151,1436,1541,2031]" pageId="21" pageNumber="22">
<emphasis id="17E5AA39B902FFDDFF70EE5EFED6EB3C" bold="true" box="[151,382,1541,1567]" pageId="21" pageNumber="22">Type information:</emphasis>
<materialsCitation id="95F97C76B902FFDDFE60EE5EFBE6EB3C" box="[391,1102,1541,1567]" collectingDate="1935-05-19" collectionCode="NHW" collectorName="F. Werner" country="Syria" location="Abde" pageId="21" pageNumber="22" specimenCount="1" typeStatus="holotype">
<typeStatus id="FA2AC889B902FFDDFE60EE5EFE5DEB3C" box="[391,501,1541,1567]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
M,
<collectingCountry id="5D8636BBB902FFDDFDCFEE5EFD2CEB3C" box="[552,644,1541,1567]" name="Syria" pageId="21" pageNumber="22">SYRIA</collectingCountry>
,
<location id="204E20F0B902FFDDFD77EE5EFD7BEB3C" LSID="urn:lsid:plazi:treatment:AD38C73DB902FFDEFF70ED15FD37EFEC:204E20F0B902FFDDFD77EE5EFD7BEB3C" box="[656,723,1541,1567]" country="Syria" name="Abde" pageId="21" pageNumber="22">Abde</location>
,
<date id="512F50EBB902FFDDFD07EE5EFCFFEB3C" box="[736,855,1541,1567]" pageId="21" pageNumber="22" value="1935-05-19">
<collectingDate id="416BA903B902FFDDFD07EE5EFCFFEB3C" box="[736,855,1541,1567]" pageId="21" pageNumber="22" value="1935-05-19">19.5.1935</collectingDate>
</date>
(
<collectorName id="886413FDB902FFDDFC8EEE5EFC49EB3C" box="[873,993,1541,1567]" pageId="21" pageNumber="22">F. Werner</collectorName>
) (
<collectionCode id="4380EEEEB902FFDDFC1EEE5EFBEEEB3C" box="[1017,1094,1541,1567]" pageId="21" pageNumber="22">NHW</collectionCode>
)
</materialsCitation>
;
<typeStatus id="FA2AC889B902FFDDFBBCEE5EFB6BEB3C" box="[1115,1219,1541,1567]" pageId="21" pageNumber="22" type="paratype">Paratype</typeStatus>
1M, same data as holotype;
<materialsCitation id="95F97C76B902FFDDFEEAEE77FC6CEB65" box="[269,964,1580,1606]" collectionCode="ZMB" country="Syria" pageId="21" pageNumber="22" specimenCount="1" typeStatus="paratype">
<typeStatus id="FA2AC889B902FFDDFEEAEE77FEDBEB65" box="[269,371,1580,1606]" pageId="21" pageNumber="22" type="paratype">Paratype</typeStatus>
1M (labelled as “Typus von
<taxonomicName id="E2910DA8B902FFDDFD20EE77FCF9EB65" authority="II" authorityName="II" box="[711,849,1580,1606]" class="Insecta" family="Tettigoniidae" genus="Pholidoptera" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="chabrieri">
<emphasis id="17E5AA39B902FFDDFD20EE77FC9CEB66" box="[711,820,1580,1605]" italics="true" pageId="21" pageNumber="22">chabrieri</emphasis>
II
</taxonomicName>
”) (
<collectionCode id="4380EEEEB902FFDDFC92EE77FC12EB65" box="[885,954,1580,1606]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="21" pageNumber="22" type="Museum">ZMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="252E762BB902FFDDFF22EE09FD13EB4F" blockId="21.[151,1436,1541,2031]" box="[197,699,1618,1644]" pageId="21" pageNumber="22">
<emphasis id="17E5AA39B902FFDDFF22EE09FE13EB4F" bold="true" box="[197,443,1618,1644]" pageId="21" pageNumber="22">Material examined:</emphasis>
<materialsCitation id="95F97C76B902FFDDFE24EE09FD10EB4F" box="[451,696,1618,1644]" collectionCode="ZMB" pageId="21" pageNumber="22" specimenCount="1" typeStatus="paratype">
<typeStatus id="FA2AC889B902FFDDFE24EE09FD82EB4F" box="[451,554,1618,1644]" pageId="21" pageNumber="22" type="paratype">Paratype</typeStatus>
1M (
<collectionCode id="4380EEEEB902FFDDFD8EEE09FD06EB4F" box="[617,686,1618,1644]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="21" pageNumber="22" type="Museum">ZMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6D8B25A0B902FFDDFF22EE23FAE9EBB0" box="[197,1345,1656,1683]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="252E762BB902FFDDFF22EE23FAE9EBB0" blockId="21.[151,1436,1541,2031]" box="[197,1345,1656,1683]" pageId="21" pageNumber="22">
