447 lines
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447 lines
92 KiB
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<document id="F1923EA7903CE5B49CF073AD860B2B7B" ID-CLB-Dataset="2759" ID-DOI="10.11646/zootaxa.5087.2.4" ID-GBIF-Dataset="7886e4d1-ce99-4dc0-b445-f03e4d06df3d" ID-ISSN="1175-5326" ID-Zenodo-Dep="5824232" ID-ZooBank="701019E0-F757-40E9-A54D-F936DBE0BCC7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1641453918897" checkinUser="plazi" docAuthor="Arimoto, Kôichi" docDate="2022" docId="03D1A92AFFAAFFC430851014FCCFE871" docLanguage="en" docName="zootaxa.5087.2.4.pdf" docOrigin="Zootaxa 5087 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ectamenogonus miyakoensis Arimoto 2022, sp. n." docType="treatment" docVersion="6" lastPageNumber="317" masterDocId="FFE8D152FFAFFFCF30121701FFCDEC44" masterDocTitle="Taxonomic revision of the genus Ectamenognonus Buysson (Coleoptera, Elateridae, Elaterinae) from Japan" masterLastPageNumber="334" masterPageNumber="306" pageNumber="311" updateTime="1699296519307" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9CA35A2098D98A64B4F07E990BCFE829">Taxonomic revision of the genus Ectamenognonus Buysson (Coleoptera, Elateridae, Elaterinae) from Japan</mods:title>
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<mods:namePart id="1232EB1BB0F18199E766D6974984219D">Arimoto, Kôichi</mods:namePart>
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<mods:title id="2B5131E683BC83C78C7BBE2305F44E6B">Zootaxa</mods:title>
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<mods:part id="F23C04582045347335EDBE2DD97A7FDD">
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<mods:date id="F13171291D6C54879B0868BA01361093">2022</mods:date>
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<mods:number id="C028B0329D26F4273ADCD0766B0919CE">2022-01-06</mods:number>
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<mods:number id="A590A9D1FB187C1846071FCC3CE943D9">5087</mods:number>
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<treatment id="03D1A92AFFAAFFC430851014FCCFE871" ID-DOI="http://doi.org/10.5281/zenodo.5824230" ID-GBIF-Taxon="191928809" ID-Zenodo-Dep="5824230" LSID="urn:lsid:plazi:treatment:03D1A92AFFAAFFC430851014FCCFE871" httpUri="http://treatment.plazi.org/id/03D1A92AFFAAFFC430851014FCCFE871" lastPageId="11" lastPageNumber="317" pageId="5" pageNumber="311">
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<subSubSection id="C3624BB7FFAAFFCA30851014FD7AEB6A" box="[151,695,1812,1839]" pageId="5" pageNumber="311" type="nomenclature">
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<paragraph id="8BC7183CFFAAFFCA30851014FD7AEB6A" blockId="5.[151,695,1812,1875]" box="[151,695,1812,1839]" pageId="5" pageNumber="311">
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<heading id="D08FAF50FFAAFFCA30851014FD7AEB6A" bold="true" box="[151,695,1812,1839]" fontSize="11" level="1" pageId="5" pageNumber="311" reason="1">
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<emphasis id="B90CC42EFFAAFFCA30851014FD7AEB6A" bold="true" box="[151,695,1812,1839]" pageId="5" pageNumber="311">
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<taxonomicName id="4C7863BFFFAAFFCA30851014FDA5EB6A" authority="Arimoto" authorityName="Arimoto" authorityYear="2022" box="[151,616,1812,1839]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="311" phylum="Arthropoda" rank="species" species="miyakoensis" status="sp. nov.">
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<emphasis id="B90CC42EFFAAFFCA30851014FE3BEB6B" bold="true" box="[151,502,1812,1839]" italics="true" pageId="5" pageNumber="311">Ectamenogonus miyakoensis</emphasis>
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Arimoto
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23F7955FFAAFFCA32671015FD7AEB6A" box="[629,695,1812,1838]" pageId="5" pageNumber="311" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3624BB7FFAAFFCA30851039FE2BEB16" box="[151,486,1848,1875]" pageId="5" pageNumber="311" type="description">
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<paragraph id="8BC7183CFFAAFFCA30851039FE2BEB16" blockId="5.[151,695,1812,1875]" box="[151,486,1848,1875]" pageId="5" pageNumber="311">
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(
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<figureCitation id="134304B9FFAAFFCA308D1039FE81EB16" box="[159,332,1848,1874]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="5" pageNumber="311">Figures 1B, 1C</figureCitation>
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,
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<figureCitation id="134304B9FFAAFFCA314B1039FEA4EB17" box="[345,361,1848,1875]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="5" pageNumber="311">5</figureCitation>
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,
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<figureCitation id="134304B9FFAAFFCA31671039FE48EB16" box="[373,389,1848,1874]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="5" pageNumber="311">6</figureCitation>
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,
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<figureCitation id="134304B9FFAAFFCA31831038FE6CEB17" box="[401,417,1849,1875]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="5" pageNumber="311">7</figureCitation>
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,
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<figureCitation id="134304B9FFAAFFCA31BF1039FE13EB16" box="[429,478,1848,1874]" captionStart="FIGURE 10" captionStartId="15.[151,250,753,777]" captionTargetBox="[151,1433,185,729]" captionTargetId="figure-30@15.[151,1436,181,729]" captionTargetPageId="15" captionText="FIGURE 10. Ectamenogonus spp., right antenna. A: E. miyakoensis, holotype, male [EMM01]; B: E. plebejus [GN. 2124]; C: E. plebejus [KEP20]; D: E. yakuensis, holotype, male (= E. plebejus)." figureDoi="http://doi.org/10.5281/zenodo.5824254" httpUri="https://zenodo.org/record/5824254/files/figure.png" pageId="5" pageNumber="311">10A</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3624BB7FFAAFFCA30851077FB4EEBE9" pageId="5" pageNumber="311" type="reference_group">
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<paragraph id="8BC7183CFFAAFFCA30851077FBA6EBCA" blockId="5.[151,1155,1910,1965]" box="[151,1131,1910,1934]" pageId="5" pageNumber="311">
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<treatmentCitationGroup id="AB683F12FFAAFFCA30851077FBA6EBCA" box="[151,1131,1910,1934]" pageId="5" pageNumber="311">
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<taxonomicName id="4C7863BFFFAAFFCA30851077FDE0EBCA" authority="(Candeze, 1873)" baseAuthorityName="Candeze" baseAuthorityYear="1873" box="[151,557,1910,1934]" class="Insecta" family="Elateridae" genus="Homotechnes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="311" phylum="Arthropoda" rank="species" species="plebejus">
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<emphasis id="B90CC42EFFAAFFCA30851077FEB6EBCA" box="[151,379,1910,1934]" italics="true" pageId="5" pageNumber="311">Homotechnes plebejus</emphasis>
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(
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<bibRefCitation id="EFE965CDFFAAFFCA319B1077FDE9EBCA" author="Candeze, E. C. A." box="[393,548,1910,1934]" pageId="5" pageNumber="311" pagination="1 - 32" refId="ref17336" refString="Candeze, E. C. A. (1873) Insects recueillis au Japon par Mr. G. Lewis. Elaterides. Memoires de la Societe Royale des Sciences de Liege, Series 2, 5, 1 - 32." type="journal article" year="1873">Candèze, 1873</bibRefCitation>
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)
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</taxonomicName>
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;
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<treatmentCitation id="0AD93E2DFFAAFFCA32251077FD16EBCA" author="Kishii, T." box="[567,731,1910,1934]" page="42" pageId="5" pageNumber="311" year="1961">
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<bibRefCitation id="EFE965CDFFAAFFCA32251077FD16EBCA" author="Kishii, T." box="[567,731,1910,1934]" pageId="5" pageNumber="311" pagination="1 - 56" refId="ref17764" refString="Kishii, T. (1961) Elateridae of Is. Tsushima " The Snappers of Island (II) " Bulletin of the Heian High School, 5, 1 - 56." type="journal article" year="1961">Kishii, 1961: 42</bibRefCitation>
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</treatmentCitation>
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(comment about distribution) [partim].
