181 lines
19 KiB
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181 lines
19 KiB
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<document id="E1EB6E81414A1BCB93298B321F73EB37" ID-DOI="10.5962/bhl.part.117818" ID-ISSN="0035-418X" ID-Zenodo-Dep="6117779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645031409249" checkinUser="carolina" docAuthor="Schwendinger, Peter J. & Ono, Hirotsugu" docDate="2011" docId="03D14E1FB049FF9F2CE918CDEE305B08" docLanguage="en" docName="RevSuisseZool.118.4.599-637.pdf.imf" docOrigin="Revue suisse de Zoologie 118 (4)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Heptathela tomokunii Ono 1997" docType="treatment" docVersion="4" lastPageNumber="613" masterDocId="FFE83667B047FF912C61182EED625E6E" masterDocTitle="On two Heptathela species from southern Vietnam, with a discussion of copulatory organs and systematics of the Liphistiidae (Araneae: Mesothelae)" masterLastPageNumber="637" masterPageNumber="599" pageNumber="613" updateTime="1699296575136" updateUser="plazi">
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<mods:titleInfo id="4A90F2E865EA044AF978336B28D9410E">
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<mods:title id="3A30D02B62411001CAF8F428FD0E1A13">On two Heptathela species from southern Vietnam, with a discussion of copulatory organs and systematics of the Liphistiidae (Araneae: Mesothelae)</mods:title>
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<mods:name id="95DFA18D5A20FE5AA024859D1CD73578" type="personal">
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<mods:namePart id="0D56D704CD040140DD8F3CCFE99AD75F">Schwendinger, Peter J.</mods:namePart>
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<mods:name id="7B9CC794C0BC097352AB8E66EE32D8C2" type="personal">
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<mods:roleTerm id="C891CEC95635C76DB2EA224AF220D7C7">Author</mods:roleTerm>
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<mods:namePart id="0A08B808F178DF357A73EEFF19D031D0">Ono, Hirotsugu</mods:namePart>
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<mods:typeOfResource id="C02D497AA3DED779319AE411EE72C391">text</mods:typeOfResource>
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<mods:titleInfo id="30561A26D5D987AE9D3D13CA0FD424E0">
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<mods:title id="7218E7E458D99B766A62C384CDF0A1B5">Revue suisse de Zoologie</mods:title>
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</mods:titleInfo>
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<mods:part id="04DF32F7DA2BA4303222473C686ACDA7">
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<mods:date id="25E629E2D9E2D19BAD10F587E94B1462">2011</mods:date>
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<mods:detail id="AB38C2F13962E84F0D284750A06AF538" type="pubDate">
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<mods:number id="0862F8E751F51977844DA146989D3915">2011-12-31</mods:number>
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</mods:detail>
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<mods:detail id="06C150F4AA8443A91565649A5244F39D" type="volume">
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<mods:number id="35E1616739E754CD3A97DA1DF3915BFB">118</mods:number>
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<mods:detail id="66C7E19BBB3ED82EDDB22FE7FB4FD132" type="issue">
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<mods:number id="5B0C8518FA37372A19BACF05A70E4CFA">4</mods:number>
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<mods:start id="ED36436B248102C09E10D1AB85840CF4">599</mods:start>
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<mods:classification id="56C92D5637BC72B33C268A150A408BB9">journal article</mods:classification>
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<mods:identifier id="8D1BCD178EEBB9178534BC01F29DFDDD" type="DOI">10.5962/bhl.part.117818</mods:identifier>
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<mods:identifier id="68E64172283A17DD19C6A3E298C40BB4" type="ISSN">0035-418X</mods:identifier>
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<mods:identifier id="97A177A00D55C9B9D951D109EF9EE235" type="Zenodo-Dep">6117779</mods:identifier>
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</mods:mods>
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<treatment id="03D14E1FB049FF9F2CE918CDEE305B08" ID-DOI="http://doi.org/10.5281/zenodo.6311855" ID-Zenodo-Dep="6311855" LSID="urn:lsid:plazi:treatment:03D14E1FB049FF9F2CE918CDEE305B08" httpUri="http://treatment.plazi.org/id/03D14E1FB049FF9F2CE918CDEE305B08" lastPageNumber="613" pageId="14" pageNumber="613">
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<subSubSection id="C362AC82B049FF9F2CE918CDE9415E95" box="[136,1059,227,251]" pageId="14" pageNumber="613" type="nomenclature">
