291 lines
29 KiB
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291 lines
29 KiB
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<document id="D7B53E6BE4F3F4E9699CC201ECC5583A" ID-DOI="10.1093/isd/ixab025" ID-ISSN="2399-3421" ID-Zenodo-Dep="7182517" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665268061086" checkinUser="felipe" docAuthor="Ward, Philip S. & Branstetter, Michael G." docDate="2022" docId="0394878EFFF5FFD8B831F964BF2FF954" docLanguage="en" docName="InsectSystDivers.6.1.4.1-31.pdf" docOrigin="Insect Systematics and Diversity (AIFB) 6 (1)" docSource="http://dx.doi.org/10.1093/isd/ixab025" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.8:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="8" docTitle="Pseudomyrmex comitator Ward & Branstetter 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="22" masterDocId="FFADFFF6FFE1FFCDBB03FFBDBE0EFF97" masterDocTitle="Species Paraphyly and Social Parasitism: Phylogenomics, Morphology, and Geography Clarify the Evolution of the Pseudomyrmex elongatulus Group (Hymenoptera:" masterLastPageNumber="31" masterPageNumber="1" pageNumber="21" updateTime="1699375543922" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2B65D8A7E37F45B5D3D1B07F07DDB708">Species Paraphyly and Social Parasitism: Phylogenomics, Morphology, and Geography Clarify the Evolution of the Pseudomyrmex elongatulus Group (Hymenoptera:</mods:title>
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<mods:name id="BEC4D783736C0C37D119FAC6AD97B8D0" type="personal">
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<mods:roleTerm id="F130BEE320B97EEE33E079CA5927B4BA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8C5BD28702665845F2994164E21B1666">Ward, Philip S.</mods:namePart>
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<mods:affiliation id="1C2F807A02E0A98BFECF5B250A10F830">Department of Entomology and Nematology, University of California, Davis, CA 95616, USA,</mods:affiliation>
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</mods:name>
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<mods:name id="8B4AF9994AF636C7C351C12C5C2A66F4" type="personal">
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</mods:role>
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<mods:namePart id="137101FF53C0F97B76606809DD962EE5">Branstetter, Michael G.</mods:namePart>
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<mods:affiliation id="4125FC6746F13F2ADDA6D46BEA353858">U. S. Department of Agriculture,</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="4CD351DEB6BBA4A00E014DF0716F70F6">text</mods:typeOfResource>
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<mods:title id="2552FB9A8A29519655FAB8E1755B8880">Insect Systematics and Diversity</mods:title>
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</mods:titleInfo>
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<mods:part id="8A5655497399D9C68F9D44A34F31A9ED">
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<mods:date id="F5D575CBF5C06C0A2A4000996C8E5048">2022</mods:date>
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<mods:detail id="0C7D1A72BA5AAA8F00162030C9AA5F56" type="series">
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<mods:title id="95ED9AF8977D26D430E139DEA7BB286E">AIFB</mods:title>
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</mods:detail>
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<mods:detail id="BD63CD02BB7E547C968C35737105F0DC" type="pubDate">
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<mods:number id="CB43E0E69AA230400CC106564CFB0405">2022-01-25</mods:number>
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</mods:detail>
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<mods:detail id="F45D42477E3CF81DF26A0BD96B29171A" type="volume">
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<mods:number id="F1D0806535030D33863A3F2F6FDCF36C">6</mods:number>
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</mods:detail>
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<mods:detail id="B2EB233FCCB5FAF2019C0BA819B23398" type="issue">
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<mods:number id="A0791BD56D4FAA959FB1CB452B885F25">1</mods:number>
