<documentid="8E9B165B21144B81132CBC314EE50022"ID-DOI="10.18590/euscorpius.2006.vol2006.iss36.1"ID-ISSN="1536-9307"ID-Zenodo-Dep="12781855"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1721360289902"checkinUser="felipe"docAuthor="Colombo, Marco"docDate="2006"docId="03F887C26C04FF946747FB6FE7E6F981"docLanguage="en"docName="Euscorpius.36.1-40.pdf"docOrigin="Euscorpius 36 (36)"docSource="https://mds.marshall.edu/euscorpius/vol2006/iss36/1/"docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article"docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A"docStyleName="Euscorpius.2002-2019.journal_article"docStyleVersion="3"docTitle="Euscorpius tergestinus"docType="treatment"docVersion="2"lastPageNumber="17"masterDocId="FFC1FFBA6C0AFF86664CFF83E60AFFCE"masterDocTitle="New data on distribution and ecology of seven species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae)"masterLastPageNumber="40"masterPageNumber="1"pageNumber="13"updateTime="1721381580369"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0">
<mods:titleid="86E6EA562B4B2FD2C1D58B48CFEFB2A6">New data on distribution and ecology of seven species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae)</mods:title>
<taxonomicNameid="4C514D576C04FF886747FB6FE4DCFACD"ID-CoL="3D94G"authority="(C. L. Koch, 1837)"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[267,726,1260,1283]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="tergestinus">
<figureCitationid="136A2A516C04FF8867C5FA84E7F0FAD0"box="[393,506,1287,1310]"captionStart-0="Figure 9"captionStart-1="Figure 10"captionStart-2="Figure 11"captionStartId-0="14.[192,268,1040,1063]"captionStartId-1="15.[192,260,1927,1948]"captionStartId-2="16.[192,268,1786,1809]"captionTargetBox-0="[192,1428,192,997]"captionTargetBox-1="[249,1370,168,1899]"captionTargetBox-2="[189,1441,277,1755]"captionTargetId-0="figure-12@14.[192,1440,192,997]"captionTargetId-1="figure-7@15.[252,1367,171,1896]"captionTargetId-2="figure-13@16.[192,1438,280,1752]"captionTargetPageId-0="14"captionTargetPageId-1="15"captionTargetPageId-2="16"captionText-0="Figure 9: Euscorpius (Euscorpius) tergestinus, adult female, Venezia (Veneto, Italy) (photo by Giorgio Colombo)."captionText-1="Figure 10: Old abandoned fortresses are favorable environments for E. tergestinus in Lubiara (Veneto, Italy) (photo by Marco Colombo)."captionText-2="Figure 11: E. tergestinus collecting sites. Emilia Romagna, Lombardy, and Veneto (Italy): 1. Cislano; 2. Lubiara; 3. Ceraino; 4. Venezia; 5. Torrechiara."figureDoi-0="http://doi.org/10.5281/zenodo.12781873"figureDoi-1="http://doi.org/10.5281/zenodo.12781875"figureDoi-2="http://doi.org/10.5281/zenodo.12781877"httpUri-0="https://zenodo.org/record/12781873/files/figure.png"httpUri-1="https://zenodo.org/record/12781875/files/figure.png"httpUri-2="https://zenodo.org/record/12781877/files/figure.png"pageId="14"pageNumber="13">Figs. 9–11</figureCitation>
” complex were elevated to species status. One of these is
<taxonomicNameid="4C514D576C04FF88668DFA3FE75FFA1D"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[193,341,1468,1491]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="tergestinus">
<bibRefCitationid="EFC04B256C04FF8864F8FA3FE721FA3F"author="VIGNOLI"pageId="14"pageNumber="13"pagination="97 - 114"refId="ref19326"refString="VIGNOLI, V., N. SALOMONE, T. CARUSO & F. BERNINI. 2005. The Euscorpius tergestinus (C. L. Koch, 1837) complex in Italy: biometrics of sympatric hidden species (Scorpiones: Euscorpiidae). Zoologischer Anzeiger, 244 (2): 97 - 114."type="journal article"year="2005">Vignoli et al. (2005)</bibRefCitation>
who separated the sibling species
<taxonomicNameid="4C514D576C04FF8864FCFA59E6EFF9DE"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="concinnus">
