treatments-xml/data/03/A3/87/03A387EDEC45FF1EFF27C372FD417A8F.xml

<document id="405F26886214AADCE0A9972D8D1A740C" ID-DOI="10.5281/zenodo.214864" ID-GBIF-Dataset="b17f3aca-8623-49fd-8f5e-d0cc601ba6cb" ID-ISSN="1175-5326" ID-Zenodo-Dep="214864" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460483566185" checkinUser="plazi" docAuthor="Carvalho, Thiago Ribeiro De, Giaretta, Ariovaldo Antonio &amp; Magrini, Leandro" docDate="2012" docId="03A387EDEC45FF1EFF27C372FD417A8F" docLanguage="en" docName="zt03321p055.pdf" docOrigin="Zootaxa 3321" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Bokermannohyla napolii Carvalho, Giaretta &amp; Magrini, 2012, sp. nov." docType="treatment" docVersion="9" lastPageNumber="46" masterDocId="FF9AFF95EC47FF17FFB0C625FFAC7D53" masterDocTitle="A new species of the Bokermannohyla circumdata group (Anura: Hylidae) from southeastern Brazil, with bioacoustic data on seven species of the genus" masterLastPageNumber="55" masterPageNumber="37" pageNumber="39" updateTime="1698308222184" updateUser="plazi">
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<mods:title id="C1A4E9847069D0DD409575077D4B12DA">A new species of the Bokermannohyla circumdata group (Anura: Hylidae) from southeastern Brazil, with bioacoustic data on seven species of the genus</mods:title>
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<mods:namePart id="D7DB667161390A76D21FA3C00F8DC32B">Carvalho, Thiago Ribeiro De</mods:namePart>
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<mods:namePart id="2CE141E661319D088431986EE807D93C">Giaretta, Ariovaldo Antonio</mods:namePart>
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<mods:namePart id="DC6B5478DA9209E5ED2BC124EA72B445">Magrini, Leandro</mods:namePart>
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<mods:date id="854E041CC1F3BA3059474352DA1DB664">2012</mods:date>
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<treatment id="03A387EDEC45FF1EFF27C372FD417A8F" ID-DOI="http://doi.org/10.5281/zenodo.5625495" ID-GBIF-Taxon="119517803" ID-Zenodo-Dep="5625495" LSID="urn:lsid:plazi:treatment:03A387EDEC45FF1EFF27C372FD417A8F" httpUri="http://treatment.plazi.org/id/03A387EDEC45FF1EFF27C372FD417A8F" lastPageId="9" lastPageNumber="46" pageId="2" pageNumber="39">
<subSubSection id="C3106570EC45FF15FF27C372FE3378C1" pageId="2" pageNumber="39" type="nomenclature">
<paragraph id="8BB536FBEC45FF15FF27C372FD867822" blockId="2.[151,554,1367,1426]" box="[151,554,1367,1393]" pageId="2" pageNumber="39">
<heading id="D0FD8197EC45FF15FF27C372FD867822" bold="true" box="[151,554,1367,1393]" fontSize="11" level="1" pageId="2" pageNumber="39" reason="1">
<emphasis id="B97EEAE9EC45FF15FF27C372FD867822" bold="true" box="[151,554,1367,1393]" pageId="2" pageNumber="39">
<taxonomicName id="4C0A4D78EC45FF15FF27C372FE697822" ID-CoL="68MY9" box="[151,453,1367,1393]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC45FF15FF27C372FE697822" bold="true" box="[151,453,1367,1393]" italics="true" pageId="2" pageNumber="39">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC45FF15FE7CC372FD867822" box="[460,554,1367,1393]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FBEC45FF15FF27C35FFE3378C1" blockId="2.[151,554,1367,1426]" box="[151,415,1402,1426]" pageId="2" pageNumber="39">
<figureCitation id="13312A7EEC45FF15FF27C35FFEAC78C1" box="[151,256,1402,1426]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1966]" captionTargetBox="[285,1300,195,1909]" captionTargetId="figure@4.[285,1300,195,1910]" captionTargetPageId="4" captionText="FIGURE 1. Adult specimens of Bokermannohyla napolii sp. nov. in life. Specimens from the Estação de Pesquisa e Desenvolvimento Ambiental Galheiro (EPDA-Galheiro), Municipality of Perdizes, State of Minas Gerais, Brazil. Upper, holotype UFMG 3333; Lower, female paratopotype ZUEC 16609." httpUri="https://zenodo.org/record/214865/files/figure.png" pageId="2" pageNumber="39">Figures 1</figureCitation>
, 
<figureCitation id="13312A7EEC45FF15FEBEC35FFEB078C1" box="[270,284,1402,1426]" captionStart="FIGURE 2" captionStartId="5.[151,250,1165,1188]" captionTargetBox="[209,1369,199,1108]" captionTargetId="figure@5.[151,1436,157,1145]" captionTargetPageId="5" captionText="FIGURE 2. Ventral view of hands of adult male specimens of Bokermannohyla napolii sp. nov. (left: ZUEC 16610) and B. luctuosa (right: AAG-UFU 0473). The arrows point to the divided tubercles of fingers III and IV in B. napolii sp. nov." httpUri="https://zenodo.org/record/214866/files/figure.png" pageId="2" pageNumber="39">2</figureCitation>
(left), and 3
</paragraph>
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<paragraph id="8BB536FBEC45FF15FF27C3E7FDB27BE6" blockId="2.[151,1437,1474,2005]" pageId="2" pageNumber="39">
<typeStatus id="54B18859EC45FF15FF27C3E7FEAF7889" box="[151,259,1474,1498]" pageId="2" pageNumber="39" type="holotype">Holotype</typeStatus>
. 
<collectionCode id="ED1BAE3EEC45FF15FEBDC3E7FEF27889" box="[269,350,1474,1498]" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="2" pageNumber="39">UFMG</collectionCode>
3333 (Ex-AAG-UFU 4767), adult male, collected at the Estação de Pesquisa e Desenvolvimento Ambiental Galheiro (EPDA-Galheiro; 19o12ʹS; 47 
<geoCoordinate id="EE3E503CEC45FF15FD64C3C3FD547B52" box="[724,760,1510,1537]" direction="west" orientation="longitude" pageId="2" pageNumber="39" precision="925" value="-0.13333334">o0 8</geoCoordinate>
ʹW, 
<quantity id="4CF29B1EEC45FF15FC97C3CCFCDD7B52" box="[807,881,1513,1537]" metricMagnitude="2" metricUnit="m" metricValue="8.45" pageId="2" pageNumber="39" unit="m" value="845.0">845 m</quantity>
a.s.l.), Municipality of Perdizes, State of Minas Gerais, southeastern 
<collectingCountry id="F31D766BEC45FF15FE32C028FE657B76" box="[386,457,1549,1573]" name="Brazil" pageId="2" pageNumber="39">Brazil</collectingCountry>
, on 
<date id="FFB4103BEC45FF15FE49C028FD167B76" box="[505,698,1549,1573]" pageId="2" pageNumber="39" value="2009-10-16">16 October 2009</date>
, by A. A. Giaretta, T. R. de Carvalho, L. B. Martins, and B. F. V. Teixeira. Paratopotypes. Five adult males: AAG-UFU 4774–4775, 
<collectionCode id="ED1BAE3EEC45FF15FC36C014FC7B7B1A" box="[902,983,1585,1609]" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="2" pageNumber="39">UFMG</collectionCode>
3334 (Ex-AAG-UFU 4770), and 
<collectionCode id="ED1BAE3EEC45FF15FAE3C014FA307B1A" box="[1363,1436,1585,1609]" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="2" pageNumber="39">ZUEC</collectionCode>
16610 (Ex-AAG-UFU 4771), collected with the 
<typeStatus id="54B18859EC45FF15FD71C070FC8A7B3E" box="[705,806,1621,1645]" pageId="2" pageNumber="39" type="holotype">holotype</typeStatus>
; AAG-UFU 0 860, on 
<date id="FFB4103BEC45FF15FB96C070FB5E7B3E" box="[1062,1266,1621,1645]" pageId="2" pageNumber="39" value="2011-12-03">3 December 2011</date>
, by A. A. Giaretta. Three adult females: AAG-UFU 4773, 
<collectionCode id="ED1BAE3EEC45FF15FD11C05CFD5E7BC2" box="[673,754,1657,1681]" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="2" pageNumber="39">UFMG</collectionCode>
3335 (Ex-AAG-UFU 4769), 
<collectionCode id="ED1BAE3EEC45FF15FBF9C05CFB3E7BC2" box="[1097,1170,1657,1681]" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="2" pageNumber="39">ZUEC</collectionCode>
16609 (Ex-AAG-UFU 4768), collected with the 
<typeStatus id="54B18859EC45FF15FE04C0B8FDB67BE6" box="[436,538,1693,1717]" pageId="2" pageNumber="39" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BB536FBEC45FF14FF77C0E4FA9D7D87" blockId="2.[151,1437,1474,2005]" lastBlockId="3.[151,1436,151,212]" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39">
Diagnosis. 
<taxonomicName id="4C0A4D78EC45FF15FEF9C0E4FDA47B8A" box="[329,520,1729,1753]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="39" phylum="Chordata" rank="genus">
<emphasis id="B97EEAE9EC45FF15FEF9C0E4FDA47B8A" box="[329,520,1729,1753]" italics="true" pageId="2" pageNumber="39">Bokermannohyla</emphasis>
</taxonomicName>
is currently supported only by molecular data, and a thorough morphological study of the genus still needs to be done (
<bibRefCitation id="EF9B4B0AEC45FF15FDDFC0C0FCC97BAE" author="Faivovich" box="[623,869,1765,1790]" pageId="2" pageNumber="39" refString="Faivovich, J., Haddad, C. F. B., Garcia, P. C. A., Frost, D. R., Campbell, J. A. &amp; Wheeler, W. C. (2005) Systematic Review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic revision. Bulletim of the American Museum Natural History, 294, 1 - 240." type="journal article" year="2005">
Faivovich 
<emphasis id="B97EEAE9EC45FF15FD59C0C2FC8E7BAD" box="[745,802,1766,1790]" italics="true" pageId="2" pageNumber="39">et al.</emphasis>
2005
</bibRefCitation>
, 
<bibRefCitation id="EF9B4B0AEC45FF15FCC1C0C0FC027BAE" author="Faivovich" box="[881,942,1765,1789]" pageId="2" pageNumber="39" refString="Faivovich, J., Lugli, L., Lourenco, A. C. C. &amp; Haddad, C. F. B. (2009) A new species of the Bokermannohyla martinsi group from central Bahia, Brazil with comments on Bokermannohyla (Anura: Hylidae). Herpetologica, 65, 303 - 310." type="journal article" year="2009">2009</bibRefCitation>
). Thus, we assign 
<taxonomicName id="4C0A4D78EC45FF15FBCFC0C2FB427BAD" box="[1151,1262,1766,1790]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC45FF15FBCFC0C2FB427BAD" box="[1151,1262,1766,1790]" italics="true" pageId="2" pageNumber="39">B. napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC45FF15FB46C0C3FAE27BAE" box="[1270,1358,1766,1789]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
to this genus on the basis of its phenotypic similarity to other species in the genus. 
<taxonomicName id="4C0A4D78EC45FF15FBB7C12CFAB37A72" box="[1031,1311,1801,1825]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC45FF15FBB7C12CFAB37A72" box="[1031,1311,1801,1825]" italics="true" pageId="2" pageNumber="39">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC45FF15FA98C12FFA2C7A72" box="[1320,1408,1802,1825]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
is assigned to the 
<taxonomicName id="4C0A4D78EC45FF15FEF3C10AFE4C7A15" box="[323,480,1838,1862]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC45FF15FEF3C10AFE4C7A15" box="[323,480,1838,1862]" italics="true" pageId="2" pageNumber="39">B. circumdata</emphasis>
</taxonomicName>
species group as it has transverse dark brown stripes on dorsum, simple (not dichotomized or anastomosed) dark brown transverse stripes on posterior surfaces of thighs and flanks, single (not bifid) large sharp prepollex, and adult males having hypertrophied forearms. The new taxon is diagnosed by the following combination of traits: (1) medium size (adult SVL 49.7–55.0 mm); (2) head wider than longer; (3) dark brown transverse stripes on dorsum; (4) dark brown vertical bars on posterior and superior surfaces of thighs; (5) vocal slits present in adult males; (6) bifid/divided distal tubercle of finger III in males, and bifid/divided distal tubercle of finger IV in males and females; (7) distinctive advertisement call pattern (see call description section).