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<paragraph id="8BC7FF09B049FF9F2CE918CDE9415E95" blockId="14.[136,1059,227,251]" box="[136,1059,227,251]" pageId="14" pageNumber="613">
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<heading id="D08F4865B049FF9F2CE918CDE9415E95" box="[136,1059,227,251]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="613" reason="2">
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<taxonomicName id="4C78848AB049FF9F2CE918CDEC905E95" authority="Ono, 1997" authorityName="Ono" authorityYear="1997" box="[136,498,227,251]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="613" phylum="Arthropoda" rank="species" species="tomokunii">
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<emphasis id="B90C231BB049FF9F2CE918CDEC185E95" bold="true" box="[136,378,227,251]" italics="true" pageId="14" pageNumber="613">Heptathela tomokunii</emphasis>
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Ono, 1997
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</taxonomicName>
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Figs 39-43
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C362AC82B049FF9F2CE91921EED75F4A" box="[136,949,271,293]" pageId="14" pageNumber="613" type="reference_group">
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<paragraph id="8BC7FF09B049FF9F2CE91921EED75F4A" blockId="14.[136,949,271,293]" box="[136,949,271,293]" pageId="14" pageNumber="613">
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<treatmentCitationGroup id="AB68D827B049FF9F2CE91921EED75F4A" box="[136,949,271,293]" pageId="14" pageNumber="613">
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<treatmentCitation id="0AD9D918B049FF9F2CE91921EF745F4B" author="ONO, H." box="[136,534,271,293]" page="26" pageId="14" pageNumber="613" year="1997">
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<taxonomicName id="4C78848AB049FF9F2CE91921EF745F4B" authority="Ono, 1997 a: 24 - 26" authorityName="Ono" authorityPageNumber="24 - 26" authorityYear="1997" box="[136,534,271,293]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="613" phylum="Arthropoda" rank="species" species="tomokunii">
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<emphasis id="B90C231BB049FF9F2CE91921EC3B5F4B" box="[136,345,271,293]" italics="true" pageId="14" pageNumber="613">Heptathela tomokunii</emphasis>
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<bibRefCitation id="EFE982F8B049FF9F2D011921EF745F4B" author="ONO, H." box="[352,534,271,293]" pageId="14" pageNumber="613" pagination="23 - 28" refId="ref21464" refString="ONO, H. 1997 a. A new species of the genus Heptathela (Araneae: Liphistiidae) from Vietnam. Acta Arachnologica 46 (1): 23 - 28." type="journal article" year="1997">Ono, 1997a: 24-26</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs 1-8 (description of male and female).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C362AC82B049FF9F2CB51919ECFD5F11" pageId="14" pageNumber="613" type="materials_examined">
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<paragraph id="8BC7FF09B049FF9F2CB51919ECFD5F11" blockId="14.[136,1059,310,383]" pageId="14" pageNumber="613">
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<materialsCitation id="3B10F554B049FF9F2CB51919EDC25F11" collectingDate="1995-10-08" collectionCode="MATERIAL, EXAMINED" country="Vietnam" elevation="900" location="Vinh Phu Province" municipality="Tam Dao" pageId="14" pageNumber="613" specimenCount="3" specimenCount-female="1" typeStatus="holotype">
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<collectionCode id="ED6967CCB049FF9F2CB51919ECD15F25" box="[212,435,311,332]" pageId="14" pageNumber="613">MATERIAL EXAMINED</collectionCode>
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: NSMT-Ar 3396-3398; male
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<typeStatus id="54C341ABB049FF9F2E921918EE255F22" box="[755,839,310,332]" pageId="14" pageNumber="613" type="holotype">holotype</typeStatus>
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(matured
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<date id="FFC6D9C9B049FF9F2FDA1918E9765F22" box="[955,1044,310,332]" pageId="14" pageNumber="613" value="1995-10-08">
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<collectingDate id="EF822021B049FF9F2FDA1918E9765F22" box="[955,1044,310,332]" pageId="14" pageNumber="613" value="1995-10-08">8.X.1995</collectingDate>
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</date>
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), female “
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<typeStatus id="54C341ABB049FF9F2C821961EC525F0B" box="[227,304,335,357]" pageId="14" pageNumber="613" type="allotype">allotype</typeStatus>
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”,