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC7183CFFAAFFCA30851097FB4EEBE9" blockId="5.[151,1155,1910,1965]" box="[151,1155,1941,1965]" pageId="5" pageNumber="311">
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<treatmentCitationGroup id="AB683F12FFAAFFCA30851097FB4EEBE9" box="[151,1155,1941,1965]" pageId="5" pageNumber="311">
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<taxonomicName id="4C7863BFFFAAFFCA30851097FD88EBE9" authority="(Candeze, 1873)" baseAuthorityName="Candeze" baseAuthorityYear="1873" box="[151,581,1941,1965]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="311" phylum="Arthropoda" rank="species" species="plebejus">
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<emphasis id="B90CC42EFFAAFFCA30851097FE5EEBE9" box="[151,403,1941,1965]" italics="true" pageId="5" pageNumber="311">Ectamenogonus plebejus</emphasis>
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(
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<bibRefCitation id="EFE965CDFFAAFFCA31B31094FDF1EBE9" author="Candeze, E. C. A." box="[417,572,1941,1965]" pageId="5" pageNumber="311" pagination="1 - 32" refId="ref17336" refString="Candeze, E. C. A. (1873) Insects recueillis au Japon par Mr. G. Lewis. Elaterides. Memoires de la Societe Royale des Sciences de Liege, Series 2, 5, 1 - 32." type="journal article" year="1873">Candèze, 1873</bibRefCitation>
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)
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</taxonomicName>
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;
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<treatmentCitation id="0AD93E2DFFAAFFCA325D1094FD3EEBE9" author="Kishii, T." box="[591,755,1941,1965]" page="56" pageId="5" pageNumber="311" year="1999">
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<bibRefCitation id="EFE965CDFFAAFFCA325D1094FD3EEBE9" author="Kishii, T." box="[591,755,1941,1965]" pageId="5" pageNumber="311" pagination="1 - 144" refId="ref18028" refString="Kishii, T. (1999) A check-list of the family Elateridae from Japan (Coleoptera). Bulletin of the Heian High School, 42, 1 - 144." type="journal article" year="1999">Kishii, 1999: 56</bibRefCitation>
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</treatmentCitation>
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(comment about distribution) [partim].
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3624BB7FFAAFFCA308510D5FC13EBAA" box="[151,990,2004,2031]" pageId="5" pageNumber="311" type="etymology">
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<paragraph id="8BC7183CFFAAFFCA308510D5FC13EBAA" blockId="5.[151,990,2004,2031]" box="[151,990,2004,2031]" pageId="5" pageNumber="311">
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<emphasis id="B90CC42EFFAAFFCA308510D5FED3EBAA" bold="true" box="[151,286,2004,2030]" pageId="5" pageNumber="311">Etymology.</emphasis>
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Specific epithet derived from Miyako Island, the
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<typeStatus id="54C3A69EFFAAFFCA335F10D4FCB0EBAB" box="[845,893,2005,2031]" pageId="5" pageNumber="311">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<caption id="DF0748B4FFA9FFC9308511EDFC2EEBFC" ID-DOI="http://doi.org/10.5281/zenodo.5824238" ID-Zenodo-Dep="5824238" httpUri="https://zenodo.org/record/5824238/files/figure.png" pageId="6" pageNumber="312" startId="6.[151,250,1772,1796]" targetBox="[151,1436,181,1740]" targetPageId="6">
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<paragraph id="8BC7183CFFA9FFC9308511EDFC2EEBFC" blockId="6.[151,1437,1772,1976]" pageId="6" pageNumber="312">
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<emphasis id="B90CC42EFFA9FFC9308511EDFEDFEB40" bold="true" box="[151,274,1772,1796]" pageId="6" pageNumber="312">FIGURE 3.</emphasis>
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<taxonomicName id="4C7863BFFFA9FFC9310A11ECFDDEEB40" baseAuthorityName="Kishii" baseAuthorityYear="1973" box="[280,531,1772,1796]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="312" phylum="Arthropoda" rank="species" species="matobai">
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<emphasis id="B90CC42EFFA9FFC9310A11ECFDDEEB40" box="[280,531,1772,1796]" italics="true" pageId="6" pageNumber="312">Ectamenogonus matobai</emphasis>
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</taxonomicName>
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, holotype, female. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum almost truncate, but slight emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge; black arrow: posterior margin of hypomeron with angle near mesial projection); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra.