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<mods:extent id="97472F2F6680FCF2103866CAE277D75A" unit="page">
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<mods:start id="F5D7550040156033308A85D7C3DED753">1</mods:start>
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<mods:end id="D79C83FF267979FF470310B55B8C409F">31</mods:end>
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<mods:location id="DE1CCF4B5C9AAA10678282D996A40CF6">
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<mods:url id="B6E3097C74CBFC5B19DCB25AC3CC04CF">http://dx.doi.org/10.1093/isd/ixab025</mods:url>
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</mods:location>
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<mods:classification id="E3215B38F2651D30D5693DCBE5ABC9D7">journal article</mods:classification>
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<mods:identifier id="245C5C609E4B05056709C735F0E75BA5" type="DOI">10.1093/isd/ixab025</mods:identifier>
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<mods:identifier id="5795483C48985CCDE1468034C304FC26" type="ISSN">2399-3421</mods:identifier>
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<mods:identifier id="76B7D632EA0D0F0C486F50651BF5C08A" type="Zenodo-Dep">7182517</mods:identifier>
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<treatment id="0394878EFFF5FFD8B831F964BF2FF954" ID-DOI="http://doi.org/10.5281/zenodo.7182513" ID-Zenodo-Dep="7182513" LSID="urn:lsid:plazi:treatment:0394878EFFF5FFD8B831F964BF2FF954" httpUri="http://treatment.plazi.org/id/0394878EFFF5FFD8B831F964BF2FF954" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="C3276513FFF5FFD9B831F964BDBAF89C" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8B823698FFF5FFD9B831F964BA62F97A" blockId="20.[818,1132,1753,1773]" box="[818,1132,1753,1773]" pageId="20" pageNumber="21">
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<heading id="D0CA81F4FFF5FFD9B831F964BA62F97A" box="[818,1132,1753,1773]" fontSize="8" level="2" pageId="20" pageNumber="21" reason="4">
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<emphasis id="B949EA8AFFF5FFD9B831F964BA62F97A" bold="true" box="[818,1132,1753,1773]" pageId="20" pageNumber="21">
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<taxonomicName id="4C3D4D1BFFF5FFD9B831F964BA2FF97A" authority="Ward & Branstetter, 2022" authorityName="Ward & Branstetter" authorityYear="2022" box="[818,1057,1753,1773]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="comitator" status="sp. nov.">
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<emphasis id="B949EA8AFFF5FFD9B831F964BA2FF97A" bold="true" box="[818,1057,1753,1773]" italics="true" pageId="20" pageNumber="21">Pseudomyrmex comitator</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A57F1FFF5FFD9BF24F964BA62F97A" box="[1063,1132,1753,1773]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B823698FFF5FFD9B831F94ABDBAF89C" blockId="20.[818,1441,1783,1861]" box="[818,948,1783,1803]" pageId="20" pageNumber="21">Figs. 9 and 24</paragraph>
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</subSubSection>
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<subSubSection id="C3276513FFF5FFD9B831F8A9BDC5F8D2" pageId="20" pageNumber="21" type="description">
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<paragraph id="8B823698FFF5FFD9B831F8A9BDC5F8D2" blockId="20.[818,1441,1783,1861]" pageId="20" pageNumber="21">
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LSID:
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<uri id="FFAC3A9AFFF5FFD9B885F8A9BDC5F8D2" pageId="20" pageNumber="21">
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urn:lsid:zoobank.org:act:
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<uuid id="FF9B0C4DFFF5FFD9BF73F8A9BDC5F8D2" pageId="20" pageNumber="21">4BAA462E-31A4-4799-BEF6- D7BC23B346F0</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3276513FFF5FFD9B831F8DFBDA5F804" pageId="20" pageNumber="21" type="reference_group">
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<paragraph id="8B823698FFF5FFD9B831F8DFBDA5F804" blockId="20.[818,1441,1889,1939]" pageId="20" pageNumber="21">
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<treatmentCitationGroup id="AB2D11B6FFF5FFD9B831F8DFBDA5F804" pageId="20" pageNumber="21">