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<specimenCount id="9D7E3480B049FF9F2D2E1961ECC95F0B" box="[335,427,335,357]" pageId="14" pageNumber="613" type="female">1 female</specimenCount>
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<typeStatus id="54C341ABB049FF9F2DDB197EEF6F5F0B" box="[442,525,336,357]" pageId="14" pageNumber="613" type="paratype">paratype</typeStatus>
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;
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<collectingCountry id="F36FBF99B049FF9F2E401961EF105F0B" box="[545,626,335,357]" name="Vietnam" pageId="14" pageNumber="613">Vietnam</collectingCountry>
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,
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<location id="8EA7A9D2B049FF9F2EE71961EE2E5F0B" LSID="urn:lsid:plazi:treatment:03D14E1FB049FF9F2CE918CDEE305B08:8EA7A9D2B049FF9F2EE71961EE2E5F0B" box="[646,844,335,357]" country="Vietnam" municipality="Tam Dao" name="Vinh Phu Province" pageId="14" pageNumber="613">Vinh Phu Province</location>
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,
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<collectingMunicipality id="6BA36573B049FF9F2F00197EEEA05F0B" box="[865,962,336,357]" pageId="14" pageNumber="613">Tam Dao</collectingMunicipality>
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,
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<quantity id="4C8052ECB049FF9F2FB71961E97E5F0B" box="[982,1052,335,357]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="613" unit="m" value="900.0">
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<elevation id="0055183AB049FF9F2FB71961E97E5F0B" box="[982,1052,335,357]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="613" unit="m" value="900.0">900 m</elevation>
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</quantity>
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; 15
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</materialsCitation>
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./
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<date id="FFC6D9C9B049FF9F2CCC1947EC785F11" box="[173,282,361,383]" pageId="14" pageNumber="613" value="1995-09-21">21.IX.1995</date>
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; leg. H. Ono.
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</paragraph>
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</subSubSection>
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<subSubSection id="C362AC82B049FF9F2CB519BFEFDB5B74" pageId="14" pageNumber="613" type="discussion">
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<paragraph id="8BC7FF09B049FF9F2CB519BFEC115A63" blockId="14.[136,1059,400,1037]" pageId="14" pageNumber="613">
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REMARKS: Diagnosis and description were given by
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<bibRefCitation id="EFE982F8B049FF9F2F2E19BEEE805FC6" author="ONO, H." box="[847,994,400,424]" pageId="14" pageNumber="613" pagination="23 - 28" refId="ref21464" refString="ONO, H. 1997 a. A new species of the genus Heptathela (Araneae: Liphistiidae) from Vietnam. Acta Arachnologica 46 (1): 23 - 28." type="journal article" year="1997">Ono (1997a)</bibRefCitation>
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. The following can be added: cymbial projection only moderately developed, quite wide (Fig. 39); paracymbium short and wide at base, at an acute angle to axis of cymbium (
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<bibRefCitation id="EFE982F8B049FF9F2CF019DFEC6D5C66" author="ONO, H." box="[145,271,497,521]" pageId="14" pageNumber="613" pagination="23 - 28" refId="ref21464" refString="ONO, H. 1997 a. A new species of the genus Heptathela (Araneae: Liphistiidae) from Vietnam. Acta Arachnologica 46 (1): 23 - 28." type="journal article" year="1997">Ono, 1997a</bibRefCitation>
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: fig. 2), similar to that of
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<taxonomicName id="4C78848AB049FF9F2E4C19DCEFA55C67" baseAuthorityName="Bristowe" baseAuthorityYear="1933" box="[557,711,497,521]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="613" phylum="Arthropoda" rank="species" species="tonkinensis">
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<emphasis id="B90C231BB049FF9F2E4C19DCEFA55C67" box="[557,711,497,521]" italics="true" pageId="14" pageNumber="613">H. tonkinensis</emphasis>
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</taxonomicName>
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; subtegulum exceptionally long (Fig. 40); contrategulum with denticles on distal edge modified into a series of parallel ribs running down to 1-2 isolated ventral denticles near proximal margin (Figs 39-40); tegulum with quite low, narrowly arched marginal apophysis and with quite short terminal apophysis (Figs 39, 41) carrying a pronounced, strongly bulging dorsal extension with a coasely dentate edge (Fig. 39;