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</paragraph>
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</caption>
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<caption id="DF0748B4FFA8FFC83085125AFE02E9FF" ID-DOI="http://doi.org/10.5281/zenodo.5824240" ID-Zenodo-Dep="5824240" httpUri="https://zenodo.org/record/5824240/files/figure.png" pageId="7" pageNumber="313" startId="7.[151,250,1371,1395]" targetBox="[151,1436,202,1311]" targetPageId="7">
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<paragraph id="8BC7183CFFA8FFC83085125AFE02E9FF" blockId="7.[151,1437,1371,1467]" pageId="7" pageNumber="313">
|
|||
|
|
<emphasis id="B90CC42EFFA8FFC83085125AFEDFE937" bold="true" box="[151,274,1371,1395]" pageId="7" pageNumber="313">FIGURE 4.</emphasis>
|
|||
|
|
<taxonomicName id="4C7863BFFFA8FFC8310A125DFDDEE937" baseAuthorityName="Kishii" baseAuthorityYear="1973" box="[280,531,1371,1395]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="313" phylum="Arthropoda" rank="species" species="matobai">
|
|||
|
|
<emphasis id="B90CC42EFFA8FFC8310A125DFDDEE937" box="[280,531,1371,1395]" italics="true" pageId="7" pageNumber="313">Ectamenogonus matobai</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, holotype, female. A: labels; B: slide; C: genitalia; D: tergite VIII, dorsal view; E: sternite VIII and tergite VIII, ventral view; F: sclerotized spines in bursa copulatrix; G: apical part of ovipositor, dorsal side; H: apical part of ovipositor, ventral side.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C3624BB7FFA8FFC830D512E8FBB7EA0F" pageId="7" pageNumber="313" type="materials_examined">
|
|||
|
|
<paragraph id="8BC7183CFFA8FFC830D512E8FBB7EA0F" blockId="7.[151,1437,1513,2008]" pageId="7" pageNumber="313">
|
|||
|
|
<emphasis id="B90CC42EFFA8FFC830D512E8FEB5EA47" bold="true" box="[199,376,1513,1539]" pageId="7" pageNumber="313">Type material.</emphasis>
|
|||
|
|
<materialsCitation id="3B101261FFA8FFC8319312E8FEB8EA08" ID-GBIF-Occurrence="3428515363" collectingDate="2015-07-06" collectionCode="EMM" collectorName="Koichi Arimoto" country="Japan" county="Male" elevation="4" latitude="24.8013" location="Miyakojima City" longLatPrecision="7" longitude="125.3174" municipality="Ryukyu" pageId="7" pageNumber="313" specimenCode="EMM01" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
|
|||
|
|
<emphasis id="B90CC42EFFA8FFC8319312E8FE38EA47" bold="true" box="[385,501,1513,1539]" pageId="7" pageNumber="313">
|
|||
|
|
<typeStatus id="54C3A69EFFA8FFC8319312E8FE3DEA47" box="[385,496,1513,1539]" pageId="7" pageNumber="313" type="holotype">Holotype</typeStatus>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
<collectingCounty id="62A660B0FFA8FFC831ED12E8FDF4EA47" box="[511,569,1513,1539]" pageId="7" pageNumber="313">Male</collectingCounty>
|
|||
|
|
[
|
|||
|
|
<specimenCode id="DBDEB047FFA8FFC8325B12E8FD7DEA40" box="[585,688,1513,1540]" collectionCode="EMM" pageId="7" pageNumber="313">EMM01</specimenCode>
|
|||
|
|
],
|
|||
|
|
<collectingCountry id="F36F58ACFFA8FFC832D612EBFCCAEA40" box="[708,775,1514,1540]" name="Japan" pageId="7" pageNumber="313">Japan</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="6BA38246FFA8FFC8330712E8FCA2EA47" box="[789,879,1513,1539]" pageId="7" pageNumber="313">Ryukyu</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="49BCD6DEFFA8FFC8336F12E8FBAFEA47" box="[893,1122,1513,1539]" country="Japan" name="Okinawa" pageId="7" pageNumber="313">Okinawa Prefecture</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<location id="8EA74EE7FFA8FFC8347D12E8FAFFEA47" LSID="urn:lsid:plazi:treatment:03D1A92AFFAAFFC430851014FCCFE871:8EA74EE7FFA8FFC8347D12E8FAFFEA47" box="[1135,1330,1513,1539]" country="Japan" county="Male" latitude="24.8013" longLatPrecision="7" longitude="125.3174" municipality="Ryukyu" name="Miyakojima City" pageId="7" pageNumber="313" stateProvince="Okinawa">Miyakojima City</location>
|
|||
|
|
(
|
|||
|
|
<location id="8EA74EE7FFA8FFC8355112E8FF2DEA6C" LSID="urn:lsid:plazi:treatment:03D1A92AFFAAFFC430851014FCCFE871:8EA74EE7FFA8FFC8355112E8FF2DEA6C" country="Japan" county="Male" latitude="24.8013" longLatPrecision="7" longitude="125.3174" municipality="Ryukyu" name="Miyako Island" pageId="7" pageNumber="313" stateProvince="Okinawa">Miyako Island</location>
|
|||
|
|
),
|
|||
|
|
<location id="8EA74EE7FFA8FFC830E0110CFDEBEA63" LSID="urn:lsid:plazi:treatment:03D1A92AFFAAFFC430851014FCCFE871:8EA74EE7FFA8FFC830E0110CFDEBEA63" box="[242,550,1549,1575]" country="Japan" county="Male" latitude="24.8013" longLatPrecision="7" longitude="125.3174" municipality="Ryukyu" name="Hirara-higashinakasonezoe" pageId="7" pageNumber="313" stateProvince="Okinawa">Hirara-higashinakasonezoe</location>
|
|||
|
|
,
|
|||
|
|
<location id="8EA74EE7FFA8FFC8323D110FFD1BEA6E" LSID="urn:lsid:plazi:treatment:03D1A92AFFAAFFC430851014FCCFE871:8EA74EE7FFA8FFC8323D110FFD1BEA6E" box="[559,726,1550,1578]" country="Japan" county="Male" latitude="24.8013" longLatPrecision="7" longitude="125.3174" municipality="Ryukyu" name="Mt. Ono-yama" pageId="7" pageNumber="313" stateProvince="Okinawa">Mt. Ôno-yama</location>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE4C7EFBFFA8FFC832F2110CFC90EA63" box="[736,861,1549,1575]" degrees="24.8013" direction="north" orientation="latitude" pageId="7" pageNumber="313" precision="5" value="24.8013">24.8013°N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE4C7EFBFFA8FFC83375110CFC3DEA6C" box="[871,1008,1549,1576]" degrees="125.3174" direction="east" orientation="longitude" pageId="7" pageNumber="313" precision="5" value="125.3174">125.3174°E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C80B5D9FFA8FFC833E8110CFBE8EA6C" box="[1018,1061,1549,1576]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="313" unit="m" value="4.0">
|
|||
|
|
<elevation id="0055FF0FFFA8FFC833E8110CFBE8EA6C" box="[1018,1061,1549,1576]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="313" unit="m" value="4.0">4 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="FFC63EFCFFA8FFC83422110CFB60EA6C" box="[1072,1197,1549,1576]" pageId="7" pageNumber="313" value="2015-07-06">
|
|||
|
|
<collectingDate id="EF82C714FFA8FFC83422110CFB60EA6C" box="[1072,1197,1549,1576]" pageId="7" pageNumber="313" value="2015-07-06">6 VII 2015</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="268D7DEAFFA8FFC834A5110CFAABEA63" box="[1207,1382,1549,1575]" pageId="7" pageNumber="313">Kôichi Arimoto</collectorName>
|
|||
|
|
leg.,
|
|||
|
|