<taxonomicNameid="4C514D576C04FF88642BFA7AE50BF9DE"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[615,769,1529,1552]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="tergestinus">
<taxonomicNameid="4C514D576C04FF886547F9D6E726F945"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="concinnus">
<bibRefCitationid="EFC04B256C04FF886762F953E7DEF929"author="FET"box="[302,468,1744,1767]"pageId="14"pageNumber="13"pagination="47 - 59"refId="ref18568"refString="FET, V., M. E. SOLEGLAD & B. GANTENBEIN. 2004. The Euroscorpion: genus Euscorpius (Scorpiones: Euscorpiidae). Proceedings of the 3 d Scorpiology Symposium, American Arachnological Society 28 th Annual Meeting, Norman, Oklahoma, 23 - 27 June 2004. Euscorpius, 17: 47 - 59."type="journal article"year="2004">Fet et al., 2004</bibRefCitation>
<taxonomicNameid="4C514D576C04FF886702F96DE7E8F8CB"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[334,482,1774,1797]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="tergestinus">
<bibRefCitationid="EFC04B256C04FF886511FBCDE24BFBAB"author="VIGNOLI"box="[861,1089,1102,1125]"pageId="14"pageNumber="13"pagination="97 - 114"refId="ref19326"refString="VIGNOLI, V., N. SALOMONE, T. CARUSO & F. BERNINI. 2005. The Euscorpius tergestinus (C. L. Koch, 1837) complex in Italy: biometrics of sympatric hidden species (Scorpiones: Euscorpiidae). Zoologischer Anzeiger, 244 (2): 97 - 114."type="journal article"year="2005">Vignoli et al. (2005)</bibRefCitation>
<taxonomicNameid="4C514D576C04FF886346FBEFE3AAFB4D"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[1290,1440,1132,1155]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="tergestinus">
Few altitudinal data can be determined from the literature, mainly because
<taxonomicNameid="4C514D576C04FF88621FFB64E2E1FB30"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[1107,1259,1255,1278]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="tergestinus">
<bibRefCitationid="EFC04B256C04FF8862C4FA85E33EFAD3"author="FET"box="[1160,1332,1286,1309]"pageId="14"pageNumber="13"pagination="255 - 256"refId="ref18388"refString="FET, V., M. KUNTNER & B. SKET. 2001. Scorpions of Slovenia: a faunistic and biogeographical survey. Pp. 255 - 256. In: Fet, V. & Selden, P. A. (eds.): Scorpions 2001. In Memoriam Gary A. Polis. British Arachnological Society: Burnham Beeches, Bucks."type="book chapter"year="2001">Fet et al. (2001)</bibRefCitation>
<taxonomicNameid="4C514D576C04FF8862D9FA5FE322FA3D"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[1173,1320,1500,1523]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="tergestinus">
<taxonomicNameid="4C514D576C04FF88627FFA78E2CCF9DC"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[1075,1222,1531,1554]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="tergestinus">
<taxonomicNameid="4C514D576C04FF886278F999E2CEF9FF"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[1076,1220,1562,1585]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="concinnus">
<bibRefCitationid="EFC04B256C04FF88622DF9BBE341F981"author="VIGNOLI"box="[1121,1355,1592,1615]"pageId="14"pageNumber="13"pagination="97 - 114"refId="ref19326"refString="VIGNOLI, V., N. SALOMONE, T. CARUSO & F. BERNINI. 2005. The Euscorpius tergestinus (C. L. Koch, 1837) complex in Italy: biometrics of sympatric hidden species (Scorpiones: Euscorpiidae). Zoologischer Anzeiger, 244 (2): 97 - 114."type="journal article"year="2005">Vignoli et al. (2005)</bibRefCitation>
, with a very high percentage (94.4%) of specimens found in abandoned houses or ruins (