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<bibRefCitation id="EFE982F8B049FF9F2EE71ABCEE615CC4" author="ONO, H." box="[646,771,658,682]" pageId="14" pageNumber="613" pagination="23 - 28" refId="ref21464" refString="ONO, H. 1997 a. A new species of the genus Heptathela (Araneae: Liphistiidae) from Vietnam. Acta Arachnologica 46 (1): 23 - 28." type="journal article" year="1997">Ono, 1997a</bibRefCitation>
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: fig. 4) not in contact with edge on marginal apophysis; genital atrium with some hairs on ventral side near posterior and lateral margins (Fig. 42); bursa copulatrix short, anterior margin widely arched; all receptacles simple and smooth [Fig. 43; also confirmed for the “
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<typeStatus id="54C341ABB049FF9F2FDB1ADDE9725D65" box="[954,1040,755,779]" pageId="14" pageNumber="613" type="allotype">allotype</typeStatus>
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”, though the corresponding illustration (
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<bibRefCitation id="EFE982F8B049FF9F2E461B3DEFC75D45" author="ONO, H." box="[551,677,787,811]" pageId="14" pageNumber="613" pagination="23 - 28" refId="ref21464" refString="ONO, H. 1997 a. A new species of the genus Heptathela (Araneae: Liphistiidae) from Vietnam. Acta Arachnologica 46 (1): 23 - 28." type="journal article" year="1997">Ono, 1997a</bibRefCitation>
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: fig. 6) indicates indistinctly cauliflower-shaped receptacles], not developed as caulifower-shaped receptacular clusters as in other
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<taxonomicName id="4C78848AB049FF9F2D641B7AECFB5D02" authorityName="Kishida" authorityYear="1923" box="[261,409,852,876]" class="Arachnida" family="Liphistiidae" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="636" phylum="Arthropoda" rank="subFamily" subFamily="Heptathelinae">Heptathelinae</taxonomicName>
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examined; larger lateral pair of receptacles sessile, slightly displaced onto dorsal wall of bursa copulatrix; smaller median pair of receptacles stalked [on common socket in
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<typeStatus id="54C341ABB049FF9F2DBF1BBBEF585DC3" box="[478,570,917,941]" pageId="14" pageNumber="613" type="paratype">paratype</typeStatus>
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(Fig. 43), on individual stalks in “
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<typeStatus id="54C341ABB049FF9F2FA01BBAE9755DC2" box="[961,1047,916,940]" pageId="14" pageNumber="613" type="allotype">allotype</typeStatus>
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” (
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<bibRefCitation id="EFE982F8B049FF9F2CF01B9BEC6C5DA3" author="ONO, H." box="[145,270,949,973]" pageId="14" pageNumber="613" pagination="23 - 28" refId="ref21464" refString="ONO, H. 1997 a. A new species of the genus Heptathela (Araneae: Liphistiidae) from Vietnam. Acta Arachnologica 46 (1): 23 - 28." type="journal article" year="1997">Ono, 1997a</bibRefCitation>
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: fig. 6)] and slightly displaced onto ventral wall (Fig. 43). “Tibial spurs” present on legs I-III of female, on last exuvia of male and also on mature male (with a few “spurs” reduced).
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</paragraph>
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<paragraph id="8BC7FF09B049FF9F2CB51C0FEFDB5B74" blockId="14.[136,1059,1057,1307]" pageId="14" pageNumber="613">
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RELATIONSHIPS: Due to similarities in the palpal organ of males (moderately developed cymbial projection; short, wide paracymbium at an acute angle to the axis of the cymbium; long subtegulum; low, arched marginal tegular apophysis; pronounced dorsal extension of terminal tegular apophysis),
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<taxonomicName id="4C78848AB049FF9F2EA11CADEE2F5AF7" authorityName="Ono" authorityYear="1997" box="[704,845,1153,1178]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="613" phylum="Arthropoda" rank="species" species="tomokunii">
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<emphasis id="B90C231BB049FF9F2EA11CADEE2F5AF7" box="[704,845,1153,1178]" italics="true" pageId="14" pageNumber="613">H. tomokunii</emphasis>
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</taxonomicName>
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appears to be more closely related to