</paragraph>
<caption id="DF756673EC44FF14FF27C721FAF97C25" ID-Table-UUID="DF756673EC44FF14FF27C721FAF97C25" httpUri="http://table.plazi.org/id/DF756673EC44FF14FF27C721FAF97C25" pageId="3" pageNumber="40" targetBox="[149,1436,398,1019]" targetIsTable="true" targetPageId="3">
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<emphasis id="B97EEAE9EC44FF14FF27C721FEAB7C4A" bold="true" box="[151,263,260,281]" pageId="3" pageNumber="40">TABLE 1.</emphasis>
Morphometric characters (mm) of adult males of 
<taxonomicName id="4C0A4D78EC44FF14FCA9C721FB897C49" box="[793,1061,260,282]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="luctuosa">
<emphasis id="B97EEAE9EC44FF14FCA9C721FB897C49" box="[793,1061,260,282]" italics="true" pageId="3" pageNumber="40">Bokermannohyla luctuosa</emphasis>
</taxonomicName>
(topotypes and specimens from the Municipalities of Cabreúva and Atibaia, State of São Paulo, Brazil); adult males of 
<taxonomicName id="4C0A4D78EC44FF14FC46C706FB2A7C6B" box="[1014,1158,290,312]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC44FF14FC46C706FB2A7C6B" box="[1014,1158,290,312]" italics="true" pageId="3" pageNumber="40">B. circumdata</emphasis>
</taxonomicName>
from the Municipalities of Petrópolis (State of Rio de Janeiro, Brazil), Arapeí and District of Paranapiacaba (State of São Paulo, Brazil), and Chiador (State of Minas Gerais, Brazil); and type series of 
<taxonomicName id="4C0A4D78EC44FF14FD23C744FD5B7C25" box="[659,759,352,374]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC44FF14FD23C744FD5B7C25" box="[659,759,352,374]" italics="true" pageId="3" pageNumber="40">B. napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC44FF14FD4DC744FCE07C25" box="[765,844,353,374]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
Mean + standard deviation (minimum–maximum).
</paragraph>
</caption>
<paragraph id="8BB536FBEC44FF14FF25C7ABFAF87EAA" pageId="3" pageNumber="40">
<table id="F90AC45BEC4400E8FF25C7ABFA307EA8" box="[149,1436,398,1019]" gridcols="4" gridrows="12" pageId="3" pageNumber="40">
<tr id="353A34B9EC4400E8FF25C7ABFA307F55" box="[149,1436,398,518]" gridrow="0" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C7ABFE3D7F55" box="[149,401,398,518]" gridcol="0" gridrow="0" pageId="3" pageNumber="40">Characters Snout-Vent Length</th>
<th id="76EB5DC5EC4400E8FE07C7ABFD227F55" box="[439,654,398,518]" gridcol="1" gridrow="0" pageId="3" pageNumber="40">
<taxonomicName id="4C0A4D78EC44FF14FE07C7AAFDA17C90" box="[439,612,399,451]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="luctuosa">
<emphasis id="B97EEAE9EC44FF14FE07C7AAFDA17C90" box="[439,612,399,451]" italics="true" pageId="3" pageNumber="40">Bokermannohyla luctuosa</emphasis>
</taxonomicName>
N=7 males 56.3+3.4 (53.4–61.9)
</th>
<th id="76EB5DC5EC4400E8FD01C7ABFC2B7F55" box="[689,903,398,518]" gridcol="2" gridrow="0" pageId="3" pageNumber="40">
<taxonomicName id="4C0A4D78EC44FF14FD01C7AAFC8F7C90" box="[689,862,399,451]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC44FF14FD01C7AAFC8F7C90" box="[689,862,399,451]" italics="true" pageId="3" pageNumber="40">Bokermannohyla circumdata</emphasis>
</taxonomicName>
N=10 males 59.0+3.4 (54.7–65.6)
</th>
<th id="76EB5DC5EC4400E8FC1AC7ABFA307F55" box="[938,1436,398,518]" gridcol="3" gridrow="0" pageId="3" pageNumber="40">
<taxonomicName id="4C0A4D78EC44FF14FC53C7AAFB727CF6" box="[995,1246,399,421]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC44FF14FC53C7AAFB727CF6" box="[995,1246,399,421]" italics="true" pageId="3" pageNumber="40">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC44FF14FB54C7AAFA9F7CF7" box="[1252,1331,399,420]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
-------------------------------------------------- N=6 males N=3 females 53.1+1.0 (52.5–55.0) 52.3+2.2 (49.7–53.7)
</th>
</tr>
<tr id="353A34B9EC4400E8FF25C438FA307F60" box="[149,1436,541,563]" gridrow="1" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C438FE3D7F60" box="[149,401,541,563]" gridcol="0" gridrow="1" pageId="3" pageNumber="40">Thigh Length</th>
<td id="76EB5DC5EC4400E8FE07C438FD227F60" box="[439,654,541,563]" gridcol="1" gridrow="1" pageId="3" pageNumber="40">30.3+1.7 (28.5–32.7)</td>
<td id="76EB5DC5EC4400E8FD01C438FC2B7F60" box="[689,903,541,563]" gridcol="2" gridrow="1" pageId="3" pageNumber="40">31.9+2.2 (28.7–36.3)</td>
<td id="76EB5DC5EC4400E8FC1AC438FA307F60" box="[938,1436,541,563]" gridcol="3" gridrow="1" pageId="3" pageNumber="40">28.9+0.5 (28.0–29.4) 28.3+0.7 (27.7–29.0)</td>
</tr>
<tr id="353A34B9EC4400E8FF25C46FFA307F32" box="[149,1436,586,609]" gridrow="2" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C46FFE3D7F32" box="[149,401,586,609]" gridcol="0" gridrow="2" pageId="3" pageNumber="40">Shank Length</th>
<td id="76EB5DC5EC4400E8FE07C46FFD227F32" box="[439,654,586,609]" gridcol="1" gridrow="2" pageId="3" pageNumber="40">30.8+1.7 (29.2–33.9)</td>
<td id="76EB5DC5EC4400E8FD01C46FFC2B7F32" box="[689,903,586,609]" gridcol="2" gridrow="2" pageId="3" pageNumber="40">32.5+2.2 (29.2–36.6)</td>
<td id="76EB5DC5EC4400E8FC1AC46FFA307F32" box="[938,1436,586,609]" gridcol="3" gridrow="2" pageId="3" pageNumber="40">29.6+0.6 (28.8–30.3) 28.8+0.7 (28.1–29.4)</td>
</tr>
<tr id="353A34B9EC4400E8FF25C45DFA307FDD" box="[149,1436,632,654]" gridrow="3" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C45DFE3D7FDD" box="[149,401,632,654]" gridcol="0" gridrow="3" pageId="3" pageNumber="40">Foot Length</th>
<td id="76EB5DC5EC4400E8FE07C45DFD227FDD" box="[439,654,632,654]" gridcol="1" gridrow="3" pageId="3" pageNumber="40">25.3+1.8 (23.7–28.8)</td>
<td id="76EB5DC5EC4400E8FD01C45DFC2B7FDD" box="[689,903,632,654]" gridcol="2" gridrow="3" pageId="3" pageNumber="40">26.8+1.7 (23.2–29.3)</td>
<td id="76EB5DC5EC4400E8FC1AC45DFA307FDD" box="[938,1436,632,654]" gridcol="3" gridrow="3" pageId="3" pageNumber="40">24.6+0.8 (23.2–25.4) 23.7+1.6 (21.8–24.8)</td>
</tr>
<tr id="353A34B9EC4400E8FF25C480FA307FE8" box="[149,1436,677,699]" gridrow="4" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C480FE3D7FE8" box="[149,401,677,699]" gridcol="0" gridrow="4" pageId="3" pageNumber="40">Hand Length</th>
<td id="76EB5DC5EC4400E8FE07C480FD227FE8" box="[439,654,677,699]" gridcol="1" gridrow="4" pageId="3" pageNumber="40">19.1+1.2 (17.9–20.8)</td>
<td id="76EB5DC5EC4400E8FD01C480FC2B7FE8" box="[689,903,677,699]" gridcol="2" gridrow="4" pageId="3" pageNumber="40">20.3+1.3 (18.3–22.5)</td>
<td id="76EB5DC5EC4400E8FC1AC480FA307FE8" box="[938,1436,677,699]" gridcol="3" gridrow="4" pageId="3" pageNumber="40">18.7+0.7 (18.1–20.0) 17.1+0.9 (16.1–17.8)</td>
</tr>
<tr id="353A34B9EC4400E8FF25C4F7FA307FBA" box="[149,1436,722,745]" gridrow="5" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C4F7FE3D7FBA" box="[149,401,722,745]" gridcol="0" gridrow="5" pageId="3" pageNumber="40">Head Length</th>
<td id="76EB5DC5EC4400E8FE07C4F7FD227FBA" box="[439,654,722,745]" gridcol="1" gridrow="5" pageId="3" pageNumber="40">20.1+1.1 (19.0–22.0)</td>
<td id="76EB5DC5EC4400E8FD01C4F7FC2B7FBA" box="[689,903,722,745]" gridcol="2" gridrow="5" pageId="3" pageNumber="40">21.5+1.3 (20.5–23.9)</td>
<td id="76EB5DC5EC4400E8FC1AC4F7FA307FBA" box="[938,1436,722,745]" gridcol="3" gridrow="5" pageId="3" pageNumber="40">19.3+0.4 (18.8–19.9) 18.5+0.7 (18.0–19.3)</td>
</tr>
<tr id="353A34B9EC4400E8FF25C525FA307E45" box="[149,1436,768,790]" gridrow="6" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C525FE3D7E45" box="[149,401,768,790]" gridcol="0" gridrow="6" pageId="3" pageNumber="40">Head Width</th>
<td id="76EB5DC5EC4400E8FE07C525FD227E45" box="[439,654,768,790]" gridcol="1" gridrow="6" pageId="3" pageNumber="40">20.7+1.1 (19.4–22.5)</td>
<td id="76EB5DC5EC4400E8FD01C525FC2B7E45" box="[689,903,768,790]" gridcol="2" gridrow="6" pageId="3" pageNumber="40">22.1+1.2 (20.9–24.2)</td>
<td id="76EB5DC5EC4400E8FC1AC525FA307E45" box="[938,1436,768,790]" gridcol="3" gridrow="6" pageId="3" pageNumber="40">20.1+0.6 (19.6–21.0) 18.9+1.0 (18.0–19.9)</td>
</tr>
<tr id="353A34B9EC4400E8FF25C508FA307E10" box="[149,1436,813,835]" gridrow="7" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C508FE3D7E10" box="[149,401,813,835]" gridcol="0" gridrow="7" pageId="3" pageNumber="40">Eye Diameter</th>
<td id="76EB5DC5EC4400E8FE07C508FD227E10" box="[439,654,813,835]" gridcol="1" gridrow="7" pageId="3" pageNumber="40">5.0+0.2 (4.7–5.3)</td>
<td id="76EB5DC5EC4400E8FD01C508FC2B7E10" box="[689,903,813,835]" gridcol="2" gridrow="7" pageId="3" pageNumber="40">5.3+0.3 (4.8–5.9)</td>
<td id="76EB5DC5EC4400E8FC1AC508FA307E10" box="[938,1436,813,835]" gridcol="3" gridrow="7" pageId="3" pageNumber="40">4.8+0.1 (4.7–5.0) 4.5+0.2 (4.3–4.6)</td>
</tr>
<tr id="353A34B9EC4400E8FF25C57FFA307E22" box="[149,1436,858,881]" gridrow="8" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C57FFE3D7E22" box="[149,401,858,881]" gridcol="0" gridrow="8" pageId="3" pageNumber="40">Tympanum Diameter</th>
<td id="76EB5DC5EC4400E8FE07C57FFD227E22" box="[439,654,858,881]" gridcol="1" gridrow="8" pageId="3" pageNumber="40">4.9+0.4 (4.1–5.3)</td>
<td id="76EB5DC5EC4400E8FD01C57FFC2B7E22" box="[689,903,858,881]" gridcol="2" gridrow="8" pageId="3" pageNumber="40">5.1+0.3 (4.6–5.3)</td>
<td id="76EB5DC5EC4400E8FC1AC57FFA307E22" box="[938,1436,858,881]" gridcol="3" gridrow="8" pageId="3" pageNumber="40">4.3+0.3 (4.0–4.7) 4.2+0.3 (3.9–4.5)</td>
</tr>
<tr id="353A34B9EC4400E8FF25C5ADFA307ECD" box="[149,1436,904,926]" gridrow="9" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C5ADFE3D7ECD" box="[149,401,904,926]" gridcol="0" gridrow="9" pageId="3" pageNumber="40">Eye-Nostril Distance</th>
<td id="76EB5DC5EC4400E8FE07C5ADFD227ECD" box="[439,654,904,926]" gridcol="1" gridrow="9" pageId="3" pageNumber="40">5.8+0.3 (5.4–6.2)</td>
<td id="76EB5DC5EC4400E8FD01C5ADFC2B7ECD" box="[689,903,904,926]" gridcol="2" gridrow="9" pageId="3" pageNumber="40">6.2+0.4 (5.6–6.7)</td>
<td id="76EB5DC5EC4400E8FC1AC5ADFA307ECD" box="[938,1436,904,926]" gridcol="3" gridrow="9" pageId="3" pageNumber="40">5.3+0.1 (5.1–5.4) 5.1+0.2 (5.0–5.3)</td>
</tr>
<tr id="353A34B9EC4400E8FF25C590FA307E9E" box="[149,1436,949,973]" gridrow="10" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C590FE3D7E9E" box="[149,401,949,973]" gridcol="0" gridrow="10" pageId="3" pageNumber="40">3rd Finger Disk Diameter</th>
<td id="76EB5DC5EC4400E8FE07C590FD227E9E" box="[439,654,949,973]" gridcol="1" gridrow="10" pageId="3" pageNumber="40">2.9+0.2 (2.7–3.3)</td>