<collectingMethod id="5239602BFFA8FFC830851130FEB8EA08" box="[151,373,1585,1612]" pageId="7" pageNumber="313">by simple light trap</collectingMethod>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="3B101261FFA8FFC831921130FBBBEA0F" ID-GBIF-Occurrence="3428515390" box="[384,1142,1585,1612]" collectingDate="2015-07-06" collectionCode="EMF" collectorName="Koichi Arimoto" country="Japan" county="Male" elevation="4" latitude="24.8013" location="Miyakojima City" longLatPrecision="7" longitude="125.3174" municipality="Ryukyu" pageId="7" pageNumber="313" specimenCode="EMF01, EMF02" specimenCount="2" specimenCount-female="2" stateProvince="Okinawa" typeStatus="paratype">
|
|||
|
|
<emphasis id="B90CC42EFFA8FFC831921130FE32EA0F" bold="true" box="[384,511,1585,1611]" pageId="7" pageNumber="313">
|
|||
|
|
<typeStatus id="54C3A69EFFA8FFC831921130FE37EA0F" box="[384,506,1585,1611]" pageId="7" pageNumber="313" type="paratype">Paratypes</typeStatus>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
<specimenCount id="9D7ED3B5FFA8FFC832171130FDBCEA0F" box="[517,625,1585,1611]" pageId="7" pageNumber="313" type="female">2 females</specimenCount>
|
|||
|
|
[
|
|||
|
|
<specimenCode id="DBDEB047FFA8FFC8326D1130FD17EA0F" box="[639,730,1585,1611]" collectionCode="EMF" pageId="7" pageNumber="313">EMF01</specimenCode>
|
|||
|
|
,
|
|||
|
|
<specimenCode id="DBDEB047FFA8FFC832F71130FC8DEA08" box="[741,832,1585,1612]" collectionCode="EMF" pageId="7" pageNumber="313">EMF02</specimenCode>
|
|||
|
|
], same data as the holotype
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3624BB7FFA8FFC830D51154FEA8EB03" pageId="7" pageNumber="313" type="diagnosis">
|
|||
|
|
<paragraph id="8BC7183CFFA8FFC830D51154FEA8EB03" blockId="7.[151,1437,1513,2008]" pageId="7" pageNumber="313">
|
|||
|
|
<emphasis id="B90CC42EFFA8FFC830D51154FE8FEA2B" bold="true" box="[199,322,1621,1647]" pageId="7" pageNumber="313">Diagnosis.</emphasis>
|
|||
|
|
Antennae not reaching pronotum posterior lateral apices by apical antennomere in male.Antennomere IV 1.2–1.3 x longer than II–III combined. Prothorax 1.2 x longer than wide. Posterior edge of scutellar shield shallowly emarginate medially. Elytra 2.4–2.5 x longer than wide, 1.9–2.0 x longer than pronotum length. Aedeagus 4.0 x longer than wide. Median lobe exceeding apices of parameres by apical 1/3. Paramere apex length 1.9–2.1 x width of parameres at expansions. Spiculum ventrale 4.0–4.1 x longer than sternite VIII. Ovipositor 1.3 x longer than length of midline of abdominal ventrites.
|
|||
|
|
<collectingRegion id="49BCD6DEFFA8FFC832A1100BFD39EB60" box="[691,756,1802,1828]" country="Turkey" name="Bursa" pageId="7" pageNumber="313">Bursa</collectingRegion>
|
|||
|
|
copulatrix 2.4–2.7 x length of uterus, with two clusters of sclerotized spines.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3624BB7FFA8FFC430D51050FAA1EEB5" lastPageId="11" lastPageNumber="317" pageId="7" pageNumber="313" type="description">
|
|||
|
|
<paragraph id="8BC7183CFFA8FFC830D51050FD83EB93" blockId="7.[151,1437,1513,2008]" pageId="7" pageNumber="313">
|
|||
|
|
<emphasis id="B90CC42EFFA8FFC830D51050FE05EB2F" bold="true" box="[199,456,1873,1899]" pageId="7" pageNumber="313">Measurements. Male.</emphasis>
|
|||
|
|
BL: 11.2, BW: 3.16, MAE: 1.88, MBE: 1.18; OI: 159, PL: 3.87, PML: 3.18, PW: 3.16, PI: 122, EL: 7.39, EW: 3.07, EI: 241, BI: 191.
|
|||
|
|
<emphasis id="B90CC42EFFA8FFC832B81074FCC5EBCB" bold="true" box="[682,776,1909,1935]" pageId="7" pageNumber="313">Female.</emphasis>
|
|||
|
|
BL: 11.5–11.6, BW: 3.10–3.35, MAE: 1.84–1.85, MBE: 1.19–1.13; OI: 155–165, PL: 3.82–3.97, PML: 3.21–3.30, PW: 3.10–3.35, PI: 119–123, EL: 7.59–7,65, EW: 3.05–3.16, EI: 242–249, BI: 193–199.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF0748B4FFA7FFC7308511EDFF39EBFC" ID-DOI="http://doi.org/10.5281/zenodo.5824242" ID-Zenodo-Dep="5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="8" pageNumber="314" startId="8.[151,250,1772,1796]" targetBox="[151,1436,181,1740]" targetPageId="8">
|
|||
|
|
<paragraph id="8BC7183CFFA7FFC7308511EDFF39EBFC" blockId="8.[151,1436,1772,1976]" pageId="8" pageNumber="314">
|
|||
|
|
<emphasis id="B90CC42EFFA7FFC7308511EDFEDEEB40" bold="true" box="[151,275,1772,1796]" pageId="8" pageNumber="314">FIGURE 5.</emphasis>
|
|||
|
|
<taxonomicName id="4C7863BFFFA7FFC7310B11ECFDF0EB40" authorityName="Arimoto" authorityYear="2022" box="[281,573,1772,1796]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="314" phylum="Arthropoda" rank="species" species="miyakoensis">
|
|||
|
|
<emphasis id="B90CC42EFFA7FFC7310B11ECFDF0EB40" box="[281,573,1772,1796]" italics="true" pageId="8" pageNumber="314">Ectamenogonus miyakoensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF0748B4FFA6FFC630851378FAF7E8F1" ID-DOI="http://doi.org/10.5281/zenodo.5824244" ID-Zenodo-Dep="5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="9" pageNumber="315" startId="9.[151,250,1145,1169]" targetBox="[151,1436,181,1121]" targetPageId="9">
|
|||
|
|
<paragraph id="8BC7183CFFA6FFC630851378FAF7E8F1" blockId="9.[151,1437,1145,1206]" pageId="9" pageNumber="315">
|
|||
|
|
<emphasis id="B90CC42EFFA6FFC630851378FEDBE8D5" bold="true" box="[151,278,1145,1169]" pageId="9" pageNumber="315">FIGURE 6.</emphasis>
|
|||
|
|
<taxonomicName id="4C7863BFFFA6FFC6310D137BFD8BE8D5" authorityName="Arimoto" authorityYear="2022" box="[287,582,1145,1169]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="315" phylum="Arthropoda" rank="species" species="miyakoensis">
|
|||
|
|
<emphasis id="B90CC42EFFA6FFC6310D137BFD8BE8D5" box="[287,582,1145,1169]" italics="true" pageId="9" pageNumber="315">Ectamenogonus miyakoensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BC7183CFFA6FFC630D513E2FBE6EA5A" blockId="9.[151,1437,1251,2034]" pageId="9" pageNumber="315">
|
|||
|
|
<emphasis id="B90CC42EFFA6FFC630D513E2FE94E8B9" bold="true" box="[199,345,1251,1277]" pageId="9" pageNumber="315">Description.</emphasis>
|
|||
|
|
Body widest around pronotum hind angles; surface with microstructures; interspaces between punctures generally as long as puncture diameter but partially smaller than puncture diameter; frons, hypomeron, scutellar shield and abdomen interspaces between punctures larger than puncture diameter (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC634B1122DFA8DE902" box="[1187,1344,1323,1350]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5A, B, G</figureCitation>
|
|||
|
|
). Color. Body brown to black-brown (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC631F11251FD91E92E" box="[483,604,1360,1386]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="9" pageNumber="315">Fig. 1B, C</figureCitation>
|
|||
|
|
). Labrum reddish (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC633231251FC47E92E" box="[817,906,1359,1386]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5C</figureCitation>