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<taxonomicName id="4C78848AB049FF9F2D311C8DEC8E5AD4" baseAuthorityName="Bristowe" baseAuthorityYear="1933" box="[336,492,1186,1210]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="613" phylum="Arthropoda" rank="species" species="tonkinensis">
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<emphasis id="B90C231BB049FF9F2D311C8DEC8E5AD4" box="[336,492,1186,1210]" italics="true" pageId="14" pageNumber="613">H. tonkinensis</emphasis>
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</taxonomicName>
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(female unknown) than to
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<taxonomicName id="4C78848AB049FF9F2F7E1C8DEEC35AD4" baseAuthorityName="Ono" baseAuthorityYear="2002" box="[799,929,1186,1210]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="613" phylum="Arthropoda" rank="species" species="australis">
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<emphasis id="B90C231BB049FF9F2F7E1C8DEEC35AD4" box="[799,929,1186,1210]" italics="true" pageId="14" pageNumber="613">H. australis</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
and
|
||
|
|
<taxonomicName id="4C78848AB049FF9F2FBD1C8DE9405AD4" authorityName="Schwendinger & Ono" authorityYear="2011" box="[988,1058,1187,1210]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="613" phylum="Arthropoda" rank="species" species="nui" status="sp. nov.">
|
||
|
|
<emphasis id="B90C231BB049FF9F2FBD1C8DE9405AD4" box="[988,1058,1187,1210]" italics="true" pageId="14" pageNumber="613">H. nui</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
<taxonomicNameLabel id="A23F9E60B049FF9F2CE91CEDEDA35AB4" box="[136,193,1219,1242]" pageId="14" pageNumber="613" rank="species">sp. n.</taxonomicNameLabel>
|
||
|
|
More or less pronounced displacements of the lateral pair of receptacles onto the dorsal wall of the bursa copulatrix in
|
||
|
|
<taxonomicName id="4C78848AB049FF9F2E4B1CCDEFDB5A94" authorityName="Ono" authorityYear="1997" box="[554,697,1250,1274]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="613" phylum="Arthropoda" rank="species" species="tomokunii">
|
||
|
|
<emphasis id="B90C231BB049FF9F2E4B1CCDEFDB5A94" box="[554,697,1250,1274]" italics="true" pageId="14" pageNumber="613">H. tomokunii</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
,
|
||
|
|
<taxonomicName id="4C78848AB049FF9F2EA91CCDEE2B5A94" baseAuthorityName="Ono" baseAuthorityYear="2002" box="[712,841,1250,1274]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="613" phylum="Arthropoda" rank="species" species="australis">
|
||
|
|
<emphasis id="B90C231BB049FF9F2EA91CCDEE2B5A94" box="[712,841,1250,1274]" italics="true" pageId="14" pageNumber="613">H. australis</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
and some
|
||
|
|
<taxonomicName id="4C78848AB049FF9F2FA51CCCEDB75B75" pageId="14" pageNumber="613">
|
||
|
|
<emphasis id="B90C231BB049FF9F2FA51CCCE9405A94" box="[964,1058,1250,1274]" italics="true" pageId="14" pageNumber="613">Ryuthela</emphasis>
|
||
|
|
species
|
||
|
|
</taxonomicName>
|
||
|
|
are considered homoplasic (see Discussion).
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="C362AC82B049FF9F2CB51D01EE305B08" pageId="14" pageNumber="613" type="distribution">
|
||
|
|
<paragraph id="8BC7FF09B049FF9F2CB51D01EE305B08" blockId="14.[136,1058,1326,1382]" pageId="14" pageNumber="613">
|
||
|
|
<materialsCitation id="3B10F554B049FF9F2CB51D01EE295B08" country="Vietnam" location="Tam Dao" pageId="14" pageNumber="613" specimenCount="1" stateProvince="Ha Noi">
|
||
|
|
DISTRIBUTION: This species is only known from the
|
||
|
|
<typeStatus id="54C341ABB049FF9F2F661D01EE575B29" box="[775,821,1327,1351]" pageId="14" pageNumber="613">type</typeStatus>
|
||
|
|
locality, the
|
||
|
|
<location id="8EA7A9D2B049FF9F2FA11D01E9405B28" LSID="urn:lsid:plazi:treatment:03D14E1FB049FF9F2CE918CDEE305B08:8EA7A9D2B049FF9F2FA11D01E9405B28" box="[960,1058,1327,1350]" country="Vietnam" name="Tam Dao" pageId="14" pageNumber="613" stateProvince="Ha Noi">Tam Dao</location>
|
||
|
|
hill station, about
|
||
|
|
<quantity id="4C8052ECB049FF9F2D2A1D60ECED5B08" box="[331,399,1358,1382]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="14" pageNumber="613" unit="km" value="85.0">85 km</quantity>
|
||
|
|
northwest of
|
||
|
|
<collectingRegion id="49BC31EBB049FF9F2E451D60EF075B08" box="[548,613,1358,1382]" country="Vietnam" name="Ha Noi" pageId="14" pageNumber="613">Hanoi</collectingRegion>
|
||
|
|
, in northern
|
||
|
|
<collectingCountry id="F36FBF99B049FF9F2E901D60EE295B08" box="[753,843,1358,1382]" name="Vietnam" pageId="14" pageNumber="613">Vietnam</collectingCountry>
|
||
|
|
</materialsCitation>
|
||
|
|
.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
|
</document>
|