<td id="76EB5DC5EC4400E8FD01C590FC2B7E9E" box="[689,903,949,973]" gridcol="2" gridrow="10" pageId="3" pageNumber="40">3.2+0.3 (2.8–3.6)</td>
<td id="76EB5DC5EC4400E8FC1AC590FA307E9E" box="[938,1436,949,973]" gridcol="3" gridrow="10" pageId="3" pageNumber="40">3.1+0.2 (2.7–3.3) 3.1+0.2 (3.0–3.3)</td>
</tr>
<tr id="353A34B9EC4400E8FF25C5C7FA307EA8" box="[149,1436,994,1019]" gridrow="11" pageId="3" pageNumber="40">
<th id="76EB5DC5EC4400E8FF25C5C7FE3D7EA8" box="[149,401,994,1019]" gridcol="0" gridrow="11" pageId="3" pageNumber="40">4th Toe Disk Diameter</th>
<td id="76EB5DC5EC4400E8FE07C5C7FD227EA8" box="[439,654,994,1019]" gridcol="1" gridrow="11" pageId="3" pageNumber="40">2.8+0.2 (2.6–3.1)</td>
<td id="76EB5DC5EC4400E8FD01C5C7FC2B7EA8" box="[689,903,994,1019]" gridcol="2" gridrow="11" pageId="3" pageNumber="40">3.0+0.3 (2.6–3.4)</td>
<td id="76EB5DC5EC4400E8FC1AC5C7FA307EA8" box="[938,1436,994,1019]" gridcol="3" gridrow="11" pageId="3" pageNumber="40">2.8+0.2 (2.5–3.1) 2.8+0.2 (2.7–3.0)</td>
</tr>
</table>
</paragraph>
<caption id="DF756673EC44FF14FF27C261FA30790A" box="[151,1436,1091,1114]" pageId="3" pageNumber="40">
<paragraph id="8BB536FBEC44FF14FF27C261FA30790A" blockId="3.[149,1436,1091,1970]" box="[151,1436,1091,1114]" pageId="3" pageNumber="40">
<emphasis id="B97EEAE9EC44FF14FF27C261FEAA790A" bold="true" box="[151,262,1092,1113]" pageId="3" pageNumber="40">TABLE 2.</emphasis>
Advertisement call variables of 
<taxonomicName id="4C0A4D78EC44FF14FDEBC261FCCB7909" box="[603,871,1092,1114]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="luctuosa">
<emphasis id="B97EEAE9EC44FF14FDEBC261FCCB7909" box="[603,871,1092,1114]" italics="true" pageId="3" pageNumber="40">Bokermannohyla luctuosa</emphasis>
</taxonomicName>
from the Municipality of Atibaia (State of São Paulo,
</paragraph>
</caption>
<paragraph id="8BB536FBEC44FF14FF27C247FA30792B" blockId="3.[149,1436,1091,1970]" box="[151,1436,1122,1144]" pageId="3" pageNumber="40">
<collectingCountry id="F31D766BEC44FF14FF27C247FF75792B" box="[151,217,1122,1144]" name="Brazil" pageId="3" pageNumber="40">Brazil</collectingCountry>
); 
<taxonomicName id="4C0A4D78EC44FF14FF5AC246FED4792B" box="[234,376,1122,1144]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC44FF14FF5AC246FED4792B" box="[234,376,1122,1144]" italics="true" pageId="3" pageNumber="40">B. circumdata</emphasis>
</taxonomicName>
from the Municipality of Chiador (State of Minas Gerais, 
<collectingCountry id="F31D766BEC44FF14FC74C247FBAA792B" box="[964,1030,1122,1144]" name="Brazil" pageId="3" pageNumber="40">Brazil</collectingCountry>
), and District of Paranapiacaba (State of
</paragraph>
<paragraph id="8BB536FBEC44FF14FF27C2A7FA3079C4" blockId="3.[149,1436,1091,1970]" box="[151,1436,1153,1175]" pageId="3" pageNumber="40">
São Paulo, 
<collectingCountry id="F31D766BEC44FF14FEBAC2A4FEE079C4" box="[266,332,1153,1175]" name="Brazil" pageId="3" pageNumber="40">Brazil</collectingCountry>
); and 
<taxonomicName id="4C0A4D78EC44FF14FE39C2A7FE4179C4" box="[393,493,1153,1175]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC44FF14FE39C2A7FE4179C4" box="[393,493,1153,1175]" italics="true" pageId="3" pageNumber="40">B. napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC44FF14FE44C2A7FDEE79C4" box="[500,578,1154,1175]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
from the 
<typeStatus id="54B18859EC44FF14FD16C2A7FD7E79C4" box="[678,722,1154,1175]" pageId="3" pageNumber="40">type</typeStatus>
locality. Mean + standard deviation (minimum–maximum). N = num-
</paragraph>
<paragraph id="8BB536FBEC44FF14FF27C285FED679E5" blockId="3.[149,1436,1091,1970]" box="[151,378,1184,1206]" pageId="3" pageNumber="40">ber of recorded males.</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C2EBFB067872" blockId="3.[149,1436,1091,1970]" pageId="3" pageNumber="40">
Variables 
<emphasis id="B97EEAE9EC44FF14FECCC2EAFCA07850" italics="true" pageId="3" pageNumber="40">
<taxonomicName id="4C0A4D78EC44FF14FECCC2EAFB8D79B6" box="[380,1057,1231,1253]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="genus">Bokermannohyla Bokermannohyla Bokermannohyla</taxonomicName>
<taxonomicName id="4C0A4D78EC44FF14FBCDC2EAFCA07850" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="subSpecies" species="luctuosa" subSpecies="circumdata">Bokermannohyla luctuosa circumdata</taxonomicName>
</emphasis>
(MG) 
<taxonomicName id="4C0A4D78EC44FF14FCC4C2C8FC4A7850" box="[884,998,1261,1283]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC44FF14FCC4C2C8FC4A7850" box="[884,998,1261,1283]" italics="true" pageId="3" pageNumber="40">circumdata</emphasis>
</taxonomicName>
(SP) 
<taxonomicName id="4C0A4D78EC44FF14FBCDC2C8FB697850" box="[1149,1221,1261,1283]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC44FF14FBCDC2C8FB697850" box="[1149,1221,1261,1283]" italics="true" pageId="3" pageNumber="40">napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC44FF14FB7BC2CBFAB67850" box="[1227,1306,1262,1283]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
N=1 N=1 N=1 N=6
</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C315FACD7815" blockId="3.[149,1436,1091,1970]" box="[149,1377,1328,1350]" pageId="3" pageNumber="40">Advertisement call 607.7+48.2 (550–708) __ __ 619.7+71.7 (460–807)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C36BFEB27836" blockId="3.[149,1436,1091,1970]" box="[149,286,1358,1381]" pageId="3" pageNumber="40">duration (ms)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C356FAF878DB" blockId="3.[149,1436,1091,1970]" box="[149,1364,1394,1417]" pageId="3" pageNumber="40">
Note 
<typeStatus id="54B18859EC44FF14FF7DC356FEAE78DB" box="[205,258,1395,1416]" pageId="3" pageNumber="40">types</typeStatus>
A/B 110.8+11.1 (97–138) __ __ 133.3+17.2 (82–167)
</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C3B4FEDB78FB" blockId="3.[149,1436,1091,1970]" box="[149,375,1425,1448]" pageId="3" pageNumber="40">internote interval (ms)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C393FAEB7898" blockId="3.[149,1436,1091,1970]" box="[149,1351,1461,1484]" pageId="3" pageNumber="40">
Note 
<typeStatus id="54B18859EC44FF14FF7DC393FF557898" box="[205,249,1462,1483]" pageId="3" pageNumber="40">type</typeStatus>
A duration 57.2+20.3 (23–93) 95.0+15.3 (71–120) __ 93.4+15.6 (53–156)
</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C3F1FF6878B9" blockId="3.[149,1436,1091,1970]" box="[149,196,1492,1514]" pageId="3" pageNumber="40">(ms)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C3DCFAC27B5E" blockId="3.[149,1436,1091,1970]" box="[149,1390,1528,1550]" pageId="3" pageNumber="40">
Note 
<typeStatus id="54B18859EC44FF14FF7CC3DCFF547B5D" box="[204,248,1529,1550]" pageId="3" pageNumber="40">type</typeStatus>
A dominant 0.57+0.09 (0.49–0.65) 0.48+0.12 (0.33– __ 0.43+0.07 (0.39–0.56) /
</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C033FACC7B7E" blockId="3.[149,1436,1091,1970]" box="[149,1376,1558,1581]" pageId="3" pageNumber="40">frequencies (kHz) / 0.66) / 0.73+0.05 0.77+0.06 (0.73–0.90)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C010FACC7B3A" blockId="3.[149,1436,1091,1970]" pageId="3" pageNumber="40">harmonics (0.70–0.80) / 1.47+0.05 (1.34–1.51) / 1.82+0.08 (1.68–1.85)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C05DFAB97BDD" blockId="3.[149,1436,1091,1970]" box="[149,1301,1656,1678]" pageId="3" pageNumber="40">Advertisement call 6.8+2.4 (4–8) __ 11.1+2.4 (8–14) 6.7+4.6 (1–14)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C0B2FEE37BFF" blockId="3.[149,1436,1091,1970]" box="[149,335,1686,1708]" pageId="3" pageNumber="40">rate (calls/minute)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C09EFACD7B83" blockId="3.[149,1436,1091,1970]" box="[149,1377,1722,1744]" pageId="3" pageNumber="40">
Note 
<typeStatus id="54B18859EC44FF14FF7DC09EFF557B83" box="[205,249,1723,1744]" pageId="3" pageNumber="40">type</typeStatus>
B duration 440.6+46.6 (371–526) 778.0+42.4 (748– 495.3+55.8 (235–544) 399.5+77.4 (242–550)
</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C0FFFD657BA3" blockId="3.[149,1436,1091,1970]" box="[149,713,1754,1776]" pageId="3" pageNumber="40">(ms) 808)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C0DBFAC27A40" blockId="3.[149,1436,1091,1970]" box="[149,1390,1789,1811]" pageId="3" pageNumber="40">
Note 
<typeStatus id="54B18859EC44FF14FF7DC0DBFF557A40" box="[205,249,1790,1811]" pageId="3" pageNumber="40">type</typeStatus>
B dominant 0.47 / 0.88+0.04 (0.82– 0.47+0.07 (0.42– 0.54+0.04 (0.52–0.61) / 0.51+0.06 (0.47–0.65) /
</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C13EFACC7A61" blockId="3.[149,1436,1091,1970]" box="[149,1376,1819,1842]" pageId="3" pageNumber="40">frequencies (kHz) / 0.90) / 1.23+0.06 (1.16– 0.52) / 0.89 1.05+0.05 (0.98–1.08) / 1.17+0.14 (0.82–1.34)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C11FFE1C7ADD" blockId="3.[149,1436,1091,1970]" pageId="3" pageNumber="40">harmonics 1.34) / 1.65+0.04 (1.59– 1.46+0.03 (1.45–1.55) / 1.57+0.07 (1.42–1.68) / 1.68) / 1.98+0.09 (1.85– 2.05+0.04 (2.04–2.11) 2.02)</paragraph>
<paragraph id="8BB536FBEC44FF14FF25C1BEFAA47AE2" blockId="3.[149,1436,1091,1970]" box="[149,1288,1947,1970]" pageId="3" pageNumber="40">Harmonic groups/note 12.2+1.3 (10–15) __ 15.7+1.6 (7–16) 2.5+1.2 (1–5)</paragraph>
<caption id="DF756673EC43FF13FF27C1B2FD747AB8" httpUri="https://zenodo.org/record/214865/files/figure.png" pageId="4" pageNumber="41" targetBox="[285,1300,195,1909]" targetPageId="4">
<paragraph id="8BB536FBEC43FF13FF27C1B2FD747AB8" blockId="4.[151,1436,1943,2027]" pageId="4" pageNumber="41">
<emphasis id="B97EEAE9EC43FF13FF27C1B2FEBF7AFE" bold="true" box="[151,275,1943,1966]" pageId="4" pageNumber="41">FIGURE 1.</emphasis>
Adult specimens of 
<taxonomicName id="4C0A4D78EC43FF13FE53C1BDFD727AFD" box="[483,734,1944,1966]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC43FF13FE53C1BDFD727AFD" box="[483,734,1944,1966]" italics="true" pageId="4" pageNumber="41">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC43FF13FD54C1BCFC9F7AFD" box="[740,819,1945,1966]" pageId="4" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
in life. Specimens from the Estação de Pesquisa e Desenvolvimento Ambiental Galheiro (EPDA-Galheiro), Municipality of Perdizes, State of Minas Gerais, Brazil. Upper, holotype UFMG 3333; Lower, female paratopotype ZUEC 16609.