|
|||
|
|
). Antennae orange (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC634781251FB09E92E" box="[1130,1220,1359,1386]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5A</figureCitation>
|
|||
|
|
). Mandible reddish but apical parts black (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC631851275FE20E9CA" box="[407,493,1395,1422]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5C</figureCitation>
|
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|
|
). Maxillary palpomeres orange (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC6334C1275FC79E9CA" box="[862,948,1395,1422]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5C</figureCitation>
|
|||
|
|
). Posterior edge of pronotum, anterior lobe of prosternum and mesial margin and posterior part of hypomeron reddish (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC633E71299FB45E9F6" box="[1013,1160,1432,1458]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="9" pageNumber="315">Figs. 1B, 1C</figureCitation>
|
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,
|
|||
|
|
<figureCitation id="134304B9FFA6FFC634841296FB20E9F6" box="[1174,1261,1431,1458]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">5C, 5H</figureCitation>
|
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|
|
). Elytra around striae blackish, orange in posterior margin (
|
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|
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<figureCitation id="134304B9FFA6FFC6328012BDFCDCE992" box="[658,785,1468,1494]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="9" pageNumber="315">Fig. 1B, C</figureCitation>
|
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|
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). Legs orange (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC633DA12BDFBE8E992" box="[968,1061,1467,1494]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5A</figureCitation>
|
|||
|
|
). Posterior margin of abdominal ventrite V orange (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC6317412E1FE0DE9BE" box="[358,448,1503,1530]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5A</figureCitation>
|
|||
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).
|
|||
|
|
<emphasis id="B90CC42EFFA6FFC631C012E1FDDBE9BE" bold="true" box="[466,534,1504,1530]" pageId="9" pageNumber="315">Male.</emphasis>
|
|||
|
|
Tergites and sternites VIII‒X and aedeagus yellow (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC6344B12E1FB51E9BE" box="[1113,1180,1503,1530]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="9" pageNumber="315">Fig. 6</figureCitation>
|
|||
|
|
).
|
|||
|
|
<emphasis id="B90CC42EFFA6FFC634A312E1FAC2E9BE" bold="true" box="[1201,1295,1504,1530]" pageId="9" pageNumber="315">Female.</emphasis>
|
|||
|
|
Tergites and sternites VIII red-brown (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC631A41105FDC3EA5A" box="[438,526,1540,1566]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="9" pageNumber="315">Fig. 7A</figureCitation>
|
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|
|
). Body covered with orange to red-brown setae.
|
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|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC7183CFFA6FFC630D51129FA40EB7A" blockId="9.[151,1437,1251,2034]" pageId="9" pageNumber="315">
|
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|
|
Head. Frons beyond eyes 0.3 x longer than wide (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC633101129FC91EA06" box="[770,860,1575,1602]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5B</figureCitation>
|
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|
|
); supra-antennal carina in ventral view emarginate medially (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC63118114DFEA8EA22" box="[266,357,1611,1638]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5C</figureCitation>
|
|||
|
|
). Antennae not reaching pronotum posterior lateral apices by apical antennomere (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC63501114DFA42EA22" box="[1299,1423,1612,1638]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="9" pageNumber="315">Fig. 1B, C</figureCitation>
|
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|
|
).
|
|||
|
|
<emphasis id="B90CC42EFFA6FFC630851171FF16EACE" bold="true" box="[151,219,1648,1674]" pageId="9" pageNumber="315">Male.</emphasis>
|
|||
|
|
Antennae extending beyond pronotum posterior margin excluding posterior angles by apical antennomere (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC630B21195FECAEAEA" box="[160,263,1684,1710]" captionStart="FIGURE 10" captionStartId="15.[151,250,753,777]" captionTargetBox="[151,1433,185,729]" captionTargetId="figure-30@15.[151,1436,181,729]" captionTargetPageId="15" captionText="FIGURE 10. Ectamenogonus spp., right antenna. A: E. miyakoensis, holotype, male [EMM01]; B: E. plebejus [GN. 2124]; C: E. plebejus [KEP20]; D: E. yakuensis, holotype, male (= E. plebejus)." figureDoi="http://doi.org/10.5281/zenodo.5824254" httpUri="https://zenodo.org/record/5824254/files/figure.png" pageId="9" pageNumber="315">Fig. 10A</figureCitation>
|
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|
|
); antennomere II 0.8 x longer than wide; III 1.1 x longer than wide; IV 1.8 x longer than wide, 1.4 x longer than II–III combined (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC6318211B9FE25EA96" box="[400,488,1719,1746]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5D</figureCitation>
|
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|
|
), 1.1 x longer than V; V 1.7 x longer than wide; XI 2.8 x longer than wide (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC6352A11B9FA42EA96" box="[1336,1423,1719,1746]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5E</figureCitation>
|
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|
|
).
|
|||
|
|
<emphasis id="B90CC42EFFA6FFC6308511DDFF38EAB2" bold="true" box="[151,245,1756,1782]" pageId="9" pageNumber="315">Female.</emphasis>
|
|||
|
|
Antennae not reaching pronotum posterior margin excluding posterior angles by apical antennomere (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC6357D11DDFF74EB5E" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="9" pageNumber="315">Fig. 1C</figureCitation>
|
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|
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); antennomere II 0.9–1.1 x longer than wide; III 1.2–1.3 x longer than wide; IV 2.0 x longer than wide, 1.2–1.3 x longer than II–III combined, 1.0–1.1 x longer than V; V 1.8 x longer than wide; XI 2.5–2.7 x longer than wide.