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<taxonomicName id="4C3D4D1BFFF5FFD9B831F8DFBDA5F8E2" ID-CoL="4P4DV" baseAuthorityName="Forel" baseAuthorityYear="1899" box="[818,939,1889,1909]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="championi">
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<emphasis id="B949EA8AFFF5FFD9B831F8DFBDA5F8E2" box="[818,939,1889,1909]" italics="true" pageId="20" pageNumber="21">P. championi</emphasis>
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</taxonomicName>
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nr;
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<treatmentCitation id="0A9C1089FFF5FFD9B8D5F8DCBAB5F8E2" author="Chomicki, G. & P. S. Ward & S. Renner" box="[982,1211,1889,1909]" page="4" pageId="20" pageNumber="21" year="2015">
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<bibRefCitation id="EFAC4B69FFF5FFD9B8D5F8DCBAB5F8E2" author="Chomicki, G. & P. S. Ward & S. Renner" box="[982,1211,1889,1909]" pageId="20" pageNumber="21" pagination="20152200" refId="ref37569" refString="Chomicki, G., P. S. Ward, and S. Renner. 2015. Macroevolutionary assembly of ant / plant symbioses: Pseudomyrmex ants and their ant-housing plants in the Neotropics. Proc. R. Soc. B. 282: 20152200." type="journal article" year="2015">Chomicki et al. 2015: 4</bibRefCitation>
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</treatmentCitation>
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. Placement in molecular phylogeny.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3276513FFF4FFD8BB91FF29BF2FF954" pageId="21" pageNumber="22" type="materials_examined">
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<paragraph id="8B823698FFF4FFD8BB91FF29BC0EFECF" blockId="21.[145,773,148,491]" pageId="21" pageNumber="22">
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<materialsCitation id="3B553CC5FFF4FFD8BB91FF29BFAFFE97" collectingDate="2007-07-21" collectionCode="UNAM" collectorName="J. Longino" elevation="1520" latitude="16.1" location="La Trinitaria" longLatPrecision="1283" longitude="-91.76667" pageId="21" pageNumber="22" specimenCode="JTLC000010310" specimenCount="1" stateProvince="MEXICO Chiapas" typeStatus="holotype">
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<typeStatus id="5486883AFFF4FFD8BB91FF29BEE7FF3F" box="[146,233,148,168]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
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dealate queen,
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<collectingRegion id="49F9F87AFFF4FFD8BA7AFF29BC27FF3F" box="[377,553,148,168]" isEnumeration="true" pageId="21" pageNumber="22">
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<collectingRegion id="49F9F87AFFF4FFD8BA7AFF29BFDAFF3F" box="[377,468,148,168]" country="Mexico" name="Mexico" pageId="21" pageNumber="22">MEXICO</collectingRegion>
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<collectingRegion id="49F9F87AFFF4FFD8BAD8FF29BC27FF3F" box="[475,553,148,168]" country="Mexico" name="Chiapas" pageId="21" pageNumber="22">Chiapas</collectingRegion>
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</collectingRegion>
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:
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<quantity id="4CC59B7DFFF4FFD8B931FF29BC60FF3F" box="[562,622,148,168]" metricMagnitude="4" metricUnit="m" metricValue="2.9" pageId="21" pageNumber="22" unit="km" value="29.0">29 km</quantity>
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E
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<location id="8EE26043FFF4FFD8B98BFF28BCF0FF3F" LSID="urn:lsid:plazi:treatment:0394878EFFF5FFD8B831F964BF2FF954:8EE26043FFF4FFD8B98BFF28BCF0FF3F" box="[648,766,148,168]" latitude="16.1" longLatPrecision="1283" longitude="-91.76667" name="La Trinitaria" pageId="21" pageNumber="22" stateProvince="MEXICO Chiapas">La Trinitaria</location>
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,
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<quantity id="4CC59B7DFFF4FFD8BB91FF0CBEECFF52" box="[146,226,177,197]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="21" pageNumber="22" unit="m" value="1520.0">