</paragraph>
</caption>
<caption id="DF756673EC42FF12FF27C2A8FA8A7991" httpUri="https://zenodo.org/record/214866/files/figure.png" pageId="5" pageNumber="42" targetBox="[209,1369,199,1108]" targetPageId="5">
<paragraph id="8BB536FBEC42FF12FF27C2A8FA8A7991" blockId="5.[151,1436,1165,1219]" pageId="5" pageNumber="42">
<emphasis id="B97EEAE9EC42FF12FF27C2A8FEBE79F0" bold="true" box="[151,274,1165,1188]" pageId="5" pageNumber="42">FIGURE 2.</emphasis>
Ventral view of hands of adult male specimens of 
<taxonomicName id="4C0A4D78EC42FF12FCBEC2ABFBA579F7" box="[782,1033,1166,1188]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC42FF12FCBEC2ABFBA579F7" box="[782,1033,1166,1188]" italics="true" pageId="5" pageNumber="42">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC42FF12FBBFC2AAFBF179F7" box="[1039,1117,1167,1188]" pageId="5" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
(left: ZUEC 16610) and 
<taxonomicName id="4C0A4D78EC42FF12FAE9C2AAFF627990" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="luctuosa">
<emphasis id="B97EEAE9EC42FF12FAE9C2AAFF627990" italics="true" pageId="5" pageNumber="42">B. luctuosa</emphasis>
</taxonomicName>
(right: AAG-UFU 0473). The arrows point to the divided tubercles of fingers III and IV in 
<taxonomicName id="4C0A4D78EC42FF12FBDDC28BFB7E7990" box="[1133,1234,1197,1219]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC42FF12FBDDC28BFB7E7990" box="[1133,1234,1197,1219]" italics="true" pageId="5" pageNumber="42">B. napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC42FF12FB68C288FA8A7991" box="[1240,1318,1197,1218]" pageId="5" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BB536FBEC42FF12FF77C2CAFCB378C4" blockId="5.[151,1437,1263,2007]" pageId="5" pageNumber="42">
Comparisons with other species. 
<taxonomicName id="4C0A4D78EC42FF12FDFDC2CAFCCB7854" box="[589,871,1263,1287]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC42FF12FDFDC2CAFCCB7854" box="[589,871,1263,1287]" italics="true" pageId="5" pageNumber="42">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC42FF12FCC3C2D5FC627854" box="[883,974,1264,1287]" pageId="5" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
(adult male SVL 52.5–55.0 mm; adult female SVL 
<quantity id="4CF29B1EEC42FF12FE98C336FE617878" box="[296,461,1299,1323]" metricMagnitude="-2" metricUnit="m" metricValue="5.17" metricValueMax="5.37" metricValueMin="4.97" pageId="5" pageNumber="42" unit="mm" value="51.7" valueMax="53.7" valueMin="49.7">49.7–53.7 mm</quantity>
) is set apart by its larger size from the small-sized (combined SVL 
<quantity id="4CF29B1EEC42FF12FB7CC336FADF7878" box="[1228,1395,1299,1323]" metricMagnitude="-2" metricUnit="m" metricValue="3.885" metricValueMax="4.76" metricValueMin="3.01" pageId="5" pageNumber="42" unit="mm" value="38.85" valueMax="47.6" valueMin="30.1">30.1–47.6 mm</quantity>
; cf. 
<bibRefCitation id="EF9B4B0AEC42FF12FF27C312FE8A781C" author="Napoli" box="[151,294,1335,1359]" pageId="5" pageNumber="42" refString="Napoli, M. F. (2000) Taxonomia, variacao morfologica e distribuicao geografica das especies do grupo de Hyla circumdata (Cope, 1870) (Amphibia, Anura, Hylidae). Doctoral thesis, unpublished. Universidade Federal do Rio de Janeiro / Museu Nacional, Rio de Janeiro, 208 pp. Available from http: // www. amphibia. ufba. br / Public / Tese _ MFNapoli. htm (accessed on 14 November 2011" type="book" year="2000">Napoli 2000</bibRefCitation>
) species of the 
<taxonomicName id="4C0A4D78EC42FF12FE6AC31DFDD5781C" box="[474,633,1335,1359]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC42FF12FE6AC31DFDD5781C" box="[474,633,1335,1359]" italics="true" pageId="5" pageNumber="42">B. circumdata</emphasis>
</taxonomicName>
group [
<taxonomicName id="4C0A4D78EC42FF12FD65C31DFB9F781C" authority="Bokermann 1967" authorityName="Bokermann" authorityYear="1967" box="[725,1075,1335,1359]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="astartea">
<emphasis id="B97EEAE9EC42FF12FD65C31DFCFE781C" box="[725,850,1336,1359]" italics="true" pageId="5" pageNumber="42">B. astartea</emphasis>
(Bokermann 1967)
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FBF0C31DFF777820" authority="Napoli &amp; Caramaschi 2004" authorityName="Napoli &amp; Caramaschi" authorityYear="2004" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="feioi">
<emphasis id="B97EEAE9EC42FF12FBF0C31DFB3F781C" box="[1088,1171,1335,1359]" italics="true" pageId="5" pageNumber="42">B. feioi</emphasis>
(
<bibRefCitation id="EF9B4B0AEC42FF12FB12C312FF7F7820" author="Napoli" pageId="5" pageNumber="42" refString="Napoli, M. F. &amp; Caramaschi, U. (2004) Two new species of the Hyla circumdata group from Serra do Mar and Serra da Mantiqueira, Southeastern Brazil, with description of the advertisement call of Hyla ibitipoca (Anura, Hylidae). Copeia, 2004 (3), 534 - 545." type="journal article" year="2004">Napoli &amp; Caramaschi 2004</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FF59C379FD1E7820" authority="Caramaschi &amp; Feio 1990" authorityName="Caramaschi &amp; Feio" authorityYear="1990" box="[233,690,1371,1395]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="ibitipoca">
<emphasis id="B97EEAE9EC42FF12FF59C379FEC37820" box="[233,367,1371,1395]" italics="true" pageId="5" pageNumber="42">B. ibitipoca</emphasis>
(
<bibRefCitation id="EF9B4B0AEC42FF12FE31C37EFD067820" author="Caramaschi" box="[385,682,1371,1395]" pageId="5" pageNumber="42" refString="Caramaschi, U. &amp; Feio, R. N. (1990) A new species of Hyla (Anura, Hylidae) from southern Minas Gerais, Brazil. Copeia, 1990 (2), 542 - 546." type="journal article" year="1990">Caramaschi &amp; Feio 1990</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FD70C379FB3C7820" authority="Jim &amp; Caramaschi 1979" authorityName="Jim &amp; Caramaschi" authorityYear="1979" box="[704,1168,1371,1395]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="izecksohni">
<emphasis id="B97EEAE9EC42FF12FD70C379FCF47820" box="[704,856,1371,1395]" italics="true" pageId="5" pageNumber="42">B. izecksohni</emphasis>
(
<bibRefCitation id="EF9B4B0AEC42FF12FCD9C37EFB247820" author="Jim" box="[873,1160,1371,1395]" pageId="5" pageNumber="42" refString="Jim, J. &amp; Caramaschi, U. (1979) Uma nova especie de Hyla da regiao de Botucatu, Sao Paulo, Brasil (Amphibia, Anura). Revista Brasileira de Biologia, 39, 717 - 719." type="journal article" year="1979">Jim &amp; Caramaschi 1979</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FB2FC379FAB37820" box="[1183,1311,1372,1395]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="nanuzae">
<emphasis id="B97EEAE9EC42FF12FB2FC379FAB37820" box="[1183,1311,1372,1395]" italics="true" pageId="5" pageNumber="42">B. nanuzae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FA80C379FDF378C4" authority="Caramaschi, Napoli &amp; Bernardes 2001" authorityName="Caramaschi, Napoli &amp; Bernardes" authorityYear="2001" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="ravida">
<emphasis id="B97EEAE9EC42FF12FA80C379FA307820" box="[1328,1436,1371,1395]" italics="true" pageId="5" pageNumber="42">B. ravida</emphasis>
(
<bibRefCitation id="EF9B4B0AEC42FF12FF2EC35AFDFB78C4" author="Caramaschi" box="[158,599,1407,1431]" pageId="5" pageNumber="42" refString="Caramaschi, U., Napoli, M. F. &amp; Bernardes, A. T. (2001) Nova especie do grupo de Hyla circumdata (Cope, 1870) do Estado de Minas Gerais, Brasil (Amphibia, Anura, Hylidae). Boletim do Museu Nacional, N. S., Zoologia, Rio de Janeiro, 457, 1 - 11." type="journal article" year="2001">Caramaschi, Napoli &amp; Bernardes 2001</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="4C0A4D78EC42FF12FD29C3A5FCA178C4" box="[665,781,1408,1431]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="sazimai">
<emphasis id="B97EEAE9EC42FF12FD29C3A5FCA178C4" box="[665,781,1408,1431]" italics="true" pageId="5" pageNumber="42">B. sazimai</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="C3106570EC42FF11FF77C386FE577C37" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42" type="reference_group">
<paragraph id="8BB536FBEC42FF12FF77C386FCE37BAC" blockId="5.[151,1437,1263,2007]" pageId="5" pageNumber="42">
<taxonomicName id="4C0A4D78EC42FF12FF77C386FE7278E8" ID-CoL="68MY9" box="[199,478,1443,1467]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC42FF12FF77C386FE7278E8" box="[199,478,1443,1467]" italics="true" pageId="5" pageNumber="42">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC42FF12FE54C381FD9778E8" box="[484,571,1444,1467]" pageId="5" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
has a wider than longer head, whereas the head in 
<taxonomicName id="4C0A4D78EC42FF12FBC9C381FE91788C" ID-CoL="68MY5" authority="Napoli &amp; Caramaschi 2004" authorityName="Napoli &amp; Caramaschi" authorityYear="2004" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="ahenea">
<emphasis id="B97EEAE9EC42FF12FBC9C381FB4578E8" box="[1145,1257,1443,1467]" italics="true" pageId="5" pageNumber="42">B. ahenea</emphasis>
(
<bibRefCitation id="EF9B4B0AEC42FF12FB48C386FE99788C" author="Napoli" pageId="5" pageNumber="42" refString="Napoli, M. F. &amp; Caramaschi, U. (2004) Two new species of the Hyla circumdata group from Serra do Mar and Serra da Mantiqueira, Southeastern Brazil, with description of the advertisement call of Hyla ibitipoca (Anura, Hylidae). Copeia, 2004 (3), 534 - 545." type="journal article" year="2004">Napoli &amp; Caramaschi 2004</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FEF8C3EDFD4C788C" ID-CoL="5WMY4" authority="Napoli and Pimenta (2009)" authorityName="Napoli and Pimenta" authorityYear="2009" box="[328,736,1479,1503]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="capra">