<emphasis id="17E5AA39B902FFDDFF22EE23FEF6EBB1" bold="true" box="[197,350,1656,1682]" pageId="21" pageNumber="22">Distribution</emphasis>
: This species is known only from the
<typeStatus id="FA2AC889B902FFDDFCFAEE22FCE7EBB0" box="[797,847,1657,1683]" pageId="21" pageNumber="22">type</typeStatus>
material from
<collectingCountry id="5D8636BBB902FFDDFC19EE22FB97EBB0" box="[1022,1087,1657,1683]" name="Syria" pageId="21" pageNumber="22">Syria</collectingCountry>
,
<collectingCountry id="5D8636BBB902FFDDFBACEE22FB1AEBB0" box="[1099,1202,1657,1683]" name="Lebanon" pageId="21" pageNumber="22">Lebanon</collectingCountry>
and
<collectingCountry id="5D8636BBB902FFDDFB0CEE22FA95EBB0" box="[1259,1341,1657,1683]" name="Jordan" pageId="21" pageNumber="22">Jordan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6D8B25A0B902FFDEFF22EEC4FD37EFEC" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="discussion">
<paragraph id="252E762BB902FFDEFF22EEC4FD08EC6F" blockId="21.[151,1436,1541,2031]" lastBlockId="22.[151,1437,152,719]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="17E5AA39B902FFDDFF22EEC4FE93EB9A" bold="true" box="[197,315,1695,1721]" pageId="21" pageNumber="22">Remarks.</emphasis>
<bibRefCitation id="41000BDAB902FFDDFEA4EEFBFE59EB99" author="Ramme, W." box="[323,497,1696,1722]" pageId="21" pageNumber="22" pagination="1 - 431" refId="ref44141" refString="Ramme, W. (1951) Zur Systematik, Faunistik und Biologie der Orthopteren von Sudost-Europa und Vorderasiens. Mitteilungen aus dem Zoologischen Museum in Berlin, 27, 1 - 431." type="journal article" year="1951">Ramme (1951)</bibRefCitation>
described
<taxonomicName id="E2910DA8B902FFDDFD8BEEFBFD4EEB9A" authorityName="Rme, Ramme" authorityYear="1951" box="[620,742,1696,1721]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B902FFDDFD8BEEFBFD4EEB9A" box="[620,742,1696,1721]" italics="true" pageId="21" pageNumber="22">E. werneri</emphasis>
</taxonomicName>
from Abde,
<collectingCountry id="5D8636BBB902FFDDFC9EEEFBFC10EB99" box="[889,952,1696,1722]" name="Syria" pageId="21" pageNumber="22">Syria</collectingCountry>
. The description was based only on males and the original description only lists males (
<typeStatus id="FA2AC889B902FFDDFD72EE9DFD55EBC3" box="[661,765,1734,1760]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
,
<typeStatus id="FA2AC889B902FFDDFCEEEE9DFCC5EBC3" box="[777,877,1734,1760]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
) under material examined. Although Ramme did not mention any female there is a figure of a female subgenital plate in
<bibRefCitation id="41000BDAB902FFDDFC58EEB6FBCEEA24" author="Ramme, W." box="[959,1126,1773,1799]" pageId="21" pageNumber="22" pagination="1 - 431" refId="ref44141" refString="Ramme, W. (1951) Zur Systematik, Faunistik und Biologie der Orthopteren von Sudost-Europa und Vorderasiens. Mitteilungen aus dem Zoologischen Museum in Berlin, 27, 1 - 431." type="journal article" year="1951">Ramme (1951</bibRefCitation>
: p. 202). In NHW there are two more specimens of
<taxonomicName id="E2910DA8B902FFDDFE57EF4FFD84EA0E" authorityName="Rme, Ramme" authorityYear="1951" box="[432,556,1812,1837]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B902FFDDFE57EF4FFD84EA0E" box="[432,556,1812,1837]" italics="true" pageId="21" pageNumber="22">E. werneri</emphasis>
</taxonomicName>
: (1F with the 4 labels (i–iv) (i) coll. Br. V. W. = coll Lederer, Beirut; (ii) 1874; (iii) 1874 Chabrieri nach Fieber, and (iv)