|
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|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC7183CFFA6FFC530D51049FE4DEE5D" blockId="9.[151,1437,1251,2034]" lastBlockId="10.[151,1437,150,538]" lastPageId="10" lastPageNumber="316" pageId="9" pageNumber="315">
|
|||
|
|
Prothorax 1.2 x longer than wide, widening posteriad, widest behind hind angles in
|
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|
|
<typeStatus id="54C3A69EFFA6FFC634931049FB2EEB26" box="[1153,1251,1864,1890]" pageId="9" pageNumber="315" type="holotype">holotype</typeStatus>
|
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|
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and at posterior lateral apices in
|
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|
|
<typeStatus id="54C3A69EFFA6FFC6315D106DFE73EBC2" box="[335,446,1900,1926]" pageId="9" pageNumber="315" type="paratype">paratypes</typeStatus>
|
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|
|
; sides rounded in anterior 1/3, straight in posterior 2/3. Pronotum posteriorly with slight median longitudinal depression. Prosternum: ventral margin weakly inclined in anterior half (at 8–15 degrees against prosternal horizontal line,
|
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|
|
<figureCitation id="134304B9FFA6FFC631A810B5FDC0EB8A" box="[442,525,1971,1998]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5F</figureCitation>
|
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|
|
); anterior lobe exceeding anterior angles of prothorax (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC6347910B5FB73EB8A" box="[1131,1214,1972,1998]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="9" pageNumber="315">Fig. 1F</figureCitation>
|
|||
|
|
). Prosternal process depressed between procoxae (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC631F110D9FDF4EBB6" box="[483,569,2007,2034]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="9" pageNumber="315">Fig. 5H</figureCitation>
|
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|
|
), in profile strongly inclined and then almost horizontal behind procoxae (
|
|||
|
|
<figureCitation id="134304B9FFA6FFC5357D10D9FF78ECF5" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" lastPageId="10" lastPageNumber="316" pageId="9" pageNumber="315">Fig. 5F</figureCitation>
|
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); sides of dorsal lobe almost straight and parallel anteriorly and then roundly narrowed; ventral margin of ventral lobe in profile straight anteriorly and rounded posteriorly; apex in ventral and lateral views rounded (
|
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|
|
<figureCitation id="134304B9FFA5FFC5350717BAFA42EC91" box="[1301,1423,186,213]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="10" pageNumber="316">Fig. 5A, F</figureCitation>
|
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|
). Hypomeron slightly grooved next to impunctate ridge (
|
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|
|
<figureCitation id="134304B9FFA5FFC5331D17DEFCA1ECBD" box="[783,876,222,249]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="10" pageNumber="316">Fig. 5H</figureCitation>
|
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: white arrow). Pronotosternal sutures moderately opened anteriorly (
|
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|
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<figureCitation id="134304B9FFA5FFC5317A1602FE76ED59" box="[360,443,258,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="10" pageNumber="316">Fig. 5H</figureCitation>
|
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). Scutellar shield 1.5–1.7 x longer than wide (
|
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<figureCitation id="134304B9FFA5FFC533A71602FBC1ED59" box="[949,1036,258,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="10" pageNumber="316">Fig. 5G</figureCitation>
|
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), widest anteriorly, convex, strongly inclined anterior-downwards, not visible in profile (
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<figureCitation id="134304B9FFA5FFC532C41626FCEAED05" box="[726,807,294,321]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="10" pageNumber="316">Fig. 5F</figureCitation>
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); anterior edge broadly rounded; sides almost straight or broadly rounded anteriorly; posterior edge shallowly emarginate medially (
|
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<figureCitation id="134304B9FFA5FFC533CA164AFBFCED21" box="[984,1073,330,357]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="10" pageNumber="316">Fig. 5G</figureCitation>
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: arrow). Mesosternum: posterior edge 0.1 x wider than mesosternum width, triangularly emarginate (
|
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<figureCitation id="134304B9FFA5FFC53383166EFC2DEDCD" box="[913,992,366,393]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="10" pageNumber="316">Fig. 5I</figureCitation>
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: arrow), with narrowly rounded angles; borders of mesosternal cavity in profile horizontal anteriorly and then roundly and abruptly inclined (
|
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<figureCitation id="134304B9FFA5FFC535241692FA5DEDE9" box="[1334,1424,402,429]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="10" pageNumber="316">Fig. 5F</figureCitation>
|
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). Elytra 2.4–2.5 x longer than wide, 1.9–2.0 x longer than pronotum length; apex emarginate; apical outer edge of each elytron rounded; apical internal edge of each elytron slightly pointed or rounded (likely due to damage) (
|
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<figureCitation id="134304B9FFA5FFC5310716FEFEA0EE5D" box="[277,365,510,537]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="10" pageNumber="316">Fig. 5J</figureCitation>
|
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).
|
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</paragraph>
|
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<caption id="DF0748B4FFA5FFC53085119FFE1BEB66" ID-DOI="http://doi.org/10.5281/zenodo.5824246" ID-Zenodo-Dep="5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="10" pageNumber="316" startId="10.[151,250,1694,1718]" targetBox="[151,1436,561,1670]" targetPageId="10">
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<paragraph id="8BC7183CFFA5FFC53085119FFE1BEB66" blockId="10.[151,1437,1694,1826]" pageId="10" pageNumber="316">
|
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<emphasis id="B90CC42EFFA5FFC53085119FFEDEEAF2" bold="true" box="[151,275,1694,1718]" pageId="10" pageNumber="316">FIGURE 7.</emphasis>
|
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<taxonomicName id="4C7863BFFFA5FFC5310B119EFDF0EAF2" authorityName="Arimoto" authorityYear="2022" box="[281,573,1694,1718]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="316" phylum="Arthropoda" rank="species" species="miyakoensis">
|
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<emphasis id="B90CC42EFFA5FFC5310B119EFDF0EAF2" box="[281,573,1694,1718]" italics="true" pageId="10" pageNumber="316">Ectamenogonus miyakoensis</emphasis>
|
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</taxonomicName>
|
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, female, paratype. A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view.
|
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</paragraph>
|
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</caption>
|
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<paragraph id="8BC7183CFFA5FFC430D51051FE31EDB1" blockId="10.[151,1437,1872,2043]" lastBlockId="11.[151,1437,151,1077]" lastPageId="11" lastPageNumber="317" pageId="10" pageNumber="316">
|
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Abdomen. Ventrite V 0.6–0.7 x longer than wide (
|
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|
|
<figureCitation id="134304B9FFA5FFC532E51051FC9CEB2F" box="[759,849,1872,1899]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="10" pageNumber="316">Fig. 5A</figureCitation>
|
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).