<emphasis id="B97EEAE9EC42FF12FEF8C3EDFE04788C" box="[328,424,1480,1503]" italics="true" pageId="5" pageNumber="42">B. capra</emphasis>
<bibRefCitation id="EF9B4B0AEC42FF12FE1EC3E2FD4C788C" author="Napoli" box="[430,736,1479,1503]" pageId="5" pageNumber="42" refString="Napoli, M. F. &amp; Pimenta, B. V. S. (2009) A new species of the Bokermannohyla circumdata group (Anura: Hylidae) from the coastal forests of Bahia, northeastern Brazil. Copeia, 2009 (4), 674 - 683." type="journal article" year="2009">Napoli and Pimenta (2009)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FD5BC3EDFC90788C" ID-CoL="3NCTV" box="[747,828,1479,1503]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="feioi">
<emphasis id="B97EEAE9EC42FF12FD5BC3EDFC90788C" box="[747,828,1479,1503]" italics="true" pageId="5" pageNumber="42">B. feioi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FCF9C3EDFC08788C" ID-CoL="68N9R" box="[841,932,1479,1503]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="hylax">
<emphasis id="B97EEAE9EC42FF12FCF9C3EDFC08788C" box="[841,932,1479,1503]" italics="true" pageId="5" pageNumber="42">B. hylax</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0A4D78EC42FF12FC51C3EDFA2F788C" ID-CoL="5WMXX" authority="Napoli &amp; Pimenta 2003" authorityName="Napoli &amp; Pimenta" authorityYear="2003" box="[993,1411,1479,1503]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="lucianae">
<emphasis id="B97EEAE9EC42FF12FC51C3EDFBCC788C" box="[993,1120,1479,1503]" italics="true" pageId="5" pageNumber="42">B. lucianae</emphasis>
(
<bibRefCitation id="EF9B4B0AEC42FF12FBDEC3E2FAD7788C" author="Napoli" box="[1134,1403,1479,1503]" pageId="5" pageNumber="42" refString="Napoli, M. F. &amp; Pimenta, B. V. S. (2003) Nova especie do grupo de Hyla circumdata (Cope, 1870) do sul da Bahia, Brasil (Amphibia, Anura, Hylidae). Arquivos do Museu Nacional, Rio de Janeiro, 61, 189 - 194." type="journal article" year="2003">Napoli &amp; Pimenta 2003</bibRefCitation>
)
</taxonomicName>
is longer than wider. The new species is also distinguished by having dark brown transverse stripes on dorsum and limbs, which are immaculate in 
<taxonomicName id="4C0A4D78EC42FF12FDA3C035FD2B7B74" ID-CoL="68MY5" box="[531,647,1551,1575]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="ahenea">
<emphasis id="B97EEAE9EC42FF12FDA3C035FD2B7B74" box="[531,647,1551,1575]" italics="true" pageId="5" pageNumber="42">B. ahenea</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FD29C035FCB47B74" ID-CoL="5WMXS" box="[665,792,1552,1575]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="astartea">
<emphasis id="B97EEAE9EC42FF12FD29C035FCB47B74" box="[665,792,1552,1575]" italics="true" pageId="5" pageNumber="42">B. astartea</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FC9BC035FC167B74" ID-CoL="68MXS" box="[811,954,1551,1575]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B97EEAE9EC42FF12FC9BC035FC167B74" box="[811,954,1551,1575]" italics="true" pageId="5" pageNumber="42">B. carvalhoi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0A4D78EC42FF12FC4FC035FA3B7B74" ID-CoL="68MY3" authority="Peixoto &amp; Cruz 1992" authorityName="Peixoto &amp; Cruz" authorityYear="1992" box="[1023,1431,1551,1575]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="gouveai">
<emphasis id="B97EEAE9EC42FF12FC4FC035FBD77B74" box="[1023,1147,1552,1575]" italics="true" pageId="5" pageNumber="42">B. gouveai</emphasis>
(
<bibRefCitation id="EF9B4B0AEC42FF12FB3EC02AFA237B74" author="Peixoto" box="[1166,1423,1551,1575]" pageId="5" pageNumber="42" refString="Peixoto, O. L. &amp; Cruz, C. A. G. (1992) Nova especie de Hyla da Serra da Mantiqueira, Itatiaia, estado do Rio de Janeiro (Amphibia, Anura, Hylidae). Memorias do Instituto Oswaldo Cruz, 87, 197 - 200." type="journal article" year="1992">Peixoto &amp; Cruz 1992</bibRefCitation>
)
</taxonomicName>
; immaculate or with weak dark brown transverse bands in 
<taxonomicName id="4C0A4D78EC42FF12FC87C011FC367B18" ID-CoL="5WMY4" box="[823,922,1588,1611]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="capra">
<emphasis id="B97EEAE9EC42FF12FC87C011FC367B18" box="[823,922,1588,1611]" italics="true" pageId="5" pageNumber="42">B. capra</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FC1CC011FADA7B18" ID-CoL="5WMY3" authority="Napoli and Junca (2006)" authorityName="Napoli and Junca" authorityYear="2006" box="[940,1398,1587,1611]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="diamantina">
<emphasis id="B97EEAE9EC42FF12FC1CC011FBE07B18" box="[940,1100,1587,1611]" italics="true" pageId="5" pageNumber="42">B. diamantina</emphasis>
<bibRefCitation id="EF9B4B0AEC42FF12FBE7C016FADA7B18" author="Napoli" box="[1111,1398,1587,1611]" pageId="5" pageNumber="42" refString="Napoli, M. F. &amp; Junca, F. A. (2006) A new species of the Bokermannohyla circumdata group (Amphibia: Anura: Hylidae) from Chapada Diamantina, State of Bahia, Brazil. Zootaxa, 1244, 57 - 68." type="journal article" year="2006">Napoli and Juncá (2006)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FA34C011FEA07B3C" ID-CoL="68MXQ" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="izecksohni">
<emphasis id="B97EEAE9EC42FF12FA34C011FEA07B3C" italics="true" pageId="5" pageNumber="42">B. izecksohni</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0A4D78EC42FF12FEFFC07DFE7E7B3C" ID-CoL="5WMXX" box="[335,466,1623,1647]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="lucianae">
<emphasis id="B97EEAE9EC42FF12FEFFC07DFE7E7B3C" box="[335,466,1623,1647]" italics="true" pageId="5" pageNumber="42">B. lucianae</emphasis>
</taxonomicName>
; and with a reticulated pattern in 
<taxonomicName id="4C0A4D78EC42FF12FCD3C07DFC767B3C" ID-CoL="68N9W" box="[867,986,1624,1647]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="sazimai">
<emphasis id="B97EEAE9EC42FF12FCD3C07DFC767B3C" box="[867,986,1624,1647]" italics="true" pageId="5" pageNumber="42">B. sazimai</emphasis>
</taxonomicName>
. Furthermore, the new taxon is distinguished by having dark brown vertical bars on posterior surface of thighs, which is immaculate in 
<taxonomicName id="4C0A4D78EC42FF12FB5AC059FA977BC0" ID-CoL="3NCTV" box="[1258,1339,1659,1683]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="feioi">
<emphasis id="B97EEAE9EC42FF12FB5AC059FA977BC0" box="[1258,1339,1659,1683]" italics="true" pageId="5" pageNumber="42">B. feioi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FAF9C059FF6A7BE4" ID-CoL="68MY3" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="gouveai">
<emphasis id="B97EEAE9EC42FF12FAF9C059FF6A7BE4" italics="true" pageId="5" pageNumber="42">B. gouveai</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FF65C085FEFF7BE4" ID-CoL="68MXX" box="[213,339,1696,1719]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="nanuzae">
<emphasis id="B97EEAE9EC42FF12FF65C085FEFF7BE4" box="[213,339,1696,1719]" italics="true" pageId="5" pageNumber="42">B. nanuzae</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0A4D78EC42FF12FE24C085FDB97BE4" ID-CoL="5WMXX" box="[404,533,1695,1719]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="lucianae">
<emphasis id="B97EEAE9EC42FF12FE24C085FDB97BE4" box="[404,533,1695,1719]" italics="true" pageId="5" pageNumber="42">B. lucianae</emphasis>
</taxonomicName>
; fragmented in 
<taxonomicName id="4C0A4D78EC42FF12FD7DC085FCF57BE4" ID-CoL="68MXS" box="[717,857,1695,1719]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B97EEAE9EC42FF12FD7DC085FCF57BE4" box="[717,857,1695,1719]" italics="true" pageId="5" pageNumber="42">B. carvalhoi</emphasis>
</taxonomicName>
, and anastomosed in 
<taxonomicName id="4C0A4D78EC42FF12FBE3C085FE957B88" ID-CoL="5WMXV" authority="Vasconcelos &amp; Giaretta 2003" authorityName="Vasconcelos &amp; Giaretta" authorityYear="2003" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="vulcaniae">
<emphasis id="B97EEAE9EC42FF12FBE3C085FB4D7BE4" box="[1107,1249,1695,1719]" italics="true" pageId="5" pageNumber="42">B. vulcaniae</emphasis>
(
<bibRefCitation id="EF9B4B0AEC42FF12FB41C0BAFE9C7B88" author="Vasconcelos" pageId="5" pageNumber="42" refString="Vasconcelos, E. G. &amp; Giaretta, A. A. (2003) A new species of Hyla (Anura: Hylidae) from southeastern Brazil. Revista Espanola de Herpetologia, 17, 21 - 27." type="journal article" year="2003">Vasconcelos &amp; Giaretta 2003</bibRefCitation>
)
</taxonomicName>
. From 
<taxonomicName id="4C0A4D78EC42FF12FE39C0E1FD797B88" ID-CoL="5WN9S" authority="Napoli 2005" authorityName="Napoli" authorityYear="2005" box="[393,725,1731,1755]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B97EEAE9EC42FF12FE39C0E1FD9E7B88" box="[393,562,1731,1755]" italics="true" pageId="5" pageNumber="42">B. caramaschii</emphasis>
(
<bibRefCitation id="EF9B4B0AEC42FF12FDF0C0E6FD627B88" author="Napoli" box="[576,718,1731,1755]" pageId="5" pageNumber="42" refString="Napoli, M. F. (2005) A new species allied to Hyla circumdata (Anura: Hylidae) from Serra da Mantiqueira, southeastern Brazil. Herpetologica, 61, 63 - 69." type="journal article" year="2005">Napoli 2005</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C0A4D78EC42FF12FCBBC0E1FC317B88" ID-CoL="68MXQ" box="[779,925,1731,1755]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="izecksohni">
<emphasis id="B97EEAE9EC42FF12FCBBC0E1FC317B88" box="[779,925,1731,1755]" italics="true" pageId="5" pageNumber="42">B. izecksohni</emphasis>
</taxonomicName>
, the new species is also distinguished by having vocal slits in adult males (absent in the other two species).