<taxonomicName id="E2910DA8B902FFDDFD2BEF60FB38EA77" authority="Rme, Ramme" authorityName="Rme, Ramme" authorityYear="1951" box="[716,1168,1850,1876]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B902FFDDFD2BEF60FC4FEA77" box="[716,999,1851,1876]" italics="true" pageId="21" pageNumber="22">Eupholidoptera werneri</emphasis>
Rme, Ramme
</taxonomicName>
det.; 1M with 2 labels (i–ii); (i) Saida Oberthuer, Coll. B. v. W, and (ii)
<taxonomicName id="E2910DA8B902FFDDFCE5EF3AFB48EA58" authority="Rme, Ramme" authorityName="Rme, Ramme" authorityYear="1951" box="[770,1248,1889,1915]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B902FFDDFCE5EF3AFB86EA59" box="[770,1070,1889,1914]" italics="true" pageId="21" pageNumber="22">Eupholidoptera werneri</emphasis>
Rme, Ramme
</taxonomicName>
det.) (Susanne Randolf; personal communication) which were identified as
<taxonomicName id="E2910DA8B902FFDDFCB4EFD3FC63EA82" authorityName="Rme, Ramme" authorityYear="1951" box="[851,971,1928,1953]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B902FFDDFCB4EFD3FC63EA82" box="[851,971,1928,1953]" italics="true" pageId="21" pageNumber="22">E. werneri</emphasis>
</taxonomicName>
by Ramme, though not mentioned in the original description.
<bibRefCitation id="41000BDAB902FFDDFE61EFF5FD98EAEB" author="Salman, S." box="[390,560,1966,1992]" pageId="21" pageNumber="22" pagination="313 - 338" refId="ref44225" refString="Salman, S. (1983) Turkish bush-crickets of the genus Euphlidoptera (Decticinae). Systematic Entomology, 8, 313 - 338." type="journal article" year="1983">Salman (1983)</bibRefCitation>
reported this species from South
<collectingCountry id="5D8636BBB902FFDDFC51EFF5FBB1EAEB" box="[950,1049,1966,1992]" name="Turkey" pageId="21" pageNumber="22">Anatolia</collectingCountry>
(
<collectingCountry id="5D8636BBB902FFDDFBCFEFF5FBD8EAEB" box="[1064,1136,1966,1992]" name="Turkey" pageId="21" pageNumber="22">Hatay</collectingCountry>
) assuming to describe the female for the first time (but see above). Later,
<bibRefCitation id="41000BDAB902FFDDFD27EF8EFC41EACC" author="Naskrecki, P. &amp; Unal M." box="[704,1001,2005,2031]" pageId="21" pageNumber="22" pagination="393 - 420" refId="ref43721" refString="Naskrecki, P. &amp; Unal M. (1995) The Orthoptera of Hatay province, S. Turkey. Beitrage zur Entomologie, 45, 393 - 420." type="journal article" year="1995">Naskrecki &amp; Ünal (1995)</bibRefCitation>
reported
<specimenCount id="3397BDA2B902FFDDFBBEEF8EFB43EACC" box="[1113,1259,2005,2031]" pageId="21" pageNumber="22" type="juvenile">six juveniles</specimenCount>
and
<specimenCount id="3397BDA2B902FFDDFAC2EF8EFA34EACC" box="[1317,1436,2005,2031]" pageId="21" pageNumber="22" type="male">two males</specimenCount>
assigned to
<taxonomicName id="E2910DA8B901FFDEFEC3E8C3FE09ED92" authorityName="Rme, Ramme" authorityYear="1951" box="[292,417,152,177]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B901FFDEFEC3E8C3FE09ED92" box="[292,417,152,177]" italics="true" pageId="22" pageNumber="23">E. werneri</emphasis>
</taxonomicName>
from the same province of
<collectingCountry id="5D8636BBB901FFDEFD0CE8C3FCE9ED91" box="[747,833,152,178]" name="Turkey" pageId="22" pageNumber="23">Turkey</collectingCountry>
. However, later Salman (personal communication) decided that the specimens from South
<collectingCountry id="5D8636BBB901FFDEFDBEE8E5FD13EDFB" box="[601,699,190,216]" name="Turkey" pageId="22" pageNumber="23">Anatolia</collectingCountry>
(
<collectingCountry id="5D8636BBB901FFDEFD2DE8E5FCB9EDFB" box="[714,785,190,216]" name="Turkey" pageId="22" pageNumber="23">Hatay</collectingCountry>
) which he identified as