<figureCitationid="136A2A516C04FF88623FF9F5E2CFF943"box="[1139,1221,1654,1677]"captionStart="Figure 10"captionStartId="15.[192,260,1927,1948]"captionTargetBox="[249,1370,168,1899]"captionTargetId="figure-7@15.[252,1367,171,1896]"captionTargetPageId="15"captionText="Figure 10: Old abandoned fortresses are favorable environments for E. tergestinus in Lubiara (Veneto, Italy) (photo by Marco Colombo)."figureDoi="http://doi.org/10.5281/zenodo.12781875"httpUri="https://zenodo.org/record/12781875/files/figure.png"pageId="14"pageNumber="13">Fig. 10</figureCitation>
); there they seemed to colonize both humid and dry rooms. A small fraction (5.6%) of
<taxonomicNameid="4C514D576C04FF8865E1F930E24BF904"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[941,1089,1715,1738]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="14"pageNumber="13"phylum="Arthropoda"rank="species"species="tergestinus">
was found in and around inhabited houses. However, the walls colonized by this species are usually humid and often covered by moss. An adult male from Ceraino (
Old abandoned fortresses are favorable environments for
<taxonomicNameid="4C514D576C05FF896522F80BE5FEF852"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[878,1012,1927,1949]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="15"pageNumber="14"phylum="Arthropoda"rank="species"species="tergestinus">
<taxonomicNameid="4C514D576C1AFF966770F97FE7CBF8DF"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[316,449,1788,1809]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="16"pageNumber="15"phylum="Arthropoda"rank="species"species="tergestinus">
<emphasisid="B925EAC66C1BFF9767D6FD7FE482FCDD"bold="true"box="[410,648,764,787]"italics="true"pageId="17"pageNumber="16">Number of specimens</emphasis>
<taxonomicNameid="4C514D576C1BFF9767C2FCB7E419FC81"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[398,531,820,847]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="17"pageNumber="16"phylum="Arthropoda"rank="species"species="tergestinus">
<tdid="76B05DEA6C1B007962D4FCB7E0A7FC9C"box="[1176,1709,820,850]"gridcol="5"gridrow="2"pageId="17"pageNumber="16">Two live females were found in cracks of the walls</td>
<tdid="76B05DEA6C1B007962D4FCD6E0A7FCA4"box="[1176,1709,853,874]"gridcol="5"gridrow="3"pageId="17"pageNumber="16">inside the fortress, in quite dark, cool, and humid</td>
<tdid="76B05DEA6C1B007962D4FCF2E0A7FC48"box="[1176,1709,881,902]"gridcol="5"gridrow="4"pageId="17"pageNumber="16">rooms; the remains of other five specimens were found</td>
<tdid="76B05DEA6C1B007962D4FC0FE0A7FC6F"box="[1176,1709,908,929]"gridcol="5"gridrow="5"pageId="17"pageNumber="16">on the ground in long and dark galleries that had no</td>
<tdid="76B05DEA6C1B007962D4FC2BE0A7FC73"box="[1176,1709,936,957]"gridcol="5"gridrow="6"pageId="17"pageNumber="16">cracks in the walls; maybe they penetrated there from</td>
<tdid="76B05DEA6C1B007962D4FC47E0A7FC17"box="[1176,1709,964,985]"gridcol="5"gridrow="7"pageId="17"pageNumber="16">the small openings on a side (observed with UV light)</td>
<taxonomicNameid="4C514D576C1BFF9767C2FC58E419FC38"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[398,531,987,1014]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="17"pageNumber="16"phylum="Arthropoda"rank="species"species="tergestinus">
<tdid="76B05DEA6C1B007964F4FC58E207FC37"box="[696,1037,987,1017]"gridcol="3"gridrow="8"pageId="17"pageNumber="16">Scala Contarini del Bovolo, Venice</td>
<tdid="76B05DEA6C1B007962D4FC58E0A7FC37"box="[1176,1709,987,1017]"gridcol="5"gridrow="8"pageId="17"pageNumber="16">Under a flower pot located in a larger flower pot full of</td>
<tdid="76B05DEA6C1B007962D4FC7FE0A7FBDF"box="[1176,1709,1020,1041]"gridcol="5"gridrow="9"pageId="17"pageNumber="16">clay balls, in a cool and quite humid courtyard</td>
<taxonomicNameid="4C514D576C1BFF9767C2FB90E419FBE0"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[398,531,1043,1070]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="17"pageNumber="16"phylum="Arthropoda"rank="species"species="tergestinus">