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<elevation id="0010D1ABFFF4FFD8BB91FF0CBEECFF52" box="[146,226,177,197]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="21" pageNumber="22" unit="m" value="1520.0">1,520 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE09505FFFF4FFD8BBEEFF0CBF44FF52" box="[237,330,177,197]" degrees="16" direction="north" minutes="06" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="16.1">
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16° 06
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<emphasis id="B949EA8AFFF4FFD8BA28FF0CBF3FFF52" box="[299,305,177,197]" italics="true" pageId="21" pageNumber="22">′</emphasis>
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N
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</geoCoordinate>
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<geoCoordinate id="EE09505FFFF4FFD8BA52FF0CBFBFFF52" box="[337,433,177,197]" degrees="91" direction="west" minutes="46" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-91.76667">
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91° 46
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<emphasis id="B949EA8AFFF4FFD8BA93FF0CBF98FF52" box="[400,406,177,197]" italics="true" pageId="21" pageNumber="22">′</emphasis>
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W
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</geoCoordinate>
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,
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<date id="FF831058FFF4FFD8BAB9FF0CBC22FF52" box="[442,556,177,197]" pageId="21" pageNumber="22" value="2007-07-21">
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<collectingDate id="EFC7E9B0FFF4FFD8BAB9FF0CBC22FF52" box="[442,556,177,197]" pageId="21" pageNumber="22" value="2007-07-21">21 Jul 2007</collectingDate>
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</date>
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, ex nest of
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<taxonomicName id="4C3D4D1BFFF4FFD8B999FF0FBD0DFF53" authorityName="Ward & Branstetter" authorityYear="2022" box="[666,771,178,196]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cognatus">
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<emphasis id="B949EA8AFFF4FFD8B999FF0FBD0DFF53" box="[666,771,178,196]" italics="true" pageId="21" pageNumber="22">P. cognatus</emphasis>
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</taxonomicName>
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in dead stick, pine oak scrubby forest,
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<collectorName id="26C8534EFFF4FFD8B936FF6DBC91FF74" box="[565,671,207,227]" pageId="21" pageNumber="22">J. Longino</collectorName>
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JTL6094 (
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<collectionCode id="ED2CAE5DFFF4FFD8BB9BFF51BEEEFE97" box="[152,224,236,256]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="21" pageNumber="22">UNAM</collectionCode>
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) (
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<specimenCode id="DB9B9EE3FFF4FFD8BBFAFF51BF95FE97" box="[249,411,236,256]" collectionCode="JTLC" pageId="21" pageNumber="22">JTLC000010310</specimenCode>
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)
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</materialsCitation>
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.
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<typeStatus id="5486883AFFF4FFD8BAB2FF50BC0CFE97" box="[433,514,237,256]" pageId="21" pageNumber="22" type="paratype">Paratype</typeStatus>
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dealate queen,
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<collectingRegion id="49F9F87AFFF4FFD8B9A4FF51BD0CFE97" box="[679,770,236,256]" country="Mexico" name="Mexico" pageId="21" pageNumber="22">MEXICO</collectingRegion>
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<collectingRegion id="49F9F87AFFF4FFD8BB91FEB4BEEEFE8A" box="[146,224,265,285]" country="Mexico" name="Chiapas" pageId="21" pageNumber="22">Chiapas</collectingRegion>
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: Lagos de Montebello,