<taxonomicName id="E2910DA8B901FFDEFBC4E8E4FB34EDFB" authorityName="Rme, Ramme" authorityYear="1951" box="[1059,1180,191,216]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B901FFDEFBC4E8E4FB34EDFB" box="[1059,1180,191,216]" italics="true" pageId="22" pageNumber="23">E. werneri</emphasis>
</taxonomicName>
in fact did belong to a new species (After examination of
<specimenCount id="3397BDA2B901FFDEFDD4E8BEFD26EDDC" box="[563,654,229,255]" pageId="22" pageNumber="23" type="male">2 males</specimenCount>
and
<specimenCount id="3397BDA2B901FFDEFD2EE8BEFC92EDDC" box="[713,826,229,255]" pageId="22" pageNumber="23" type="female">2 females</specimenCount>
of these specimens we support his opinion) and the occurrence of
<taxonomicName id="E2910DA8B901FFDEFEDFE957FE18EC06" authorityName="Rme, Ramme" authorityYear="1951" box="[312,432,268,293]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B901FFDEFEDFE957FE18EC06" box="[312,432,268,293]" italics="true" pageId="22" pageNumber="23">E. werneri</emphasis>
</taxonomicName>
in
<collectingCountry id="5D8636BBB901FFDEFE35E957FD8CEC05" box="[466,548,268,294]" name="Turkey" pageId="22" pageNumber="23">Turkey</collectingCountry>
remains doubtful. The specimens from
<bibRefCitation id="41000BDAB901FFDEFC01E957FAAFEC05" author="Naskrecki, P. &amp; Unal M." box="[998,1287,268,294]" pageId="22" pageNumber="23" pagination="393 - 420" refId="ref43721" refString="Naskrecki, P. &amp; Unal M. (1995) The Orthoptera of Hatay province, S. Turkey. Beitrage zur Entomologie, 45, 393 - 420." type="journal article" year="1995">Naskrecki &amp; Ünal (1995)</bibRefCitation>
were used to describe
<taxonomicName id="E2910DA8B901FFDEFF1BE969FD33EC6F" authority="Unal &amp; Naskrecki, 2002" authorityName="Unal &amp; Naskrecki" authorityYear="2002" box="[252,667,306,332]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="akdeniz">
<emphasis id="17E5AA39B901FFDEFF1BE969FEDBEC68" box="[252,371,306,331]" italics="true" pageId="22" pageNumber="23">E. akdeniz</emphasis>
<bibRefCitation id="41000BDAB901FFDEFE9DE969FD33EC6F" author="Unal, M. &amp; Naskrecki, P." box="[378,667,306,332]" pageId="22" pageNumber="23" pagination="1 - 5" refId="ref44719" refString="Unal, M. &amp; Naskrecki, P. (2002) A new bushcricket from southernmost part of Turkey. Miscellaneous papers, 87, 1 - 5." type="journal article" year="2002">Ünal &amp; Naskrecki, 2002</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="252E762BB901FFDEFF22E902FD37EFEC" blockId="22.[151,1437,152,719]" pageId="22" pageNumber="23">
<taxonomicName id="E2910DA8B901FFDEFF22E902FEE2EC51" authorityName="Rme, Ramme" authorityYear="1951" box="[197,330,345,370]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B901FFDEFF22E902FEE2EC51" box="[197,330,345,370]" italics="true" pageId="22" pageNumber="23">E. werneri</emphasis>
</taxonomicName>
and
<taxonomicName id="E2910DA8B901FFDEFE6BE902FDA8EC51" authorityName="Ramme" authorityYear="1951" box="[396,512,345,370]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cypria">
<emphasis id="17E5AA39B901FFDEFE6BE902FDA8EC51" box="[396,512,345,370]" italics="true" pageId="22" pageNumber="23">E. cypria</emphasis>
</taxonomicName>
are two closely related species both of which are described in the same publication by
<bibRefCitation id="41000BDAB901FFDEFEA9E9DBFE54ECB9" author="Ramme, W." box="[334,508,384,410]" pageId="22" pageNumber="23" pagination="1 - 431" refId="ref44141" refString="Ramme, W. (1951) Zur Systematik, Faunistik und Biologie der Orthopteren von Sudost-Europa und Vorderasiens. Mitteilungen aus dem Zoologischen Museum in Berlin, 27, 1 - 431." type="journal article" year="1951">Ramme, 1951</bibRefCitation>
.