|
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<emphasis id="B90CC42EFFA5FFC533761051FC65EB2E" bold="true" box="[868,936,1872,1898]" pageId="10" pageNumber="316">Male.</emphasis>
|
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Tergite VIII 1.3 x longer than wide, rounded apically (
|
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|
|
<figureCitation id="134304B9FFA5FFC530EC1075FE95EBCB" box="[254,344,1908,1935]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="10" pageNumber="316">Fig. 6A</figureCitation>
|
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|
|
). Sternite VIII with deep and rounded median notch (
|
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|
|
<figureCitation id="134304B9FFA5FFC533BB1075FBCCEBCB" box="[937,1025,1908,1935]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="10" pageNumber="316">Fig. 6B</figureCitation>
|
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|
|
). Tergite IX 0.9 x length of tergite X, with median notch rounded (
|
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|
|
<figureCitation id="134304B9FFA5FFC531C81099FDFFEBF7" box="[474,562,1944,1971]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="10" pageNumber="316">Fig. 6C</figureCitation>
|
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|
|
). Tergite X 1.5 x longer than wide, rounded apically (
|
|||
|
|
<figureCitation id="134304B9FFA5FFC534981099FB2EEBF7" box="[1162,1251,1944,1971]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="10" pageNumber="316">Fig. 6C</figureCitation>
|
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|
|
). Sternite IX 3.5 x longer than wide, rounded apically (
|
|||
|
|
<figureCitation id="134304B9FFA5FFC5322C10BDFD55EB93" box="[574,664,1980,2007]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="10" pageNumber="316">Fig. 6D</figureCitation>
|
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|
|
). Aedeagus 4.0 x longer than wide (
|
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|
|
<figureCitation id="134304B9FFA5FFC5343410BDFB54EB93" box="[1062,1177,1980,2007]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="10" pageNumber="316">Fig. 6E, F</figureCitation>
|
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|
|
). Phallobase 0.5 x total length of aedeagus, 2.0 x longer than wide. Median lobe exceeding apices of parameres by apical 1/3; basal struts 0.4 x total length of median lobe. Parameres fused ventrally; preapical expansions not projecting laterally beyond side of apex; apex beyond preapical expansions blade-like shaped (
|
|||
|
|
<figureCitation id="134304B9FFA4FFC433B917BAFBC5EC92" box="[939,1032,187,214]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="11" pageNumber="317">Fig. 6G</figureCitation>
|
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), rounded laterally, with two setae dorsally, with three setae ventrally; apex length 1.9–2.1 x width of parameres at expansions in ventral side.
|
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|
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<emphasis id="B90CC42EFFA4FFC4352C17DEFA51ECBD" bold="true" box="[1342,1436,223,249]" pageId="11" pageNumber="317">Female.</emphasis>
|
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|
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Terigite VIII 1.5 x longer than wide (
|
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|
|
<figureCitation id="134304B9FFA4FFC432231602FD44ED59" box="[561,649,259,285]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7B</figureCitation>
|
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|
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). Sternite VIII 1.2–1.3 x longer than wide (
|
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|
|
<figureCitation id="134304B9FFA4FFC4347B1602FB0CED59" box="[1129,1217,259,285]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7C</figureCitation>
|
|||
|
|
); spiculum ventrale 4.0–4.1 x longer than sternite VIII (
|
|||
|
|
<figureCitation id="134304B9FFA4FFC432361626FDB3ED05" box="[548,638,295,321]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7A</figureCitation>
|
|||
|
|
). Ovipositor 1.3 x longer than length of midline of abdominal ventrites; coxites two segmented at ventral side (
|
|||
|
|
<figureCitation id="134304B9FFA4FFC4325D164AFD64ED21" box="[591,681,331,357]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7H</figureCitation>
|
|||
|
|
), with some setae; stylus with setae around apex (
|
|||
|
|
<figureCitation id="134304B9FFA4FFC434CD164AFAF7ED21" box="[1247,1338,331,357]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7H</figureCitation>
|
|||
|
|
). Vagina long; uterus globular (
|
|||
|
|
<figureCitation id="134304B9FFA4FFC43183166EFE21EDCD" box="[401,492,367,393]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7A</figureCitation>
|
|||
|
|
); bursa copulatrix U-shaped (
|
|||
|
|
<figureCitation id="134304B9FFA4FFC43328166EFC59EDCD" box="[826,916,367,393]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7A</figureCitation>
|
|||
|
|
), long, 2.4–2.7 x length of uterus, with 61–71 sclerotized spines, of which 25–26 spines forming cluster in first half (
|
|||
|
|
<figureCitation id="134304B9FFA4FFC433A11692FBC1EDE9" box="[947,1036,403,429]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7D</figureCitation>
|
|||
|
|
) and 35–46 spines forming rows in latter half (
|
|||
|
|
<figureCitation id="134304B9FFA4FFC4310116B6FEA7ED95" box="[275,362,439,465]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7E</figureCitation>
|
|||
|
|
); long thin sac extending from apex of bursa copulatrix (
|
|||
|
|
<figureCitation id="134304B9FFA4FFC433F716B6FBF7ED95" box="[997,1082,439,465]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7F</figureCitation>
|
|||
|
|
), with thin coiled sac near apex of bursa copulatrix (
|
|||
|
|
<figureCitation id="134304B9FFA4FFC4316816DAFE3DEDB1" box="[378,496,475,501]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.A–C, F–H: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="11" pageNumber="317">Fig. 7F, G</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC7183CFFA4FFC430D516FFFAA1EEB5" blockId="11.[151,1437,151,1077]" pageId="11" pageNumber="317">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC430D516FFFE6AEE5D" bold="true" box="[199,423,510,537]" pageId="11" pageNumber="317">Comparative note.</emphasis>
|
|||
|
|
<taxonomicName id="4C7863BFFFA4FFC431A216FEFD39EE5D" authorityName="Arimoto" authorityYear="2022" box="[432,756,511,537]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="317" phylum="Arthropoda" rank="species" species="miyakoensis">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC431A216FEFD39EE5D" box="[432,756,511,537]" italics="true" pageId="11" pageNumber="317">Ectamenogonus miyakoensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is similar to
|
|||
|
|
<taxonomicName id="4C7863BFFFA4FFC4339E16FEFBC7EE5D" baseAuthorityName="Candeze" baseAuthorityYear="1873" box="[908,1034,511,537]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="317" phylum="Arthropoda" rank="species" species="plebejus">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC4339E16FEFC50EE5C" box="[908,925,511,536]" italics="true" pageId="11" pageNumber="317">E</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC433BF16FEFBC7EE5D" box="[941,1034,511,537]" italics="true" pageId="11" pageNumber="317">plebejus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in the proportions of the prothorax and elytra and shape of the aedeagus, but it is distinguished from
|
|||
|
|
<taxonomicName id="4C7863BFFFA4FFC4337B1522FC28EE79" baseAuthorityName="Candeze" baseAuthorityYear="1873" box="[873,997,547,573]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="317" phylum="Arthropoda" rank="species" species="plebejus">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC4337B1522FCB7EE78" box="[873,890,547,572]" italics="true" pageId="11" pageNumber="317">E</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC4339A1522FC28EE79" box="[904,997,547,573]" italics="true" pageId="11" pageNumber="317">plebejus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the following contrasting characters (