</paragraph>
<paragraph id="8BB536FBEC42FF12FF77C12EFBC37A38" blockId="5.[151,1437,1263,2007]" pageId="5" pageNumber="42">
The distal bifid/divided subarticular tubercles of fingers III and IV in males, and finger IV in males and females (fig. 2) diagnose 
<taxonomicName id="4C0A4D78EC42FF12FE71C114FD9E7A14" ID-CoL="68MY9" box="[449,562,1839,1864]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC42FF12FE71C114FD9E7A14" box="[449,562,1839,1864]" italics="true" pageId="5" pageNumber="42">B. napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC42FF12FD8CC115FD3A7A14" box="[572,662,1840,1863]" pageId="5" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
from 
<taxonomicName id="4C0A4D78EC42FF12FD50C114FCF87A14" ID-CoL="68MY5" box="[736,852,1839,1864]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="ahenea">
<emphasis id="B97EEAE9EC42FF12FD50C114FCF87A14" box="[736,852,1839,1864]" italics="true" pageId="5" pageNumber="42">B. ahenea</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FCD4C114FC4F7A1B" ID-CoL="5WMXS" box="[868,995,1841,1864]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="astartea">
<emphasis id="B97EEAE9EC42FF12FCD4C114FC4F7A1B" box="[868,995,1841,1864]" italics="true" pageId="5" pageNumber="42">B. astartea</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FC44C114FB2D7A14" ID-CoL="68MXS" box="[1012,1153,1839,1864]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B97EEAE9EC42FF12FC44C114FB2D7A14" box="[1012,1153,1839,1864]" italics="true" pageId="5" pageNumber="42">B. carvalhoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FB23C114FB4B7A14" ID-CoL="3NCTV" box="[1171,1255,1839,1864]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="feioi">
<emphasis id="B97EEAE9EC42FF12FB23C114FB4B7A14" box="[1171,1255,1839,1864]" italics="true" pageId="5" pageNumber="42">B. feioi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FB48C114FADF7A14" ID-CoL="68MY3" box="[1272,1395,1840,1864]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="gouveai">
<emphasis id="B97EEAE9EC42FF12FB48C114FADF7A14" box="[1272,1395,1840,1864]" italics="true" pageId="5" pageNumber="42">B. gouveai</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FA34C114FF787A38" ID-CoL="68N9R" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="hylax">
<emphasis id="B97EEAE9EC42FF12FA34C114FF787A38" italics="true" pageId="5" pageNumber="42">B. hylax</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FF51C171FEC87A38" ID-CoL="5WMY2" box="[225,356,1875,1899]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="ibitipoca">
<emphasis id="B97EEAE9EC42FF12FF51C171FEC87A38" box="[225,356,1875,1899]" italics="true" pageId="5" pageNumber="42">B. ibitipoca</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FEC2C171FDA97A38" ID-CoL="68MXQ" box="[370,517,1875,1899]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="izecksohni">
<emphasis id="B97EEAE9EC42FF12FEC2C171FDA97A38" box="[370,517,1875,1899]" italics="true" pageId="5" pageNumber="42">B. izecksohni</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FDA4C171FD3E7A38" ID-CoL="5WMY9" box="[532,658,1875,1899]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="luctuosa">
<emphasis id="B97EEAE9EC42FF12FDA4C171FD3E7A38" box="[532,658,1875,1899]" italics="true" pageId="5" pageNumber="42">B. luctuosa</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FD10C171FCB07A38" ID-CoL="68MXX" box="[672,796,1876,1899]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="nanuzae">
<emphasis id="B97EEAE9EC42FF12FD10C171FCB07A38" box="[672,796,1876,1899]" italics="true" pageId="5" pageNumber="42">B. nanuzae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FC9AC171FC327A38" ID-CoL="68N9W" box="[810,926,1876,1899]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="sazimai">
<emphasis id="B97EEAE9EC42FF12FC9AC171FC327A38" box="[810,926,1876,1899]" italics="true" pageId="5" pageNumber="42">B. sazimai</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0A4D78EC42FF12FC6CC171FBC47A38" ID-CoL="5WMXV" box="[988,1128,1875,1899]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="vulcaniae">
<emphasis id="B97EEAE9EC42FF12FC6CC171FBC47A38" box="[988,1128,1875,1899]" italics="true" pageId="5" pageNumber="42">B. vulcaniae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB536FBEC42FF11FF77C152FE577C37" blockId="5.[151,1437,1263,2007]" lastBlockId="6.[151,1437,151,1981]" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42">
The advertisement call of 
<taxonomicName id="4C0A4D78EC42FF12FE5AC152FD537ADC" ID-CoL="68MY9" box="[490,767,1911,1935]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC42FF12FE5AC152FD537ADC" box="[490,767,1911,1935]" italics="true" pageId="5" pageNumber="42">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC42FF12FCB6C15DFCF07ADC" box="[774,860,1912,1935]" pageId="5" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
differs from that of all species of the 
<taxonomicName id="4C0A4D78EC42FF12FB4EC15DFA377ADC" ID-CoL="68MY4" box="[1278,1435,1911,1935]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC42FF12FB4EC15DFA377ADC" box="[1278,1435,1911,1935]" italics="true" pageId="5" pageNumber="42">B. circumdata</emphasis>
</taxonomicName>
group with calls described so far (unknown advertisement calls: 
<taxonomicName id="4C0A4D78EC42FF12FCC6C1B9FC447AE0" ID-CoL="68MY5" box="[886,1000,1947,1971]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="ahenea">
<emphasis id="B97EEAE9EC42FF12FCC6C1B9FC447AE0" box="[886,1000,1947,1971]" italics="true" pageId="5" pageNumber="42">B. ahenea</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FC47C1B9FB0E7AE0" ID-CoL="5WN9S" box="[1015,1186,1947,1971]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B97EEAE9EC42FF12FC47C1B9FB0E7AE0" box="[1015,1186,1947,1971]" italics="true" pageId="5" pageNumber="42">B. caramaschii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FB01C1B9FA877AE0" ID-CoL="68MY3" box="[1201,1323,1948,1971]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="gouveai">
<emphasis id="B97EEAE9EC42FF12FB01C1B9FA877AE0" box="[1201,1323,1948,1971]" italics="true" pageId="5" pageNumber="42">B. gouveai</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FA8AC1B9FF787A84" ID-CoL="68MXQ" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="izecksohni">
<emphasis id="B97EEAE9EC42FF12FA8AC1B9FF787A84" italics="true" pageId="5" pageNumber="42">B. izecksohni</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC42FF12FF51C1E5FEE57A84" ID-CoL="5WMY8" box="[225,329,1983,2007]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="ravida">
<emphasis id="B97EEAE9EC42FF12FF51C1E5FEE57A84" box="[225,329,1983,2007]" italics="true" pageId="5" pageNumber="42">B. ravida</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0A4D78EC42FF12FE36C1E5FDBD7A84" ID-CoL="5WMXV" box="[390,529,1983,2007]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="vulcaniae">
<emphasis id="B97EEAE9EC42FF12FE36C1E5FDBD7A84" box="[390,529,1983,2007]" italics="true" pageId="5" pageNumber="42">B. vulcaniae</emphasis>
</taxonomicName>
), except 
<taxonomicName id="4C0A4D78EC42FF12FDC8C1E5FCB97A84" ID-CoL="68MY4" box="[632,789,1983,2007]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC42FF12FDC8C1E5FCB97A84" box="[632,789,1983,2007]" italics="true" pageId="5" pageNumber="42">B. circumdata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A4D78EC42FF12FCFBC1E5FC657A84" ID-CoL="5WMY9" box="[843,969,1983,2007]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="luctuosa">
<emphasis id="B97EEAE9EC42FF12FCFBC1E5FC657A84" box="[843,969,1983,2007]" italics="true" pageId="5" pageNumber="42">B. luctuosa</emphasis>
</taxonomicName>
, due to its harmonic structure. The advertisement call of the new species (figs. 6 A–B) most resembles the call pattern of 
<taxonomicName id="4C0A4D78EC41FF11FB87C6BCFB797DE3" ID-CoL="68MY4" box="[1079,1237,152,176]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC41FF11FB87C6BCFB797DE3" box="[1079,1237,152,176]" italics="true" pageId="6" pageNumber="43">B. circumdata</emphasis>
</taxonomicName>
(fig. 6 C) and 
<taxonomicName id="4C0A4D78EC41FF11FA34C6BCFF5A7D87" ID-CoL="5WMY9" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="luctuosa">
<emphasis id="B97EEAE9EC41FF11FA34C6BCFF5A7D87" italics="true" pageId="6" pageNumber="43">B. luctuosa</emphasis>
</taxonomicName>
(fig. 6 D). Advertisement calls of 
<taxonomicName id="4C0A4D78EC41FF11FD33C698FC8E7D87" ID-CoL="68MY4" box="[643,802,188,212]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC41FF11FD33C698FC8E7D87" box="[643,802,188,212]" italics="true" pageId="6" pageNumber="43">B. circumdata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC41FF11FC82C698FC1D7D87" ID-CoL="5WMY9" box="[818,945,188,212]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="luctuosa">
<emphasis id="B97EEAE9EC41FF11FC82C698FC1D7D87" box="[818,945,188,212]" italics="true" pageId="6" pageNumber="43">B. luctuosa</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0A4D78EC41FF11FC42C698FBCF7D87" ID-CoL="68MY9" box="[1010,1123,188,212]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC41FF11FC42C698FBCF7D87" box="[1010,1123,188,212]" italics="true" pageId="6" pageNumber="43">B. napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC41FF11FBDBC698FB6F7D87" box="[1131,1219,189,212]" pageId="6" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
are composed of a short note followed by a long note. The first two have long notes composed of several disjunctive and/or juxtaposed harmonic groups, whereas the long note of 
<taxonomicName id="4C0A4D78EC41FF11FD73C720FC9D7C4F" ID-CoL="68MY9" box="[707,817,260,284]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC41FF11FD73C720FC9D7C4F" box="[707,817,260,284]" italics="true" pageId="6" pageNumber="43">B. napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC41FF11FC88C720FC227C4F" box="[824,910,261,284]" pageId="6" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
has fewer well-defined harmonic groups (1–5) with more sound intensity at the beginning of the note, or ill-defined harmonic groups with increasingly weaker sound intensity along its extent.
</paragraph>
</subSubSection>
<subSubSection id="C3106570EC41FF1EFF77C74AFD417A8F" lastPageId="9" lastPageNumber="46" pageId="6" pageNumber="43" type="description">
<paragraph id="8BB536FBEC41FF11FF77C74AFD8E7E9E" blockId="6.[151,1437,151,1981]" pageId="6" pageNumber="43">
Description of 
<typeStatus id="54B18859EC41FF11FEDEC74AFE787CD4" box="[366,468,367,391]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
. Adult male (UFMG 3333; figs. 1, 2(left), and 3). Body robust; head wider than longer. Snout rounded in dorsal and lateral views (figs. 3 A-B, respectively); nostrils directed laterally, much closer to the tip of snout than to the eye; canthus rostralis distinct; loreal region slightly concave; tympanum circular, 83% of the eye diameter; supra-tympanic fold present; pupil horizontal; vocal sac single, subgular, expanded; vocal slits present; tongue round; vomerine teeth in two almost contiguous, slightly arched series between choanae. Crenulated ridge on outer surface of forearms; forearm hypertrophied; finger lengths when put together I&lt;II&lt;IV&lt;III (fig. 3 C); disks of fingers II, III, and IV larger than that of I; disk of finger 
<date id="FFB4103BEC41FF11FCD6C46DFC097F0C" box="[870,933,583,607]" pageId="6" pageNumber="43" value="1975-03">III 75</date>
% of the tympanum diameter; distal subarticular tubercles of fingers I and II ovoid, those of fingers III and IV bifid/divided; nuptial pads present; pollical tubercle large, elongated; prepollex well-developed, spine curved, completely covered by skin; inner metacarpal tubercle region formed by several smaller granular tubercles; palmar tubercle developed, elongated, divided; supernumerary tubercles present on the palm of hand; hand webbing formula 
<date id="FFB4103BEC41FF11FC0DC4FEFB8A7FA1" box="[957,1062,730,754]" pageId="6" pageNumber="43">I 2–3 II 2</date>
-–2 2/3 
<date id="FFB4103BEC41FF11FBC5C4FEFB097FA1" box="[1141,1189,730,754]" pageId="6" pageNumber="43">III 2</date>
1/2– 
<date id="FFB4103BEC41FF11FB6DC4FFFAB87FA1" box="[1245,1300,730,754]" pageId="6" pageNumber="43">2 IV.</date>
Toe lengths when put together I&lt;II&lt;V&lt;III&lt;IV (fig. 3 D); disks of toes III, IV, and V larger than those of I and II; disk of toe 
<date id="FFB4103BEC41FF11FACFC525FF187E6E" pageId="6" pageNumber="43" value="1968-04">IV 68</date>
% of the tympanum diameter; toe webbing formula 
<date id="FFB4103BEC41FF11FCBAC503FCEB7E6E" box="[778,839,805,829]" pageId="6" pageNumber="43">I 1–2</date>
- 
<date id="FFB4103BEC41FF11FCE1C503FC437E6E" box="[849,1007,805,829]" pageId="6" pageNumber="43">II 1–2 III 1–2</date>
+ 
<date id="FFB4103BEC41FF11FC4DC503FB9E7E6E" box="[1021,1074,805,829]" pageId="6" pageNumber="43">IV 2</date>
+– 
<date id="FFB4103BEC41FF11FBFCC500FBDA7E6E" box="[1100,1142,805,829]" pageId="6" pageNumber="43">1 V</date>
; inner metatarsal tubercle ovoid, outer metatarsal tubercle barely distinct; toes fringed; supernumerary tubercles on the sole of foot; subarticular tubercles of toes rounded; inner tarsal fold weak, extending along full length of the tarsus; outer surface of tarsi. Dorsal surfaces and flanks smooth; belly granular; supra-vent granular ridge defined; discrete pointed dermal protuberance on outer heel present.