<bibRefCitation id="41000BDAB901FFDEFDEFE9DBFD14ECB9" author="Salman, S." box="[520,700,384,410]" pageId="22" pageNumber="23" pagination="313 - 338" refId="ref44225" refString="Salman, S. (1983) Turkish bush-crickets of the genus Euphlidoptera (Decticinae). Systematic Entomology, 8, 313 - 338." type="journal article" year="1983">Salman (1983)</bibRefCitation>
described the mainland population of
<taxonomicName id="E2910DA8B901FFDEFB74E9DBFAAAECBA" authorityName="Ramme" authorityYear="1951" box="[1171,1282,384,409]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cypria">
<emphasis id="17E5AA39B901FFDEFB74E9DBFAAAECBA" box="[1171,1282,384,409]" italics="true" pageId="22" pageNumber="23">E. cypria</emphasis>
</taxonomicName>
from
<collectingCountry id="5D8636BBB901FFDEFAA9E9DBFA3FECB9" box="[1358,1431,384,410]" name="Turkey" pageId="22" pageNumber="23">Hatay</collectingCountry>
,
<collectingCountry id="5D8636BBB901FFDEFF70E9FDFF43ECE3" box="[151,235,422,448]" name="Turkey" pageId="22" pageNumber="23">Turkey</collectingCountry>
as a separate subspecies (
<taxonomicName id="E2910DA8B901FFDEFDFBE9FCFD48ECE3" authorityName="Salman" authorityYear="1983" box="[540,736,423,448]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="cypria" subSpecies="turcica">
<emphasis id="17E5AA39B901FFDEFDFBE9FCFD48ECE3" box="[540,736,423,448]" italics="true" pageId="22" pageNumber="23">E. cypria turcica</emphasis>
</taxonomicName>
) and this subspecies was later recorded from
<collectingRegion id="E755B8C9B901FFDEFB10E9FDFAFAECE3" box="[1271,1362,422,448]" country="Syria" name="Al Ladhiqiyah" pageId="22" pageNumber="23">Latakia</collectingRegion>
,
<collectingCountry id="5D8636BBB901FFDEFAB9E9FDFA34ECE3" box="[1374,1436,422,448]" name="Syria" pageId="22" pageNumber="23">Syria</collectingCountry>
(see above). The main differences between
<taxonomicName id="E2910DA8B901FFDEFD7FE996FCBCECC5" authorityName="Rme, Ramme" authorityYear="1951" box="[664,788,461,486]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B901FFDEFD7FE996FCBCECC5" box="[664,788,461,486]" italics="true" pageId="22" pageNumber="23">E. werneri</emphasis>
</taxonomicName>
and its close relative
<taxonomicName id="E2910DA8B901FFDEFBF1E996FB2AECC5" authorityName="Ramme" authorityYear="1951" box="[1046,1154,461,486]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cypria">
<emphasis id="17E5AA39B901FFDEFBF1E996FB2AECC5" box="[1046,1154,461,486]" italics="true" pageId="22" pageNumber="23">E. cypria</emphasis>
</taxonomicName>
are the shape of female subgenital plate (compare
<figureCitation id="BDAA6AAEB901FFDEFE3AE9AFFD07EF2D" box="[477,687,500,526]" captionStart="FIGURES 189–200" captionStartId="45.[151,270,1867,1891]" captionTargetBox="[264,1348,186,1844]" captionTargetId="figure-96@45.[254,1350,176,1861]" captionTargetPageId="45" captionText="FIGURES 189–200. Female subgenital plate of Eupholidoptera species (L— lateral view, V— ventral view) (Scale=2 mm). 189— E. peneri, 190— E. pallipes, 191—E. annulipes (A and B, different forms from the range), 192—E. marashensis, 193—E. uvarovi, 194—E. palaestinensis, 195—E. lyra, 196—E. ledereri, 197—E. werneri, 198—E. cypria (A, Turkey; B, Cyprus), 199— E. helina, 200— E. smyrnensis (Of the figures given above: 190 from Willemse &amp; Heller, 2001; 196 from Ramme, 1951)." pageId="22" pageNumber="23">Figures 197, 198</figureCitation>
) and the long unfused apical arms of the titillators (compare