<tdid="76B05DEA6C1B007962D4FBB6E0A7FB84"box="[1176,1709,1077,1098]"gridcol="5"gridrow="11"pageId="17"pageNumber="16">doubtful identification due to small size and premature</td>
<taxonomicNameid="4C514D576C1BFF9767C2FBEBE419FB4D"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[398,531,1128,1155]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="17"pageNumber="16"phylum="Arthropoda"rank="species"species="tergestinus">
<tdid="76B05DEA6C1B007962D4FBEBE0A7FB48"box="[1176,1709,1128,1158]"gridcol="5"gridrow="13"pageId="17"pageNumber="16">Behind a door in a warm and dry room of a castle</td>
<taxonomicNameid="4C514D576C1BFF9767C2FB3FE419FB19"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[398,531,1212,1239]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="17"pageNumber="16"phylum="Arthropoda"rank="species"species="tergestinus">
<tdid="76B05DEA6C1B007962D4FB3FE0A7FB14"box="[1176,1709,1212,1242]"gridcol="5"gridrow="16"pageId="17"pageNumber="16">The male was found in a wall crack, in a humid, dark</td>
<tdid="76B05DEA6C1B007962D4FB5EE0A7FB3C"box="[1176,1709,1245,1266]"gridcol="5"gridrow="17"pageId="17"pageNumber="16">and cool room; the juvenile was found under a stone in</td>
<tdid="76B05DEA6C1B007967C2FB7BE491FAC3"box="[398,667,1272,1293]"gridcol="2"gridrow="18"pageId="17"pageNumber="16">adult females, remains of at</td>
<tdid="76B05DEA6C1B007962D4FB7BE0A7FAC3"box="[1176,1709,1272,1293]"gridcol="5"gridrow="18"pageId="17"pageNumber="16">an underground room, cool and humid; dead specimens</td>
<tdid="76B05DEA6C1B007962D4FA97E0A7FAE7"box="[1176,1709,1300,1321]"gridcol="5"gridrow="19"pageId="17"pageNumber="16">were found all over the fortress, on the ground; only</td>
<tdid="76B05DEA6C1B007962D4FAB3E0A7FA8B"box="[1176,1709,1328,1349]"gridcol="5"gridrow="20"pageId="17"pageNumber="16">some remains were found in the wall crack under the</td>
<tdid="76B05DEA6C1B007962D4FAC8E0A7FAAE"box="[1176,1709,1355,1376]"gridcol="5"gridrow="21"pageId="17"pageNumber="16">one inhabited by the living male (cannibalism?)</td>
<taxonomicNameid="4C514D576C1BFF9767C2FAFDE419FA57"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[398,531,1406,1433]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="17"pageNumber="16"phylum="Arthropoda"rank="species"species="tergestinus">
<tdid="76B05DEA6C1B007962D4FA38E0A7FA1E"box="[1176,1709,1467,1488]"gridcol="5"gridrow="24"pageId="17"pageNumber="16">Inside the fortress, rooms have different degree of light</td>
<tdid="76B05DEA6C1B007962D4FA54E0A7FA22"box="[1176,1709,1495,1516]"gridcol="5"gridrow="25"pageId="17"pageNumber="16">and humidity, due to their geographic exposure: the</td>
<tdid="76B05DEA6C1B007962D4FA71E0A7F9C9"box="[1176,1709,1522,1543]"gridcol="5"gridrow="26"pageId="17"pageNumber="16">room with the scorpion (that was found nearly dead</td>
<tdid="76B05DEA6C1B007962D4F98DE0A7F9ED"box="[1176,1709,1550,1571]"gridcol="5"gridrow="27"pageId="17"pageNumber="16">maybe due to the low temperature, 1°C) was not very</td>
<taxonomicNameid="4C514D576C1BFF976684F9DCE7AEF9B4"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[200,420,1631,1658]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="17"pageNumber="16"phylum="Arthropoda"rank="species"species="tergestinus">
<bibRefCitationid="EFC04B256C18FF9466BCFF44E7DBFF10"author="VIGNOLI"box="[240,465,199,222]"pageId="18"pageNumber="17"pagination="97 - 114"refId="ref19326"refString="VIGNOLI, V., N. SALOMONE, T. CARUSO & F. BERNINI. 2005. The Euscorpius tergestinus (C. L. Koch, 1837) complex in Italy: biometrics of sympatric hidden species (Scorpiones: Euscorpiidae). Zoologischer Anzeiger, 244 (2): 97 - 114."type="journal article"year="2005">Vignoli et al. (2005)</bibRefCitation>