</paragraph>
<paragraph id="8BB536FBEC41FF11FF77C5FFFD56796A" blockId="6.[151,1437,151,1981]" pageId="6" pageNumber="43">
Measurements of 
<typeStatus id="54B18859EC41FF11FE22C5FCFE547EA2" box="[402,504,985,1009]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
. Morphometric characters (mm) and ratios (%) in relation to SVL (
<quantity id="4CF29B1EEC41FF11FB46C5FCFACE7EA2" box="[1270,1378,985,1009]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" pageId="6" pageNumber="43" unit="mm" value="52.5">52.5 mm</quantity>
): HL 19.2 (36.6), HW 19.6 (37.3), ED 4.8 (9.1), TD 4.0 (7.6), 
<collectingCountry id="F31D766BEC41FF11FC93C5DBFCF17946" box="[803,861,1022,1045]" name="India" pageId="6" pageNumber="43">END</collectingCountry>
5.1 (9.7), TL 28.5 (54.3), SL 29.0 (55.2), FL 24.5 (46.7), HAL 18.6 (35.4), 3FD 3.0 (5.7), 4TD 2.7 (5.1).
</paragraph>
<paragraph id="8BB536FBEC41FF11FF77C260FDAB7866" blockId="6.[151,1437,151,1981]" pageId="6" pageNumber="43">
Color of 
<typeStatus id="54B18859EC41FF11FE9CC260FE22790E" box="[300,398,1093,1117]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
in life (fig. 1). Dorsal surfaces uniformly pale cream, with subtly darker transverse stripes; a few white dots and blotches on dorsal surfaces and flanks; background color of the hidden parts of thighs and arms bluish and violet, with 8–9 dark brown vertical bars on posterior and superior surfaces; dorsal surface of shanks with 4–6 dark brown transverse stripes; flanks with subtle violet bars on a yellow marked region; inguinal region tending to the same color (violet) of the hidden parts of thighs, with a few dark brown transverse bars; supra-vent crenulated ridge white; crenulated ridge on outer forearm white; throat, chest, and belly light cream; iris gold, pupil black; tympanum tends to violet.
</paragraph>
<paragraph id="8BB536FBEC41FF11FF77C364FBAE7B2A" blockId="6.[151,1437,151,1981]" pageId="6" pageNumber="43">
Color of 
<typeStatus id="54B18859EC41FF11FE80C364FE3E780A" box="[304,402,1345,1369]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
in preservative. Dorsum dark grayish brown with some faded parts on dorsum and limbs, and dark brown transverse stripes barely distinct on dorsum; a few white blotches scattered on dorsum, limbs, and flanks; hidden parts of thighs and arms light gray, their posterior and superior surfaces with 8–9 dark brown vertical bars; anterior surface of thighs immaculate; dorsal surface of shanks with 4–6 dark brown transverse stripes, sometimes fragmented; dark brown transverse stripes on dorsal surface of forearms indistinct; background color of flanks light gray, with a few dark brown reticulations, especially on inguinal region; supra-vent crenulated ridge white; crenulated ridge on outer forearm white; throat, chest, belly, and ventral surfaces of limbs cream, gular region with dark brown punctuations bordering lower jaw, region of tubercles and webbing of hands and feet slightly gray; tympanum brown, slightly lighter than dorsal coloration pattern.
</paragraph>
<paragraph id="8BB536FBEC41FF11FF77C0A0FC947AEE" blockId="6.[151,1437,151,1981]" pageId="6" pageNumber="43">
Variation. Dorsum varies in color from medium brown (females) to dark gray (males) in ethanol preserved specimens. Number of dark brown vertical bars on dorsal surfaces of forearms (3–4), thighs (6–10), shanks (4–7), and tarsi (3–4). Dark brown transverse stripes on dorsal surfaces may be barely distinct. The male specimen AAG- UFU 0 860 has no distinctive transverse stripes on dorsal surfaces. The presence and extent of white blotches on dorsum and limbs are variable, being completely absent in AAG-UFU 4775. 
<typeStatus id="54B18859EC41FF11FBB0C133FB947A7D" box="[1024,1080,1814,1838]" pageId="6" pageNumber="43">Type</typeStatus>
series, except the 
<typeStatus id="54B18859EC41FF11FABAC130FAC07A7E" box="[1290,1388,1813,1837]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
and AAG-UFU 0 860, has a dark brown vertebral stripe, sometimes restricted to the first half of dorsal length. One female (AAG-UFU 4773) has distal subarticular tubercles of finger III divided; one male (AAG-UFU 4774) has distal subarticular tubercles fingers III and IV cordiform. Two male specimens (AAG-UFU 4775 and ZUEC 16610) have distal subarticular tubercles of finger I divided.
</paragraph>
<caption id="DF756673EC40FF10FF27C066FD4A7B2A" httpUri="https://zenodo.org/record/214867/files/figure.png" pageId="7" pageNumber="44" targetBox="[155,1412,218,1564]" targetPageId="7">
<paragraph id="8BB536FBEC40FF10FF27C066FD4A7B2A" blockId="7.[151,1436,1603,1657]" pageId="7" pageNumber="44">
<emphasis id="B97EEAE9EC40FF10FF27C066FEBF7B0A" bold="true" box="[151,275,1603,1626]" pageId="7" pageNumber="44">FIGURE 3.</emphasis>
<taxonomicName id="4C0A4D78EC40FF10FEABC061FDBB7B09" box="[283,535,1604,1626]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC40FF10FEABC061FDBB7B09" box="[283,535,1604,1626]" italics="true" pageId="7" pageNumber="44">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC40FF10FDAEC060FDDC7B09" box="[542,624,1605,1626]" pageId="7" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (UFMG 3333), adult male. Dorsal (A) and lateral (B) views of head; ventral view of hand (C) and foot (D). Scale bar = 10 mm.
</paragraph>
</caption>
<paragraph id="8BB536FBEC40FF1FFF77C085FE7E7C6C" blockId="7.[151,1436,1696,2009]" lastBlockId="8.[151,1437,151,320]" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44">
Natural History. Despite extensive fieldworks since 2001, this species was only heard (found) in 2009, during an explosive breeding event. Afterwards, a few males were heard once at the end of 2011. Males started calling late at night (around 01:00 AM), and a few males were heard still calling at nearly midday on the next day. They gathered around a muddy slow-watered section of a stream at the forest border. Males (N = 10) were calling from the ground or perched on low (&lt;
<quantity id="4CF29B1EEC40FF10FE6EC115FDA57A1B" box="[478,521,1840,1864]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="44" unit="m" value="1.0">1 m</quantity>
high) vegetation. Egg clutches (N = 4; fig. 4) were laid 
<quantity id="4CF29B1EEC40FF10FBCCC115FB717A1B" box="[1148,1245,1840,1864]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="7" pageNumber="44" unit="cm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 cm</quantity>
deep and hidden among dead broad leaves under water. Eggs were adhered to each other so that the entire egg mass (fresh, &lt;6 hours) could be removed from water using both hands to scoop it up. Number of eggs per clutch ranged from 373 to 742 (mean 637, SD = 176, N = 
<specimenCount id="9D0CFD72EC40FF10FDA1C1B9FDE27AE6" box="[529,590,1948,1973]" pageId="7" pageNumber="44" type="egg">4 egg</specimenCount>
clutches); yolk diameter ranged from 
<quantity id="4CF29B1EEC40FF10FC48C1B9FB3F7AE7" box="[1016,1171,1948,1972]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.3" metricValueMin="1.2" pageId="7" pageNumber="44" unit="mm" value="1.75" valueMax="2.3" valueMin="1.2">1.2 to 2.3 mm</quantity>
(mean 1.8, SD = 0.3, N = 
<specimenCount id="9D0CFD72EC40FF10FF1EC1E5FF477A8A" box="[174,235,1984,2009]" pageId="7" pageNumber="44" type="egg">3 egg</specimenCount>
clutches); gelatinous capsules ranged from 
<quantity id="4CF29B1EEC40FF10FD69C1E5FCDB7A8B" box="[729,887,1984,2008]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.4" metricValueMin="3.8" pageId="7" pageNumber="44" unit="mm" value="4.6" valueMax="5.4" valueMin="3.8">3.8 to 5.4 mm</quantity>
(mean 4.4, SD = 0.6, N = 
<specimenCount id="9D0CFD72EC40FF10FB12C1E5FB4D7A8A" box="[1186,1249,1984,2009]" pageId="7" pageNumber="44" type="egg">4 egg</specimenCount>
clutches). Three collected females released ovocytes inside plastic bags: yolk diameter ranged from 1.5 to 2.2 (mean 2.0, SD = 0.6); gelatinous capsules ranged from 
<quantity id="4CF29B1EEC4FFF1FFDB5C699FD337D87" box="[517,671,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.4" metricValueMin="3.1" pageId="8" pageNumber="45" unit="mm" value="4.25" valueMax="5.4" valueMin="3.1">3.1 to 5.4 mm</quantity>
(mean 4.2, SD = 0.5). Some males had their backs scratched (fig. 1), probably as a result of agonistic interactions. The new species occurs simpatrically with 
<taxonomicName id="4C0A4D78EC4FFF1FFBC9C6C5FA397DAB" box="[1145,1429,224,248]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="sazimai">
<emphasis id="B97EEAE9EC4FFF1FFBC9C6C5FA397DAB" box="[1145,1429,224,248]" italics="true" pageId="8" pageNumber="45">Bokermannohyla sazimai</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC4FFF1FFF27C721FDB57C4F" authority="Wied-Neuwied" authorityName="Wied-Neuwied" box="[151,537,260,284]" class="Amphibia" family="Hylidae" genus="Hypsiboas" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="faber">
<emphasis id="B97EEAE9EC4FFF1FFF27C721FEFD7C4F" box="[151,337,260,284]" italics="true" pageId="8" pageNumber="45">Hypsiboas faber</emphasis>
(Wied-Neuwied)
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC4FFF1FFD96C721FCE37C4F" authority="Lutz" authorityName="Lutz" box="[550,847,260,284]" class="Amphibia" family="Hylidae" genus="Hypsiboas" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="goianus">
<emphasis id="B97EEAE9EC4FFF1FFD96C721FD527C4F" box="[550,766,260,284]" italics="true" pageId="8" pageNumber="45">Hypsiboas goianus</emphasis>
(Lutz)
</taxonomicName>
, 
<taxonomicName id="4C0A4D78EC4FFF1FFCECC721FB187C4F" authority="Burmeister" authorityName="Burmeister" box="[860,1204,260,284]" class="Amphibia" family="Hylidae" genus="Hypsiboas" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="lundii">
<emphasis id="B97EEAE9EC4FFF1FFCECC721FBB17C4F" box="[860,1053,260,284]" italics="true" pageId="8" pageNumber="45">Hypsiboas lundii</emphasis>
(Burmeister)
</taxonomicName>
, and 
<taxonomicName id="4C0A4D78EC4FFF1FFB46C720FE637C6C" authority="Cardoso &amp; Haddad" authorityName="Cardoso &amp; Haddad" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="canastrensis">
<emphasis id="B97EEAE9EC4FFF1FFB46C720FF757C13" italics="true" pageId="8" pageNumber="45">Scinax canastrensis</emphasis>
(Cardoso &amp; Haddad)
</taxonomicName>
.
</paragraph>
<caption id="DF756673EC4FFF1FFF27C2C5FC237860" httpUri="https://zenodo.org/record/214868/files/figure.png" pageId="8" pageNumber="45" targetBox="[151,1436,373,1226]" targetPageId="8">
<paragraph id="8BB536FBEC4FFF1FFF27C2C5FC237860" blockId="8.[151,1436,1248,1331]" pageId="8" pageNumber="45">
<emphasis id="B97EEAE9EC4FFF1FFF27C2C5FEBF79A6" bold="true" box="[151,275,1248,1271]" pageId="8" pageNumber="45">FIGURE 4.</emphasis>
An egg clutch of 
<taxonomicName id="4C0A4D78EC4FFF1FFE7BC2C5FD6A79A5" box="[459,710,1248,1270]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC4FFF1FFE7BC2C5FD6A79A5" box="[459,710,1248,1270]" italics="true" pageId="8" pageNumber="45">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC4FFF1FFD7DC2C4FCB079A5" box="[717,796,1249,1270]" pageId="8" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
Eggs masses were laid hidden among dead broad leaves under water at the forest border. The egg clutch was exposed in order to take the picture. Clutches had an average of 637 eggs, each egg with a mean of 4.4 mm in diameter considering the gelatinous capsule.