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<quantity id="4CC59B7DFFF4FFD8BACBFEB4BC15FE8A" box="[456,539,265,285]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="21" pageNumber="22" unit="m" value="1520.0">1,520 m</quantity>
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,
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<geoCoordinate id="EE09505FFFF4FFD8B92AFEB4BC80FE8A" box="[553,654,265,285]" degrees="16" direction="north" minutes="08" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="16.133333">
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16° 08
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<emphasis id="B949EA8AFFF4FFD8B96FFEB4BC7CFE8A" box="[620,626,265,285]" italics="true" pageId="21" pageNumber="22">′</emphasis>
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N
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</geoCoordinate>
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<geoCoordinate id="EE09505FFFF4FFD8B99BFEB4BD0FFE8A" box="[664,769,265,285]" degrees="91" direction="west" minutes="44" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-91.73333">
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91° 44
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<emphasis id="B949EA8AFFF4FFD8B9D8FEB4BCEFFE8A" box="[731,737,265,285]" italics="true" pageId="21" pageNumber="22">′</emphasis>
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W
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</geoCoordinate>
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,
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<date id="FF831058FFF4FFD8BB91FE9ABF0BFEAC" box="[146,261,295,315]" pageId="21" pageNumber="22" value="2007-07-21">21 Jul 2007</date>
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, on ground, pine oak
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|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BADFFE9ABC5BFEAC" box="[476,597,295,315]" class="Magnoliopsida" family="Altingiaceae" genus="Liquidambar" kingdom="Plantae" order="Saxifragales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BADFFE9ABC5BFEAC" box="[476,597,295,315]" italics="true" pageId="21" pageNumber="22">Liquidambar</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
forest, J. Longino JTL6093-s (
|
|||
|
|
<collectionCode id="ED2CAE5DFFF4FFD8BA03FEF9BF4FFECF" box="[256,321,324,344]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="21" pageNumber="22" type="Museum">UCDC</collectionCode>
|
|||
|
|
) (
|
|||
|
|
<collectionCode id="ED2CAE5DFFF4FFD8BA50FEF9BF87FECF" box="[339,393,324,344]" pageId="21" pageNumber="22">JTLC</collectionCode>
|
|||
|
|
000010349).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B823698FFF4FFD8BBB1FEDCBCFFFEE2" blockId="21.[145,773,148,491]" box="[178,753,353,373]" pageId="21" pageNumber="22">
|
|||
|
|
Other material examined. Known only from the
|
|||
|
|
<typeStatus id="5486883AFFF4FFD8B972FEDFBC96FEE2" box="[625,664,354,373]" pageId="21" pageNumber="22">type</typeStatus>
|
|||
|
|
material.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B823698FFF4FFD8BBB1FEC2BCDCFE7C" blockId="21.[145,773,148,491]" pageId="21" pageNumber="22">
|
|||
|
|
Queen measurements (
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BA84FE3DBF9DFE05" box="[391,403,384,402]" italics="true" pageId="21" pageNumber="22">n</emphasis>
|
|||
|
|
= 2, with
|
|||
|
|
<typeStatus id="5486883AFFF4FFD8BAF4FEC2BC46FE04" box="[503,584,383,403]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
|||
|
|
measurement given first). HW 0.81, 0.77; HL 1.01, 1.00; MFC
|
|||
|
|
<geoCoordinate id="EE09505FFFF4FFD8B922FE21BC56FE27" box="[545,600,412,432]" degrees="0.038" direction="north" orientation="latitude" pageId="21" pageNumber="22" precision="55" value="0.038">0.038</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE09505FFFF4FFD8B962FE21BC97FE27" box="[609,665,412,432]" degrees="0.046" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="55" value="0.046">0.046</geoCoordinate>
|
|||
|
|
; LHT 0.70, 0.69; CI 0.80, 0.77; FCI
|
|||
|
|
<geoCoordinate id="EE09505FFFF4FFD8BA7BFE04BFA1FE5A" box="[376,431,441,461]" degrees="0.047" direction="north" orientation="latitude" pageId="21" pageNumber="22" precision="55" value="0.047">0.047</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE09505FFFF4FFD8BAB9FE04BFFCFE5A" box="[442,498,441,461]" degrees="0.060" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="55" value="0.06">0.060</geoCoordinate>