<figureCitation id="BDAA6AAEB901FFDEFF70EA41FE82EF17" box="[151,298,538,564]" captionStart="FIGURES 149–153" captionStartId="33.[151,269,1857,1881]" captionTargetBox="[208,1376,199,1825]" captionTargetId="figure-59@33.[188,1398,116,1953]" captionTargetPageId="33" captionText="FIGURES 149–153. Titillators of Eupholidoptera species (d— Dorsal view, L— lateral view, V— ventral view) (Scale= 2 mm). 149—E. uvarovi, 150—E. palaestinensis, 151—E. lyra, 152—E. ledereri, 153—E. werneri (Of the figures given above 152, 153 from Ramme, 1951)." pageId="22" pageNumber="23">Figures 153</figureCitation>
and
<figureCitation id="BDAA6AAEB901FFDEFE8EEA41FE34EF17" box="[361,412,538,564]" captionStart="FIGURES 154–156" captionStartId="34.[151,269,1629,1653]" captionTargetBox="[210,1387,242,1603]" captionTargetId="figure-130@34.[190,1400,220,1654]" captionTargetPageId="34" captionText="FIGURES 154–156. Titillators of Eupholidoptera species (d— Dorsal view, L— lateral view, V— ventral view) (Scale = 2 mm). 154—E. cypria, 155— E. helina, 156—E. smyrnensis (Of the figures given above 156C, 156C from Willemse, 1980)." pageId="22" pageNumber="23">154</figureCitation>
). Since there were no females of
<taxonomicName id="E2910DA8B901FFDEFCA7EA41FC6AEF10" authorityName="Rme, Ramme" authorityYear="1951" box="[832,962,538,563]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B901FFDEFCA7EA41FC6AEF10" box="[832,962,538,563]" italics="true" pageId="22" pageNumber="23">E. werneri</emphasis>
</taxonomicName>
available during this study it was not possible to determine if the female subgenital plates are different as figured by
<bibRefCitation id="41000BDAB901FFDEFBDFEA1AFB44EF78" author="Ramme, W." box="[1080,1260,577,603]" pageId="22" pageNumber="23" pagination="1 - 431" refId="ref44141" refString="Ramme, W. (1951) Zur Systematik, Faunistik und Biologie der Orthopteren von Sudost-Europa und Vorderasiens. Mitteilungen aus dem Zoologischen Museum in Berlin, 27, 1 - 431." type="journal article" year="1951">Ramme (1951)</bibRefCitation>
. Since
<bibRefCitation id="41000BDAB901FFDEFAA7EA1AFF4DEFA1" author="Ramme, W." pageId="22" pageNumber="23" pagination="1 - 431" refId="ref44141" refString="Ramme, W. (1951) Zur Systematik, Faunistik und Biologie der Orthopteren von Sudost-Europa und Vorderasiens. Mitteilungen aus dem Zoologischen Museum in Berlin, 27, 1 - 431." type="journal article" year="1951">Ramme (1951)</bibRefCitation>
described both
<taxonomicName id="E2910DA8B901FFDEFE44EA33FDB8EFA2" authorityName="Ramme" authorityYear="1951" box="[419,528,616,641]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cypria">
<emphasis id="17E5AA39B901FFDEFE44EA33FDB8EFA2" box="[419,528,616,641]" italics="true" pageId="22" pageNumber="23">E. cypria</emphasis>
</taxonomicName>
and
<taxonomicName id="E2910DA8B901FFDEFDACEA33FD60EFA2" authorityName="Rme, Ramme" authorityYear="1951" box="[587,712,616,641]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B901FFDEFDACEA33FD60EFA2" box="[587,712,616,641]" italics="true" pageId="22" pageNumber="23">E. werneri</emphasis>
</taxonomicName>
in the same publication the second species is treated here on the basis of his figures and description. However, the confirmation of
<taxonomicName id="E2910DA8B901FFDEFC2CEAD4FBEEEF8B" authorityName="Rme, Ramme" authorityYear="1951" box="[971,1094,655,680]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="17E5AA39B901FFDEFC2CEAD4FBEEEF8B" box="[971,1094,655,680]" italics="true" pageId="22" pageNumber="23">E. werneri</emphasis>
</taxonomicName>
as a distinct separate species requires new material from the
<typeStatus id="FA2AC889B901FFDEFDE0EAEEFD91EFEC" box="[519,569,693,719]" pageId="22" pageNumber="23">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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