</paragraph>
</caption>
<paragraph id="8BB536FBEC4FFF1FFF77C345FAFD782B" blockId="8.[151,1437,1376,1976]" box="[199,1361,1376,1401]" pageId="8" pageNumber="45">
Geographic distribution. 
<taxonomicName id="4C0A4D78EC4FFF1FFE6FC345FD58782B" box="[479,756,1376,1400]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC4FFF1FFE6FC345FD58782B" box="[479,756,1376,1400]" italics="true" pageId="8" pageNumber="45">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC4FFF1FFD4CC344FCFF782B" box="[764,851,1377,1400]" pageId="8" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
is known only from the 
<typeStatus id="54B18859EC4FFF1FFBD5C344FB39782A" box="[1125,1173,1377,1401]" pageId="8" pageNumber="45">type</typeStatus>
locality. (fig. 5).
</paragraph>
<paragraph id="8BB536FBEC4FFF1FFF77C3A1FD197893" blockId="8.[151,1437,1376,1976]" pageId="8" pageNumber="45">
Etymology. The name is a noun in the genitive case honoring Marcelo F. Napoli for his extensive contribution to the knowledge of the genus 
<taxonomicName id="4C0A4D78EC4FFF1FFE5EC38DFD017893" box="[494,685,1448,1472]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="B97EEAE9EC4FFF1FFE5EC38DFD017893" box="[494,685,1448,1472]" italics="true" pageId="8" pageNumber="45">Bokermannohyla</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB536FBEC4FFF1FFF77C3E9FAA47AEB" blockId="8.[151,1437,1376,1976]" pageId="8" pageNumber="45">
Vocalizations of 
<taxonomicName id="4C0A4D78EC4FFF1FFE32C3E9FD3478B7" box="[386,664,1484,1508]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC4FFF1FFE32C3E9FD3478B7" box="[386,664,1484,1508]" italics="true" pageId="8" pageNumber="45">Bokermannohyla napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC4FFF1FFD2EC3E8FD5978B7" box="[670,757,1485,1508]" pageId="8" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
Six males recorded (N = 76 advertisement call samples; N = 43 aggressive call samples). Quantitative traits are summarized in 
<tableCitation id="C6880340EC4FFF1FFC32C3D5FC757B5B" box="[898,985,1520,1544]" captionStart="TABLE 2" captionStartId="3.[151,235,1092,1113]" captionText="TABLE 2. Advertisement call variables of Bokermannohyla luctuosa from the Municipality of Atibaia (State of São Paulo," pageId="8" pageNumber="45">Table 2</tableCitation>
. Advertisement call is composed of two note 
<typeStatus id="54B18859EC4FFF1FFF7EC030FEA57B7E" box="[206,265,1557,1581]" pageId="8" pageNumber="45">types</typeStatus>
(referred herein to as note 
<typeStatus id="54B18859EC4FFF1FFD88C030FDDF7B7E" box="[568,627,1557,1581]" pageId="8" pageNumber="45">types</typeStatus>
A and B; figs. 6 A-B) emitted consecutively at a rate of 1–14 calls/minute (mean 6.7, SD = 4.6). Additional notes B may eventually be emitted just after a typical A/B sequence though. Advertisement call duration (notes A + B) was 460–807 ms (mean 619.7, SD = 71.7). Note A is a short note (53– 156 ms; mean 93.4, SD = 15.6) with 5–6 harmonics. Dominant frequencies from 0.39–0.56 kHz (mean 0.43, SD = 0.07), from 0.73–0.90 kHz (mean 0.77, SD = 0.06), from 1.34–1.51 kHz (mean 1.47, SD = 0.05), and from 1.68– 1.85 kHz (mean 1.82, SD = 0.08), which correspond to the first four harmonics. Note B is a long note (242–550 ms; mean 399.5, SD = 77.4) with a harmonic structure (5–6 harmonics; sometimes ill-defined), and sound intensity concentrated the most on the first part of the note, and sound intensity modulations forming 1–5 disjunctive or juxtaposed harmonic groups (mean 2.5, SD = 1.2) along its extent, or with ill-defined harmonic structure (N = 8 note samples). A/B internote interval from 82–167 ms (mean 133.3, SD = 17.2). Dominant frequencies from 0.47–0.65 kHz (mean 0.51, SD = 0.06), from 0.82–1.34 kHz (mean 1.17, SD = 0.14), from 1.42–1.68 kHz (mean 1.57, SD = 0.07), and from 2.02–2.11 kHz (mean 2.05, SD = 0.04), which correspond to the first four harmonics.
</paragraph>
<caption id="DF756673EC4EFF1EFF27C020FEA67B24" httpUri="https://zenodo.org/record/214869/files/figure.png" pageId="9" pageNumber="46" targetBox="[160,1428,198,1514]" targetPageId="9">
<paragraph id="8BB536FBEC4EFF1EFF27C020FEA67B24" blockId="9.[151,1437,1541,1655]" pageId="9" pageNumber="46">
<emphasis id="B97EEAE9EC4EFF1EFF27C020FEBE7B48" bold="true" box="[151,274,1541,1564]" pageId="9" pageNumber="46">FIGURE 5.</emphasis>
Type localities (based on original descriptions) of medium and large-sized species of the 
<taxonomicName id="4C0A4D78EC4EFF1EFB2BC020FF687B69" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC4EFF1EFB2BC020FF687B69" italics="true" pageId="9" pageNumber="46">Bokermannohyla circumdata</emphasis>
</taxonomicName>
species group on a topographic map: 
<taxonomicName id="4C0A4D78EC4EFF1EFDFCC000FD1E7B69" box="[588,690,1572,1594]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="napolii" status="sp. nov.">
<emphasis id="B97EEAE9EC4EFF1EFDFCC000FD1E7B69" box="[588,690,1572,1594]" italics="true" pageId="9" pageNumber="46">B. napolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5792EC4EFF1EFD09C000FCA57B69" box="[697,777,1573,1594]" pageId="9" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
(star); 
<taxonomicName id="4C0A4D78EC4EFF1EFCE4C000FC787B69" box="[852,980,1572,1594]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="vulcaniae">
<emphasis id="B97EEAE9EC4EFF1EFCE4C000FC787B69" box="[852,980,1572,1594]" italics="true" pageId="9" pageNumber="46">B. vulcaniae</emphasis>
</taxonomicName>
(1); 
<taxonomicName id="4C0A4D78EC4EFF1EFBB7C000FBD77B69" box="[1031,1147,1572,1594]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="luctuosa">
<emphasis id="B97EEAE9EC4EFF1EFBB7C000FBD77B69" box="[1031,1147,1572,1594]" italics="true" pageId="9" pageNumber="46">B. luctuosa</emphasis>
</taxonomicName>
(2); 
<taxonomicName id="4C0A4D78EC4EFF1EFB1EC000FAAE7B69" box="[1198,1282,1572,1594]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="hylax">
<emphasis id="B97EEAE9EC4EFF1EFB1EC000FAAE7B69" box="[1198,1282,1572,1594]" italics="true" pageId="9" pageNumber="46">B. hylax</emphasis>
</taxonomicName>
(3); 
<taxonomicName id="4C0A4D78EC4EFF1EFA85C000FA307B69" box="[1333,1436,1572,1594]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="ahenea">
<emphasis id="B97EEAE9EC4EFF1EFA85C000FA307B69" box="[1333,1436,1572,1594]" italics="true" pageId="9" pageNumber="46">B. ahenea</emphasis>
</taxonomicName>
(4); 
<taxonomicName id="4C0A4D78EC4EFF1EFF72C066FE957B0B" box="[194,313,1603,1624]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="gouveaei">
<emphasis id="B97EEAE9EC4EFF1EFF72C066FE957B0B" box="[194,313,1603,1624]" italics="true" pageId="9" pageNumber="46">B. gouveaei</emphasis>
</taxonomicName>
(5); 
<taxonomicName id="4C0A4D78EC4EFF1EFEDBC066FE447B0B" box="[363,488,1602,1624]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B97EEAE9EC4EFF1EFEDBC066FE447B0B" box="[363,488,1602,1624]" italics="true" pageId="9" pageNumber="46">B. carvalhoi</emphasis>
</taxonomicName>
(6); 
<taxonomicName id="4C0A4D78EC4EFF1EFDAAC066FD047B0B" box="[538,680,1602,1624]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="circumdata">
<emphasis id="B97EEAE9EC4EFF1EFDAAC066FD047B0B" box="[538,680,1602,1624]" italics="true" pageId="9" pageNumber="46">B. circumdata</emphasis>
</taxonomicName>
(7)*; 
<taxonomicName id="4C0A4D78EC4EFF1EFD56C066FC2C7B0B" box="[742,896,1602,1624]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B97EEAE9EC4EFF1EFD56C066FC2C7B0B" box="[742,896,1602,1624]" italics="true" pageId="9" pageNumber="46">B. caramaschii</emphasis>
</taxonomicName>
(8); 
<taxonomicName id="4C0A4D78EC4EFF1EFC01C066FBEC7B0B" box="[945,1088,1602,1624]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="diamantina">
<emphasis id="B97EEAE9EC4EFF1EFC01C066FBEC7B0B" box="[945,1088,1602,1624]" italics="true" pageId="9" pageNumber="46">B. diamantina</emphasis>
</taxonomicName>
(9); and 
<taxonomicName id="4C0A4D78EC4EFF1EFB2DC066FB587B0B" box="[1181,1268,1603,1624]" class="Amphibia" family="Hylidae" genus="Bokermannohyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="capra">
<emphasis id="B97EEAE9EC4EFF1EFB2DC066FB587B0B" box="[1181,1268,1603,1624]" italics="true" pageId="9" pageNumber="46">B. capra</emphasis>
</taxonomicName>
(10). *Based on Frost 2011.
</paragraph>
</caption>
<paragraph id="8BB536FBEC4EFF1EFF77C080FD417A8F" blockId="9.[151,1436,1700,2013]" pageId="9" pageNumber="46">
We have also identified a third note 
<typeStatus id="54B18859EC4EFF1EFDD6C080FD3A7BEE" box="[614,662,1701,1725]" pageId="9" pageNumber="46">type</typeStatus>
(referred herein to as note 
<typeStatus id="54B18859EC4EFF1EFC7FC080FC537BEE" box="[975,1023,1701,1725]" pageId="9" pageNumber="46">type</typeStatus>
C; fig. 7 A) that should possibly be attributed to a kind of aggressive call (
<emphasis id="B97EEAE9EC4EFF1EFDFAC0ECFD247BB3" box="[586,648,1737,1760]" italics="true" pageId="9" pageNumber="46">sensu</emphasis>
<bibRefCitation id="EF9B4B0AEC4EFF1EFD20C0EDFCBD7BB3" author="Wells" box="[656,785,1736,1760]" pageId="9" pageNumber="46" refString="Wells, K. D. (2007) Anuran vocal communication. In: Wells, K. D. (Eds.), The Ecology and Behavior of Amphibians. University of Chicago Press, Chicago &amp; London, pp. 268 - 337." type="book chapter" year="2007">Wells 2007</bibRefCitation>
) because it has an ill-defined lower sound intensity structure with remarkable differences in temporal traits (longer duration) in comparison with note B, yet similar in spectral traits (dominant frequencies). Besides, this aggressive call has no distinguishable harmonic structure as in notes A and B of the advertisement call. Note C is emitted at a rate of 12–26 notes/minute (mean 20.0, SD = 5.7) alone or just after a note A. The emission rate of advertisement calls of a given male was inversely proportional to the emission rate of aggressive calls. Note C duration was 437–1,120 ms (mean 763.9, SD = 210.4) with A/C internote interval from 90–128 ms (mean 115.3, SD = 14.6). Despite the absence of well-defined harmonic structure, dominant frequencies coincided with those of note 
<typeStatus id="54B18859EC4EFF1EFD2AC1E0FD667A8E" box="[666,714,1989,2013]" pageId="9" pageNumber="46">type</typeStatus>
A.
</paragraph>
</subSubSection>
</treatment>
</document>