reported the unusual case of a specimen of
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that would occupy anthropogenic habitats. Larger species usually occupy most favorable habitats against smaller species, then segregated to harder environmental conditions (Polis & McCormick, 1987; quoted after
<bibRefCitationid="EFC04B256C18FF94654CFE1DE780FE1D"author="VIGNOLI"pageId="18"pageNumber="17"pagination="97 - 114"refId="ref19326"refString="VIGNOLI, V., N. SALOMONE, T. CARUSO & F. BERNINI. 2005. The Euscorpius tergestinus (C. L. Koch, 1837) complex in Italy: biometrics of sympatric hidden species (Scorpiones: Euscorpiidae). Zoologischer Anzeiger, 244 (2): 97 - 114."type="journal article"year="2005">Vignoli et al., 2005</bibRefCitation>
). This kind of interaction, observed with other pairs of sympatric and sometimes syntopic species, was noticed once by the author in Ceraino (
<taxonomicNameid="4C514D576C18FF9467D5FD9BE426FDE1"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[409,556,536,559]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="18"pageNumber="17"phylum="Arthropoda"rank="species"species="tergestinus">
was found under a stone next to the path leading to the fortress. We can only suggest that other specimens could be inhabiting cracks in the rocky cliff; however, no other scorpions were found under stones. In this case the larger species,
, would occupy the most favorable habitat, considered by
<bibRefCitationid="EFC04B256C18FF94668EFC8DE7AEFCEB"author="VIGNOLI"box="[194,420,782,805]"pageId="18"pageNumber="17"pagination="97 - 114"refId="ref19326"refString="VIGNOLI, V., N. SALOMONE, T. CARUSO & F. BERNINI. 2005. The Euscorpius tergestinus (C. L. Koch, 1837) complex in Italy: biometrics of sympatric hidden species (Scorpiones: Euscorpiidae). Zoologischer Anzeiger, 244 (2): 97 - 114."type="journal article"year="2005">Vignoli et al. (2005)</bibRefCitation>
as the anthropogenic one. It could be that rocky cliffs are more ecologically favorable than abandoned buildings. The cliffs are heated considerably to convection by the stone, but they are also drier due to their exposure to sunlight. On the contrary abandoned buildings are not sufficiently heated (especially in underground portions); however, they have a quite stable range of temperatures and humidity during the year. Protection from predators should be similar in both habitats, mainly due to the similar way of life of scorpions, which occupy cracks in both cases.
<bibRefCitationid="EFC04B256C18FF9467EEFBDCE448FBB8"author="CRUCITTI"box="[418,578,1119,1142]"pageId="18"pageNumber="17"pagination="281 - 300"refId="ref17937"refString="CRUCITTI, P. 1993. Distribution and diversity of Italian scorpions. Redia, 76 (2): 281 - 300."type="journal article"year="1993">Crucitti (1993)</bibRefCitation>
In this case it could be that this larger species occupied its most favorable habitat, the hotter and drier one (rocky cliffs), while the smaller one (
<taxonomicNameid="4C514D576C18FF946547FB7EE73FFAFD"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="18"pageNumber="17"phylum="Arthropoda"rank="species"species="tergestinus">
Regarding intraspecific relations, inside the same fortress some scorpion remains (
<taxonomicNameid="4C514D576C18FF946470FAF8E4DCFA5C"baseAuthorityName="C. L. Koch"baseAuthorityYear="1837"box="[572,726,1403,1426]"class="Arachnida"family="Euscorpiidae"genus="Euscorpius"kingdom="Animalia"order="Scorpiones"pageId="18"pageNumber="17"phylum="Arthropoda"rank="species"species="tergestinus">
) were found on the ground under the shelter of an adult male (maybe the result of cannibalism). During this study, males were not found together with live females; only, in a fortress in Ceraino (