|
|||
|
|
<taxonomicName id="4C7863BFFFA4FFC430B31546FED1EE25" baseAuthorityName="Candeze" baseAuthorityYear="1873" box="[161,284,583,609]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="317" phylum="Arthropoda" rank="species" species="plebejus">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC430B31546FF7FEE24" box="[161,178,583,608]" italics="true" pageId="11" pageNumber="317">E</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC430AD1546FED1EE25" box="[191,284,583,609]" italics="true" pageId="11" pageNumber="317">plebejus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in parentheses): antennae not reaching pronotum posterior lateral apices by apical antennomere in male (antennae just reaching or extending beyond pronotum posterior lateral apices by apical antennomere in male); posterior edge of scutellar shield shallowly emarginate medially (posterior edge of scutellar shield rounded but truncate in some); elytra 1.9–2.0 x longer than pronotum length, BI: 191–199 (elytra 2.0–2.3 x longer than pronotum length, BI: 202–229); bursa copulatrix 2.4–2.7 x length of uterus (bursa copulatrix 1.7–2.3 x length of uterus).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3624BB7FFA4FFC430D515FBFC28EF8D" pageId="11" pageNumber="317" type="discussion">
|
|||
|
|
<paragraph id="8BC7183CFFA4FFC430D515FBFC28EF8D" blockId="11.[151,1437,151,1077]" pageId="11" pageNumber="317">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC430D515FBFE81EF50" bold="true" box="[199,332,762,788]" pageId="11" pageNumber="317">Discussion.</emphasis>
|
|||
|
|
<bibRefCitation id="EFE965CDFFA4FFC4314A15FAFE3EEF52" author="Kishii, T." box="[344,499,763,790]" pageId="11" pageNumber="317" pagination="1 - 56" refId="ref17764" refString="Kishii, T. (1961) Elateridae of Is. Tsushima " The Snappers of Island (II) " Bulletin of the Heian High School, 5, 1 - 56." type="journal article" year="1961">Kishii (1961)</bibRefCitation>
|
|||
|
|
included Miyako Island in the distribution of
|
|||
|
|
<taxonomicName id="4C7863BFFFA4FFC4343315FAFB6FEF51" baseAuthorityName="Candeze" baseAuthorityYear="1873" box="[1057,1186,763,789]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="317" phylum="Arthropoda" rank="species" species="plebejus">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC4343315FAFBFFEF50" box="[1057,1074,763,788]" italics="true" pageId="11" pageNumber="317">E</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC4345715FAFB6FEF51" box="[1093,1186,763,789]" italics="true" pageId="11" pageNumber="317">plebejus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, but he did not give specimen information. This study did not find specimens that served as evidence of the distribution information in
|
|||
|
|
<bibRefCitation id="EFE965CDFFA4FFC430851442FEFDEF1A" author="Kishii, T." box="[151,304,835,862]" pageId="11" pageNumber="317" pagination="1 - 56" refId="ref17764" refString="Kishii, T. (1961) Elateridae of Is. Tsushima " The Snappers of Island (II) " Bulletin of the Heian High School, 5, 1 - 56." type="journal article" year="1961">Kishii (1961)</bibRefCitation>
|
|||
|
|
. Specimens newly collected from Miyako Island (EMM01 and EMF01–02) are distinguished from
|
|||
|
|
<taxonomicName id="4C7863BFFFA4FFC435961442FF39EFC5" baseAuthorityName="Candeze" baseAuthorityYear="1873" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="317" phylum="Arthropoda" rank="species" species="plebejus">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC435961442FA58EF18" box="[1412,1429,835,860]" italics="true" pageId="11" pageNumber="317">E</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC430851466FF39EFC5" box="[151,244,871,897]" italics="true" pageId="11" pageNumber="317">plebejus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the antenna length, proportion of body, shape of the scutellar shield, and length of the bursa copulatrix (see comparative note of
|
|||
|
|
<taxonomicName id="4C7863BFFFA4FFC431AA148AFDACEFE1" authorityName="Arimoto" authorityYear="2022" box="[440,609,907,933]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="317" phylum="Arthropoda" rank="species" species="miyakoensis">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC431AA148AFE04EFE0" box="[440,457,907,932]" italics="true" pageId="11" pageNumber="317">E</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC431CA148AFDACEFE1" box="[472,609,907,933]" italics="true" pageId="11" pageNumber="317">miyakoensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
). Therefore, the specimens from Miyako Island were determined to be a new species,
|
|||
|
|
<taxonomicName id="4C7863BFFFA4FFC4313E14AEFE19EF8D" authorityName="Arimoto" authorityYear="2022" box="[300,468,943,969]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="317" phylum="Arthropoda" rank="species" species="miyakoensis">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC4313E14AEFEF0EF8C" box="[300,317,943,968]" italics="true" pageId="11" pageNumber="317">E</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC4315914AEFE19EF8D" box="[331,468,943,969]" italics="true" pageId="11" pageNumber="317">miyakoensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and removed
|
|||
|
|
<taxonomicName id="4C7863BFFFA4FFC4326814AEFD3BEF8D" baseAuthorityName="Candeze" baseAuthorityYear="1873" box="[634,758,943,969]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="317" phylum="Arthropoda" rank="species" species="plebejus">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC4326814AEFD46EF8C" box="[634,651,943,968]" italics="true" pageId="11" pageNumber="317">E</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC4328B14AEFD3BEF8D" box="[665,758,943,969]" italics="true" pageId="11" pageNumber="317">plebejus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from Miyako Island.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3624BB7FFA4FFC430D514D3FB52EFAA" box="[199,1183,978,1006]" pageId="11" pageNumber="317" type="distribution">
|
|||
|
|
<paragraph id="8BC7183CFFA4FFC430D514D3FB52EFAA" blockId="11.[151,1437,151,1077]" box="[199,1183,978,1006]" pageId="11" pageNumber="317">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC430D514D3FE94EFA8" bold="true" box="[199,345,978,1004]" pageId="11" pageNumber="317">Distribution</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="134304B9FFA4FFC4317B14D2FE1DEFA9" box="[361,464,979,1005]" captionStart="FIGURE 20" captionStartId="26.[151,250,1786,1810]" captionTargetBox="[155,1431,1285,1759]" captionTargetId="figure-534@26.[151,1436,1281,1763]" captionTargetPageId="26" captionText="FIGURE 20. Map of the study area showing the location of the islands mentioned in this study. A: East Asia; B: Ryukyu Islands." figureDoi="http://doi.org/10.5281/zenodo.5824278" httpUri="https://zenodo.org/record/5824278/files/figure.png" pageId="11" pageNumber="317">Fig. 20B</figureCitation>
|
|||
|
|
).
|
|||
|
|
<collectingCountry id="F36F58ACFFA4FFC431F114D2FDEAEFA9" box="[483,551,979,1005]" name="Japan" pageId="11" pageNumber="317">Japan</collectingCountry>
|
|||
|
|
: Miyako Islands of the Ryukyu Islands (Miyako Island).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3624BB7FFA4FFC430D514F6FCCFE871" pageId="11" pageNumber="317" type="biology_ecology">
|
|||
|
|
<paragraph id="8BC7183CFFA4FFC430D514F6FCCFE871" blockId="11.[151,1437,151,1077]" pageId="11" pageNumber="317">
|
|||
|
|
<emphasis id="B90CC42EFFA4FFC430D514F6FEE1E855" bold="true" box="[199,300,1015,1041]" pageId="11" pageNumber="317">Ecology.</emphasis>
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The
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<typeStatus id="54C3A69EFFA4FFC4317E14F6FE51E855" box="[364,412,1015,1041]" pageId="11" pageNumber="317">type</typeStatus>
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specimens were collected at night using simple light traps made by combining a flight interception trap with a 4 W chemical fluorescent light.
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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