|
|||
|
|
; REL 0.40, 0.42; REL2 0.51, 0.54; FI 0.40, 0.43; PLI 0.67, 0.83; PWI 0.56, 0.80; MSC 4, 7.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B823698FFF4FFD8BB91FDA8BF0AFC1F" blockId="21.[145,770,532,904]" pageId="21" pageNumber="22">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BB91FDA8BF37FDB0" box="[146,313,533,552]" italics="true" pageId="21" pageNumber="22">Queen Diagnosis.</emphasis>
|
|||
|
|
Small species (see HW, HL, and LHT measurements), with elongate head (
|
|||
|
|
<collectionCode id="ED2CAE5DFFF4FFD8BA9DFD8CBFBBFDD2" box="[414,437,561,581]" country="USA" lsid="urn:lsid:biocol.org:col:15344" name="Carnegie Institution of Washington" pageId="21" pageNumber="22" type="Herbarium">CI</collectionCode>
|
|||
|
|
0.77–0.80); upper surface of mandible finely reticulate with scattered punctures, lacking striae; eyes relatively short (
|
|||
|
|
<collectionCode id="ED2CAE5DFFF4FFD8BA28FDD1BF5BFD17" box="[299,341,620,640]" pageId="21" pageNumber="22">REL</collectionCode>
|
|||
|
|
0.40–0.42, REL2 0.51–0.54); frontal carinae separated by slightly less than basal scape width; petiole stout and short, the height and width of petiole about 0.8× the length (less so in the
|
|||
|
|
<typeStatus id="5486883AFFF4FFD8BBB1FD79BF0BFD4F" box="[178,261,708,728]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
|
|||
|
|
); in profile, petiole with flat to convex anterodorsal face, ascending to summit in posterior quarter of node, then rounding into more steeply descending posterior face; profemur slender;
|
|||
|
|
<materialsCitation id="3B553CC5FFF4FFD8B9B5FD42BF0EFC1F" collectionCode="LHT" pageId="21" pageNumber="22" specimenCode="MSC 4-7" specimenCount="1">
|
|||
|
|
hind leg relatively short (
|
|||
|
|
<collectionCode id="ED2CAE5DFFF4FFD8BA2AFCA1BF58FCA7" box="[297,342,796,816]" country="Finland" lsid="urn:lsid:biocol.org:col:13463" name="Lahti City Museum" pageId="21" pageNumber="22" type="Herbarium">LHT</collectionCode>
|
|||
|
|
/HL 0.69). Head sublucid, densely punctulatecoriarious. Standing pilosity sparse and short on most of body (
|
|||
|
|
<specimenCode id="DB9B9EE3FFF4FFD8B9D1FC84BEB8FCFC" collectionCode="MSC" country="USA" lsid="urn:lsid:biocol.org:col:15587" name="Michigan State University" pageId="21" pageNumber="22" type="Herbarium">MSC 4–7</specimenCode>
|
|||
|
|
). Dark brown to brownish-black, head and gaster darker than rest of body
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B823698FFF4FFD8BB91FC0FBC61FA0C" blockId="21.[144,771,945,1435]" pageId="21" pageNumber="22">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BB91FC0FBEF2FC52" box="[146,252,946,965]" italics="true" pageId="21" pageNumber="22">Comments.</emphasis>
|
|||
|
|
This species, known only from two dealate queens, bears some resemblance to
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BA5AFC6DBFCFFC75" authorityName="Ward & Branstetter" authorityYear="2022" box="[345,449,976,994]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cognatus">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BA5AFC6DBFCFFC75" box="[345,449,976,994]" italics="true" pageId="21" pageNumber="22">P. cognatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, but differs in ways that suggest it represents a workerless inquiline (see also notes on Distribution and Biology below). The
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BA5AFBB7BFC2FB8B" authorityName="Ward & Branstetter" authorityYear="2022" box="[345,460,1034,1052]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="comitator">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BA5AFBB7BFC2FB8B" box="[345,460,1034,1052]" italics="true" pageId="21" pageNumber="22">P. comitator</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
queens are notably smaller than those of
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BBE6FB95BF41FBAD" authorityName="Ward & Branstetter" authorityYear="2022" box="[229,335,1064,1082]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cognatus">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BBE6FB95BF41FBAD" box="[229,335,1064,1082]" italics="true" pageId="21" pageNumber="22">P. cognatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="ED2CAE5DFFF4FFD8BA58FB9ABF8DFBAC" box="[347,387,1063,1083]" pageId="21" pageNumber="22">HW</collectionCode>
|
|||
|
|
0.77–0.81, HL 1.00–1.01, vs HW 0.99– 1.11, HL
|
|||
|
|
<quantity id="4CC59B7DFFF4FFD8BBF1FBF9BF65FBCF" box="[242,363,1092,1112]" metricMagnitude="-2" metricUnit="m" metricValue="3.429" metricValueMax="3.6068" metricValueMin="3.2512" pageId="21" pageNumber="22" unit="in" value="1.35" valueMax="1.42" valueMin="1.28">1.28–1.42 in</quantity>
|
|||
|
|
queens of
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BADAFBF8BC4AFBC0" authorityName="Ward & Branstetter" authorityYear="2022" box="[473,580,1093,1111]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cognatus">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BADAFBF8BC4AFBC0" box="[473,580,1093,1111]" italics="true" pageId="21" pageNumber="22">P. cognatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
), lack striae on the mandibles, have reduced mesosomal pilosity (
|
|||
|
|
<collectionCode id="ED2CAE5DFFF4FFD8B934FBDCBC69FBE2" box="[567,615,1121,1141]" country="USA" lsid="urn:lsid:biocol.org:col:15587" name="Michigan State University" pageId="21" pageNumber="22" type="Herbarium">MSC</collectionCode>
|
|||
|
|
4–7 vs
|
|||
|
|
<quantity id="4CC59B7DFFF4FFD8B9ACFBDCBD0CFBE2" box="[687,770,1121,1141]" metricMagnitude="-1" metricUnit="m" metricValue="4.699" metricValueMax="5.588" metricValueMin="3.81" pageId="21" pageNumber="22" unit="in" value="18.5" valueMax="22.0" valueMin="15.0">15–22 in</quantity>
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BB91FB3DBEF7FB05" authorityName="Ward & Branstetter" authorityYear="2022" box="[146,249,1152,1170]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cognatus">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BB91FB3DBEF7FB05" box="[146,249,1152,1170]" italics="true" pageId="21" pageNumber="22">P. cognatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
queens), and possess an oddly swollen and foreshortened petiole (
|
|||
|
|
<collectionCode id="ED2CAE5DFFF4FFD8BBD8FB21BEF2FB27" box="[219,252,1180,1200]" pageId="21" pageNumber="22">PLI</collectionCode>
|
|||
|
|
0.67–0.83, vs
|
|||
|
|
<quantity id="4CC59B7DFFF4FFD8BA86FB21BFF5FB27" box="[389,507,1180,1200]" metricMagnitude="-2" metricUnit="m" metricValue="1.2446" metricValueMax="1.3462" metricValueMin="1.143" pageId="21" pageNumber="22" unit="in" value="0.49" valueMax="0.53" valueMin="0.45">0.45–0.53 in</quantity>
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8B900FB20BC62FB38" authorityName="Ward & Branstetter" authorityYear="2022" box="[515,620,1181,1199]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cognatus">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8B900FB20BC62FB38" box="[515,620,1181,1199]" italics="true" pageId="21" pageNumber="22">P. cognatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
queens) (Figs. 8 and 9). The
|
|||
|
|
<specimenCount id="9D3BFD11FFF4FFD8BA0BFB07BF9EFB5A" box="[264,400,1209,1229]" pageId="21" pageNumber="22" type="generic">two specimens</specimenCount>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BAB7FB07BC28FB5B" authorityName="Ward & Branstetter" authorityYear="2022" box="[436,550,1210,1228]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="comitator">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BAB7FB07BC28FB5B" box="[436,550,1210,1228]" italics="true" pageId="21" pageNumber="22">P. comitator</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
do not show the same degree of modification of the petiole (compare PLI and PWI values), with the
|
|||
|
|
<typeStatus id="5486883AFFF4FFD8BBEBFB49BF37FA9F" box="[232,313,1268,1288]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
|||
|
|
being more extreme, indicating some instability in the expression of this character. Both individuals of
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8B992FAAFBD0DFAB3" authorityName="Ward & Branstetter" authorityYear="2022" box="[657,771,1298,1316]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="comitator">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8B992FAAFBD0DFAB3" box="[657,771,1298,1316]" italics="true" pageId="21" pageNumber="22">P. comitator</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
were sequenced; they are sister taxa in our UCE tree and embedded phylogenetically within the putative host species (Fig. 1). They are more closely related to
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BA6AFAD7BFDFFAEB" authorityName="Ward & Branstetter" authorityYear="2022" box="[361,465,1386,1404]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cognatus">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BA6AFAD7BFDFFAEB" box="[361,465,1386,1404]" italics="true" pageId="21" pageNumber="22">P. cognatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
populations from
|
|||
|
|
<collectingCountry id="F32A7608FFF4FFD8B97EFAD4BCD4FAEA" box="[637,730,1385,1405]" name="Honduras" pageId="21" pageNumber="22">Honduras</collectingCountry>
|
|||
|
|
and
|
|||
|
|
<collectingCountry id="F32A7608FFF4FFD8BB91FA3ABEFAFA0C" box="[146,244,1415,1435]" name="Nicaragua" pageId="21" pageNumber="22">Nicaragua</collectingCountry>
|
|||
|
|
, however, than to samples from Chiapas.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B823698FFF4FFD8BB91FA79BF2FF954" blockId="21.[146,771,1476,1731]" pageId="21" pageNumber="22">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BB91FA79BC76FA40" box="[146,632,1476,1496]" italics="true" pageId="21" pageNumber="22">
|
|||
|
|
Distribution and Biology.
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BA8AFA79BC76FA40" authorityName="Ward & Branstetter" authorityYear="2022" box="[393,632,1476,1496]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="comitator">Pseudomyrmex comitator</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
is an apparent workerless social parasite of
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BAAAFA5FBC1BFA63" authorityName="Ward & Branstetter" authorityYear="2022" box="[425,533,1506,1524]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cognatus">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BAAAFA5FBC1BFA63" box="[425,533,1506,1524]" italics="true" pageId="21" pageNumber="22">P. cognatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and is currently known only from two adjacent sites in high-elevation forest (
|
|||
|
|
<quantity id="4CC59B7DFFF4FFD8B9ABFA42BCF2F984" box="[680,764,1535,1555]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="21" pageNumber="22" unit="m" value="1520.0">1,520 m</quantity>
|
|||
|
|
) of
|
|||
|
|
<collectingRegion id="49F9F87AFFF4FFD8BBAEF9A1BEF4F9A7" box="[173,250,1564,1584]" country="Mexico" name="Chiapas" pageId="21" pageNumber="22">Chiapas</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F32A7608FFF4FFD8BA05F9A1BF41F9A7" box="[262,335,1564,1584]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
|
|||
|
|
. The
|
|||
|
|
<typeStatus id="5486883AFFF4FFD8BA86F9A1BFD8F9A7" box="[389,470,1564,1584]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
|||
|
|
was collected by Jack Longino in a nest of
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BA04F987BF7CF9DB" authorityName="Ward & Branstetter" authorityYear="2022" box="[263,370,1594,1612]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cognatus">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BA04F987BF7CF9DB" box="[263,370,1594,1612]" italics="true" pageId="21" pageNumber="22">P. cognatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
that contained workers, alate males, alate queens, and brood (larvae, prepupae) of the presumed host. Only a single dealate queen of
|
|||
|
|
<taxonomicName id="4C3D4D1BFFF4FFD8BA85F9C8BFF4F910" authorityName="Ward & Branstetter" authorityYear="2022" box="[390,506,1653,1671]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="comitator">
|
|||
|
|
<emphasis id="B949EA8AFFF4FFD8BA85F9C8BFF4F910" box="[390,506,1653,1671]" italics="true" pageId="21" pageNumber="22">P. comitator</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
was found in the nest. The
|
|||
|
|
<typeStatus id="5486883AFFF4FFD8BB91F92FBEEDF932" box="[146,227,1682,1701]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
|
|||
|
|
queen was encountered as a stray on the ground, in a nearby locality.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|